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<document id="4D36CF289FC52C706760832287F0B34F" ID-DOI="10.5281/zenodo.11512244" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512244" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1717719192119" checkinUser="felipe" docAuthor="Zug, George R." docDate="2015" docId="03856357CC4DFFECFFD05CD1FBDC6541" docLanguage="en" docName="ProcCalifAcadSci.62.5.135-190.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (5)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Kalophrynus interlineatus Striped Sticky Frog" docType="treatment" docVersion="5" lastPageNumber="164" masterDocId="FFBC1B2FCC57FFF1FFAB5C0AFF8C611C" masterDocTitle="Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs" masterLastPageNumber="190" masterPageNumber="135" pageNumber="161" updateTime="1721744699758" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A188A89821E108A6B0B50EA2162464CC">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
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<taxonomicName id="4C2CA9C2CC4DFFEBFFD05CD1FB0261EF" ID-CoL="6NK4V" authority="(Blyth, 1855) Striped Sticky Frog" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[123,1166,219,243]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4DFFEBFFD05CD1FB0261EF" bold="true" box="[123,1166,219,243]" pageId="26" pageNumber="161">
<emphasis id="B9580E53CC4DFFEBFFD05CD1FE1761EF" bold="true" box="[123,411,219,243]" italics="true" pageId="26" pageNumber="161">Kalophrynus interlineatus</emphasis>
(
<bibRefCitation id="EFBDAFB0CC4DFFEBFE075CD1FDA161EF" author="BLYTH, E." box="[428,557,219,243]" pageId="26" pageNumber="161" pagination="713 - 723" refId="ref36957" refString="BLYTH, E. 1855. Various notabilia contained in the collections presented by Capt. Berdmore and Mr. Theobald. Proceedings of the Asiatic Society of Bengal 1855 (7): 713 - 723." type="journal article" year="1855">Blyth, 1855</bibRefCitation>
) Striped Sticky Frog
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C33681CACC4DFFEBFFD05CF1FE83600D" box="[123,271,251,273]" pageId="26" pageNumber="161" type="description">
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<figureCitation id="1317CEC4CC4DFFEBFFD05CF1FF59600D" box="[123,213,251,273]" captionStart="FIGURE" captionStartId="27.[539,551,1549,1568]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-382@27.[504,1176,216,1531]" captionTargetPageId="27" captionText="FIGURE 7. Images of lectotype of Kalophrynus interlineatus (NHMUK/BMNH 1947. 2.31.26, female) in (A) dorsal and (B) ventral views, and of paralectotype (NHMUK/BMNH 1947. 2.31.27, male) in (C) dorsal view [photographer G.R. Zug]." figureDoi="http://doi.org/10.5281/zenodo.11512256" httpUri="https://zenodo.org/record/11512256/files/figure.png" pageId="26" pageNumber="161">Figures 7</figureCitation>
,
<figureCitation id="1317CEC4CC4DFFEBFF495CF6FE85600D" box="[226,265,252,273]" captionStart="FIGURE" captionStartId="44.[155,167,1351,1370]" captionTargetBox="[169,1129,216,1336]" captionTargetId="figure-124@44.[169,1129,215,1336]" captionTargetPageId="44" captionText="FIGURE 11. Coloration of living representatives of pleurostigma-interlineatus group. (A) Dorsolateral view of a K. anya (USNM 520321) from the Chatthin Wildlife Sanctuary, Sagaing, Myanmar [photographer, C. Hansen]; (B) dorsolateral view of a K. anya (USNM 523966) from the Chatthin Wildlife Sanctuary, Sagaing, Myanmar [photographer, G.R. Zug]; (C) dorsolateral view of a K. interlineatus (CAS) from the Tanintharyi National Park, Tanintharyi, Myanmar [photograph- er, J.V. Vindum]; (D) dorsolateral view of a K. meizon (FMNH 242796) from Poring Station, Mt. Kinabulu Park, Ranua District, Sabah [photographer, R.F. Inger]; (E) dorsolateral view of a K. sinensis (UK 333148) from Agusan del Norte, Mindanao, Philippines [photographer, R.M. Brown]; (F) dorsolateral view of a K. pleurostigma (UK 333148) from Gunung Kunyit, Jambi, Sumatra [photographer, E.N. Smith]." figureDoi="http://doi.org/10.5281/zenodo.11512262" httpUri="https://zenodo.org/record/11512262/files/figure.png" pageId="26" pageNumber="161">11C</figureCitation>
.
</paragraph>
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<taxonomicName id="4C2CA9C2CC4DFFEBFFD05D29FD8A6024" ID-CoL="39TSX" authority="Blyth, 1855" authorityName="Blyth" authorityYear="1855" box="[123,518,290,312]" class="Amphibia" family="Microhylidae" genus="Engystoma" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="genus">
<emphasis id="B9580E53CC4DFFEBFFD05D29FF6A6024" box="[123,230,291,312]" italics="true" pageId="26" pageNumber="161">Engystoma</emphasis>
(?)
<emphasis id="B9580E53CC4DFFEBFEA65D28FE02602B" box="[269,398,290,311]" italics="true" pageId="26" pageNumber="161">interlineatum</emphasis>
<materialsCitation id="3B44D81CCC4DFFEBFE3F5D28FDD96044" country="Myanmar" location="Asiatic Society of Bengal" pageId="26" pageNumber="161" specimenCount="1" stateProvince="Bago">
<bibRefCitation id="EFBDAFB0CC4DFFEBFE3F5D28FD8A6024" author="BLYTH, E." box="[404,518,290,312]" pageId="26" pageNumber="161" pagination="713 - 723" refId="ref36957" refString="BLYTH, E. 1855. Various notabilia contained in the collections presented by Capt. Berdmore and Mr. Theobald. Proceedings of the Asiatic Society of Bengal 1855 (7): 713 - 723." type="journal article" year="1855">Blyth, 1855</bibRefCitation>
,
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Journal of the
<location id="8EF3849ACC4DFFEBFD0B5D28FC1D602B" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4DFFEBFD0B5D28FC1D602B" box="[672,913,290,311]" country="Myanmar" name="Asiatic Society of Bengal" pageId="26" pageNumber="161" stateProvince="Bago">Asiatic Society of Bengal</location>
</emphasis>
[1854] 23:732 [
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locality: “
<collectingRegion id="49E81CA3CC4DFFEBFF665D49FF716044" box="[205,253,323,344]" country="Myanmar" name="Bago" pageId="26" pageNumber="161">Pegu</collectingRegion>
”,
<collectingCountry id="F33B92D1CC4DFFEBFEBF5D49FEFD6044" box="[276,369,323,344]" name="Myanmar" pageId="26" pageNumber="161">Myanmar</collectingCountry>
; see Comments below]
</materialsCitation>
</taxonomicName>
.
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<materialsCitation id="3B44D81CCC4DFFEBFF005D7FFC4F60B1" collectingDate="1947-02-31" collectionCode="TYPE, MATERIAL" country="United Kingdom" location="Natural History Museum" municipality="British Museum of Natural History" pageId="26" pageNumber="161" specimenCount="1" typeStatus="lectotype">
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.—
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:
<location id="8EF3849ACC4DFFEBFDA35D7EFC9B6090" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4DFFEBFDA35D7EFC9B6090" box="[520,791,372,396]" country="United Kingdom" municipality="British Museum of Natural History" name="Natural History Museum" pageId="26" pageNumber="161">Natural History Museum</location>
<collectingCountry id="F33B92D1CC4DFFEBFC8A5D7EFC5B6090" box="[801,983,372,396]" name="United Kingdom" pageId="26" pageNumber="161">United Kingdom</collectingCountry>
, formerly
<collectingMunicipality id="6BF7483BCC4DFFEBFBE65D7EFE5D60B1" pageId="26" pageNumber="161">British Museum of Natural History</collectingMunicipality>
68.4.3.128, recataloged as
<date id="FF92F481CC4DFFEBFC995D9FFC2460B0" box="[818,936,405,429]" pageId="26" pageNumber="161" value="1947-02-31">
<collectingDate id="EFD60D69CC4DFFEBFC995D9FFC2460B0" box="[818,936,405,429]" pageId="26" pageNumber="161" value="1947-02-31">1947.2.31.</collectingDate>
</date>
26
</materialsCitation>
.
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<typeStatus id="54976CE3CC4DFFEBFC755D9FFB0160B0" box="[990,1165,405,428]" pageId="26" pageNumber="161" type="paralectotype">PARALECTOTYPE</typeStatus>
</smallCapsWord>
:
<collectionCode id="ED3D4A84CC4DFFEBFFD05DBCFF6C60D1" box="[123,224,438,461]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="161" type="Museum">NHMUK</collectionCode>
/
<specimenCode id="DB8A7A3ACC4DFFEBFF4C5DBCFED660D1" box="[231,346,437,461]" collectionCode="BMNH" pageId="26" pageNumber="161">BMNH 68</specimenCode>
.4.3. 129, recataloged as
<date id="FF92F481CC4DFFEBFDC25DBFFD5760D1" box="[617,731,437,461]" pageId="26" pageNumber="161" value="1947-02-31">
<collectingDate id="EFD60D69CC4DFFEBFDC25DBFFD5760D1" box="[617,731,437,461]" pageId="26" pageNumber="161" value="1947-02-31">1947.2.31.</collectingDate>
</date>
27.
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discussion on the assignment of type specimens in the
<collectingMunicipality id="6BF7483BCC4DFFEBFEC15DDCFE5260F2" box="[362,478,470,494]" pageId="26" pageNumber="161">Comments</collectingMunicipality>
section below
</materialsCitation>
.
</paragraph>
</subSubSection>
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.
</emphasis>
<materialsCitation id="3B44D81CCC4DFFEBFD565DFCFF2F6373" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="1" specimenCount-female="1">
Moderate-sized adults, not sexually dimorphic although females average larger (mean 41.5, 35.046.
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<collectionCode id="ED3D4A84CC4DFFEBFC3B5E1DFC4E6333" box="[912,962,535,559]" pageId="26" pageNumber="161">SVL</collectionCode>
) than males (39.2, 33.744.
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). These differences are reflected in the other measurements: HeadL 11.215.
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<specimenCount id="9D2A19C8CC4DFFEBFFD05E52FF2F6373" box="[123,163,600,623]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
</materialsCitation>
, 10.313.
<quantity id="4CD47FA4CC4DFFEBFEA55E52FEC76373" box="[270,331,600,624]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="161" unit="mm" value="4.0">4 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFEF85E52FEF76373" box="[339,379,600,623]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; HeadW 12.715.
<quantity id="4CD47FA4CC4DFFEBFD945E52FDF06373" box="[575,636,600,624]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="161" unit="mm" value="1.0">1 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFD2F5E52FD206373" box="[644,684,600,623]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 11.313.
<quantity id="4CD47FA4CC4DFFEBFCBC5E5DFCD86373" box="[791,852,599,623]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="161" unit="mm" value="5.0">5 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFCF05E52FC0F6373" box="[859,899,600,623]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; SnEye 4.35.
<quantity id="4CD47FA4CC4DFFEBFB895E52FBD36373" box="[1058,1119,600,623]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="161" unit="mm" value="7.0">7 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFBCC5E52FB036373" box="[1127,1167,600,623]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 3.8
<quantity id="4CD47FA4CC4DFFEBFF025E72FF766393" box="[169,250,632,656]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="26" pageNumber="161" unit="mm" value="5.3">5.3 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFEA85E72FEA76393" box="[259,299,632,655]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; NarEye 2.8
<quantity id="4CD47FA4CC4DFFEBFE695E72FD9F6393" box="[450,531,632,656]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="26" pageNumber="161" unit="mm" value="3.7">3.7 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFDB75E72FDC86393" box="[540,580,632,655]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 2.3
<quantity id="4CD47FA4CC4DFFEBFD2A5E72FD5E6393" box="[641,722,632,656]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="26" pageNumber="161" unit="mm" value="3.8">3.8 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFD715E72FC8E6393" box="[730,770,632,655]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; EyeD 3.6
<quantity id="4CD47FA4CC4DFFEBFC2E5E72FC5A6393" box="[901,982,632,656]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="161" unit="mm" value="4.5">4.5 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFC745E72FB8B6393" box="[991,1031,632,655]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 3.4
<quantity id="4CD47FA4CC4DFFEBFBEF5E72FB196393" box="[1092,1173,632,656]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="161" unit="mm" value="4.5">4.5 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFFD05E93FF2F63AC" box="[123,163,665,688]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; Tymp 3.13.
<quantity id="4CD47FA4CC4DFFEBFEE95E92FEF363AC" box="[322,383,664,688]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="161" unit="mm" value="8.0">8 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFE275E93FE3863AC" box="[396,436,665,688]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 2.6
<quantity id="4CD47FA4CC4DFFEBFE5E5E92FDCA63AC" box="[501,582,664,688]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="161" unit="mm" value="3.5">3.5 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFDF85E93FDF763AC" box="[595,635,665,688]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; ForarmL 9.013.
<quantity id="4CD47FA4CC4DFFEBFCE35E92FC0963AC" box="[840,901,664,688]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="161" unit="mm" value="2.0">2 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFC395E93FC3663AC" box="[914,954,665,688]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 8.712.
<quantity id="4CD47FA4CC4DFFEBFBB75E92FBD563AC" box="[1052,1113,664,688]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="161" unit="mm" value="2.0">2 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFBCD5E93FB0263AC" box="[1126,1166,665,688]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; HandL 8.411.
<quantity id="4CD47FA4CC4DFFEBFEB25EB3FEDA63CC" box="[281,342,697,721]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="161" unit="mm" value="3.0">3 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFEF75EB3FE0863CC" box="[348,388,697,720]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 7.710.0 mm
<specimenCount id="9D2A19C8CC4DFFEBFD8E5EB3FDC163CC" box="[549,589,697,720]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; ThghL 13.718.
<quantity id="4CD47FA4CC4DFFEBFCAE5EB3FCCE63CC" box="[773,834,697,721]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="161" unit="mm" value="6.0">6 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFCE35EB3FCFC63CC" box="[840,880,697,720]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 14.117.
<quantity id="4CD47FA4CC4DFFEBFC705EB3FB9463CC" box="[987,1048,697,720]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="161" unit="mm" value="7.0">7 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFBB55EB3FBCA63CC" box="[1054,1094,697,720]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; CrusL 13.016.
<quantity id="4CD47FA4CC4DFFEBFF735ED3FE9963ED" box="[216,277,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="161" unit="mm" value="2.0">2 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFE8E5ED0FEC163ED" box="[293,333,730,753]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 12.015.
<quantity id="4CD47FA4CC4DFFEBFE6B5ED3FE7163ED" box="[448,509,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="161" unit="mm" value="6.0">6 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFDA65ED0FDB963ED" box="[525,565,730,753]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; TarsL 8.510.
<quantity id="4CD47FA4CC4DFFEBFD435ED3FCA963ED" box="[744,805,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="161" unit="mm" value="4.0">4 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFC9E5ED0FCD163ED" box="[821,861,730,753]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 7.910.
<quantity id="4CD47FA4CC4DFFEBFC685ED3FB8C63ED" box="[963,1024,729,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="161" unit="mm" value="8.0">8 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFBBB5ED0FBB463ED" box="[1040,1080,730,753]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
; HndfL 13.016.
<quantity id="4CD47FA4CC4DFFEBFF735EF3FE99620D" box="[216,277,761,785]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="161" unit="mm" value="6.0">6 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFEB75EF0FEC8620D" box="[284,324,762,785]" count="1" pageId="26" pageNumber="161" type="female">♀♀</specimenCount>
, 11.715.
<quantity id="4CD47FA4CC4DFFEBFE055EF0FE67620D" box="[430,491,762,786]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="161" unit="mm" value="8.0">8 mm</quantity>
<specimenCount id="9D2A19C8CC4DFFEBFE595EF0FD96620D" box="[498,538,762,785]" count="1" pageId="26" pageNumber="161" type="male">♂♂</specimenCount>
.
<materialsCitation id="3B44D81CCC4DFFEBFD8C5EF3FCF7622D" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="36" specimenCount-female="36">
Body proportion means differ between females and males, although not greatly so (all values are percent): HeadL/
<collectionCode id="ED3D4A84CC4DFFEBFD7B5F10FC8E622E" box="[720,770,794,818]" pageId="26" pageNumber="161">SVL</collectionCode>
29
<specimenCount id="9D2A19C8CC4DFFEBFC9A5F10FCF7622D" box="[817,891,794,818]" count="36" pageId="26" pageNumber="161" type="female">36 ♀♀</specimenCount>
</materialsCitation>
, 28
<specimenCount id="9D2A19C8CC4DFFEBFC195F10FC70622D" box="[946,1020,794,818]" count="31" pageId="26" pageNumber="161" type="male">31 ♂♂</specimenCount>
;
<materialsCitation id="3B44D81CCC4DFFEBFBA05F10FF61624E" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="36" specimenCount-female="36">
HeadW/
<collectionCode id="ED3D4A84CC4DFFEBFBCF5F10FB1A622E" box="[1124,1174,794,818]" pageId="26" pageNumber="161">SVL</collectionCode>
31
<specimenCount id="9D2A19C8CC4DFFEBFF085F30FF61624E" box="[163,237,826,850]" count="36" pageId="26" pageNumber="161" type="female">36 ♀♀</specimenCount>
</materialsCitation>
, 29
<specimenCount id="9D2A19C8CC4DFFEBFE8F5F30FEE3624E" box="[292,367,826,850]" count="36" pageId="26" pageNumber="161" type="male">36 ♂♂</specimenCount>
; HeadW/HeadL 95
<specimenCount id="9D2A19C8CC4DFFEBFDE55F30FD14624E" box="[590,664,826,850]" count="11" pageId="26" pageNumber="161" type="female">11 ♀♀</specimenCount>
, 95
<specimenCount id="9D2A19C8CC4DFFEBFD645F30FCA9624E" box="[719,805,826,850]" count="116" pageId="26" pageNumber="161" type="male">116 ♂♂</specimenCount>
; SnEye/HeadL 36
<specimenCount id="9D2A19C8CC4DFFEBFC525F30FBC8624E" box="[1017,1092,826,850]" count="41" pageId="26" pageNumber="161" type="female">41 ♀♀</specimenCount>
, 36
<specimenCount id="9D2A19C8CC4DFFEBFBD05F30FF2F626E" count="45" pageId="26" pageNumber="161" type="male">45 ♂♂</specimenCount>
; NarEye/SnEye 56
<specimenCount id="9D2A19C8CC4DFFEBFE2C5F51FE58626E" box="[391,468,859,883]" count="66" pageId="26" pageNumber="161" type="female">66 ♀♀</specimenCount>
, 54
<specimenCount id="9D2A19C8CC4DFFEBFDA55F51FDD7626E" box="[526,603,859,883]" count="78" pageId="26" pageNumber="161" type="male">78 ♂♂</specimenCount>
; EyeD/HeadL 28
<specimenCount id="9D2A19C8CC4DFFEBFC865F51FCF6626E" box="[813,890,859,883]" count="36" pageId="26" pageNumber="161" type="female">36 ♀♀</specimenCount>
, 30
<specimenCount id="9D2A19C8CC4DFFEBFC1F5F51FB8D626E" box="[948,1025,859,883]" count="37" pageId="26" pageNumber="161" type="male">37 ♂♂</specimenCount>
; Tymp/EyeD 72
<specimenCount id="9D2A19C8CC4DFFEBFF085F71FF70628F" box="[163,252,891,915]" count="100" pageId="26" pageNumber="161" type="female">100 ♀♀</specimenCount>
, 71
<specimenCount id="9D2A19C8CC4DFFEBFE9E5F71FE0D628F" box="[309,385,891,915]" count="79" pageId="26" pageNumber="161" type="male">79 ♂♂</specimenCount>
;
<materialsCitation id="3B44D81CCC4DFFEBFE395F76FD1B628F" box="[402,663,891,915]" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="31" specimenCount-female="31">
Forarm/
<collectionCode id="ED3D4A84CC4DFFEBFE435F71FD96628F" box="[488,538,891,915]" pageId="26" pageNumber="161">SVL</collectionCode>
25
<specimenCount id="9D2A19C8CC4DFFEBFDE05F71FD1B628F" box="[587,663,891,915]" count="31" pageId="26" pageNumber="161" type="female">31 ♀♀</specimenCount>
</materialsCitation>
, 24
<specimenCount id="9D2A19C8CC4DFFEBFD645F71FC90628F" box="[719,796,891,915]" count="31" pageId="26" pageNumber="161" type="male">31 ♂♂</specimenCount>
; Forarm/CrusL 69
<specimenCount id="9D2A19C8CC4DFFEBFC5D5F71FBCE628F" box="[1014,1090,891,915]" count="83" pageId="26" pageNumber="161" type="female">83 ♀♀</specimenCount>
, 65
<specimenCount id="9D2A19C8CC4DFFEBFBD05F71FF2F62AF" count="86" pageId="26" pageNumber="161" type="male">86 ♂♂</specimenCount>
;
<materialsCitation id="3B44D81CCC4DFFEBFF185F91FE4562AF" box="[179,457,923,948]" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="147" specimenCount-female="147">
HndlL/
<collectionCode id="ED3D4A84CC4DFFEBFEAB5F96FEBE62A8" box="[256,306,924,948]" pageId="26" pageNumber="161">SVL</collectionCode>
127
<specimenCount id="9D2A19C8CC4DFFEBFEDB5F96FE4562AF" box="[368,457,924,948]" count="147" pageId="26" pageNumber="161" type="female">147 ♀♀</specimenCount>
</materialsCitation>
, 122
<specimenCount id="9D2A19C8CC4DFFEBFDA55F96FDEB62AF" box="[526,615,924,948]" count="148" pageId="26" pageNumber="161" type="male">148 ♂♂</specimenCount>
;
<materialsCitation id="3B44D81CCC4DFFEBFDD35F96FCFC62AF" box="[632,880,924,948]" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="40" specimenCount-female="40">
CrusL/
<collectionCode id="ED3D4A84CC4DFFEBFD695F96FD7862A8" box="[706,756,924,948]" pageId="26" pageNumber="161">SVL</collectionCode>
34
<specimenCount id="9D2A19C8CC4DFFEBFC8F5F96FCFC62AF" box="[804,880,924,948]" count="40" pageId="26" pageNumber="161" type="female">40 ♀♀</specimenCount>
</materialsCitation>
, 30
<specimenCount id="9D2A19C8CC4DFFEBFC0C5F96FC7F62AF" box="[935,1011,924,948]" count="39" pageId="26" pageNumber="161" type="male">39 ♂♂</specimenCount>
; CrusL/ThghL 83
<specimenCount id="9D2A19C8CC4DFFEBFF085FB6FF6162CF" box="[163,237,956,980]" count="95" pageId="26" pageNumber="161" type="female">95 ♀♀</specimenCount>
, 81
<specimenCount id="9D2A19C8CC4DFFEBFE885FB6FEE262CF" box="[291,366,956,980]" count="96" pageId="26" pageNumber="161" type="male">96 ♂♂</specimenCount>
; TarsL/ThghL 51
<specimenCount id="9D2A19C8CC4DFFEBFD925FB6FD0F62CF" box="[569,643,956,980]" count="62" pageId="26" pageNumber="161" type="female">62 ♀♀</specimenCount>
, 53
<specimenCount id="9D2A19C8CC4DFFEBFD115FB6FC8862CF" box="[698,772,956,980]" count="63" pageId="26" pageNumber="161" type="male">63 ♂♂</specimenCount>
;
<materialsCitation id="3B44D81CCC4DFFEBFCB85FB6FB8162CF" box="[787,1037,956,980]" collectionCode="SVL" pageId="26" pageNumber="161" specimenCount="40" specimenCount-female="40">
HndfL/
<collectionCode id="ED3D4A84CC4DFFEBFCC95FB6FC1862C8" box="[866,916,956,980]" pageId="26" pageNumber="161">SVL</collectionCode>
30
<specimenCount id="9D2A19C8CC4DFFEBFC685FB6FB8162CF" box="[963,1037,956,980]" count="40" pageId="26" pageNumber="161" type="female">40 ♀♀</specimenCount>
</materialsCitation>
, 30
<specimenCount id="9D2A19C8CC4DFFEBFBEF5FB6FB0262CF" box="[1092,1166,956,980]" count="40" pageId="26" pageNumber="161" type="male">40 ♂♂</specimenCount>
; HndfL/ThghL 84
<specimenCount id="9D2A19C8CC4DFFEBFEEB5FD6FE0662E8" box="[320,394,988,1012]" count="95" pageId="26" pageNumber="161" type="female">95 ♀♀</specimenCount>
, 80
<specimenCount id="9D2A19C8CC4DFFEBFE6B5FD6FD8662E8" box="[448,522,988,1012]" count="93" pageId="26" pageNumber="161" type="male">93 ♂♂</specimenCount>
.
</paragraph>
<paragraph id="8B93D241CC4DFFEBFF005FF7FB9F65AA" blockId="26.[123,1175,372,1726]" pageId="26" pageNumber="161">
I noted earlier in the Morphometric section that the
<collectingRegion id="49E81CA3CC4DFFEBFD4B5FF7FCD76509" box="[736,859,1021,1045]" country="Myanmar" name="Taninthayi" pageId="26" pageNumber="161">Tanintharyi</collectingRegion>
sample displayed no significant sexual dimorphism and that females were significantly larger in the SE Asia sample. When the samples are combined, the overall size dimorphism declines, although a few traits remain dimorphic. Females have larger heads than males; HeadL, HeadW, and SnEye average larger, but none of the other head metrics do, and there are no proportional differences in the head or body metrics. There is strong dimorphism in ForarmL and HndfL with females being larger in both.
</paragraph>
<paragraph id="8B93D241CC4DFFEBFF0058CAFCE06780" blockId="26.[123,1175,372,1726]" pageId="26" pageNumber="161">
In their amphibian atlas,
<bibRefCitation id="EFBDAFB0CC4DFFEBFE6E58B5FDF065CB" author="FEI, L. &amp; C. YE &amp; J. JIANG" box="[453,636,1215,1239]" pageId="26" pageNumber="161" refId="ref37499" refString="FEI, L., C. YE, AND J. JIANG. 2010. Zhongguo liang qi dong wu cai se tu jian [Colored Atlas of Chinese Amphibians]. Sichuan Publishing Group, Chengdu." type="book" year="2010">Fei et al. (2010)</bibRefCitation>
reported Chinese males as
<quantity id="4CD47FA4CC4DFFEBFC1E58B5FBAB65CB" box="[949,1063,1215,1239]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="26" pageNumber="161" unit="mm" value="35.0" valueMax="38.0" valueMin="32.0">3238 mm</quantity>
<collectionCode id="ED3D4A84CC4DFFEBFB9858B5FBE965CB" box="[1075,1125,1215,1239]" pageId="26" pageNumber="161">SVL</collectionCode>
and females to
<quantity id="4CD47FA4CC4DFFEBFF5958EAFEB065EB" box="[242,316,1248,1272]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="161" unit="mm" value="40.0">40 mm</quantity>
. These adult lengths are smaller, likely significantly so, than my sample which is comprised of frogs from more southern locations.
<bibRefCitation id="EFBDAFB0CC4DFFEBFD11590BFC1F6404" author="KARSEN, S. J. &amp; LAU, AND &amp; BOGADEK" box="[698,915,1280,1304]" pageId="26" pageNumber="161" refId="ref37741" refString="KARSEN, S. J., M. W. LAU, AND BOGADEK. 1986. Hong Kong Amphibians and Reptiles. Urban Council, Hong Kong." type="book" year="1986">Karsen et al. (1986)</bibRefCitation>
did not give a range or mean for
<collectingCountry id="F33B92D1CC4DFFEBFF42592BFEE66425" box="[233,362,1313,1337]" name="Hong Kong" pageId="26" pageNumber="161">Hong Kong</collectingCountry>
<taxonomicName id="4C2CA9C2CC4DFFEBFEDE592BFD936424" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[373,543,1312,1336]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4DFFEBFEDE592BFD936424" box="[373,543,1312,1336]" italics="true" pageId="26" pageNumber="161">K. interlineatus</emphasis>
</taxonomicName>
and his “up to
<quantity id="4CD47FA4CC4DFFEBFD7E592AFC826424" box="[725,782,1312,1336]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="161" unit="cm" value="6.0">6 cm</quantity>
long”, contrasts sharply with Feis lengths and is much larger than the maximum
<collectionCode id="ED3D4A84CC4DFFEBFDDA594BFD2F6445" box="[625,675,1345,1369]" pageId="26" pageNumber="161">SVL</collectionCode>
in our sample or that reported for this species elsewhere.
<bibRefCitation id="EFBDAFB0CC4DFFEBFF5F5968FE196465" author="BOURRET, R." box="[244,405,1377,1401]" pageId="26" pageNumber="161" pagination="1 - 547" refId="ref37093" refString="BOURRET, R. 1942. Les Batraciens de l'Indochine. Memoires de l'Institut Oceanographique de l'Indochine 6: 1 - 547." type="journal article" year="1942">Bourret (1942)</bibRefCitation>
gave single value adult sizes of
<quantity id="4CD47FA4CC4DFFEBFD51596BFCC86465" box="[762,836,1377,1401]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="26" pageNumber="161" unit="mm" value="38.0">38 mm</quantity>
<collectionCode id="ED3D4A84CC4DFFEBFCE6596BFCF36465" box="[845,895,1377,1401]" pageId="26" pageNumber="161">SVL</collectionCode>
for males and
<quantity id="4CD47FA4CC4DFFEBFB88596BFBE26465" box="[1059,1134,1377,1401]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="26" pageNumber="161" unit="mm" value="44.0">44 mm</quantity>
for females.
<bibRefCitation id="EFBDAFB0CC4DFFEBFF765988FEE56486" author="BERRY, P. Y." box="[221,361,1410,1434]" pageId="26" pageNumber="161" refId="ref36936" refString="BERRY, P. Y. 1975. The Amphibian Fauna of Peninsular Malaysia. Tropical Press, Kuala Lumpur." type="book" year="1975">Berry (1975)</bibRefCitation>
gave total lengths of Peninsular
<collectingCountry id="F33B92D1CC4DFFEBFD7A5988FCBF6486" box="[721,819,1410,1434]" name="Malaysia" pageId="26" pageNumber="161">Malaysia</collectingCountry>
<taxonomicName id="4C2CA9C2CC4DFFEBFC975988FC686486" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[828,996,1410,1434]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4DFFEBFC975988FC686486" box="[828,996,1410,1434]" italics="true" pageId="26" pageNumber="161">K. interlineatus</emphasis>
</taxonomicName>
as
<quantity id="4CD47FA4CC4DFFEBFBA15988FBF16485" box="[1034,1149,1410,1434]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.8" metricValueMin="4.7" pageId="26" pageNumber="161" unit="mm" value="52.5" valueMax="58.0" valueMin="47.0">4758 mm</quantity>
. I interpret his total length as
<collectionCode id="ED3D4A84CC4DFFEBFE0A59A8FE5F64A6" box="[417,467,1442,1466]" pageId="26" pageNumber="161">SVL</collectionCode>
, hence his size data are greater than the range for my Tanintharyi and Thai-Indochina samples and more closely match the size of
<taxonomicName id="4C2CA9C2CC4DFFEBFC8959C9FC0164C6" authorityName="Zug" authorityYear="2015" box="[802,909,1475,1498]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="species" species="meizon">
<emphasis id="B9580E53CC4DFFEBFC8959C9FC0164C6" box="[802,909,1475,1498]" italics="true" pageId="26" pageNumber="161">K. meizon</emphasis>
</taxonomicName>
. Although I do not reject his data, I am uncertain how to interpret it and have not included his values in
<tableCitation id="C6AEE7FACC4DFFEBFC7C59E9FBAB64E7" box="[983,1063,1507,1531]" captionStart="TABLE" captionStartId="13.[134,147,468,534]" captionTargetId="graphics-1066@13.[83,1597,560,1521]" captionTargetPageId="13" captionText="TABLE3.SelectcharacteristicofcurrentlyrecognizedspeciesofKalophrynus." pageId="26" pageNumber="161">Table 3</tableCitation>
. Manthey and Grossmann (1997), in contrast, gave
<quantity id="4CD47FA4CC4DFFEBFDEE5A09FD3B6707" box="[581,695,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="26" pageNumber="161" unit="mm" value="38.0" valueMax="41.0" valueMin="35.0">3541 mm</quantity>
<collectionCode id="ED3D4A84CC4DFFEBFD6A5A09FD7F6707" box="[705,755,1539,1563]" pageId="26" pageNumber="161">SVL</collectionCode>
for males and
<quantity id="4CD47FA4CC4DFFEBFC365A09FB836707" box="[925,1039,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.6" metricValueMin="3.8" pageId="26" pageNumber="161" unit="mm" value="42.0" valueMax="46.0" valueMin="38.0">3846 mm</quantity>
for females from Peninsular
<collectingCountry id="F33B92D1CC4DFFEBFE9A5A2EFE1F6720" box="[305,403,1572,1596]" name="Malaysia" pageId="26" pageNumber="161">Malaysia</collectingCountry>
, matching my adult size range for the Tanintharyi and Thai-Indochina samples. A more recent study of Malaysian
<taxonomicName id="4C2CA9C2CC4DFFEBFDF85A4FFD766740" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[595,762,1604,1628]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="26" pageNumber="161" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4DFFEBFDF85A4FFD766740" box="[595,762,1604,1628]" italics="true" pageId="26" pageNumber="161">K. interlineatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBDAFB0CC4DFFEBFCA15A4EFC366740" author="CHAN, K. O. &amp; L. L. GRISMER &amp; J. GRISMER" box="[778,954,1604,1628]" pageId="26" pageNumber="161" pagination="60 - 68" refId="ref37116" refString="CHAN, K. O., L. L. GRISMER, AND J. GRISMER. 2011. A new insular endemic frog of the genus Kalophrynus Tschudi, 1838 (Anura: Microhylidae) from Tioman Island, Pahang, Peninsular Malaysia. Zootaxa 3123: 60 - 68." type="journal article" year="2011">Chan et al, 2011</bibRefCitation>
) reported 33.738.
<quantity id="4CD47FA4CC4DFFEBFB235A4EFF286760" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="161" unit="mm" value="1.0">1 mm</quantity>
<collectionCode id="ED3D4A84CC4DFFEBFF075A6FFF526761" box="[172,222,1637,1661]" pageId="26" pageNumber="161">SVL</collectionCode>
for adult males (
<emphasis id="B9580E53CC4DFFEBFE315A6FFE4C6760" box="[410,448,1637,1660]" italics="true" pageId="26" pageNumber="161">n =</emphasis>
7) and 41.147.
<quantity id="4CD47FA4CC4DFFEBFDD95A6FFD236760" box="[626,687,1637,1661]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="161" unit="mm" value="3.0">3 mm</quantity>
<collectionCode id="ED3D4A84CC4DFFEBFD1C5A6FFD656761" box="[695,745,1637,1661]" pageId="26" pageNumber="161">SVL</collectionCode>
for adult females (
<emphasis id="B9580E53CC4DFFEBFC115A6FFC4B6760" box="[954,967,1637,1660]" italics="true" pageId="26" pageNumber="161">n</emphasis>
=10); these values also match my values for the more northern Thai-Indochinese sample.
</paragraph>
<paragraph id="8B93D241CC4DFFEAFF005AACFED3624E" blockId="26.[123,1175,372,1726]" lastBlockId="27.[123,478,219,1715]" lastPageId="27" lastPageNumber="162" pageId="26" pageNumber="161">
Tongue is broadly obovate, posterior edge smooth; vomerine teeth are absent; palatal fold morphology appears relatively uniform among individuals although these data are not quantified. Vomerine folds are smoothedged flaps, one adjacent to each choanae and widely separated from its opposite. Postorbital folds are more variable; in Tanintharyi frogs, a pair of folds is separated on the midline, and each side has two or three. Variation in the Thai and Indochina frogs is described in the morphology section. Buccal fold is a continuous lobular fold with abutting lobes and each lobe a low, round-edged rectangle. See Fei et al. (2005) for an illustration of the buccal cavity of a Chinese
<emphasis id="B9580E53CC4CFFEAFF1A5F36FED4624F" box="[177,344,827,851]" italics="true" pageId="27" pageNumber="162">K. interlineatus</emphasis>
.
</paragraph>
<paragraph id="8B93D241CC4CFFEAFF005F51FEB1676F" blockId="27.[123,478,219,1715]" pageId="27" pageNumber="162">Fingers lack webbing. Both finger and toe tips are bluntly rounded. Subarticular tubercle are well developed on the digits; only third finger bears a subarticular tubercle on free portion of digit; all fingers have a tubercle at their base and another row between a large, circular, nearly medial outer palmar tubercle. For the hindfoot, each toe has a basal subarticular tubercle, third and fifth toes have an addition tubercle on free portion of digit, two tubercles on fourth toe. Inner and outer metatarsal tubercles are present; inner is large, nearly circular to elliptical; outer small to nearly absent and circular. Toes modestly webbed WebIII2 median 2.0 (1.03.0), WebIV1 1.0 (0.52.0). Digit lengths nearly constant for fore- and hindfeet; finger formula 3&gt;2≈1&gt;4; toe formula 4&gt;3&gt;5&gt;2&gt;1.</paragraph>
<caption id="DF5382C9CC4CFFEAFDB05A07FC886770" ID-DOI="http://doi.org/10.5281/zenodo.11512256" ID-Zenodo-Dep="11512256" httpUri="https://zenodo.org/record/11512256/files/figure.png" pageId="27" pageNumber="162" startId="27.[539,551,1549,1568]" targetBox="[123,1176,216,1715]" targetPageId="27" targetType="figure">
<paragraph id="8B93D241CC4CFFEAFDB05A07FC886770" blockId="27.[507,1174,1548,1644]" pageId="27" pageNumber="162">
<smallCapsWord id="8D75449DCC4CFFEAFDB05A07FDD76702" baselines="1563,1562" box="[539,603,1549,1568]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="27" pageNumber="162">FIGURE</smallCapsWord>
7. Images of lectotype of
<taxonomicName id="4C2CA9C2CC4CFFEAFC365A06FB196703" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[925,1173,1548,1567]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="27" pageNumber="162" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4CFFEAFC365A06FB196703" box="[925,1173,1548,1567]" italics="true" pageId="27" pageNumber="162">Kalophrynus interlineatus</emphasis>
</taxonomicName>
(NHMUK/BMNH 1947. 2.31.26, female) in (A) dorsal and (B) ventral views, and of paralectotype (NHMUK/BMNH 1947. 2.31.27, male) in (C) dorsal view [photographer G.R. Zug].
</paragraph>
</caption>
<paragraph id="8B93D241CC4CFFEDFF005A71FCCA6307" blockId="27.[123,478,219,1715]" lastBlockId="28.[123,1174,219,1726]" lastPageId="28" lastPageNumber="163" pageId="27" pageNumber="162">
Color pattern variation statistics for entire sample of juvenile and adults are (median and range): HeadMid 1, 02; HeadPsag 0, 02; DorsNap 1, 02; DorsPsag 1, 02; IngSpt 2, 02; HndlBr 2, 02; DlatSt 2, 12; Loreal 2, 02; LatTrnk 2, 02;
<collectingRegion id="49E81CA3CC4BFFEDFBED5CF6FBF66008" box="[1094,1146,252,276]" country="Myanmar" name="Chin" pageId="28" pageNumber="163">Chin</collectingRegion>
2, 02; Chest 1, 02. In summary,
<taxonomicName id="4C2CA9C2CC4BFFEDFE655D15FDF8602A" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[462,628,286,310]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFE655D15FDF8602A" box="[462,628,286,310]" italics="true" pageId="28" pageNumber="163">K. interlineatus</emphasis>
</taxonomicName>
usually has a faded to distinct middorsal stripe and no parasagittal stripes on head, distinct nape stripe in about half of the individuals continuing into bilateral (parasagittal) stripes, inguinal spot almost always present and usually as ocelli, dorsolateral narrow white stripe rarely absent or indistinct, hindlimbs almost always distinctly barred, loreal area and lateral trunk usually uniformly dark, chin and throat always dusky and usually with pair of darker longitudinal bars, and chest commonly dusky. Dorsal ground color is variable, ranging from a light beige or tan to medium or rufous brown, occasional individuals are rose to pink; sides from lore to inguinal are usually shades of medium to dark brown.
</paragraph>
<paragraph id="8B93D241CC4BFFEDFF005E2FFE3465B3" blockId="28.[123,1174,219,1726]" pageId="28" pageNumber="163">
<bibRefCitation id="EFBDAFB0CC4BFFEDFF005E2FFEEE6321" author="FEI, L. &amp; C. YE &amp; J. JIANG" box="[171,354,549,573]" pageId="28" pageNumber="163" refId="ref37499" refString="FEI, L., C. YE, AND J. JIANG. 2010. Zhongguo liang qi dong wu cai se tu jian [Colored Atlas of Chinese Amphibians]. Sichuan Publishing Group, Chengdu." type="book" year="2010">Fei et al. (2010)</bibRefCitation>
provided color images of five living individuals whose dorsal coloration ranged from unicolor [
<emphasis id="B9580E53CC4BFFEDFEC75E4CFEF56341" box="[364,377,582,605]" italics="true" pageId="28" pageNumber="163">n</emphasis>
= 1 individual] to strongly patterned with all six of my dorsal pattern characters strongly developed [
<emphasis id="B9580E53CC4BFFEDFE335E63FE29639C" box="[408,421,617,640]" italics="true" pageId="28" pageNumber="163">n</emphasis>
= 5]. The dorsal stripe pattern of Chinese
<taxonomicName id="4C2CA9C2CC4BFFEDFCCF5E63FB85639C" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[868,1033,616,640]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFCCF5E63FB85639C" box="[868,1033,616,640]" italics="true" pageId="28" pageNumber="163">K. interlineatus</emphasis>
</taxonomicName>
has the same layout/arrangement; however, the stripes differ markedly in being broader with strongly scalloped edges and fragmented, especially the DorsPsag ones.
<bibRefCitation id="EFBDAFB0CC4BFFEDFD6A5EA1FC1663DF" author="KARSEN, S. J. &amp; LAU, AND &amp; BOGADEK" box="[705,922,682,707]" pageId="28" pageNumber="163" refId="ref37741" refString="KARSEN, S. J., M. W. LAU, AND BOGADEK. 1986. Hong Kong Amphibians and Reptiles. Urban Council, Hong Kong." type="book" year="1986">Karsen et al. (1986)</bibRefCitation>
pictured a
<collectingCountry id="F33B92D1CC4BFFEDFBBD5EA1FB1963DF" box="[1046,1173,683,707]" name="Hong Kong" pageId="28" pageNumber="163">Hong Kong</collectingCountry>
striped individual that matched those of Fei et al. Additionally Feis photographs showed well-developed white dorsolateral stripes that were strongly speculate in all individuals. The single individual depicted for
<collectingCountry id="F33B92D1CC4BFFEDFEE75F06FE366238" box="[332,442,780,804]" name="Cambodia" pageId="28" pageNumber="163">Cambodia</collectingCountry>
(Thy and Holden 2008) had a well-developed dorsal pattern similar to the western populations, although the white dorsolateral stripe was narrow and appeared interrupted. Ohler and Grosjean (2005) reported distinct pattern difference between “western” and Vietnamese frogs. The former depicted by a
<collectingCountry id="F33B92D1CC4BFFEDFD955F65FDFE629A" box="[574,626,879,902]" name="Laos" pageId="28" pageNumber="163">Laos</collectingCountry>
individual (O&amp;G 2005: figs. 1a,c,e) has straight-edged dorsal stripes, the latter (2005: figs. 1b,d,f) has the scalloped and fragmented stripes of the Chinese individuals. The preceding summary confirms Ohlers and Grosjeans observations of color pattern differences between west (
<collectingCountry id="F33B92D1CC4BFFEDFD865FDAFD0062F4" box="[557,652,976,1000]" name="Thailand" pageId="28" pageNumber="163">Thailand</collectingCountry>
) and east (
<collectingCountry id="F33B92D1CC4BFFEDFCAD5FDAFCEC62F4" box="[774,864,976,1000]" name="Vietnam" pageId="28" pageNumber="163">Vietnam</collectingCountry>
) and, as they noted, hints at the possibility of speciation between eastern and western populations. Chan and collaborators (2011) noted that half of their sample of Malaysian
<taxonomicName id="4C2CA9C2CC4BFFEDFD06581EFCD86537" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[685,852,1043,1067]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFD06581EFCD86537" box="[685,852,1043,1067]" italics="true" pageId="28" pageNumber="163">K. interlineatus</emphasis>
</taxonomicName>
(= their
<taxonomicName id="4C2CA9C2CC4BFFEDFC185819FBB36537" authorityName="Tschudi" authorityYear="1838" box="[947,1087,1043,1067]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="pleurostigma">
<emphasis id="B9580E53CC4BFFEDFC185819FBB36537" box="[947,1087,1043,1067]" italics="true" pageId="28" pageNumber="163">pleurostigma</emphasis>
</taxonomicName>
) lacked inguinal spots on one or both sides and concluded that “populations in Peninsular
<collectingCountry id="F33B92D1CC4BFFEDFC4F583CFBCA6552" box="[996,1094,1078,1102]" name="Malaysia" pageId="28" pageNumber="163">Malaysia</collectingCountry>
are not conspecific” with mainland Asia populations. Their interpretation may be correct, although I suspect that there is no genetic discontinuity between these Malaysian population and those of peninsular
<collectingCountry id="F33B92D1CC4BFFEDFF1D5892FE9265AC" box="[182,286,1176,1200]" name="Myanmar" pageId="28" pageNumber="163">Myanmar</collectingCountry>
and
<collectingCountry id="F33B92D1CC4BFFEDFEF85892FE3E65AC" box="[339,434,1176,1200]" name="Thailand" pageId="28" pageNumber="163">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC4BFFEDFF0058B1FB8D6409" pageId="28" pageNumber="163" type="etymology">
<paragraph id="8B93D241CC4BFFEDFF0058B1FB8D6409" blockId="28.[123,1174,219,1726]" pageId="28" pageNumber="163">
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<emphasis id="B9580E53CC4BFFEDFF0058B1FEB965CD" bold="true" box="[171,309,1211,1235]" pageId="28" pageNumber="163">ETYMOLOGY</emphasis>
</smallCapsWord>
.—
<bibRefCitation id="EFBDAFB0CC4BFFEDFEF558B0FE6B65CF" author="BLYTH, E." box="[350,487,1210,1235]" pageId="28" pageNumber="163" pagination="713 - 723" refId="ref36957" refString="BLYTH, E. 1855. Various notabilia contained in the collections presented by Capt. Berdmore and Mr. Theobald. Proceedings of the Asiatic Society of Bengal 1855 (7): 713 - 723." type="journal article" year="1855">Blyth (1855)</bibRefCitation>
offered no explanation for his name, presumably using
<taxonomicName id="4C2CA9C2CC4BFFEDFB9558B0FF3F65E8" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFB9558B0FF3F65E8" italics="true" pageId="28" pageNumber="163">interlineatus</emphasis>
</taxonomicName>
to note the longitudinal lines on the dorsum, hence a derivation from the Latin
<emphasis id="B9580E53CC4BFFEDFC5C58D7FBA565E8" box="[1015,1065,1245,1268]" italics="true" pageId="28" pageNumber="163">inter</emphasis>
, between, and
<emphasis id="B9580E53CC4BFFEDFF0258F7FF716409" box="[169,253,1277,1301]" italics="true" pageId="28" pageNumber="163">lineatus</emphasis>
, from Latin
<emphasis id="B9580E53CC4BFFEDFE2D58F7FE376409" box="[390,443,1277,1301]" italics="true" pageId="28" pageNumber="163">lineo</emphasis>
, drawing lines, and
<emphasis id="B9580E53CC4BFFEDFD3D58F7FCC56409" box="[662,841,1277,1301]" italics="true" pageId="28" pageNumber="163">linearis, lineatus</emphasis>
, of a line, linear.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC4BFFECFF005914FB8460CA" lastPageId="29" lastPageNumber="164" pageId="28" pageNumber="163" type="distribution">
<paragraph id="8B93D241CC4BFFEDFF005914FBF8648B" blockId="28.[123,1174,219,1726]" pageId="28" pageNumber="163">
<smallCapsWord id="8D75449DCC4BFFEDFF005914FECF6428" baselines="1328,1327" box="[171,323,1310,1334]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="28" pageNumber="163">
<emphasis id="B9580E53CC4BFFEDFF005914FECF6428" bold="true" box="[171,323,1310,1334]" pageId="28" pageNumber="163">DISTRIBUTION</emphasis>
</smallCapsWord>
.—
<taxonomicName id="4C2CA9C2CC4BFFEDFEC05914FD0E642A" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[363,642,1310,1334]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFEC05914FD0E642A" box="[363,642,1310,1334]" italics="true" pageId="28" pageNumber="163">Kalophrynus interlineatus</emphasis>
</taxonomicName>
as here defined has the broadest geographic range of the
<taxonomicName id="4C2CA9C2CC4BFFEDFF145934FEC7644A" authorityName="Tschudi" authorityYear="1838" box="[191,331,1342,1366]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="pleurostigma">
<emphasis id="B9580E53CC4BFFEDFF145934FEC7644A" box="[191,331,1342,1366]" italics="true" pageId="28" pageNumber="163">pleurostigma</emphasis>
</taxonomicName>
group, extending eastward from northern-most Peninsular
<collectingCountry id="F33B92D1CC4BFFEDFC695935FBA6644A" box="[962,1066,1343,1366]" name="Myanmar" pageId="28" pageNumber="163">Myanmar</collectingCountry>
, eastward through northern
<collectingCountry id="F33B92D1CC4BFFEDFE9E5955FE18646B" box="[309,404,1375,1399]" name="Thailand" pageId="28" pageNumber="163">Thailand</collectingCountry>
,
<collectingCountry id="F33B92D1CC4BFFEDFE0B5955FE58646A" box="[416,468,1375,1398]" name="Laos" pageId="28" pageNumber="163">Laos</collectingCountry>
,
<collectingCountry id="F33B92D1CC4BFFEDFE4A5955FDC3646B" box="[481,591,1375,1399]" name="Cambodia" pageId="28" pageNumber="163">Cambodia</collectingCountry>
, and
<collectingCountry id="F33B92D1CC4BFFEDFD225955FD6F646B" box="[649,739,1375,1399]" name="Vietnam" pageId="28" pageNumber="163">Vietnam</collectingCountry>
, southward through Peninsula
<collectingCountry id="F33B92D1CC4BFFEDFB865955FB03646B" box="[1069,1167,1375,1399]" name="Malaysia" pageId="28" pageNumber="163">Malaysia</collectingCountry>
, and also in southern
<collectingCountry id="F33B92D1CC4BFFEDFEF1598AFE166484" box="[346,410,1408,1432]" name="China" pageId="28" pageNumber="163">China</collectingCountry>
from southeastern
<collectingRegion id="49E81CA3CC4BFFEDFDC2598AFD36648B" box="[617,698,1408,1431]" country="China" name="Yunnan" pageId="28" pageNumber="163">Yunnan</collectingRegion>
to
<collectingCountry id="F33B92D1CC4BFFEDFD75598AFCD76484" box="[734,859,1408,1432]" name="Hong Kong" pageId="28" pageNumber="163">Hong Kong</collectingCountry>
and adjacent Guangdong.
</paragraph>
<paragraph id="8B93D241CC4BFFECFF0059AAFC49600F" blockId="28.[123,1174,219,1726]" lastBlockId="29.[123,1174,219,1118]" lastPageId="29" lastPageNumber="164" pageId="28" pageNumber="163">
<bibRefCitation id="EFBDAFB0CC4BFFEDFF0059AAFED164A4" author="BOURRET, R." box="[171,349,1440,1464]" pageId="28" pageNumber="163" pagination="1 - 547" refId="ref37093" refString="BOURRET, R. 1942. Les Batraciens de l'Indochine. Memoires de l'Institut Oceanographique de l'Indochine 6: 1 - 547." type="journal article" year="1942">Bourrets (1942)</bibRefCitation>
concept of
<taxonomicName id="4C2CA9C2CC4BFFEDFE7459ABFD0A64A4" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[479,646,1440,1464]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFE7459ABFD0A64A4" box="[479,646,1440,1464]" italics="true" pageId="28" pageNumber="163">K. interlineatus</emphasis>
</taxonomicName>
was as a subspecies with a northern distribution from
<collectingCountry id="F33B92D1CC4BFFEDFF1D59CBFE9264C5" box="[182,286,1473,1497]" name="Myanmar" pageId="28" pageNumber="163">Myanmar</collectingCountry>
through northern
<collectingCountry id="F33B92D1CC4BFFEDFE4859CBFDCE64C5" box="[483,578,1473,1497]" name="Thailand" pageId="28" pageNumber="163">Thailand</collectingCountry>
to
<collectingCountry id="F33B92D1CC4BFFEDFDCD59CBFD4C64C5" box="[614,704,1473,1497]" name="Vietnam" pageId="28" pageNumber="163">Vietnam</collectingCountry>
and adjacent
<collectingCountry id="F33B92D1CC4BFFEDFCFC59CBFC1B64C5" box="[855,919,1473,1497]" name="China" pageId="28" pageNumber="163">China</collectingCountry>
to
<collectingCountry id="F33B92D1CC4BFFEDFC1059CBFBB564C5" box="[955,1081,1473,1497]" name="Hong Kong" pageId="28" pageNumber="163">Hong Kong</collectingCountry>
. He had only a single male specimen (
<quantity id="4CD47FA4CC4BFFEDFE1159E8FD8864E5" box="[442,516,1506,1530]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="28" pageNumber="163" unit="mm" value="38.0">38 mm</quantity>
<collectionCode id="ED3D4A84CC4BFFEDFDA759E8FDB264E6" box="[524,574,1506,1530]" pageId="28" pageNumber="163">SVL</collectionCode>
) of questionable locality (probably
<collectingCountry id="F33B92D1CC4BFFEDFC1559EBFB8564E5" box="[958,1033,1505,1529]" name="Vietnam" pageId="28" pageNumber="163">Tonkin</collectingCountry>
,
<collectingCountry id="F33B92D1CC4BFFEDFBBC59EBFBFD64E5" box="[1047,1137,1505,1529]" name="Vietnam" pageId="28" pageNumber="163">Vietnam</collectingCountry>
) in Indochina. He listed four Burmese localities: Bhamo, Teinzo, Palon, Toungo (credited to Oates [Toungo] and Fea [diverse localities]). The first two records are potential localities for
<taxonomicName id="4C2CA9C2CC4BFFEDFBBE5A2EFBE46727" authorityName="Zug" authorityYear="2015" box="[1045,1128,1572,1595]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="anya">
<emphasis id="B9580E53CC4BFFEDFBBE5A2EFBE46727" box="[1045,1128,1572,1595]" italics="true" pageId="28" pageNumber="163">K. anya</emphasis>
</taxonomicName>
, the latter two are potential localities for
<taxonomicName id="4C2CA9C2CC4BFFEDFDA25A4EFD3D6747" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[521,689,1603,1627]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFDA25A4EFD3D6747" box="[521,689,1603,1627]" italics="true" pageId="28" pageNumber="163">K. interlineatus</emphasis>
</taxonomicName>
. Toungo (= Taungo,
<geoCoordinate id="EE18B486CC4BFFEDFC3C5A4EFC7F6747" box="[919,1011,1603,1628]" degrees="20" direction="north" minutes="56" orientation="latitude" pageId="28" pageNumber="163" precision="925" value="20.933332">20°56´N</geoCoordinate>
<geoCoordinate id="EE18B486CC4BFFEDFC575A49FBD96747" box="[1020,1109,1603,1628]" degrees="95" direction="east" minutes="24" orientation="longitude" pageId="28" pageNumber="163" precision="925" value="95.4">95°24´E</geoCoordinate>
) is in the upper Sittaung River valley.
<collectionCode id="ED3D4A84CC4BFFEDFE735A6FFDA56760" box="[472,553,1637,1660]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="163">MNHN</collectionCode>
1893.492 is identified as
<taxonomicName id="4C2CA9C2CC4BFFEDFCE95A6EFBD66760" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[834,1114,1636,1660]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="28" pageNumber="163" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4BFFEDFCE95A6EFBD66760" box="[834,1114,1636,1660]" italics="true" pageId="28" pageNumber="163">Kalophrynus interlineatus</emphasis>
</taxonomicName>
from Palon; I did not examine this specimen to confirm its presence or identification. Palon (
<geoCoordinate id="EE18B486CC4BFFEDFBEC5A8FFB196781" box="[1095,1173,1669,1693]" degrees="7" direction="north" minutes="41" orientation="latitude" pageId="28" pageNumber="163" precision="925" value="7.6833334">7°41´N</geoCoordinate>
<geoCoordinate id="EE18B486CC4BFFEDFFD05AACFF5867A1" box="[123,212,1701,1726]" degrees="97" direction="east" minutes="31" orientation="longitude" pageId="28" pageNumber="163" precision="925" value="97.51667">97°31´E</geoCoordinate>
), Kayin State, is identified as a Pegu locality in the British natural history museum specimen register but represented by only one gecko and one
<taxonomicName id="4C2CA9C2CC4AFFECFD5F5CD1FC4461EF" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[756,968,219,243]" class="Amphibia" family="Microhylidae" genus="Micryletta" kingdom="Animalia" order="Anura" pageId="29" pageNumber="164" phylum="Chordata" rank="species" species="inornata">
<emphasis id="B9580E53CC4AFFECFD5F5CD1FC4461EF" box="[756,968,219,243]" italics="true" pageId="29" pageNumber="164">Micryletta inornata</emphasis>
</taxonomicName>
. This record highlights the broad geographic concept of Pegu by the British in late 19
<superScript id="7C597F09CC4AFFECFCF15CF1FCE56010" attach="left" box="[858,873,251,268]" fontSize="7" pageId="29" pageNumber="164">th</superScript>
century.
</paragraph>
<paragraph id="8B93D241CC4AFFECFF005D16FE5C6048" blockId="29.[123,1174,219,1118]" pageId="29" pageNumber="164">
Thy and Holden (2008) stated that
<taxonomicName id="4C2CA9C2CC4AFFECFD835D17FD5C6028" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[552,720,284,308]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="29" pageNumber="164" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC4AFFECFD835D17FD5C6028" box="[552,720,284,308]" italics="true" pageId="29" pageNumber="164">K. interlineatus</emphasis>
</taxonomicName>
was a common frog and suggested that it occurred throughout
<collectingCountry id="F33B92D1CC4AFFECFEF75D37FE466049" box="[348,458,317,341]" name="Cambodia" pageId="29" pageNumber="164">Cambodia</collectingCountry>
.
</paragraph>
<paragraph id="8B93D241CC4AFFECFF005D57FB8460CA" blockId="29.[123,1174,219,1118]" pageId="29" pageNumber="164">
<smallCapsWord id="8D75449DCC4AFFECFF005D57FE9D6068" baselines="367,367" box="[171,273,349,373]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Natural" pageId="29" pageNumber="164">
<emphasis id="B9580E53CC4AFFECFF005D57FE9D6068" bold="true" box="[171,273,349,373]" pageId="29" pageNumber="164">NATURAL</emphasis>
</smallCapsWord>
<smallCapsWord id="8D75449DCC4AFFECFEBC5D57FEF4606F" baselines="367,366" box="[279,376,349,373]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="History" pageId="29" pageNumber="164">
<emphasis id="B9580E53CC4AFFECFEBC5D57FEF4606F" bold="true" box="[279,376,349,373]" pageId="29" pageNumber="164">HISTORY</emphasis>
</smallCapsWord>
.— In
<collectingCountry id="F33B92D1CC4AFFECFE165D57FDA76069" box="[445,555,349,373]" name="Cambodia" pageId="29" pageNumber="164">Cambodia</collectingCountry>
, Thy and Holden (2008) reported it as a common species living in grassland, scrub forest adjacent to villages, and deciduous forest. The recent Burmese individuals derived from the soil surface in mixed deciduous-evergreen secondary growth forest (Mon) and from evergreen forest abutting a clear-cut (weedy) pipeline (Tanintharyi).
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC4AFFECFF005DD4FBDC6541" pageId="29" pageNumber="164" type="discussion">
<paragraph id="8B93D241CC4AFFECFF005DD4FBE063A4" blockId="29.[123,1174,219,1118]" pageId="29" pageNumber="164">
<smallCapsWord id="8D75449DCC4AFFECFF005DD4FEA460E9" baselines="496,496" box="[171,296,478,502]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="29" pageNumber="164">
<emphasis id="B9580E53CC4AFFECFF005DD4FEA460E9" bold="true" box="[171,296,478,502]" pageId="29" pageNumber="164">COMMENTS</emphasis>
</smallCapsWord>
.—
<collectorName id="26D9B797CC4AFFECFEE45DD4FEF460EA" box="[335,376,478,502]" pageId="29" pageNumber="164">The</collectorName>
<typeStatus id="54976CE3CC4AFFECFE2A5DD5FE2360EB" box="[385,431,479,503]" pageId="29" pageNumber="164">type</typeStatus>
locality “Pegu” is commonly interpreted by biologists as equivalent to the present
<collectingCountry id="F33B92D1CC4AFFECFEBA5DF5FEF5630B" box="[273,377,511,535]" name="Myanmar" pageId="29" pageNumber="164">Myanmar</collectingCountry>
division of
<collectingRegion id="49E81CA3CC4AFFECFE525DF5FDBD630B" box="[505,561,511,535]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Bago</collectingRegion>
(formerly called
<collectingRegion id="49E81CA3CC4AFFECFD405DF5FCAC630B" box="[747,800,511,535]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
).
<collectorName id="26D9B797CC4AFFECFC9C5DF5FC20630B" box="[823,940,511,535]" pageId="29" pageNumber="164">Nineteenth</collectorName>
century
<collectingRegion id="49E81CA3CC4AFFECFBA25DF5FBB2630B" box="[1033,1086,511,535]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
encompassed a much broader area than the present political division.
<location id="8EF3849ACC4AFFECFC855E15FCDB632B" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4AFFECFC855E15FCDB632B" box="[814,855,543,567]" country="Myanmar" name="The" pageId="29" pageNumber="164" stateProvince="Bago">The</location>
older
<collectingRegion id="49E81CA3CC4AFFECFC0B5E2AFC596324" box="[928,981,544,568]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
encompassed the area from the
<collectorName id="26D9B797CC4AFFECFEBF5E4AFEDB6344" box="[276,343,576,600]" pageId="29" pageNumber="164">Arkan</collectorName>
(roughly equivalent to the present state of
<collectingRegion id="49E81CA3CC4AFFECFC9A5E4AFC066344" box="[817,906,576,600]" country="Myanmar" name="Rakhine" pageId="29" pageNumber="164">Rakhine</collectingRegion>
) eastward to the
<collectingMunicipality id="6BF7483BCC4AFFECFBE15E4AFF3A6364" pageId="29" pageNumber="164">Sittang River</collectingMunicipality>
drainage.
<location id="8EF3849ACC4AFFECFE8F5E6AFEDF6364" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4AFFECFE8F5E6AFEDF6364" box="[292,339,608,632]" country="Myanmar" municipality="Sittang River" name="This" pageId="29" pageNumber="164" stateProvince="Rakhine">This</location>
broad
<collectingRegion id="49E81CA3CC4AFFECFE375E6AFE5D6364" box="[412,465,608,632]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
likely encompassed northernmost
<collectorName id="26D9B797CC4AFFECFCE95E6AFC236364" box="[834,943,608,632]" pageId="29" pageNumber="164">Mon State</collectorName>
, hence my placement of the
<typeStatus id="54976CE3CC4AFFECFF655E8BFF706385" box="[206,252,641,665]" pageId="29" pageNumber="164">type</typeStatus>
locality straddling the
<collectingMunicipality id="6BF7483BCC4AFFECFDA45E8BFD0F6385" box="[527,643,641,665]" pageId="29" pageNumber="164">Bago-Mon</collectingMunicipality>
border.
<location id="8EF3849ACC4AFFECFD425E8AFC946384" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4AFFECFD425E8AFC946384" box="[745,792,640,664]" country="Myanmar" municipality="Bago-Mon" name="This" pageId="29" pageNumber="164" stateProvince="Bago">This</location>
broad
<collectingRegion id="49E81CA3CC4AFFECFCDA5E8BFC2A6385" box="[881,934,641,665]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
is emphasized by a
<collectionCode id="ED3D4A84CC4AFFECFFD05EABFF6C63A4" box="[123,224,673,696]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="164" type="Museum">NHMUK</collectionCode>
/
<collectionCode id="ED3D4A84CC4AFFECFF4C5EABFEBB63A4" box="[231,311,673,696]" pageId="29" pageNumber="164">BMNH</collectionCode>
specimen from
<location id="8EF3849ACC4AFFECFE4E5EABFDAE63A5" LSID="urn:lsid:plazi:treatment:03856357CC4DFFECFFD05CD1FBDC6541:8EF3849ACC4AFFECFE4E5EABFDAE63A5" box="[485,546,673,697]" country="Myanmar" name="Palon" pageId="29" pageNumber="164" stateProvince="Kayin">Palon</location>
,
<collectingRegion id="49E81CA3CC4AFFECFD9B5EABFD2663A4" box="[560,682,673,697]" country="Myanmar" name="Kayin" pageId="29" pageNumber="164">Kayin state</collectingRegion>
, that was geographically labeled as
<collectingRegion id="49E81CA3CC4AFFECFB845EABFBE863A5" box="[1071,1124,673,697]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
.
</paragraph>
<paragraph id="8B93D241CC4AFFECFF005ECBFBDC6541" blockId="29.[123,1174,219,1118]" pageId="29" pageNumber="164">
Ohler and Grosjean (2005) reported examining the
<typeStatus id="54976CE3CC4AFFECFD5D5ECBFCDF63C5" box="[758,851,705,729]" pageId="29" pageNumber="164" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2CA9C2CC4AFFECFC285EC8FF3A63E5" authority="Blyth" authorityName="Blyth" authorityYear="1855" class="Amphibia" family="Microhylidae" genus="Engystoma" kingdom="Animalia" order="Anura" pageId="29" pageNumber="164" phylum="Chordata" rank="species" species="interlineatum">
<emphasis id="B9580E53CC4AFFECFC285EC8FB1A63C5" box="[899,1174,705,729]" italics="true" pageId="29" pageNumber="164">Engystoma interlineatum</emphasis>
Blyth
</taxonomicName>
, a specimen (
<collectionCode id="ED3D4A84CC4AFFECFEF25EEBFE0D63E5" box="[345,385,737,761]" pageId="29" pageNumber="164">ZSI</collectionCode>
9853) from Mergui,
<collectingCountry id="F33B92D1CC4AFFECFDC15EE8FD5E63E6" box="[618,722,738,762]" name="Myanmar" pageId="29" pageNumber="164">Myanmar</collectingCountry>
. The British Museum has
<specimenCount id="9D2A19C8CC4AFFECFC5D5EE8FB1963E5" box="[1014,1173,737,761]" count="2" pageId="29" pageNumber="164" type="generic">two specimens</specimenCount>
(
<collectionCode id="ED3D4A84CC4AFFECFF2F5F08FF656205" box="[132,233,770,793]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="164" type="Museum">NHMUK</collectionCode>
/
<collectionCode id="ED3D4A84CC4AFFECFF5B5F08FECC6205" box="[240,320,770,793]" pageId="29" pageNumber="164">BMNH</collectionCode>
68.4.3.128129, recataloged as
<date id="FF92F481CC4AFFECFD005F08FC906205" box="[683,796,770,794]" pageId="29" pageNumber="164" value="1947-02-31">1947.2.31.</date>
2627, an adult female and male, respectively) collected by Theobald in
<collectingRegion id="49E81CA3CC4AFFECFD9E5F29FDE66227" box="[565,618,803,827]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
. I believe that the latter two
<collectingRegion id="49E81CA3CC4AFFECFC685F29FC746227" box="[963,1016,803,827]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
specimens of Theobalds (
<figureCitation id="1317CEC4CC4AFFECFEAA5F48FECC6246" box="[257,320,834,858]" captionStart="FIGURE" captionStartId="27.[539,551,1549,1568]" captionTargetBox="[123,1176,216,1715]" captionTargetId="figure-382@27.[504,1176,216,1531]" captionTargetPageId="27" captionText="FIGURE 7. Images of lectotype of Kalophrynus interlineatus (NHMUK/BMNH 1947. 2.31.26, female) in (A) dorsal and (B) ventral views, and of paralectotype (NHMUK/BMNH 1947. 2.31.27, male) in (C) dorsal view [photographer G.R. Zug]." figureDoi="http://doi.org/10.5281/zenodo.11512256" httpUri="https://zenodo.org/record/11512256/files/figure.png" pageId="29" pageNumber="164">Fig. 7</figureCitation>
) are the individuals on which Blyth based his description of
<taxonomicName id="4C2CA9C2CC4AFFECFC7A5F48FBED6246" authorityName="Blyth" authorityYear="1855" box="[977,1121,834,858]" class="Amphibia" family="Microhylidae" genus="Engystoma" kingdom="Animalia" order="Anura" pageId="29" pageNumber="164" phylum="Chordata" rank="species" species="interlineatum">
<emphasis id="B9580E53CC4AFFECFC7A5F48FBED6246" box="[977,1121,834,858]" italics="true" pageId="29" pageNumber="164">interlineatum</emphasis>
</taxonomicName>
, and they were subsequently sent to London. The precision of
<collectionCode id="ED3D4A84CC4AFFECFD495F69FC866267" box="[738,778,867,891]" pageId="29" pageNumber="164">ZSI</collectionCode>
locality indicates that someone arbitrarily selected a
<taxonomicName id="4C2CA9C2CC4AFFECFE845F89FE356287" authorityName="Tschudi" authorityYear="1838" box="[303,441,899,923]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="29" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B9580E53CC4AFFECFE845F89FE356287" box="[303,441,899,923]" italics="true" pageId="29" pageNumber="164">Kalophrynus</emphasis>
</taxonomicName>
specimen and designated it as a
<collectionCode id="ED3D4A84CC4AFFECFCBB5F89FCB46287" box="[784,824,899,923]" pageId="29" pageNumber="164">ZSI</collectionCode>
<typeStatus id="54976CE3CC4AFFECFC955F89FC176287" box="[830,923,899,923]" pageId="29" pageNumber="164" type="holotype">holotype</typeStatus>
. Mergui was never recognized as part of
<collectingRegion id="49E81CA3CC4AFFECFEEB5FAEFEF962A0" box="[320,373,932,956]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
and is in Tenasserim. Because both
<bibRefCitation id="EFBDAFB0CC4AFFECFD525FA9FC3D62A0" author="BLYTH, E." box="[761,945,931,956]" pageId="29" pageNumber="164" pagination="713 - 723" refId="ref36957" refString="BLYTH, E. 1855. Various notabilia contained in the collections presented by Capt. Berdmore and Mr. Theobald. Proceedings of the Asiatic Society of Bengal 1855 (7): 713 - 723." type="journal article" year="1855">Blyth (1855:720)</bibRefCitation>
and
<bibRefCitation id="EFBDAFB0CC4AFFECFC4F5FA9FF2062C0" author="THEOBALD, W." pageId="29" pageNumber="164" refId="ref38585" refString="THEOBALD, W. 1882. Burma, Its People and Productions; Notes on Fauna, Flora and Minerals of Tenasserim, Pegu and Burma by Rev. F. Mason, D. D., M. R. A. S. Vol. I. Geology, Mineralogy and Zoology. Stephen Austin &amp; Sons, Hertford." type="book" year="1882">Theobald (1882: 192)</bibRefCitation>
specifically noted having
<specimenCount id="9D2A19C8CC4AFFECFE665FCEFDE062C0" box="[461,620,964,988]" count="2" pageId="29" pageNumber="164" type="generic">two specimens</specimenCount>
, I recommend recognizing one of them as a
<typeStatus id="54976CE3CC4AFFECFBF25FCEFF2562E1" pageId="29" pageNumber="164" type="lectotype">lectotype</typeStatus>
and select
<collectionCode id="ED3D4A84CC4AFFECFE835FEFFEF462E0" box="[296,376,997,1020]" pageId="29" pageNumber="164">BMNH</collectionCode>
<date id="FF92F481CC4AFFECFE2A5FEEFE7F62E0" box="[385,499,996,1020]" pageId="29" pageNumber="164" value="1947-02-31">1947.2.31.</date>
26, the female, because this specimen is presently in the best physical condition.
<collectionCode id="ED3D4A84CC4AFFECFEE4580FFE136500" box="[335,415,1029,1052]" pageId="29" pageNumber="164">BMNH</collectionCode>
<date id="FF92F481CC4AFFECFE0D580FFD946500" box="[422,536,1029,1053]" pageId="29" pageNumber="164" value="1947-02-31">1947.2.31.</date>
27 is then a
<typeStatus id="54976CE3CC4AFFECFD36580FFCA16501" box="[669,813,1029,1053]" pageId="29" pageNumber="164" type="paralectotype">paralectotype</typeStatus>
. Because
<collectingRegion id="49E81CA3CC4AFFECFC37580FFC5D6501" box="[924,977,1029,1053]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Pegu</collectingRegion>
is not sufficiently delimited, I recommend a more precise, yet not overly restrictive, type locality:
<collectingRegion id="49E81CA3CC4AFFECFC44582FFBAB6521" box="[1007,1063,1061,1085]" country="Myanmar" name="Bago" pageId="29" pageNumber="164">Bago</collectingRegion>
DivisionMon State border in the lower Sittaung River valley,
<collectingCountry id="F33B92D1CC4AFFECFD1D584CFC926542" box="[694,798,1094,1118]" name="Myanmar" pageId="29" pageNumber="164">Myanmar</collectingCountry>
(~
<geoCoordinate id="EE18B486CC4AFFECFC97584FFC376541" box="[828,955,1093,1117]" degrees="17" direction="north" minutes="35" orientation="latitude" pageId="29" pageNumber="164" precision="15" seconds="24" value="17.59">17°35´24˝N</geoCoordinate>
<geoCoordinate id="EE18B486CC4AFFECFC68584FFBB36541" box="[963,1087,1093,1117]" degrees="96" direction="east" minutes="53" orientation="longitude" pageId="29" pageNumber="164" precision="15" seconds="33" value="96.892494">96°53´33˝E</geoCoordinate>
).
</paragraph>
</subSubSection>
</treatment>
</document>