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<document id="5C4838C985076D69F2E6AD9A1EDF6AEA" ID-DOI="10.5281/zenodo.207722" ID-GBIF-Dataset="72761a61-6909-4725-a05e-24cb71274e7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="207722" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460329843254" checkinUser="plazi" docAuthor="Zhang, Jun, Jiang, Ke, Vogel, Gernot &amp; Rao, Dingqi" docDate="2011" docId="03F62777F24DFF8DFF63FC78FE22DB97" docLanguage="en" docName="zt02982p068.pdf" docOrigin="Zootaxa 2982" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lycodon liuchengchaoi Zhang, Jiang, Vogel &amp; Rao, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="FFCF5F0FF24CFF88FFF4FF8FFF96DC16" masterDocTitle="A new species of the genus Lycodon (Squamata, Colubridae) from Sichuan Province, China" masterLastPageNumber="68" masterPageNumber="59" pageNumber="60" updateTime="1698258478942" updateUser="plazi">
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<mods:title id="681E77BFA3EB1C9A98DC4821967DC24F">A new species of the genus Lycodon (Squamata, Colubridae) from Sichuan Province, China</mods:title>
</mods:titleInfo>
<mods:name id="9D181DEA04F4EAA2D8D87B136E11922B" type="personal">
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<mods:namePart id="9CA022B6F6384E384083DDCC76C94C4C">Zhang, Jun</mods:namePart>
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<mods:namePart id="3AA80B306F02684F152D3B2C150A3FFE">Jiang, Ke</mods:namePart>
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<mods:namePart id="C3EED7DA375F605CE04F6441D91FC586">Vogel, Gernot</mods:namePart>
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<mods:roleTerm id="C3F878209F12B64F97CE7A8EE9EAD0A2">Author</mods:roleTerm>
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<mods:namePart id="2DC4D2B8622C44C586800BA0FED3141B">Rao, Dingqi</mods:namePart>
</mods:name>
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<mods:date id="911FFA1D78241821E094382C643B5394">2011</mods:date>
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<treatment id="03F62777F24DFF8DFF63FC78FE22DB97" ID-DOI="http://doi.org/10.5281/zenodo.6194841" ID-GBIF-Taxon="119404334" ID-Zenodo-Dep="6194841" LSID="urn:lsid:zoobank.org:act:4EA4AEED-C616-418C-94C7-312A1B1EEC67" httpUri="http://treatment.plazi.org/id/03F62777F24DFF8DFF63FC78FE22DB97" lastPageId="5" lastPageNumber="64" pageId="1" pageNumber="60">
<subSubSection id="C345C5EAF24DFF89FF63FC78FE0CD824" pageId="1" pageNumber="60" type="nomenclature">
<paragraph id="8BE09661F24DFF89FF63FC78FD81D807" blockId="1.[151,535,1015,1074]" box="[151,535,1015,1041]" pageId="1" pageNumber="60">
<heading id="D0A8210DF24DFF89FF63FC78FD81D807" bold="true" box="[151,535,1015,1041]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<emphasis id="B92B4A73F24DFF89FF63FC78FD81D807" bold="true" box="[151,535,1015,1041]" pageId="1" pageNumber="60">
<taxonomicName id="4C5FEDE2F24DFF89FF63FC78FE24D807" box="[151,434,1015,1041]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="liuchengchaoi" status="sp. nov.">
<emphasis id="B92B4A73F24DFF89FF63FC78FE24D807" bold="true" box="[151,434,1015,1041]" italics="true" pageId="1" pageNumber="60">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218F708F24DFF89FE4DFC78FD81D807" box="[441,535,1015,1041]" pageId="1" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE09661F24DFF89FF63FB95FE0CD824" blockId="1.[151,535,1015,1074]" box="[151,410,1050,1074]" pageId="1" pageNumber="60">
(
<figureCitation id="13648AE4F24DFF89FF6BFB95FE9DD824" box="[159,267,1050,1074]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1089,1112]" captionStartId-1="4.[151,250,1988,2011]" captionStartId-2="5.[151,250,1434,1457]" captionStartId-3="6.[151,250,1073,1096]" captionStartId-4="6.[151,250,1709,1732]" captionTargetBox-0="[511,1091,223,1048]" captionTargetBox-1="[192,1381,861,1950]" captionTargetBox-2="[194,1390,236,1376]" captionTargetBox-3="[433,1162,210,1048]" captionTargetBox-4="[441,1145,1175,1687]" captionTargetId-0="figure@2.[495,1091,195,1065]" captionTargetId-1="figure@4.[151,1436,831,1967]" captionTargetId-2="figure@5.[151,1436,193,1413]" captionTargetId-3="figure@6.[367,1220,195,1059]" captionTargetId-4="figure@6.[441,1145,1175,1687]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="FIGURE 1. Head of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov., in dorsal (a); lateral (b); ventral (c) views. Photographs by Ke Jiang." captionText-1="FIGURE 2. Dorsal view of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. Photograph by Ke Jiang." captionText-2="FIGURE 3. Ventral view of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. Photograph by Ke Jiang." captionText-3="FIGURE 4. Hemipenis of the holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. (a) and Lycodon fasciatus (b). Drawings by Ke Jiang." captionText-4="FIGURE 5. Map showing the two collection sites (black spots) of Lycodon liuchengchaoi sp. nov. in Sichuan province. Tangjiahe National Nature Reserve (type locality), Qingchuan County (a); Mouping (now Baoxing County) (b)." httpUri-0="https://zenodo.org/record/207723/files/figure.png" httpUri-1="https://zenodo.org/record/207724/files/figure.png" httpUri-2="https://zenodo.org/record/207725/files/figure.png" httpUri-3="https://zenodo.org/record/207726/files/figure.png" httpUri-4="https://zenodo.org/record/207727/files/figure.png" pageId="1" pageNumber="60">Figs. 15</figureCitation>
; Tables 12)
</paragraph>
</subSubSection>
<subSubSection id="C345C5EAF24DFF89FF63FBECFD37D938" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="8BE09661F24DFF89FF63FBECFE87D8D4" blockId="1.[151,1437,1122,2010]" pageId="1" pageNumber="60">
<emphasis id="B92B4A73F24DFF89FF63FBECFE9DD86D" bold="true" box="[151,267,1123,1147]" pageId="1" pageNumber="60">
<typeStatus id="54E428C3F24DFF89FF63FBECFE90D86D" box="[151,262,1123,1147]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED4E0EA4F24DFF89FEEDFBEDFEE6D86C" box="[281,368,1122,1146]" pageId="1" pageNumber="60">CWNU</collectionCode>
867001 (
<figureCitation id="13648AE4F24DFF89FE2DFBEDFDDDD86C" box="[473,587,1122,1146]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1089,1112]" captionStartId-1="4.[151,250,1988,2011]" captionStartId-2="5.[151,250,1434,1457]" captionTargetBox-0="[511,1091,223,1048]" captionTargetBox-1="[192,1381,861,1950]" captionTargetBox-2="[194,1390,236,1376]" captionTargetId-0="figure@2.[495,1091,195,1065]" captionTargetId-1="figure@4.[151,1436,831,1967]" captionTargetId-2="figure@5.[151,1436,193,1413]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Head of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov., in dorsal (a); lateral (b); ventral (c) views. Photographs by Ke Jiang." captionText-1="FIGURE 2. Dorsal view of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. Photograph by Ke Jiang." captionText-2="FIGURE 3. Ventral view of holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. Photograph by Ke Jiang." httpUri-0="https://zenodo.org/record/207723/files/figure.png" httpUri-1="https://zenodo.org/record/207724/files/figure.png" httpUri-2="https://zenodo.org/record/207725/files/figure.png" pageId="1" pageNumber="60">Figs. 13</figureCitation>
), an adult male from Tangjiahe National Nature Reserve (
<geoCoordinate id="EE6BF0A6F24DFF89FAEDFBEDFA01D86C" box="[1305,1431,1122,1146]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="5" value="32.5439">32.5439°N</geoCoordinate>
,
<geoCoordinate id="EE6BF0A6F24DFF89FF63FB09FE89D888" box="[151,287,1158,1182]" direction="east" orientation="longitude" pageId="1" pageNumber="60" precision="5" value="104.8322">104.8322°E</geoCoordinate>
, elevation
<quantity id="4CA73B84F24DFF89FE6EFB09FE7AD888" box="[410,492,1158,1182]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="60" unit="m" value="1360.0">1360m</quantity>
), Qingchuan County, Sichuan Province, P. R.
<collectingCountry id="F348D6F1F24DFF89FC01FB09FBAAD888" box="[1013,1084,1158,1182]" name="China" pageId="1" pageNumber="60">China</collectingCountry>
. Collected by Qixiang Deng in July, 1986.
</paragraph>
<paragraph id="8BE09661F24DFF89FF31FB40FD37D938" blockId="1.[151,1437,1122,2010]" pageId="1" pageNumber="60">
<emphasis id="B92B4A73F24DFF89FF31FB40FED2D8F1" bold="true" box="[197,324,1231,1255]" pageId="1" pageNumber="60">
<typeStatus id="54E428C3F24DFF89FF31FB40FEA9D8F1" box="[197,319,1231,1255]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED4E0EA4F24DFF89FEBEFB41FE34D8F0" box="[330,418,1230,1254]" pageId="1" pageNumber="60">CWNU</collectionCode>
84002, an adult female, collected from an elevation of
<quantity id="4CA73B84F24DFF89FBF7FB41FBC1D8F0" box="[1027,1111,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="60" unit="m" value="1230.0">1230 m</quantity>
in the Tangjiahe, by Qixiang Deng on the same date as
<typeStatus id="54E428C3F24DFF89FE3FFB7DFDA7D91C" box="[459,561,1266,1290]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED4E0EA4F24DFF89FDCAFB7CFD19D91C" LSID="urn:lsid:biocol.org:col:34795" box="[574,655,1267,1290]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="60">FMNH</collectionCode>
15148, Monping, Szechwan (=Baoxing County, Sichuan Province), collected by Herbert Stevens,
<date id="FFE1B0A1F24DFF89FE13FA99FD0BD938" box="[487,669,1302,1327]" pageId="1" pageNumber="60" value="1929-08-28">28 August 1929</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C345C5EAF24DFF89FF31FAB5FCF4DAC8" pageId="1" pageNumber="60" type="diagnosis">
<paragraph id="8BE09661F24DFF89FF31FAB5FF2ADA10" blockId="1.[151,1437,1122,2010]" pageId="1" pageNumber="60">
<emphasis id="B92B4A73F24DFF89FF31FAB5FEA9D945" bold="true" box="[197,319,1338,1363]" pageId="1" pageNumber="60">Diagnosis.</emphasis>
<taxonomicName id="4C5FEDE2F24DFF89FEB2FAB4FDDDD945" box="[326,587,1339,1363]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="liuchengchaoi" status="sp. nov.">
<emphasis id="B92B4A73F24DFF89FEB2FAB4FDDDD945" box="[326,587,1339,1363]" italics="true" pageId="1" pageNumber="60">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218F708F24DFF89FDA5FAB4FD3DD945" box="[593,683,1339,1363]" pageId="1" pageNumber="60" rank="species">
<emphasis id="B92B4A73F24DFF89FDA5FAB4FD3DD945" bold="true" box="[593,683,1339,1363]" pageId="1" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from all other species of
<taxonomicName id="4C5FEDE2F24DFF89FB89FAB4FB4BD945" box="[1149,1245,1339,1363]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="B92B4A73F24DFF89FB89FAB4FB4BD945" box="[1149,1245,1339,1363]" italics="true" pageId="1" pageNumber="60">Lycodon</emphasis>
</taxonomicName>
by the following combination of characters: (1) 17 dorsal scale rows at midbody, several median rows feebly keeled; (2) 78 supralabials, the third and fourth or the third to fifth entering eye; (3) loreal entering orbit but not in contact with internasals; (4) anal divided; (5) more than 40 well-defined yellow rings evenly spaced along the entire length of the black body, and more than 10 yellow rings evenly spaced along the black tail; (6) hemipenis not forked at the tip.
</paragraph>
<paragraph id="8BE09661F24DFF89FF31F99CFCF4DAC8" blockId="1.[151,1437,1122,2010]" pageId="1" pageNumber="60">
In terms of pattern,
<taxonomicName id="4C5FEDE2F24DFF89FE56F99BFDC8DA3D" box="[418,606,1555,1579]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="liuchengchaoi">
<emphasis id="B92B4A73F24DFF89FE56F99BFDC8DA3D" box="[418,606,1555,1579]" italics="true" pageId="1" pageNumber="60">L. liuchengchaoi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C5FEDE2F24DFF89FCD8F99BFC38DA3D" box="[812,942,1555,1579]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24DFF89FCD8F99BFC38DA3D" box="[812,942,1555,1579]" italics="true" pageId="1" pageNumber="60">L. fasciatus</emphasis>
</taxonomicName>
except for the fact that the bands are yellow instead of whitish-brown or cream, but differs by the following traits: seven supralabials in two of the three specimens, the third and fourth entering orbit (in one case the third, fourth and fifth) (eight supralabials in all
<specimenCount id="9D595DE8F24DFF89FAC1F9D5FF44DA80" pageId="1" pageNumber="60" type="generic">72 specimens</specimenCount>
of
<taxonomicName id="4C5FEDE2F24DFF89FF0DF90FFEEDDA80" box="[249,379,1662,1687]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24DFF89FF0DF90FFEEDDA80" box="[249,379,1662,1687]" italics="true" pageId="1" pageNumber="60">L. fasciatus</emphasis>
</taxonomicName>
examined by us except one, which has 9 on one side, with the third, fourth and fifth entering eye); usually 8 infralabials (
<taxonomicName id="4C5FEDE2F24DFF89FE25F92BFDC4DAAD" box="[465,594,1699,1723]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24DFF89FE25F92BFDC4DAAD" box="[465,594,1699,1723]" italics="true" pageId="1" pageNumber="60">L. fasciatus</emphasis>
</taxonomicName>
usually 9); anal divided (anal entire in
<taxonomicName id="4C5FEDE2F24DFF89FBFDF92BFB1CDAAD" box="[1033,1162,1699,1723]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24DFF89FBFDF92BFB1CDAAD" box="[1033,1162,1699,1723]" italics="true" pageId="1" pageNumber="60">L. fasciatus</emphasis>
</taxonomicName>
); hemipenis has no nick at the tip (hemipenis of
<taxonomicName id="4C5FEDE2F24DFF89FE56F947FDB4DAC8" box="[418,546,1734,1759]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24DFF89FE56F947FDB4DAC8" box="[418,546,1734,1759]" italics="true" pageId="1" pageNumber="60">L. fasciatus</emphasis>
</taxonomicName>
has a nick at the tip,
<figureCitation id="13648AE4F24DFF89FCE5F949FCC3DAC8" box="[785,853,1734,1758]" captionStart="FIGURE 4" captionStartId="6.[151,250,1073,1096]" captionTargetBox="[433,1162,210,1048]" captionTargetId="figure@6.[367,1220,195,1059]" captionTargetPageId="6" captionText="FIGURE 4. Hemipenis of the holotype (CWNU 867001) of Lycodon liuchengchaoi sp. nov. (a) and Lycodon fasciatus (b). Drawings by Ke Jiang." httpUri="https://zenodo.org/record/207726/files/figure.png" pageId="1" pageNumber="60">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C345C5EAF24DFF8CFF31F965FE94DCA2" lastPageId="4" lastPageNumber="63" pageId="1" pageNumber="60" type="description">
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<emphasis id="B92B4A73F24DFF89FF31F965FD8FDB15" bold="true" box="[197,537,1770,1795]" pageId="1" pageNumber="60">
Description of the
<typeStatus id="54E428C3F24DFF89FE5FF964FD82DB15" box="[427,532,1771,1795]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male; SVL:
<quantity id="4CA73B84F24DFF89FD09F965FCF7DB14" box="[765,865,1770,1794]" metricMagnitude="-1" metricUnit="m" metricValue="5.95" pageId="1" pageNumber="60" unit="mm" value="595.0">595 mm</quantity>
, TaL:
<quantity id="4CA73B84F24DFF89FC5AF965FB87DB14" box="[942,1041,1770,1794]" metricMagnitude="-1" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="60" unit="mm" value="152.0">152 mm</quantity>
, ratio TaL/TL: 0.203; head length
<quantity id="4CA73B84F24DFF89FF63F881FE9BDB30" box="[151,269,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.435" pageId="1" pageNumber="60" unit="mm" value="14.35">14.35 mm</quantity>
, head width
<quantity id="4CA73B84F24DFF89FE57F881FD9ADB30" box="[419,524,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="60" unit="mm" value="10.1">10.1 mm</quantity>
; head distinct from neck, distinctly flattened; snout projects beyond lower jaw; body subcylindrical; vertebral ridge poorly developed; maxillary teeth 8, last group with 2 teeth and enlarged. Rostral large, triangular, about 1.5 times as broad as deep; its portion visible from above measuring a little less than 1/ 4 of its distance from frontal; internasals subtriangular and slightly broader than long; laterally in contact with nasal; prefrontals 1.1 times wider than long, not in contact with orbit but with loreal; prefrontal suture obviously longer than internasal suture; frontal subtriangular, long and wide slightly equal; parietals about 1.65 times longer than broad, parietal suture obviously longer than frontal suture; nasal divided, nostril in the anterior nasal; loreals rectangular, about 1.9 times longer than broad, contacting orbit; preocular 1/1, upward; distance between eye and snout tip 1.9 times eye diameter; pupil elliptical; postoculars 2; temporals 2+2/2+2; supralabials 7/7, first supralabial in contact with rostral and nasal, second supralabial in contact with nasal and loreal, third and fourth entering eye, fifth in contact with postocular and anterior temporal, sixth in contact with lower anterior and lower posterior temporal, seventh in contact with posterior temporal; infralabial 8/8, first pair in contact along midline, first five infralabials in contact with chin shield; posterior chin shield slightly shorter and narrower than anterior, in broad contact. Dorsal scale rows 17-17-15; dorsal scales small and feebly keeled in median several rows close with the ridge (invisible on neck); ventrals 204, angulated weakly; anal divided; subcaudals 68/68+1; The dorsal scale rows reduction formula is:
</paragraph>
<caption id="DF20C6E9F24EFF8AFF63FBCEFE0DD861" httpUri="https://zenodo.org/record/207723/files/figure.png" pageId="2" pageNumber="61" targetBox="[511,1091,223,1048]" targetPageId="2">
<paragraph id="8BE09661F24EFF8AFF63FBCEFE0DD861" blockId="2.[151,1436,1089,1143]" pageId="2" pageNumber="61">
<emphasis id="B92B4A73F24EFF8AFF63FBCEFE85D841" bold="true" box="[151,275,1089,1112]" pageId="2" pageNumber="61">FIGURE 1.</emphasis>
Head of holotype (CWNU867001) of
<taxonomicName id="4C5FEDE2F24EFF8AFD6AFBCDFC1CD84E" box="[670,906,1090,1112]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="liuchengchaoi" status="sp. nov.">
<emphasis id="B92B4A73F24EFF8AFD6AFBCDFC1CD84E" box="[670,906,1090,1112]" italics="true" pageId="2" pageNumber="61">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218F708F24EFF8AFC66FBCDFC74D841" box="[914,994,1090,1111]" pageId="2" pageNumber="61" rank="species">
<emphasis id="B92B4A73F24EFF8AFC66FBCDFC74D841" bold="true" box="[914,994,1090,1111]" pageId="2" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
, in dorsal (a); lateral (b); ventral (c) views. Photographs by Ke Jiang.
</paragraph>
</caption>
<paragraph id="8BE09661F24EFF8AFF3BF9A0FEFEDA51" blockId="2.[151,623,1583,1715]" box="[207,360,1583,1607]" pageId="2" pageNumber="61">3+4⊣3 (145)</paragraph>
<paragraph id="8BE09661F24EFF8AFF63F9DCFEF7DA99" blockId="2.[151,623,1583,1715]" pageId="2" pageNumber="61">17 ———————— 15 3+4⊣3 (147)</paragraph>
<paragraph id="8BE09661F24EFF8AFF63F914FDF9DAA5" blockId="2.[151,623,1583,1715]" box="[151,623,1691,1715]" pageId="2" pageNumber="61">The scale reduction formula on the tail as:</paragraph>
<paragraph id="8BE09661F24EFF8AFF4EF96CFD97DAED" blockId="2.[151,557,1763,1859]" box="[186,513,1763,1787]" pageId="2" pageNumber="61">3+4⊣3 (18) 2+3⊣2 (30)</paragraph>
<paragraph id="8BE09661F24EFF8AFF63F888FD9FDB55" blockId="2.[151,557,1763,1859]" pageId="2" pageNumber="61">8 —————— 6 —————— 4 3+4⊣3 (17) 2+3⊣2 (31)</paragraph>
<paragraph id="8BE09661F24EFF8AFF31F8FCFCAEDBC5" blockId="2.[151,1436,1907,2004]" pageId="2" pageNumber="61">
Hemipenis is fairly short and thin, extends to the eighth subcaudal, and the origin of
<emphasis id="B92B4A73F24EFF8AFB82F8FAFA0ADB9A" box="[1142,1436,1909,1932]" italics="true" pageId="2" pageNumber="61">m. retractor penis magnus</emphasis>
to the twenty-fifth subcaudal. Hemipenis spinose, with small spines all over the organ, spines enlarged at the base of organ. The simple sulcus is prominent. No nick at the tip.
</paragraph>
<paragraph id="8BE09661F24FFF8BFF31FF18FF4DDD72" blockId="3.[151,1436,151,428]" pageId="3" pageNumber="62">
<emphasis id="B92B4A73F24FFF8BFF31FF18FED0DCA6" bold="true" box="[197,326,151,176]" pageId="3" pageNumber="62">Coloration</emphasis>
(in formalin): Crown largely black, except for a yellow crossband across the occipital region, faint or incomplete along the midline; underside of head pale yellow, except for more or less black on mental, anterior chin-shield, and first five infralabials. Dorsal surface of body and tail black with 40 jagged-edged, brownish-yellow rings on the body, and 10 on the tail. All the rings are 2 to 3 dorsal scales wide, except for the first which is only a scale-length in width. The yellow rings extend across the belly as wide as 2 to 4 ventral plates, other ventrals are black.
</paragraph>
<paragraph id="8BE09661F24FFF8BFF31FEE0FF67DDBA" blockId="3.[151,1436,151,428]" pageId="3" pageNumber="62">
<emphasis id="B92B4A73F24FFF8BFF31FEE0FEAFDD9E" bold="true" box="[197,313,367,392]" pageId="3" pageNumber="62">Variation.</emphasis>
The
<typeStatus id="54E428C3F24FFF8BFE8EFEE0FE4ADD91" box="[378,476,367,391]" pageId="3" pageNumber="62" type="holotype">holotype</typeStatus>
and the
<typeStatus id="54E428C3F24FFF8BFDB5FEFFFD3ADD9E" box="[577,684,368,392]" pageId="3" pageNumber="62" type="paratype">paratypes</typeStatus>
are similar in general aspects. The differences are summarised in Table 1.
</paragraph>
<paragraph id="8BE09661F24FFF8BFF63FE58FB15DEAE" blockId="3.[151,1286,470,1192]" pageId="3" pageNumber="62">
<table id="F95F64C1F24F0077FF63FE59FA90DEAE" box="[151,1286,470,696]" gridcols="4" gridrows="3" pageId="3" pageNumber="62">
<tr id="356F9423F24F0077FF63FE59FA90DDFB" box="[151,1286,470,493]" gridrow="0" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FE59FEE0DDFB" box="[151,374,470,493]" gridcol="0" gridrow="0" pageId="3" pageNumber="62">
<emphasis id="B92B4A73F24FFF8BFF63FE58FE92DDFA" bold="true" box="[151,260,471,492]" pageId="3" pageNumber="62">TABLE 1.</emphasis>
Characters
</th>
<th id="76BEFD5FF24F0077FE89FE59FE66DDFB" box="[381,496,470,493]" gridcol="1" gridrow="0" pageId="3" pageNumber="62">of the three</th>
<th id="76BEFD5FF24F0077FE03FE59FCB5DDFB" box="[503,803,470,493]" gridcol="2" gridrow="0" pageId="3" pageNumber="62">
known specimens of
<taxonomicName id="4C5FEDE2F24FFF8BFD39FE58FCB5DDFB" box="[717,803,471,493]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="genus">
<emphasis id="B92B4A73F24FFF8BFD39FE58FCB5DDFB" box="[717,803,471,493]" italics="true" pageId="3" pageNumber="62">Lycodon</emphasis>
</taxonomicName>
</th>
<th id="76BEFD5FF24F0077FCDEFE59FA90DDFB" box="[810,1286,470,493]" gridcol="3" gridrow="0" pageId="3" pageNumber="62">
<emphasis id="B92B4A73F24FFF8BFCDEFE58FC56DDFB" box="[810,960,471,493]" italics="true" pageId="3" pageNumber="62">
<taxonomicName id="4C5FEDE2F24FFF8BFCDEFE58FC2BDDFB" box="[810,957,471,493]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="liuchengchaoi">liuchengchaoi</taxonomicName>
.
</emphasis>
Dimensions in mm.
</th>
</tr>
<tr id="356F9423F24F0077FF63FD87FA90DE2A" box="[151,1286,520,572]" gridrow="1" pageId="3" pageNumber="62" rowspan-1="1">
<th id="76BEFD5FF24F0077FF63FD87FEE0DE2A" box="[151,374,520,572]" gridcol="0" gridrow="1" pageId="3" pageNumber="62">Characters</th>
<td id="76BEFD5FF24F0077FE03FD87FCB5DE2A" box="[503,803,520,572]" gridcol="2" gridrow="1" pageId="3" pageNumber="62">CWNU867001 holotype</td>
<td id="76BEFD5FF24F0077FCDEFD87FA90DE2A" box="[810,1286,520,572]" gridcol="3" gridrow="1" pageId="3" pageNumber="62">CWNU84002 FMNH 15148 paratype paratype</td>
</tr>
<tr id="356F9423F24F0077FF63FDDFFA90DEAE" box="[151,1286,592,696]" gridrow="2" pageId="3" pageNumber="62" rowspan-1="1">
<th id="76BEFD5FF24F0077FF63FDDFFEE0DEAE" box="[151,374,592,696]" gridcol="0" gridrow="2" pageId="3" pageNumber="62">Sex Snout-vent length Tail length</th>
<td id="76BEFD5FF24F0077FE03FDDFFCB5DEAE" box="[503,803,592,696]" gridcol="2" gridrow="2" pageId="3" pageNumber="62">adult male 595 152</td>
<td id="76BEFD5FF24F0077FCDEFDDFFA90DEAE" box="[810,1286,592,696]" gridcol="3" gridrow="2" pageId="3" pageNumber="62">adult female adult male 481 676 134?</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE09661F24FFF8BFF54FD44FB15D8BE" blockId="3.[151,1286,470,1192]" pageId="3" pageNumber="62">
<table id="F95F64C1F24F0077FF63FD44FA90D8BE" box="[151,1286,715,1192]" gridcols="4" gridrows="5" pageId="3" pageNumber="62">
<tr id="356F9423F24F0077FF63FD44FA90DEF7" box="[151,1286,715,737]" gridrow="0" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FD44FE24DEF7" box="[151,434,715,737]" gridcol="0" gridrow="0" pageId="3" pageNumber="62">Relative tail length</th>
<th id="76BEFD5FF24F0077FD95FD44FD0CDEF7" box="[609,666,715,737]" gridcol="1" gridrow="0" pageId="3" pageNumber="62">0.203</th>
<th id="76BEFD5FF24F0077FC9EFD44FC56DEF7" box="[874,960,715,737]" gridcol="2" gridrow="0" pageId="3" pageNumber="62">0.218</th>
<th id="76BEFD5FF24F0077FB8CFD44FA90DEF7" box="[1144,1286,715,737]" gridcol="3" gridrow="0" pageId="3" pageNumber="62">?</th>
</tr>
<tr id="356F9423F24F0077FF63FD7AFA90DF22" box="[151,1286,757,820]" gridrow="1" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FD7AFE24DF22" box="[151,434,757,820]" gridcol="0" gridrow="1" pageId="3" pageNumber="62">Anterior dorsal scale rows Midbody dorsal scale rows</th>
<td id="76BEFD5FF24F0077FD95FD7AFD0CDF22" box="[609,666,757,820]" gridcol="1" gridrow="1" pageId="3" pageNumber="62">17 17</td>
<td id="76BEFD5FF24F0077FC9EFD7AFC56DF22" box="[874,960,757,820]" gridcol="2" gridrow="1" pageId="3" pageNumber="62">17 17</td>
<td id="76BEFD5FF24F0077FB8CFD7AFA90DF22" box="[1144,1286,757,820]" gridcol="3" gridrow="1" pageId="3" pageNumber="62">17 17</td>
</tr>
<tr id="356F9423F24F0077FF63FCC7FA90DFCF" box="[151,1286,840,985]" gridrow="2" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FCC7FE24DFCF" box="[151,434,840,985]" gridcol="0" gridrow="2" pageId="3" pageNumber="62">Ventrals Subcaudals Loreal enters eye Dorsal bands</th>
<td id="76BEFD5FF24F0077FD95FCC7FD0CDFCF" box="[609,666,840,985]" gridcol="1" gridrow="2" pageId="3" pageNumber="62">204 68 yes 40</td>
<td id="76BEFD5FF24F0077FC9EFCC7FC56DFCF" box="[874,960,840,985]" gridcol="2" gridrow="2" pageId="3" pageNumber="62">206 77 yes 45</td>
<td id="76BEFD5FF24F0077FB8CFCC7FA90DFCF" box="[1144,1286,840,985]" gridcol="3" gridrow="2" pageId="3" pageNumber="62">202 69+ yes 45</td>
</tr>
<tr id="356F9423F24F0077FF63FC62FA90D83A" box="[151,1286,1005,1068]" gridrow="3" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FC62FE24D83A" box="[151,434,1005,1068]" gridcol="0" gridrow="3" pageId="3" pageNumber="62">Tail bands Upper labials</th>
<td id="76BEFD5FF24F0077FD95FC62FD0CD83A" box="[609,666,1005,1068]" gridcol="1" gridrow="3" pageId="3" pageNumber="62">10 7</td>
<td id="76BEFD5FF24F0077FC9EFC62FC56D83A" box="[874,960,1005,1068]" gridcol="2" gridrow="3" pageId="3" pageNumber="62">11 7</td>
<td id="76BEFD5FF24F0077FB8CFC62FA90D83A" box="[1144,1286,1005,1068]" gridcol="3" gridrow="3" pageId="3" pageNumber="62">15 8</td>
</tr>
<tr id="356F9423F24F0077FF63FBCFFA90D8BE" box="[151,1286,1088,1192]" gridrow="4" pageId="3" pageNumber="62">
<th id="76BEFD5FF24F0077FF63FBCFFE24D8BE" box="[151,434,1088,1192]" gridcol="0" gridrow="4" pageId="3" pageNumber="62">Temporals Infralabials Maxillary teeth</th>
<td id="76BEFD5FF24F0077FD95FBCFFD0CD8BE" box="[609,666,1088,1192]" gridcol="1" gridrow="4" pageId="3" pageNumber="62">2+2 8 / 8 8</td>
<td id="76BEFD5FF24F0077FC9EFBCFFC56D8BE" box="[874,960,1088,1192]" gridcol="2" gridrow="4" pageId="3" pageNumber="62">2+2/1+2 7 / 8 9</td>
<td id="76BEFD5FF24F0077FB8CFBCFFA90D8BE" box="[1144,1286,1088,1192]" gridcol="3" gridrow="4" pageId="3" pageNumber="62">1+2 8 / 9?</td>
</tr>
</table>
</paragraph>
<caption id="DF20C6E9F24FFF8BFF63FB62FE3FD968" pageId="3" pageNumber="62">
<paragraph id="8BE09661F24FFF8BFF63FB62FE3FD968" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
<emphasis id="B92B4A73F24FFF8BFF63FB62FE90D914" bold="true" box="[151,262,1261,1282]" pageId="3" pageNumber="62">TABLE 2.</emphasis>
A Comparison of the main morphological characters in species of the
<taxonomicName id="4C5FEDE2F24FFF8BFC13FB62FB36D915" box="[999,1184,1261,1283]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24FFF8BFC13FB62FB36D915" box="[999,1184,1261,1283]" italics="true" pageId="3" pageNumber="62">Lycodon fasciatus</emphasis>
</taxonomicName>
group. Values in square brackets are exceptional (observed frequency of &lt;20%). Abbreviations: MSR midbody scale rows; Lo touch loreal shield touches the eye; IL—Infrralabials; SL—Supralabials. + there are still several lineages combined under this name. ++ one specimen is an outlier with 43 bands, the mean value is 29.9 S = 4.9. * Data from Pauwels
<emphasis id="B92B4A73F24FFF8BFBF5FAC5FBB8D949" box="[1025,1070,1353,1375]" italics="true" pageId="3" pageNumber="62">et al</emphasis>
. (2005). ** In Pauwels
<emphasis id="B92B4A73F24FFF8BFAEBFAC5FADAD949" box="[1311,1356,1353,1375]" italics="true" pageId="3" pageNumber="62">et al</emphasis>
. (2005) these values are erroneous.
</paragraph>
</caption>
<paragraph id="8BE09661F24FFF8BFF6BFA15FB13D9B9" blockId="3.[151,1436,1261,2010]" box="[159,1157,1433,1455]" pageId="3" pageNumber="62">Taxon Maximum total Relative tail length Ventrals</paragraph>
<paragraph id="8BE09661F24FFF8BFE4AFA37FDAAD9D8" blockId="3.[151,1436,1261,2010]" box="[446,572,1464,1486]" pageId="3" pageNumber="62">length [mm]</paragraph>
<paragraph id="8BE09661F24FFF8BFE4AFA6EFAF3DA29" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
Males Females Males Females Males Females
<emphasis id="B92B4A73F24FFF8BFF6BF985FEFFDA36" bold="true" box="[159,361,1546,1568]" pageId="3" pageNumber="62">
<taxonomicName id="4C5FEDE2F24FFF8BFF6BF985FECDDA36" box="[159,347,1546,1568]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B92B4A73F24FFF8BFF6BF985FECDDA36" bold="true" box="[159,347,1546,1568]" italics="true" pageId="3" pageNumber="62">Lycodon fasciatus</emphasis>
</taxonomicName>
+
</emphasis>
894 679 0.1980.225 0.1900.219 190211 182219 (n = 67) (? = 0.208) (? = 0.204) (? = 202,3) (? = 205.3)
</paragraph>
<paragraph id="8BE09661F24FFF8BFD58F9C7FB34DAB0" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
(
<emphasis id="B92B4A73F24FFF8BFD40F9C6FD28DA48" box="[692,702,1609,1630]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 0.007) (
<emphasis id="B92B4A73F24FFF8BFC83F9C6FC17DA48" box="[887,897,1609,1630]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 0.009) (
<emphasis id="B92B4A73F24FFF8BFBCEF9C6FBD2DA48" box="[1082,1092,1609,1630]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 5.3) (
<emphasis id="B92B4A73F24FFF8BFB09F9C6FA91DA48" box="[1277,1287,1609,1630]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 8.6)
<taxonomicName id="4C5FEDE2F24FFF8BFF6BF9FEFEC8DA91" box="[159,350,1649,1671]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="gongshan">
<emphasis id="B92B4A73F24FFF8BFF6BF9FEFEC8DA91" box="[159,350,1649,1671]" italics="true" pageId="3" pageNumber="62">Lycodon gongshan</emphasis>
</taxonomicName>
963 762 0.2310.232 0.215 210216 215 (n = 4) (? = 0.232) (? = 212.7)
</paragraph>
<paragraph id="8BE09661F24FFF8BFD58F920FB11DAD3" blockId="3.[151,1436,1261,2010]" box="[684,1159,1711,1733]" pageId="3" pageNumber="62">
(
<emphasis id="B92B4A73F24FFF8BFD40F93FFD28DAD3" box="[692,702,1712,1733]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 0.0002) (
<emphasis id="B92B4A73F24FFF8BFBCEF93FFBD2DAD3" box="[1082,1092,1712,1733]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 3.1)
</paragraph>
<paragraph id="8BE09661F24FFF8BFF6BF957FAC4DB1B" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
<taxonomicName id="4C5FEDE2F24FFF8BFF6BF957FEC2DAF8" box="[159,340,1752,1774]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="synaptor">
<emphasis id="B92B4A73F24FFF8BFF6BF957FEC2DAF8" box="[159,340,1752,1774]" italics="true" pageId="3" pageNumber="62">Lycodon synaptor</emphasis>
</taxonomicName>
unknown 487 unknown 0.1890.192 unknown 201203 (n = 2) (? = 0.190) (? = 202)
</paragraph>
<paragraph id="8BE09661F24FFF8BFC9BF899FF72DB62" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
(
<emphasis id="B92B4A73F24FFF8BFC83F899FC17DB3D" box="[887,897,1814,1835]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 0.002) (
<emphasis id="B92B4A73F24FFF8BFB09F899FA91DB3D" box="[1277,1287,1814,1835]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 1.4)
<taxonomicName id="4C5FEDE2F24FFF8BFF6BF8B0FE1DDB43" box="[159,395,1855,1877]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="liuchengchaoi">
<emphasis id="B92B4A73F24FFF8BFF6BF8B0FE1DDB43" box="[159,395,1855,1877]" italics="true" pageId="3" pageNumber="62">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
676 615 0.203 0.218 202204 206 (n = 3)
</paragraph>
<paragraph id="8BE09661F24FFF8BFF6BF809FB34DBAD" blockId="3.[151,1436,1261,2010]" pageId="3" pageNumber="62">
<taxonomicName id="4C5FEDE2F24FFF8BFF6BF809FE37DB8A" box="[159,417,1926,1948]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="62" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="B92B4A73F24FFF8BFF6BF809FE37DB8A" bold="true" box="[159,417,1926,1948]" italics="true" pageId="3" pageNumber="62">Lycodon cardamomensis</emphasis>
</taxonomicName>
* 893 545 0.2030.206** 0.200** 215222 223 (n = 4) (? = 0.205) (? = 218.5)
</paragraph>
<paragraph id="8BE09661F24FFF8BFD58F84BFB11DBCC" blockId="3.[151,1436,1261,2010]" box="[684,1159,1988,2010]" pageId="3" pageNumber="62">
(
<emphasis id="B92B4A73F24FFF8BFD40F84AFD28DBCC" box="[692,702,1989,2010]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 0.002) (
<emphasis id="B92B4A73F24FFF8BFBCEF84AFBD2DBCC" box="[1082,1092,1989,2010]" italics="true" pageId="3" pageNumber="62">s</emphasis>
= 4.9)
</paragraph>
<paragraph id="8BE09661F248FF8CFF63FF11FE94DCA2" blockId="4.[151,258,158,180]" box="[151,258,158,180]" pageId="4" pageNumber="63">continued.</paragraph>
</subSubSection>
<paragraph id="8BE09661F248FF8CFF6BFF5FFAFCDEC5" pageId="4" pageNumber="63">
<table id="F95F64C1F2480077FF63FF40FA17DF07" box="[151,1409,207,785]" gridcols="8" gridrows="6" pageId="4" pageNumber="63">
<tr id="356F9423F2480077FF63FF40FA17DD19" box="[151,1409,207,271]" gridrow="0" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FF40FE4EDD19" box="[151,472,207,271]" gridcol="0" gridrow="0" pageId="4" pageNumber="63">Taxon Subcaudals Males</th>
<th id="76BEFD5FF2480077FE00FF40FDC1DD19" box="[500,599,207,271]" gridcol="1" gridrow="0" pageId="4" pageNumber="63">Females</th>
<th id="76BEFD5FF2480077FD70FF40FD2CDD19" box="[644,698,207,271]" gridcol="2" gridrow="0" pageId="4" pageNumber="63">MSR</th>
<th id="76BEFD5FF2480077FD2EFF40FCB7DD19" box="[730,801,207,271]" gridcol="3" gridrow="0" pageId="4" pageNumber="63">Bands body</th>
<th id="76BEFD5FF2480077FCB9FF40FC1BDD19" box="[845,909,207,271]" gridcol="4" gridrow="0" pageId="4" pageNumber="63">Bands tail</th>
<th id="76BEFD5FF2480077FC42FF40FC78DD19" box="[950,1006,207,271]" gridcol="5" gridrow="0" pageId="4" pageNumber="63">Lo touch</th>
<th id="76BEFD5FF2480077FBFBFF40FB0BDD19" box="[1039,1181,207,271]" gridcol="6" gridrow="0" pageId="4" pageNumber="63">IL SL</th>
<th id="76BEFD5FF2480077FB3CFF40FA17DD19" box="[1224,1409,207,271]" gridcol="7" gridrow="0" pageId="4" pageNumber="63">Anal Colour of shields bands</th>
</tr>
<tr id="356F9423F2480077FF63FEADFA17DD60" box="[151,1409,290,374]" gridrow="1" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FEADFE4EDD60" box="[151,472,290,374]" gridcol="0" gridrow="1" pageId="4" pageNumber="63">
<emphasis id="B92B4A73F248FF8CFF6BFEADFED2DD2E" bold="true" box="[159,324,290,312]" italics="true" pageId="4" pageNumber="63">
<taxonomicName id="4C5FEDE2F248FF8CFF6BFEADFF6EDD2E" box="[159,248,290,312]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="genus">Lycodon</taxonomicName>
fascia-
</emphasis>
7488
<emphasis id="B92B4A73F248FF8CFF6BFECDFF5ADD41" bold="true" box="[159,204,322,343]" pageId="4" pageNumber="63">
<emphasis id="B92B4A73F248FF8CFF6BFECDFF28DD41" bold="true" box="[159,190,322,343]" italics="true" pageId="4" pageNumber="63">tus</emphasis>
+
</emphasis>
(? = 81.2) (n = 67) (
<emphasis id="B92B4A73F248FF8CFE98FEEEFEE0DD60" box="[364,374,353,374]" italics="true" pageId="4" pageNumber="63">s</emphasis>
= 4.3)
</th>
<td id="76BEFD5FF2480077FE00FEADFDC1DD60" box="[500,599,290,374]" gridcol="1" gridrow="1" pageId="4" pageNumber="63">
6588 (? = 79.3) (
<emphasis id="B92B4A73F248FF8CFE08FEEEFD90DD60" box="[508,518,353,374]" italics="true" pageId="4" pageNumber="63">s</emphasis>
= 6.1)
</td>
<td id="76BEFD5FF2480077FD70FEADFD2CDD60" box="[644,698,290,374]" gridcol="2" gridrow="1" pageId="4" pageNumber="63">17</td>
<td id="76BEFD5FF2480077FD2EFEADFCB7DD60" box="[730,801,290,374]" gridcol="3" gridrow="1" pageId="4" pageNumber="63">
1937 [43]
<emphasis id="B92B4A73F248FF8CFCF0FECDFCB7DD41" bold="true" box="[772,801,322,343]" pageId="4" pageNumber="63">++</emphasis>
</td>
<td id="76BEFD5FF2480077FCB9FEADFC1BDD60" box="[845,909,290,374]" gridcol="4" gridrow="1" pageId="4" pageNumber="63">721</td>
<td id="76BEFD5FF2480077FC42FEADFC78DD60" box="[950,1006,290,374]" gridcol="5" gridrow="1" pageId="4" pageNumber="63">1</td>
<td id="76BEFD5FF2480077FBFBFEADFB0BDD60" box="[1039,1181,290,374]" gridcol="6" gridrow="1" pageId="4" pageNumber="63">9 [8,10] 8</td>
<td id="76BEFD5FF2480077FB3CFEADFA17DD60" box="[1224,1409,290,374]" gridcol="7" gridrow="1" pageId="4" pageNumber="63">1 whitish</td>
</tr>
<tr id="356F9423F2480077FF63FE06FA17DDCB" box="[151,1409,393,477]" gridrow="2" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FE06FE4EDDCB" box="[151,472,393,477]" gridcol="0" gridrow="2" pageId="4" pageNumber="63">
<emphasis id="B92B4A73F248FF8CFF6BFE06FEA1DD89" box="[159,311,393,415]" italics="true" pageId="4" pageNumber="63">
<taxonomicName id="4C5FEDE2F248FF8CFF6BFE06FF63DD89" box="[159,245,393,415]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="genus">Lycodon</taxonomicName>
gong-
</emphasis>
9596
<emphasis id="B92B4A73F248FF8CFF6BFE28FF59DDAB" box="[159,207,423,445]" italics="true" pageId="4" pageNumber="63">shan</emphasis>
(? = 95.5) (n = 4) (
<emphasis id="B92B4A73F248FF8CFE98FE48FEE0DDCA" box="[364,374,455,476]" italics="true" pageId="4" pageNumber="63">s</emphasis>
= 0.7)
</th>
<td id="76BEFD5FF2480077FE00FE06FDC1DDCB" box="[500,599,393,477]" gridcol="1" gridrow="2" pageId="4" pageNumber="63">92</td>
<td id="76BEFD5FF2480077FD70FE06FD2CDDCB" box="[644,698,393,477]" gridcol="2" gridrow="2" pageId="4" pageNumber="63">17</td>
<td id="76BEFD5FF2480077FD2EFE06FCB7DDCB" box="[730,801,393,477]" gridcol="3" gridrow="2" pageId="4" pageNumber="63">3240</td>
<td id="76BEFD5FF2480077FCB9FE06FC1BDDCB" box="[845,909,393,477]" gridcol="4" gridrow="2" pageId="4" pageNumber="63">1316</td>
<td id="76BEFD5FF2480077FC42FE06FC78DDCB" box="[950,1006,393,477]" gridcol="5" gridrow="2" pageId="4" pageNumber="63">1</td>
<td id="76BEFD5FF2480077FBFBFE06FB0BDDCB" box="[1039,1181,393,477]" gridcol="6" gridrow="2" pageId="4" pageNumber="63">8 8</td>
<td id="76BEFD5FF2480077FB3CFE06FA17DDCB" box="[1224,1409,393,477]" gridcol="7" gridrow="2" pageId="4" pageNumber="63">1 whitish</td>
</tr>
<tr id="356F9423F2480077FF63FE60FA17DE55" box="[151,1409,495,579]" gridrow="3" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FE60FE4EDE55" box="[151,472,495,579]" gridcol="0" gridrow="3" pageId="4" pageNumber="63">
<taxonomicName id="4C5FEDE2F248FF8CFF6BFE60FEC5DE10" box="[159,339,495,518]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="synaptor">
<emphasis id="B92B4A73F248FF8CFF6BFE60FEC5DE10" box="[159,339,495,518]" italics="true" pageId="4" pageNumber="63">Lycodon synaptor</emphasis>
</taxonomicName>
unknown (n = 2)
</th>
<td id="76BEFD5FF2480077FE00FE60FDC1DE55" box="[500,599,495,579]" gridcol="1" gridrow="3" pageId="4" pageNumber="63">
6869 (? = 68.5) (
<emphasis id="B92B4A73F248FF8CFE08FDA1FD90DE55" box="[508,518,558,579]" italics="true" pageId="4" pageNumber="63">s</emphasis>
=0.7)
</td>
<td id="76BEFD5FF2480077FD70FE60FD2CDE55" box="[644,698,495,579]" gridcol="2" gridrow="3" pageId="4" pageNumber="63">17</td>
<td id="76BEFD5FF2480077FD2EFE60FCB7DE55" box="[730,801,495,579]" gridcol="3" gridrow="3" pageId="4" pageNumber="63">3031</td>
<td id="76BEFD5FF2480077FCB9FE60FC1BDE55" box="[845,909,495,579]" gridcol="4" gridrow="3" pageId="4" pageNumber="63">9</td>
<td id="76BEFD5FF2480077FC42FE60FC78DE55" box="[950,1006,495,579]" gridcol="5" gridrow="3" pageId="4" pageNumber="63">0</td>
<td id="76BEFD5FF2480077FBFBFE60FB0BDE55" box="[1039,1181,495,579]" gridcol="6" gridrow="3" pageId="4" pageNumber="63">8 8</td>
<td id="76BEFD5FF2480077FB3CFE60FA17DE55" box="[1224,1409,495,579]" gridcol="7" gridrow="3" pageId="4" pageNumber="63">1 whitish</td>
</tr>
<tr id="356F9423F2480077FF63FDD9FA17DEBC" box="[151,1409,598,682]" gridrow="4" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FDD9FE4EDEBC" box="[151,472,598,682]" gridcol="0" gridrow="4" pageId="4" pageNumber="63">
<taxonomicName id="4C5FEDE2F248FF8CFF6BFDD9FF63DE7A" box="[159,245,598,620]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="genus">
<emphasis id="B92B4A73F248FF8CFF6BFDD9FF63DE7A" box="[159,245,598,620]" italics="true" pageId="4" pageNumber="63">Lycodon</emphasis>
</taxonomicName>
68
<taxonomicName id="4C5FEDE2F248FF8CFF6BFDFAFEB8DE9D" box="[159,302,629,651]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="liuchengchaoi">
<emphasis id="B92B4A73F248FF8CFF6BFDFAFEB8DE9D" box="[159,302,629,651]" italics="true" pageId="4" pageNumber="63">liuchengchaoi</emphasis>
</taxonomicName>
(n = 3)
</th>
<td id="76BEFD5FF2480077FE00FDD9FDC1DEBC" box="[500,599,598,682]" gridcol="1" gridrow="4" pageId="4" pageNumber="63">77</td>
<td id="76BEFD5FF2480077FD70FDD9FD2CDEBC" box="[644,698,598,682]" gridcol="2" gridrow="4" pageId="4" pageNumber="63">17</td>
<td id="76BEFD5FF2480077FD2EFDD9FCB7DEBC" box="[730,801,598,682]" gridcol="3" gridrow="4" pageId="4" pageNumber="63">4045</td>
<td id="76BEFD5FF2480077FCB9FDD9FC1BDEBC" box="[845,909,598,682]" gridcol="4" gridrow="4" pageId="4" pageNumber="63">1015</td>
<td id="76BEFD5FF2480077FC42FDD9FC78DEBC" box="[950,1006,598,682]" gridcol="5" gridrow="4" pageId="4" pageNumber="63">1</td>
<td id="76BEFD5FF2480077FBFBFDD9FB0BDEBC" box="[1039,1181,598,682]" gridcol="6" gridrow="4" pageId="4" pageNumber="63">8 [7,9] 78</td>
<td id="76BEFD5FF2480077FB3CFDD9FA17DEBC" box="[1224,1409,598,682]" gridcol="7" gridrow="4" pageId="4" pageNumber="63">2 yellow</td>
</tr>
<tr id="356F9423F2480077FF63FD32FA17DF07" box="[151,1409,701,785]" gridrow="5" pageId="4" pageNumber="63">
<th id="76BEFD5FF2480077FF63FD32FE4EDF07" box="[151,472,701,785]" gridcol="0" gridrow="5" pageId="4" pageNumber="63">
<emphasis id="B92B4A73F248FF8CFF6BFD32FED4DEC5" bold="true" box="[159,322,701,723]" italics="true" pageId="4" pageNumber="63">
<taxonomicName id="4C5FEDE2F248FF8CFF6BFD32FF6EDEC5" box="[159,248,701,723]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="genus">Lycodon</taxonomicName>
carda-
</emphasis>
9293
<emphasis id="B92B4A73F248FF8CFF6BFD54FE91DEE7" bold="true" box="[159,263,731,753]" italics="true" pageId="4" pageNumber="63">momensis</emphasis>
* (? = 92.5) (n = 4) (
<emphasis id="B92B4A73F248FF8CFE98FD74FEE0DF06" box="[364,374,763,784]" italics="true" pageId="4" pageNumber="63">s</emphasis>
= 0.7)
</th>
<td id="76BEFD5FF2480077FE00FD32FDC1DF07" box="[500,599,701,785]" gridcol="1" gridrow="5" pageId="4" pageNumber="63">92</td>
<td id="76BEFD5FF2480077FD70FD32FD2CDF07" box="[644,698,701,785]" gridcol="2" gridrow="5" pageId="4" pageNumber="63">17</td>
<td id="76BEFD5FF2480077FD2EFD32FCB7DF07" box="[730,801,701,785]" gridcol="3" gridrow="5" pageId="4" pageNumber="63">1213</td>
<td id="76BEFD5FF2480077FCB9FD32FC1BDF07" box="[845,909,701,785]" gridcol="4" gridrow="5" pageId="4" pageNumber="63">9</td>
<td id="76BEFD5FF2480077FC42FD32FC78DF07" box="[950,1006,701,785]" gridcol="5" gridrow="5" pageId="4" pageNumber="63">0</td>
<td id="76BEFD5FF2480077FBFBFD32FB0BDF07" box="[1039,1181,701,785]" gridcol="6" gridrow="5" pageId="4" pageNumber="63">10 8</td>
<td id="76BEFD5FF2480077FB3CFD32FA17DF07" box="[1224,1409,701,785]" gridcol="7" gridrow="5" pageId="4" pageNumber="63">1 whitish</td>
</tr>
</table>
</paragraph>
<caption id="DF20C6E9F248FF8CFF63F84BFAB3DBCD" box="[151,1317,1988,2011]" httpUri="https://zenodo.org/record/207724/files/figure.png" pageId="4" pageNumber="63" targetBox="[192,1381,861,1950]" targetPageId="4">
<paragraph id="8BE09661F248FF8CFF63F84BFAB3DBCD" blockId="4.[151,1317,1988,2011]" box="[151,1317,1988,2011]" pageId="4" pageNumber="63">
<emphasis id="B92B4A73F248FF8CFF63F84BFE85DBCC" bold="true" box="[151,275,1988,2011]" pageId="4" pageNumber="63">FIGURE 2.</emphasis>
Dorsal view of holotype (CWNU867001) of
<taxonomicName id="4C5FEDE2F248FF8CFD14F84AFC5ADBCD" box="[736,972,1989,2011]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="liuchengchaoi" status="sp. nov.">
<emphasis id="B92B4A73F248FF8CFD14F84AFC5ADBCD" box="[736,972,1989,2011]" italics="true" pageId="4" pageNumber="63">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218F708F248FF8CFC27F84AFBB5DBCC" box="[979,1059,1989,2010]" pageId="4" pageNumber="63" rank="species">
<emphasis id="B92B4A73F248FF8CFC27F84AFBB5DBCC" bold="true" box="[979,1059,1989,2010]" pageId="4" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
Photograph by Ke Jiang.
</paragraph>
</caption>
<caption id="DF20C6E9F249FF8DFF63FA15FABDD9A7" box="[151,1323,1434,1457]" httpUri="https://zenodo.org/record/207725/files/figure.png" pageId="5" pageNumber="64" targetBox="[194,1390,236,1376]" targetPageId="5">
<paragraph id="8BE09661F249FF8DFF63FA15FABDD9A7" blockId="5.[151,1323,1434,1457]" box="[151,1323,1434,1457]" pageId="5" pageNumber="64">
<emphasis id="B92B4A73F249FF8DFF63FA15FE84D9A6" bold="true" box="[151,274,1434,1457]" pageId="5" pageNumber="64">FIGURE 3.</emphasis>
Ventral view of holotype (CWNU867001) of
<taxonomicName id="4C5FEDE2F249FF8DFD12FA14FC45D9A7" box="[742,979,1435,1457]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="liuchengchaoi" status="sp. nov.">
<emphasis id="B92B4A73F249FF8DFD12FA14FC45D9A7" box="[742,979,1435,1457]" italics="true" pageId="5" pageNumber="64">Lycodon liuchengchaoi</emphasis>
</taxonomicName>
<emphasis id="B92B4A73F249FF8DFC2CFA14FBBFD9A6" bold="true" box="[984,1065,1435,1456]" pageId="5" pageNumber="64">
<taxonomicNameLabel id="A218F708F249FF8DFC2CFA14FBBFD9A6" box="[984,1065,1435,1456]" pageId="5" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Photograph by Ke Jiang.
</paragraph>
</caption>
<subSubSection id="C345C5EAF249FF8DFF31FA51FB4CDA90" pageId="5" pageNumber="64" type="etymology">
<paragraph id="8BE09661F249FF8DFF31FA51FB4CDA90" blockId="5.[151,1437,1502,1921]" pageId="5" pageNumber="64">
<emphasis id="B92B4A73F249FF8DFF31FA51FEDDD9E0" bold="true" box="[197,331,1502,1526]" pageId="5" pageNumber="64">Etymology.</emphasis>
The species is named after Dr. Cheng-chao Liu (now spelled Chengzhao Liu;
<date id="FFE1B0A1F249FF8DFB4BFA51FAD7D9E0" box="[1215,1345,1502,1526]" pageId="5" pageNumber="64" value="1900" valueMax="1976">19001976</date>
), one of the founders of modern Chinese herpetology. Since 1938, Dr. Liu taught in West
<collectingCountry id="F348D6F1F249FF8DFBB4F98EFB15DA0F" box="[1088,1155,1537,1561]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
Union University (West
<collectingCountry id="F348D6F1F249FF8DFF63F9A9FF4CDA28" box="[151,218,1574,1598]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
University of Medical Science later) until the end of his life, and established the Department of Herpetology in Chengdu Institute of Biology in the 1960s. His best-known book “Amphibians of Western
<collectingCountry id="F348D6F1F249FF8DFB53F9C6FB7ADA77" box="[1191,1260,1609,1633]" name="China" pageId="5" pageNumber="64">China</collectingCountry>
” was published in 1950. He contributed substantially to the taxonomy and life history of amphibians and reptiles.
</paragraph>
</subSubSection>
<subSubSection id="C345C5EAF249FF8DFF31F91EFD0BDB00" pageId="5" pageNumber="64" type="biology_ecology">
<paragraph id="8BE09661F249FF8DFF31F91EFD0BDB00" blockId="5.[151,1437,1502,1921]" pageId="5" pageNumber="64">
<emphasis id="B92B4A73F249FF8DFF31F91EFEB3DABC" bold="true" box="[197,293,1681,1706]" pageId="5" pageNumber="64">Biology.</emphasis>
The
<typeStatus id="54E428C3F249FF8DFE96F91EFE52DABF" box="[354,452,1681,1705]" pageId="5" pageNumber="64" type="holotype">holotype</typeStatus>
(CWNU867001) and one of the
<typeStatus id="54E428C3F249FF8DFCB4F91DFC3ADABC" box="[832,940,1682,1706]" pageId="5" pageNumber="64" type="paratype">paratypes</typeStatus>
(CWNU84002) were found on the floor of broad-leaved deciduous forest at night. They were associated with random riprap and duff, not far from a forest stream. Many scales of
<taxonomicName id="4C5FEDE2F249FF8DFE6BF955FCD6DAE7" authority="Gray, 1853" authorityName="Gray" authorityYear="1853" box="[415,832,1753,1778]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="indicus">
<emphasis id="B92B4A73F249FF8DFE6BF955FD3DDAE4" box="[415,683,1754,1778]" italics="true" pageId="5" pageNumber="64">Sphenomorphus indicus</emphasis>
(Gray, 1853)
</taxonomicName>
were found in the intestine of CWNU84002. This lizard species was common at the collecting site.
</paragraph>
</subSubSection>
<subSubSection id="C345C5EAF249FF8DFF31F8AEFE22DB97" pageId="5" pageNumber="64" type="distribution">
<paragraph id="8BE09661F249FF8DFF31F8AEFE22DB97" blockId="5.[151,1437,1502,1921]" pageId="5" pageNumber="64">
<emphasis id="B92B4A73F249FF8DFF31F8AEFEC8DB2C" bold="true" box="[197,350,1825,1850]" pageId="5" pageNumber="64">Distribution.</emphasis>
Known only from the localities of the
<typeStatus id="54E428C3F249FF8DFCB7F8AEFC33DB2F" box="[835,933,1825,1849]" pageId="5" pageNumber="64" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E428C3F249FF8DFC1CF8ADFBC0DB2C" box="[1000,1110,1826,1850]" pageId="5" pageNumber="64" type="paratype">paratypes</typeStatus>
, Tangjiahe National Nature Reserve on the northern edge of Sichuan Basin, in Mount Min System and Monping (Baoxing County) in Mount Qionglai System (
<figureCitation id="13648AE4F249FF8DFE96F8E6FE30DB97" box="[354,422,1897,1921]" captionStart="FIGURE 5" captionStartId="6.[151,250,1709,1732]" captionTargetBox="[441,1145,1175,1687]" captionTargetId="figure@6.[441,1145,1175,1687]" captionTargetPageId="6" captionText="FIGURE 5. Map showing the two collection sites (black spots) of Lycodon liuchengchaoi sp. nov. in Sichuan province. Tangjiahe National Nature Reserve (type locality), Qingchuan County (a); Mouping (now Baoxing County) (b)." httpUri="https://zenodo.org/record/207727/files/figure.png" pageId="5" pageNumber="64">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>