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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e104969" ID-Pensoft-Pub="1864-8312-81-611" ID-Pensoft-UUID="D6555CEAD4F355FF9A3D61D126ABBDE6" ID-ZooBank="8C3E7D063B2548429F28A76926D4A741" ModsDocID="1864-8312-81-611" checkinTime="1689969740977" checkinUser="pensoft" docAuthor="Patitucci, Luciano Damian, Mulieri, Pablo Ricardo, Couri, Marcia Souto &amp; Dominguez, Martha Cecilia" docDate="2023" docId="5AD8CADC9809584CA1BE46D184212A33" docLanguage="en" docName="ArthropodSystPhyl 81: 611-653" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-07-21" docSource="http://dx.doi.org/10.3897/asp.81.e104969" docTitle="Coenosia delneneo Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez 2023, sp. nov." docType="treatment" docUuid="1863B9FA-097C-4465-9289-DE7B93737D43" docUuidSource="ZooBank" docVersion="1" id="D6555CEAD4F355FF9A3D61D126ABBDE6" lastPageNumber="611" masterDocId="D6555CEAD4F355FF9A3D61D126ABBDE6" masterDocTitle="Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae)" masterLastPageNumber="653" masterPageNumber="611" pageNumber="611" updateTime="1689969740977" updateUser="pensoft">
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<mods:title id="663F5A709E06AFD17445A993EA627CA7">Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae)</mods:title>
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<mods:namePart id="8DC17ACA9D21F2B73F300CE71441F754">Patitucci, Luciano Damian</mods:namePart>
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<mods:affiliation id="82BE17C20807B473A22C94B055DAC70A">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
<mods:nameIdentifier id="CA5FAF0554B0969443247927E3B96789" type="email">lpatitu@yahoo.com.ar</mods:nameIdentifier>
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<mods:namePart id="30FDCC52E971F596517CD0334B236B66">Mulieri, Pablo Ricardo</mods:namePart>
<mods:affiliation id="2675FFCD2FBEB9C6FEB45F5F1120087E">Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; (MACN), Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="A728347516A0077EF4D987A6BBF258CE">Couri, Marcia Souto</mods:namePart>
<mods:affiliation id="111A95FBE0210CBAE0BFC633B3FDA1B0">Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brazil &amp; Conselho Nacional de Desenvolvimento Cientifico e Tecnologico (CNPq), Brazil</mods:affiliation>
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<mods:namePart id="C650CAA68B1756D55358BB672499A9E4">Dominguez, Martha Cecilia</mods:namePart>
<mods:affiliation id="78EF4D1408A68A1D44187E65876814D2">Laboratorio de Entomologia, Instituto Argentino de Investigaciones de Zonas Aridas (IADIZA-CONICET) Mendoza, Mendoza, Argentina</mods:affiliation>
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<mods:title id="C94F449EC0958034841B1C9A2385F439">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="8D5F4A544019FA188C75B3572DF9248D">2023</mods:date>
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<mods:number id="D1550BE0AA144A28418544154CF6369F">81</mods:number>
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<treatment id="5AD8CADC9809584CA1BE46D184212A33" LSID="urn:lsid:zoobank.org:act:1863B9FA-097C-4465-9289-DE7B93737D43" httpUri="http://treatment.plazi.org/id/5AD8CADC9809584CA1BE46D184212A33" lastPageNumber="611" pageId="0" pageNumber="611">
<subSubSection id="992C65BE14F68B8C7BD060BC585416F6" pageId="0" pageNumber="611" type="nomenclature">
<paragraph id="D7C530B0B9149ACE918949CBA7AD570F" pageId="0" pageNumber="611">
<taxonomicName id="0D70419E3E335636D1CDAD98BA8DF281" LSID="https://zoobank.org/1863B9FA-097C-4465-9289-DE7B93737D43" authority="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez, 2023" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia delneneo" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="delneneo" status="sp. nov.">Coenosia delneneo</taxonomicName>
<taxonomicNameLabel id="D369ECD177DF418123002A6ED2ED0A8D" pageId="0" pageNumber="611">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9871510E890EE7973926237B19202BDC" pageId="0" pageNumber="611" type="male (fig. 8a)">
<paragraph id="8EDDDA84783FB4120F6E7D4BC3B2CF19" pageId="0" pageNumber="611">Male (Fig. 8A).</paragraph>
<paragraph id="8421E46ADF37C00185D97CA7CF37F187" pageId="0" pageNumber="611">
Length. Body: 5.56-5.90 mm, wing: 4.1-4.69 mm.
<emphasis id="84A44D03B52A76CE6E66244A3D3B952F" bold="true" italics="true" pageId="0" pageNumber="611">Head</emphasis>
: (Fig.
<figureCitation id="FE5AA8F2DB49A53F192C35AFB787B100" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coenosia delneneo sp. nov., male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, G cercus and surstylus, lateral view, H detail of cercus, distal tip (SEM), I phallic complex, lateral view, J detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K)." figureDoi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611">8B</figureCitation>
). Frons at vertex about one third of the head width. Frons, fronto-orbital plate, parafacial and gena black with yellow-grey pollinosity. Fronto-orbital plate with 1-3 little setulae, close to parafacialia. Antenna dark brown, apex of pedicel with yellow pollinosity; arista brown with its longest microtrichia hardly longer than its basal diameter. Palpus black.
<emphasis id="5FF9085080E428D89708D34CFBF66050" bold="true" italics="true" pageId="0" pageNumber="611">Thorax</emphasis>
: (Fig.
<figureCitation id="1154323F36DBF8B00DF20146C488D394" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coenosia delneneo sp. nov., male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, G cercus and surstylus, lateral view, H detail of cercus, distal tip (SEM), I phallic complex, lateral view, J detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K)." figureDoi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611">8C</figureCitation>
). Grey with light brown pollinosity. Chaetotaxy: acr s strong and biseriate; anterior presutural acr s shorter than the eaDC; dorsocentrals 1+3, eaDC is half (or more) of length of the aDC. Anepisternum with a series of 4-5 strong setae; katepisternum with 2-3 setula.
<emphasis id="704766F76F3D8895D6805C452312B7A0" bold="true" italics="true" pageId="0" pageNumber="611">Wing</emphasis>
: Both calypters whitish hyaline with white margins.
<emphasis id="4B6484CDF530FEE8B83D1CF02AB4D4A8" bold="true" italics="true" pageId="0" pageNumber="611">Legs</emphasis>
: Coxae grey. Fore femur yellow with a black large spot on dorsal to posterior surface. Trochanters, femora, tibiae, and 1-4 tarsomeres yellow. Fifth tarsomere black. Fore femur with a row of strong pd setae, and a row of strong pv setae. Mid femur with a row of strong setae on ventral surface, and 5-6 setae on anterior surface; mid tibia with one ad and one pd median setae with the same length, ad seta positioned below of the pd seta. Hind femur with a complete row of ad and a complete row of av setae, two setae on anterior surface at middle, and a row of setae on ventral surface, hind tibia with 3 preapical setae (ad, dorsal, ventral).
<emphasis id="E51563B52BBEF2FE05414FC43ECD4810" bold="true" italics="true" pageId="0" pageNumber="611">Abdomen</emphasis>
: (Fig.
<figureCitation id="AF3D6D244DCDFA5250BFAE664494267D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coenosia delneneo sp. nov., male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, G cercus and surstylus, lateral view, H detail of cercus, distal tip (SEM), I phallic complex, lateral view, J detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K)." figureDoi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611">8D</figureCitation>
). Grey without spots, tergite 6 and epand brighter than the other tergites. Sternite 5 with apical margin concave (
<normalizedToken id="6EE27A09212B661236C3EE52FAAA078D" originalValue="“U”">&quot;U&quot;</normalizedToken>
shape) and membrane; setae concentrated on the lobes and some long and strong on apical margin (Fig.
<figureCitation id="2B72B2B871D97F5B50DCA5AE12CB2A84" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coenosia delneneo sp. nov., male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, G cercus and surstylus, lateral view, H detail of cercus, distal tip (SEM), I phallic complex, lateral view, J detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K)." figureDoi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611">8E</figureCitation>
).
<emphasis id="79C9D36923F9C12C9A3521C680347274" bold="true" italics="true" pageId="0" pageNumber="611">Terminalia</emphasis>
: Cercus with strong spines on apical middle (Fig.
<figureCitation id="41085EADF57DFBD53A41D852E8FE4FF7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coenosia delneneo sp. nov., male. A Lateral view, B head, frontal view, C thorax, dorsal view, D abdomen, dorsal view, E fifth sternite, F cercus, G cercus and surstylus, lateral view, H detail of cercus, distal tip (SEM), I phallic complex, lateral view, J detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K)." figureDoi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611">8F, G</figureCitation>
).
</paragraph>
<caption id="7AD7B233C61153B7E78494B725CC8E3E" doi="10.3897/asp.81.e104969.figure8" httpUri="https://binary.pensoft.net/fig/882076" pageId="0" pageNumber="611" start="Figure 8" startId="F8">
<paragraph id="F5D3E0E3810373A03EEB5325484BF55D" pageId="0" pageNumber="611">
<emphasis id="FAD8CD1F6CD46334CCCA788E97CBB097" bold="true" pageId="0" pageNumber="611">Figure 8.</emphasis>
<taxonomicName id="0AF0F762423C06ADE7130E741A21AD6E" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia delneneo" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="delneneo">
<emphasis id="3F96A2DC05B7115A009E5E15A6945899" italics="true" pageId="0" pageNumber="611">Coenosia delneneo</emphasis>
</taxonomicName>
<emphasis id="18CF2F24546169C7802FDEAD6F710A79" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
, male.
<emphasis id="CCFC3D4581555145EC8270F8322BE7B5" bold="true" pageId="0" pageNumber="611">A</emphasis>
Lateral view,
<emphasis id="052C1CFE6EA3D3A590B975E5E82984E9" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view,
<emphasis id="E569CFE33C9125CB167C604E04E05D37" bold="true" pageId="0" pageNumber="611">C</emphasis>
thorax, dorsal view,
<emphasis id="D10A82515BC34FA228C1F52AD5B6B4FF" bold="true" pageId="0" pageNumber="611">D</emphasis>
abdomen, dorsal view,
<emphasis id="F1B66FED70FD985D665E80663277779B" bold="true" pageId="0" pageNumber="611">E</emphasis>
fifth sternite,
<emphasis id="243BF02628E450F88439189E9190EF8C" bold="true" pageId="0" pageNumber="611">F</emphasis>
cercus,
<emphasis id="960665813319D1187D7947A6841D0D84" bold="true" pageId="0" pageNumber="611">G</emphasis>
cercus and surstylus, lateral view,
<emphasis id="1F6D92D15F681CAD45937A1E551378AA" bold="true" pageId="0" pageNumber="611">H</emphasis>
detail of cercus, distal tip (SEM),
<emphasis id="100C699FC2EE52F66D0AEC78E60DB0F9" bold="true" pageId="0" pageNumber="611">I</emphasis>
phallic complex, lateral view,
<emphasis id="B3EBBD57DE62B9CE8F406821EB86229E" bold="true" pageId="0" pageNumber="611">J</emphasis>
detail of phallic complex, distal tip. Scale bar: 1 mm (A), 0.5 mm (B-E), 0.2 mm (F, G), 0.02 mm (H), 0.01 mm (J, K).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="51226712F9E6F25B2CAB07CE599127B5" pageId="0" pageNumber="611" type="female (fig. 9a, b)">
<paragraph id="BF414E0C4E11A68ECA4F660F29509BD1" pageId="0" pageNumber="611">Female (Fig. 9A, B).</paragraph>
<paragraph id="02AD79121D17C906CC570D9F9EB5D318" pageId="0" pageNumber="611">
Length. Body: 5.70-6.00 mm, wing: 4.8-5.12 mm. Differs from male as follows:
<emphasis id="87AAC583923D534A871ABF29DD637CFA" bold="true" italics="true" pageId="0" pageNumber="611">Legs</emphasis>
: Mid tibia with one ad and one pd median setae, both setae with the same length and position.
<emphasis id="765228B4E098A46E2505D4EA31D96864" bold="true" italics="true" pageId="0" pageNumber="611">Ovipositor</emphasis>
: Segments longer than wide. Tergites 7 and 8 with two long and wide sclerotized plates (Fig.
<figureCitation id="2C7307706005FC432E5711F1B0C7CB57" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coenosia delneneo sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae, F details of spermathecae (SEM). Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E), 0.05 mm (F). Sixth and seventh sternites - red arrows, spermathecae small prominence - white arrow." figureDoi="10.3897/asp.81.e104969.figure9" httpUri="https://binary.pensoft.net/fig/882077" pageId="0" pageNumber="611">9C</figureCitation>
). Sternites 6 and 7 with one short and thin plates; sternite 8 divided into two small and sclerotized plates each with 4-5 setae on distal margin (Fig.
<figureCitation id="F250FA2542C34834AB94C9BE3C92DE60" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coenosia delneneo sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae, F details of spermathecae (SEM). Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E), 0.05 mm (F). Sixth and seventh sternites - red arrows, spermathecae small prominence - white arrow." figureDoi="10.3897/asp.81.e104969.figure9" httpUri="https://binary.pensoft.net/fig/882077" pageId="0" pageNumber="611">9D</figureCitation>
). Three spermathecae, oval shaped, with a small prominence at the opposite end of the insertion of the tracheola (Fig.
<figureCitation id="5042401675DE8D9B9521AA72624612AF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coenosia delneneo sp. nov., female. A Lateral view, B head, frontal view, C ovipositor, dorsal view, D ovipositor, ventral view, E spermathecae, F details of spermathecae (SEM). Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E), 0.05 mm (F). Sixth and seventh sternites - red arrows, spermathecae small prominence - white arrow." figureDoi="10.3897/asp.81.e104969.figure9" httpUri="https://binary.pensoft.net/fig/882077" pageId="0" pageNumber="611">9E, F</figureCitation>
).
</paragraph>
<caption id="AEAA59892626F8673E419323B8A76388" doi="10.3897/asp.81.e104969.figure9" httpUri="https://binary.pensoft.net/fig/882077" pageId="0" pageNumber="611" start="Figure 9" startId="F9">
<paragraph id="44FF3ED290C7046575AC5BC575F708FB" pageId="0" pageNumber="611">
<emphasis id="D2D438FA21876A40645E5DE6B2AE48BA" bold="true" pageId="0" pageNumber="611">Figure 9.</emphasis>
<taxonomicName id="BA4D9AF7EFAB624B229907879C113595" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia delneneo" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="delneneo">
<emphasis id="8EE78C6E2BB4712B876B57B73FE7291C" italics="true" pageId="0" pageNumber="611">Coenosia delneneo</emphasis>
</taxonomicName>
<emphasis id="CFA8A436B4C3469F76FC4183EC4C0CF7" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
, female.
<emphasis id="E755DEB556D119995563AA4BCCD2719F" bold="true" pageId="0" pageNumber="611">A</emphasis>
Lateral view,
<emphasis id="D0001008CCE0C2E6EAD02859657618CD" bold="true" pageId="0" pageNumber="611">B</emphasis>
head, frontal view,
<emphasis id="118575239C60C803944D7A79E01FEFB0" bold="true" pageId="0" pageNumber="611">C</emphasis>
ovipositor, dorsal view,
<emphasis id="E5CF907E6B793781D33A47D13CD1F42B" bold="true" pageId="0" pageNumber="611">D</emphasis>
ovipositor, ventral view,
<emphasis id="DFA2F398C5A2C13CDB78378DA1AD75B3" bold="true" pageId="0" pageNumber="611">E</emphasis>
spermathecae,
<emphasis id="5F8FADE00C4F5301E700F46BF440826B" bold="true" pageId="0" pageNumber="611">F</emphasis>
details of spermathecae (SEM). Scale bar: 1 mm (A), 0.5 mm (B-D), 0.2 mm (E), 0.05 mm (F). Sixth and seventh sternites - red arrows, spermathecae small prominence - white arrow.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D4ED52F436139CCE105C190BDB48B3C0" pageId="0" pageNumber="611" type="type material">
<paragraph id="E9EB323621F2620C6AD1EE395F27F408" pageId="0" pageNumber="611">Type material.</paragraph>
<paragraph id="C31ECA0D8FB807626C1518AF1E92C4E5" pageId="0" pageNumber="611">
<emphasis id="6C1DB96541965F0B80E6CFEB744A0471" bold="true" pageId="0" pageNumber="611">
<emphasis id="640B724766554FC5D2EEFC20F9E12D6E" italics="true" pageId="0" pageNumber="611">Holotype</emphasis>
.
</emphasis>
Male; MACN [pinned]. Original labels: &quot;Arg.
<normalizedToken id="DFC079B3E4F8D80FB0CC74934B0F70DF" originalValue="Neuquén">Neuquen</normalizedToken>
,
<normalizedToken id="E19DC4632C08E24552344D00A4619AA3" originalValue="Aluminé">Alumine</normalizedToken>
/
<normalizedToken id="B68A529934C59B7E4B4640B897CA6540" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="0FAEB297865D8295E53ECDC6944CF6DA" originalValue="Aluminé">Alumine</normalizedToken>
896 m /
<geoCoordinate id="BAB4F271647EAF33739E2A2C5CC08B8F" degrees="39" direction="south" minutes="14" orientation="latitude" precision="1" seconds="26.03" value="-39.240562">
39°14
<normalizedToken id="68DBB54A01D6FA07C7BC9EAC0719E0EE" originalValue="26.03">'26.03''</normalizedToken>
S
</geoCoordinate>
<geoCoordinate id="DEFF7D43C8DBB7C7AA0FCD0840906E36" degrees="70" direction="west" minutes="54" orientation="longitude" precision="1" seconds="43.93" value="-70.9122">
70°54
<normalizedToken id="0BD07B830DE1C49205B9156FCCE5455B" originalValue="43.93">'43.93''</normalizedToken>
W
</geoCoordinate>
/ JAN2013 Olea, Mulieri &amp; Patitucci leg.&quot; print. on white paper; &quot;MACN-En / 34728&quot; print. on white paper.;
<normalizedToken id="E2926D9FF2FA9CFB8F119746FFA065A1" originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
print. on red paper, black frame. -
<emphasis id="F3AC87F45D077F555D7C72A94ABD36CA" bold="true" pageId="0" pageNumber="611">
<emphasis id="15D1558BAC9D54DFC0A092E837B93622" italics="true" pageId="0" pageNumber="611">Paratypes</emphasis>
.
</emphasis>
One male; MACN [pinned]. Original labels: &quot;Arg.
<normalizedToken id="35EADA3AA137AB2DD7013D5FEC3A3765" originalValue="Neuquén">Neuquen</normalizedToken>
,
<normalizedToken id="4E967BE9EAC811BDFAD2B07027A367F7" originalValue="Aluminé">Alumine</normalizedToken>
/
<normalizedToken id="2461483E59DB7F8CFE5EAEBD1623D965" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="FFAECB5222208D6CCAD4D1BAFD68052A" originalValue="Aluminé">Alumine</normalizedToken>
896 m /
<geoCoordinate id="7D1344422875E582C9D918BCAA6925EB" degrees="39" direction="south" minutes="14" orientation="latitude" precision="1" seconds="26.03" value="-39.240562">
39°14
<normalizedToken id="B0FEFAE39814F1FCF685517ADAFBEB93" originalValue="26.03">'26.03''</normalizedToken>
S
</geoCoordinate>
<geoCoordinate id="28BA205348E750482DCBBAB535F37BF2" degrees="70" direction="west" minutes="54" orientation="longitude" precision="1" seconds="43.93" value="-70.9122">
70°54
<normalizedToken id="23B2021FA4F3D8E3F9A087C2B252CE45" originalValue="43.93">'43.93''</normalizedToken>
W
</geoCoordinate>
/ JAN2013 Olea, Mulieri &amp; Patitucci leg.&quot; print. on white paper; &quot;MACN-En / 34729&quot; print. on white paper.;
<normalizedToken id="524F015E18927BBD42BE22492ECF84C2" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. Seven females; MACN [pinned]. Original labels: &quot;Arg.
<normalizedToken id="90350C0857A3697A37FF832E05EF19AB" originalValue="Neuquén">Neuquen</normalizedToken>
,
<normalizedToken id="25C86E9092805107BDE5A32E169F71C6" originalValue="Aluminé">Alumine</normalizedToken>
/
<normalizedToken id="CF994D93A458F5B10BDED01B347EB946" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="AC53C8D477D3D8A6CFEBC5BA719CF35B" originalValue="Aluminé">Alumine</normalizedToken>
896 m /
<geoCoordinate id="42CD4B90D1A92B6654E48A1791382857" degrees="39" direction="south" minutes="14" orientation="latitude" precision="1" seconds="26.03" value="-39.240562">
39°14
<normalizedToken id="3B8B8D820BCA8089AED5C3A60A522A62" originalValue="26.03">'26.03''</normalizedToken>
S
</geoCoordinate>
<geoCoordinate id="27FFEAF8307DE17ADA9C4D688DE38CC7" degrees="70" direction="west" minutes="54" orientation="longitude" precision="1" seconds="43.93" value="-70.9122">
70°54
<normalizedToken id="99E2D78FA7488F294D17AD756B80CEE7" originalValue="43.93">'43.93''</normalizedToken>
W
</geoCoordinate>
/ JAN2013 Olea, Mulieri &amp; Patitucci leg.&quot; print. on white paper; &quot;MACN-En / 34721 [to] 37427&quot; print. on white paper.;
<normalizedToken id="AFB656035FFF50853AF6537E3754719A" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. Two males and one female; MLP [pinned]. Original labels: &quot;Arg.
<normalizedToken id="0FB3A550E5F0900A21987CF2CB455F3A" originalValue="Neuquén">Neuquen</normalizedToken>
, PN
<normalizedToken id="1E4E5840F01011E63ACD9BD502276626" originalValue="Lanín">Lanin</normalizedToken>
/ Huechulafquen 1070 m (transecta) /
<geoCoordinate id="9C804ADA34C1E28C2F1647C425882EB5" degrees="39" direction="south" minutes="47" orientation="latitude" precision="1" seconds="33.25" value="-39.79257">
39°47
<normalizedToken id="54FC26F01D3EB35521CFA2CCEB48B9CD" originalValue="33.25">'33.25''</normalizedToken>
S
</geoCoordinate>
<geoCoordinate id="67F2E29A4FC5482A627F193BD7C830E4" degrees="71" direction="west" minutes="12" orientation="longitude" precision="1" seconds="56.26" value="-71.21562">
71°12
<normalizedToken id="23AC981980D19465C90F65A6D475FEA9" originalValue="56.26">'56.26''</normalizedToken>
W
</geoCoordinate>
/ 13-XII-2013 Mulieri leg.&quot; print. on white paper,
<normalizedToken id="C5AB6D778E2E3C9C40651AB04801C3FC" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame. One male and 2 females; IFML [pinned]. Original labels: &quot;Arg.
<normalizedToken id="6782203C20B275439BCD5D01E6763476" originalValue="Neuquén">Neuquen</normalizedToken>
, Ruta 13 / Pampa de Lonco Luan / 1522 m a.s.l. transecta /
<geoCoordinate id="D0253B391D9317BCEC1D7E705A376597" degrees="38.897919" direction="south" orientation="latitude" precision="1" value="-38.89792">-38.897919</geoCoordinate>
<geoCoordinate id="973989ED0D64D8B25BD04606371B0010" degrees="70.890376" direction="west" orientation="longitude" precision="1" value="-70.89037">-70.890376</geoCoordinate>
/ 09.II.2018 Patitucci leg.&quot; print. on white paper,
<normalizedToken id="65F67A8C63AD88922998504C9259F7F2" originalValue="“Paratype”">&quot;Paratype&quot;</normalizedToken>
print. on red paper, black frame.
</paragraph>
</subSubSection>
<subSubSection id="B155561C4E7FE88B9909F6B383276F95" pageId="0" pageNumber="611" type="materials_examined">
<paragraph id="9CBCF388365DF34C901D49D192375177" pageId="0" pageNumber="611">Additional material examined.</paragraph>
<paragraph id="A210CB4352DB4DC0993738252DF7627C" pageId="0" pageNumber="611">
<materialsCitation id="521B533957A0A2B97C8C6394EBF389C5" collectingDate="2012-01-12" collectorName="El Bolson, Mulieri, Patitucci" country="ARGENTINA" elevation="1190" latitude="-41.972378" location="Piltriquitron" longLatPrecision="1" longitude="-71.478714" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio Negro province">
<collectingCountry id="615C4246ADB8A96EB1E170BE035BAA65" name="Argentina">ARGENTINA</collectingCountry>
-
<emphasis id="63B89222449F36B0796DD11CEA1BF018" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="4D17C409F393DA4F2BAEE04EAE913D0C" country="Argentina" name="Rio Negro">
<normalizedToken id="F749A58AAECADF1B36A86E22500332B3" originalValue="Río">Rio</normalizedToken>
Negro province
</collectingRegion>
</emphasis>
<specimenCount id="2E38A3B37D31F9A1024B4DC93919078B" type="female">1 female</specimenCount>
,
<specimenCount id="A3E3BE0B95BC65EFB564E159D18E5F6E" type="male">1male</specimenCount>
;
<collectorName id="84E92D1784127CF5C8E06917A8C94313">El Bolson</collectorName>
,
<location id="06180AD20BB6FBFDEFD29AB351317038" LSID="urn:lsid:plazi:treatment:5AD8CADC9809584CA1BE46D184212A33:06180AD20BB6FBFDEFD29AB351317038" country="ARGENTINA" latitude="-41.972378" longLatPrecision="1" longitude="-71.478714" name="Piltriquitron" stateProvince="Rio Negro province">Piltriquitron</location>
;
<geoCoordinate id="DBE07C5E154665AA52A04340F4663ECA" degrees="41.972377" direction="south" orientation="latitude" precision="1" value="-41.972378">-41.972377</geoCoordinate>
,
<geoCoordinate id="9AB7069FFC75DB93CAA1D079DFDFC207" degrees="71.478712" direction="west" orientation="longitude" precision="1" value="-71.478714">-71.478712</geoCoordinate>
;
<elevation id="4C7124C344086D660D34F61B67C99452" metricMagnitude="3" metricUnit="m" metricValue="1.19" unit="m" value="1190.0">
<quantity id="EE9D523FB65AB1350E380A4B7108440B" metricMagnitude="3" metricUnit="m" metricValue="1.19" unit="m" value="1190.0">1190 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="B4B882009D31088A22CFAF48A3CFC3BF" value="2012-01-12">12 Jan 2012</collectingDate>
;
<collectorName id="9CED65F05B3416DA8419C82314FB6B22">Mulieri</collectorName>
&amp;
<collectorName id="CC3F44517288634AE24F87962BA070C0">Patitucci</collectorName>
leg.; MACN
</materialsCitation>
. -
<materialsCitation id="E2DB99827D3ECA54F06EE5AB28112634" collectingDate="2018-02-07" collectorName="Mulieri, Patitucci" country="Argentina" elevation="1240" latitude="-38.85935" location="Ruta" longLatPrecision="1" longitude="-70.47257" municipality="Arroyo Guyapa" specimenCount="2" specimenCount-female="2" stateProvince="Neuquen province">
<emphasis id="68A0A4783F82646851941CC002316D6B" bold="true" pageId="0" pageNumber="611">
<collectingRegion id="D35BDC98A2975DFFC2AC66CF5430EB10" country="Argentina" name="Neuquen">
<normalizedToken id="DABD4DD121CEC4D964F4974D377EDAE8" originalValue="Neuquén">Neuquen</normalizedToken>
province
</collectingRegion>
</emphasis>
<specimenCount id="47F7B7906F64B3D09EC95C556FAA0C54" type="female">2 females</specimenCount>
;
<collectingMunicipality id="2190998051E9AE81B8C89D60D13A2639">Arroyo Guyapa</collectingMunicipality>
,
<location id="6DB38F6A4C46A30870177325B52EF938" LSID="urn:lsid:plazi:treatment:5AD8CADC9809584CA1BE46D184212A33:6DB38F6A4C46A30870177325B52EF938" country="Argentina" latitude="-38.85935" longLatPrecision="1" longitude="-70.47257" municipality="Arroyo Guyapa" name="Ruta" stateProvince="Neuquen province">Ruta</location>
13,
<geoCoordinate id="6219CA8CD1C4C876833CC541984F847B" degrees="38.859349" direction="south" orientation="latitude" precision="1" value="-38.85935">-38.859349</geoCoordinate>
,
<geoCoordinate id="5982336E035AB06104C61CEA8DFB5771" degrees="70.472570" direction="west" orientation="longitude" precision="1" value="-70.47257">-70.472570</geoCoordinate>
;
<elevation id="7684C6C672D98A6699914CA8C530097B" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">
<quantity id="EA017A732C5780EF7315FB7D3854445F" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">1240 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="80833C98D7E36D5F5556D9507A370B46" value="2018-02-07">07 Feb 2018</collectingDate>
;
<collectorName id="B1231A642307079C510FAFB7344C7A59">Mulieri</collectorName>
&amp;
<collectorName id="0516057AD4E29963DE5C8174042AF75F">Patitucci</collectorName>
leg.; MACN
</materialsCitation>
<materialsCitation id="DB915F88721BC9E44CFA3F3AB53D3078" collectingDate="2018-02-08" collectorName="Mulieri" country="Argentina" elevation="1876" latitude="-38.891693" location="Portezuelo La Atravezada" longLatPrecision="1" longitude="-70.65219" municipality="Primeros Pinos" specimenCount="1" specimenCount-male="1" stateProvince="Neuquen">
<specimenCount id="87F1C73123A32540EA8C362772F417F8" type="male">1 male</specimenCount>
;
<collectingMunicipality id="44750C6F2C307BDDB576DE492C1D0668">Primeros Pinos</collectingMunicipality>
,
<location id="892FBE0498882D614D431B7D992B0D79" LSID="urn:lsid:plazi:treatment:5AD8CADC9809584CA1BE46D184212A33:892FBE0498882D614D431B7D992B0D79" country="Argentina" latitude="-38.891693" longLatPrecision="1" longitude="-70.65219" municipality="Primeros Pinos" name="Portezuelo La Atravezada" stateProvince="Neuquen">Portezuelo La Atravezada</location>
;
<geoCoordinate id="990B2A89B607647CD10F017B4EA8B315" degrees="38.891693" direction="south" orientation="latitude" precision="1" value="-38.891693">-38.891693</geoCoordinate>
,
<geoCoordinate id="542992AA25E3C5C91BC3F84417696863" degrees="70.65219" direction="west" orientation="longitude" precision="1" value="-70.65219">-70.65219</geoCoordinate>
2;
<elevation id="9E9E03C1787DBBB7A3A7196209FAFA61" metricMagnitude="3" metricUnit="m" metricValue="1.876" unit="m" value="1876.0">
<quantity id="EA7438287626B62D30C3366D0CDAAD2F" metricMagnitude="3" metricUnit="m" metricValue="1.876" unit="m" value="1876.0">1876 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="09D445FA3DE2BB8B22C99E9E9979531E" value="2018-02-08">08 Feb 2018</collectingDate>
;
<collectorName id="617AE0ED79FC864146B1C816E2A207C5">Mulieri</collectorName>
leg.; MACN
</materialsCitation>
<materialsCitation id="D39DBDA8167E6B2CF5CE6CC1B8CDBAA2" collectingDate="2013-12-16" collectorName="Mulieri, Patitucci" country="Argentina" latitude="-39.809776" location="Ruta" longLatPrecision="1" longitude="-71.03066" specimenCount="3" specimenCount-female="3" stateProvince="Neuquen">
<specimenCount id="A4F7828B38C4346259D926BF11ED69C8" type="female">3 females</specimenCount>
;
<location id="DDB455A07615C671CD2E0ABD85BE6CB7" LSID="urn:lsid:plazi:treatment:5AD8CADC9809584CA1BE46D184212A33:DDB455A07615C671CD2E0ABD85BE6CB7" country="Argentina" latitude="-39.809776" longLatPrecision="1" longitude="-71.03066" name="Ruta" stateProvince="Neuquen">Ruta</location>
23;
<geoCoordinate id="E402A8FBA51D970C427CA18065D83072" degrees="39.809776" direction="south" orientation="latitude" precision="1" value="-39.809776">-39.809776</geoCoordinate>
,
<geoCoordinate id="2EF321BCC12013E281D278B529650B31" degrees="71.030665" direction="west" orientation="longitude" precision="1" value="-71.03066">-71.030665</geoCoordinate>
;
<collectingDate id="6C7211FFFE4DF4FFBB5F2CFACC43752E" value="2013-12-16">16 Dec 2013</collectingDate>
;
<collectorName id="CC0B3A0BE004711DAC0AA6EFBE0E77D0">Mulieri</collectorName>
&amp;
<collectorName id="DF357AFE9952FB0708110C695FA97DE0">Patitucci</collectorName>
leg.; MACN
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43639623811FAE9CEB1A45C4624611F8" pageId="0" pageNumber="611" type="distribution">
<paragraph id="01561222E70463ED2EEFAEF607E96294" pageId="0" pageNumber="611">Distribution (Fig. 12B).</paragraph>
<paragraph id="23123848EEC5E977C1B8C3D3B7460330" pageId="0" pageNumber="611">
ARGENTINA:
<normalizedToken id="F9D046AA13EB42D2293E9F85C51CD651" originalValue="Neuquén">Neuquen</normalizedToken>
,
<normalizedToken id="594B9897CD2639808A0EBA3BCA390427" originalValue="Río">Rio</normalizedToken>
Negro.
</paragraph>
</subSubSection>
<subSubSection id="D6D00F4435D028C75BA76012FB2CD5B6" pageId="0" pageNumber="611" type="etymology">
<paragraph id="BF7DC08D01931B7D4587A79249746AB0" pageId="0" pageNumber="611">Etymology.</paragraph>
<paragraph id="F2EFB5F9F5856389941541FE6D239B6E" pageId="0" pageNumber="611">
The name
<normalizedToken id="4E5AF478C91FAB4D0B38DF5ABE8091D7" originalValue="“delneneo”">&quot;delneneo&quot;</normalizedToken>
refers to an expression in Spanish to the vulgar name (neneo) of the plant
<taxonomicName id="BF8C012A91042F78C181C4CB75F35E5E" class="Magnoliopsida" family="Apiaceae" genus="Mulinum" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="Mulinum spinosum" order="Apiales" pageId="0" pageNumber="611" phylum="Magnoliophyta" rank="species" species="spinosum">
<emphasis id="82959E3AA566F22052A46344D374462A" italics="true" pageId="0" pageNumber="611">Mulinum spinosum</emphasis>
</taxonomicName>
Pers., where several specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="26B8751046C906C1422D070DCDD84139" pageId="0" pageNumber="611" type="remarks">
<paragraph id="ED2F2A630B3AE331287F037FE8A83720" pageId="0" pageNumber="611">Remarks.</paragraph>
<paragraph id="0C8FEAA7AFCA9DC6AC01968A6A32FB1F" pageId="0" pageNumber="611">
<taxonomicName id="391738ECFBE7957659987CE54A4851A6" authorityName="Patitucci &amp; Mulieri &amp; Couri &amp; Domínguez" authorityYear="2023" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia delneneo" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="delneneo">
<emphasis id="7B9236B59A174C9FEA238685E57D3746" italics="true" pageId="0" pageNumber="611">Coenosia delneneo</emphasis>
</taxonomicName>
<emphasis id="D68D318F97D92E13A37FB84B46E16953" bold="true" pageId="0" pageNumber="611">sp. nov.</emphasis>
is distinguished from its congeners by the following combination of characters: gena height greater than the width of the postpedicel, thorax and abdomen gray without vitta or spots, fore femur yellow with a black large spot on dorsal to posterior surface, fifth tarsomere black, dorsocentrals 1+3, with an extra short presutural pair which is half (or more) of the length of the presutural seta, hind tibia with one long ad seta, a long pd seta, and one fine av setae, and cercus longer than broad and with strong spines. Male: tergite 6 and epandrium visible and brighter, distiph with a brush of fine hair on ventral surface. Female: ovipositor in tergites 6 with two long and wide sclerotized plates and two square strongly sclerotized plates, and epiproct with
<normalizedToken id="A1E2061AD7A3ED63B1C33F2FAA385640" originalValue="“boomerang”">&quot;boomerang&quot;</normalizedToken>
shape. In
<normalizedToken id="53A2A0C8F73638BF115D9731DE492981" originalValue="Steins">Stein's</normalizedToken>
key (1911),
<taxonomicName id="9076E6DBD556F104604898AF099908C1" lsidName="C. delneneo" pageId="0" pageNumber="611" rank="species" species="delneneo">
<emphasis id="85C95840C6F304E75B18AC3ED06F3332" italics="true" pageId="0" pageNumber="611">C. delneneo</emphasis>
</taxonomicName>
comes close to
<taxonomicName id="DF82855F92C839648450F635472B4760" authorityName="Stein" authorityYear="1911" class="Insecta" family="Muscidae" genus="Coenosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coenosia pilitibia" order="Diptera" pageId="0" pageNumber="611" phylum="Arthropoda" rank="species" species="pilitibia">
<emphasis id="27353B9D83C91D735146093828D49957" italics="true" pageId="0" pageNumber="611">Coenosia pilitibia</emphasis>
</taxonomicName>
<bibRefCitation id="26EF9E8A6555AB4B23A0688F06E89189" author="Stein, P" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="0" pageNumber="611" pagination="61 - 189" refId="B67" refString="Stein, P, 1911. Die von Schnuse in Suedamerika gefangenen Anthomyiden. Archiv fuer Naturgeschichte 77: 61 - 189" title="Die von Schnuse in Suedamerika gefangenen Anthomyiden." volume="77" year="1911">Stein 1911</bibRefCitation>
, but differs in the chaetotaxy of the mid tibia. In
<normalizedToken id="8802C0A96104AF9E81EAFD9EBE1E2D84" originalValue="Malloch">Malloch'</normalizedToken>
key (1934),
<taxonomicName id="F379A890FDB710E17F848860C6E9894C" lsidName="C. delneneo" pageId="0" pageNumber="611" rank="species" species="delneneo">
<emphasis id="D57AC2E52EABBB1F3E6AFB31F0EAC99A" italics="true" pageId="0" pageNumber="611">C. delneneo</emphasis>
</taxonomicName>
comes close to
<taxonomicName id="1EC46C1CB6254B974E632011CE966EEC" lsidName="C. ignobilis" pageId="0" pageNumber="611" rank="species" species="ignobilis">
<emphasis id="4ACB0A9F857AF3370185FDABD34361F2" italics="true" pageId="0" pageNumber="611">C. ignobilis</emphasis>
</taxonomicName>
Stein, 1911, but the coloration of the fore femur and the shape of the abdomen can separate it from the latter.
</paragraph>
</subSubSection>
</treatment>
</document>