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<document id="07C7824CCCC52069FD96BB5E02982FB0" ID-CLB-Dataset="22377" ID-DOI="10.5852/ejt.2018.433" ID-GBIF-Dataset="6e9cff4b-1933-4022-99e2-b20ac41a65f1" ID-Zenodo-Dep="3787853" ID-ZooBank="F4E9A74F-FAAE-4CE4-A959-D86C9633882D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1588706205841" checkinUser="valdenar" docAuthor="Kantor, Yuri I., Fedosov, Alexander E., Snyder, Martin Avery &amp; Bouchet, Philippe" docDate="2018" docId="C84287C40C1BEB49FE6625FF19F97E67" docLanguage="en" docName="ejt-433_kantor_fedosov_snyder.pdf.imf" docOrigin="European Journal of Taxonomy 433" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Vermeijius retiarius Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet 2018, gen. et comb. nov." docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="347BFFBC0C2AEB7BFFB726081B367D67" masterDocTitle="Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="50" updateTime="1698838646784" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0B4C9C846E68AD41BA4FAF45C2D636B6">Pseudolatirus Bellardi, 1884 revisited, with the description of two new genera and five new species (Neogastropoda: Fasciolariidae)</mods:title>
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<mods:namePart id="ED7C91BAA87122E240ECC8F984AF1E6B">Kantor, Yuri I.</mods:namePart>
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<mods:affiliation id="2D2B099DD35CB6DD6D9EF0B8B7BD0097">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski prospect 33, 119071, Moscow, Russian Federation. Institut de Systématique, Evolution, Biodiversité ISYEB - UMR 7205 - CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 26, F- 75005, Paris, France. Academy of Natural Sciences of Drexel University, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103 - 1195, USA.</mods:affiliation>
<mods:nameIdentifier id="5B26EBCAB2F65154D9D9F2B9B6015DF1" type="email">kantor.yuri1956@gmail.com</mods:nameIdentifier>
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<mods:namePart id="75D1FD1639471F7C120D728E76BF15F2">Fedosov, Alexander E.</mods:namePart>
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<mods:affiliation id="E500C5B890E403B19812579EFB893A31">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski prospect 33, 119071, Moscow, Russian Federation. Institut de Systématique, Evolution, Biodiversité ISYEB - UMR 7205 - CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 26, F- 75005, Paris, France. Academy of Natural Sciences of Drexel University, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103 - 1195, USA.</mods:affiliation>
<mods:nameIdentifier id="B8B9AC0496222B53A79362714EAD6C8E" type="email">fedosovalexander@gmail.com</mods:nameIdentifier>
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<mods:namePart id="442BA1ED6514DB007AB2351F8B7D8967">Snyder, Martin Avery</mods:namePart>
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<mods:title id="DF7F007A20A0AD4A03F5C3F19655B6F8">European Journal of Taxonomy</mods:title>
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<mods:date id="BA8596907A6D04BBD3448FF3A57F31EF">2018</mods:date>
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<heading id="1B1C81BE0C1BEB4AFE6625FF1F557975" box="[465,1123,1015,1042]" centered="true" fontSize="11" level="2" pageId="49" pageNumber="50" reason="2">
<taxonomicName id="87EB4D510C1BEB4AFE6625FF18B37975" ID-CoL="5B3RS" authority="(Martens, 1901)" authorityName="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet" authorityYear="2018" baseAuthorityName="Martens" baseAuthorityYear="1901" box="[465,901,1015,1042]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="retiarius" status="gen. et comb. nov.">
<emphasis id="729FEAC00C1BEB4AFE6625FF19F67976" bold="true" box="[465,704,1015,1041]" italics="true" pageId="49" pageNumber="50">Vermeijius retiarius</emphasis>
(
<bibRefCitation id="247A4B230C1BEB4AFD7825F0184A7975" author="Martens E. von" box="[719,892,1016,1042]" pageId="49" pageNumber="50" pagination="14 - 26" refId="ref35411" refString="Martens E. von. 1901. Neue Meer-Conchylien von der deutschen Tiefsee-Expedition unter der Leitung von Prof. Chun. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin: 14 - 26." type="book chapter" year="1901">Martens, 1901</bibRefCitation>
)
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<taxonomicNameLabel id="69AC57BB0C1BEB4AFC3B25F01F557975" box="[908,1123,1015,1042]" pageId="49" pageNumber="50" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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<subSubSection id="08F165590C1BEB4AFD512214187B7951" box="[742,845,1052,1078]" pageId="49" pageNumber="50" type="description">
<paragraph id="405436D20C1BEB4AFD512214187B7951" blockId="49.[465,1123,1015,1078]" box="[742,845,1052,1078]" pageId="49" pageNumber="50">
<figureCitation id="D8D02A570C1BEB4AFD512214187B7951" box="[742,845,1052,1078]" captionStart="Fig" captionStartId="46.[189,231,1613,1639]" captionTargetBox="[246,1341,265,1590]" captionTargetId="figure@46.[246,1341,265,1590]" captionTargetPageId="46" captionText="Fig. 11. Species of Vermeijius gen. nov. A–E. Vermeijius pallidus (Kuroda &amp; Habe, 1961) gen. et comb. nov. A–B. South China Sea, DongSha 2014, stn CP4128, 20°42′ N, 116°08′ E, 420–444 m. A. SL 50.6 mm (MNHN IM-2013-50173). B. SL 56.6 mm (MNHN IM-2013-50174). C. Solomon Islands, SALOMON 2, stn CP2290, 08°40′ S, 157°32′ E, 384–418 m, SL 59.9 mm (MNHN IM-2007-32537). D–E. New Caledonia, CONCALIS, stn CP2962, Grand Passage, 18°59′ S, 163°08′ E, 580–647 m, SL 45.2 mm (MNHN IM-2007-35093). F–J. Vermeijius virginiae (Hadorn &amp; Fraussen, 2002) gen. et comb. nov. F. Holotype, SL 58.9 mm (MNHN IM-2000-9626). G–H. South Madagascar, ATIMO VATAE, stn CP3585, 25°32′ S; 44°16′ E, 549–576 m. G. SL 53.8 mm (MNHN IM-2009-15025). H. SL 52.6 mm (MNHN IM-2009-15026). I–J. South Madagascar, ATIMO VATAE, stn CP3591, 25°03′ S, 43°58′ E, 400–402 m, SL 56 mm (MNHN IM-2009-15021). K. Vermeijius retiarius (Martens, 1901) gen. et comb. nov., South Madagascar, ATIMO VATAE, stn CP3595, 25°35′ S, 44°15′ E, 821–910 m, SL 47.5 mm (MNHN IM-2009-15087). All specimens sequenced, except F. Figures A–D, F–I, K to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3787877" httpUri="https://zenodo.org/record/3787877/files/figure.png" pageId="49" pageNumber="50">Fig. 11K</figureCitation>
</paragraph>
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<paragraph id="405436D20C1BEB4AFF0A22571954791E" blockId="49.[189,610,1119,1145]" box="[189,610,1119,1145]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="60FB11FC0C1BEB4AFF0A22571954791E" box="[189,610,1119,1145]" pageId="49" pageNumber="50">
<emphasis id="729FEAC00C1BEB4AFF0A22571A24791E" box="[189,274,1119,1145]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="87EB4D510C1BEB4AFF0A22571A31791E" ID-CoL="831e346a-9e8f-4b98-8df3-fbd29be19e1a" box="[189,263,1119,1145]" class="Gastropoda" family="Fasciolariidae" genus="Fusus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="genus">Fusus</taxonomicName>
?
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<treatmentCitation id="C14A10C30C1BEB4AFEAE22571968791E" author="Martens E. von" box="[281,606,1119,1145]" page="22" pageId="49" pageNumber="50" year="1901">
<taxonomicName id="87EB4D510C1BEB4AFEAE22571968791E" ID-CoL="5B3RS" authority="Martens, 1901: 22" authorityName="Martens" authorityPageNumber="22" authorityYear="1901" box="[281,606,1119,1145]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="retiarius">
<emphasis id="729FEAC00C1BEB4AFEAE22571A4B791F" box="[281,381,1119,1144]" italics="true" pageId="49" pageNumber="50">retiarius</emphasis>
<bibRefCitation id="247A4B230C1BEB4AFE3322571968791E" author="Martens E. von" box="[388,606,1119,1145]" pageId="49" pageNumber="50" pagination="14 - 26" refId="ref35411" refString="Martens E. von. 1901. Neue Meer-Conchylien von der deutschen Tiefsee-Expedition unter der Leitung von Prof. Chun. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin: 14 - 26." type="book chapter" year="1901">Martens, 1901: 22</bibRefCitation>
</taxonomicName>
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.
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<treatmentCitationGroup id="60FB11FC0C1BEB4AFF0A22AA187179DC" box="[189,839,1185,1212]" pageId="49" pageNumber="50">
<emphasis id="729FEAC00C1BEB4AFF0A22AA1A4B79DC" box="[189,381,1186,1212]" italics="true" pageId="49" pageNumber="50">
<taxonomicName id="87EB4D510C1BEB4AFF0A22AA1A3179DB" ID-CoL="831e346a-9e8f-4b98-8df3-fbd29be19e1a" box="[189,263,1186,1212]" class="Gastropoda" family="Fasciolariidae" genus="Fusus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="genus">Fusus</taxonomicName>
?
<taxonomicName id="87EB4D510C1BEB4AFEAE22AA1A4B79DC" ID-CoL="5B3RS" authorityName="Martens" authorityYear="1901" box="[281,381,1186,1211]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="retiarius">retiarius</taxonomicName>
</emphasis>
–
<treatmentCitation id="C14A10C30C1BEB4AFE2D22AA198379DB" box="[410,693,1185,1212]" pageId="49" pageNumber="50">Martens 1904: 104–105</treatmentCitation>
, pl. 2, fig. 4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="405436D20C1BEB4AFF0A22CE1E6D7987" blockId="49.[189,1371,1185,1248]" box="[189,1371,1221,1248]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="60FB11FC0C1BEB4AFF0A22CE1E6D7987" box="[189,1371,1221,1248]" pageId="49" pageNumber="50">
<taxonomicName id="87EB4D510C1BEB4AFF0A22CE1AB379B8" ID-CoL="6K5SX" baseAuthorityName="Martens" baseAuthorityYear="1901" box="[189,389,1222,1247]" class="Gastropoda" family="Fasciolariidae" genus="Fusinus" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="retiarius">
<emphasis id="729FEAC00C1BEB4AFF0A22CE1AB379B8" box="[189,389,1222,1247]" italics="true" pageId="49" pageNumber="50">Fusinus retiarius</emphasis>
</taxonomicName>
–
<treatmentCitation id="C14A10C30C1BEB4AFE1422CD19CC79B8" author="Hadorn R. &amp; Fraussen K." box="[419,762,1221,1248]" page="64" pageId="49" pageNumber="50" year="2002">
<bibRefCitation id="247A4B230C1BEB4AFE1422CD19CC79B8" author="Hadorn R. &amp; Fraussen K." box="[419,762,1221,1248]" pageId="49" pageNumber="50" pagination="63 - 72" refId="ref34699" refString="Hadorn R. &amp; Fraussen K. 2002. Two new Fusinus from East Africa (Gastropoda: Fasciolariidae). [Dos nuevos Fusinus de Africa oriental (Gastropoda: Fasciolariidae)]. Iberus 20: 63 - 72." type="journal article" year="2002">Hadorn &amp; Fraussen 2002: 64</bibRefCitation>
</treatmentCitation>
, 66, 68, figs 3–8, 21. —
<treatmentCitation id="C14A10C30C1BEB4AFBAC22CD1E607987" author="Mallard D. &amp; Robin A." box="[1051,1366,1221,1248]" page="81" pageId="49" pageNumber="50" year="2017">
<bibRefCitation id="247A4B230C1BEB4AFBAC22CD1E607987" author="Mallard D. &amp; Robin A." box="[1051,1366,1221,1248]" pageId="49" pageNumber="50" refId="ref35336" refString="Mallard D. &amp; Robin A. 2017. Recent Fasciolariidae. Association Francaise de Conchiologie, Paris and Conchbooks, Harxheim." type="book" year="2017">Mallard &amp; Robin 2017: 81</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="08F165590C1BEB4AFF0A230019C07B55" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="405436D20C1BEB4AFF0A23001A5A7845" blockId="49.[189,364,1288,1314]" box="[189,364,1288,1314]" pageId="49" pageNumber="50">
<heading id="1B1C81BE0C1BEB4AFF0A23001A5A7845" bold="true" box="[189,364,1288,1314]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="729FEAC00C1BEB4AFF0A23001A5A7845" bold="true" box="[189,364,1288,1314]" pageId="49" pageNumber="50">Type material</emphasis>
</heading>
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<emphasis id="729FEAC00C1BEB4AFF5D23301A507835" bold="true" box="[234,358,1336,1362]" pageId="49" pageNumber="50">
<typeStatus id="9F5088700C1BEB4AFF5D23301A507835" box="[234,358,1336,1362]" pageId="49" pageNumber="50" type="lectotype">Lectotype</typeStatus>
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<paragraph id="405436D20C1BEB4AFF0A2354186378FC" blockId="49.[189,1399,1336,1435]" pageId="49" pageNumber="50">
<materialsCitation id="F0833C8F0C1BEB4AFF0A2354186378FC" ID-GBIF-Occurrence="2610407301" collectionCode="ZMB" country="Somalia" elevation="1134" latitude="1.8166666" location="southern Somalia" longLatPrecision="1308" longitude="45.483334" pageId="49" pageNumber="50" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="38FC76420C1BEB4AFF0A23541A717811" box="[189,327,1372,1398]" name="Somalia" pageId="49" pageNumber="50">SOMALIA</collectingCountry>
: dd, 38.5 × 15.7 mm, off
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,
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,
<geoCoordinate id="25DF50150C1BEB4AFC7C23541F057810" box="[971,1075,1370,1399]" degrees="45" direction="east" minutes="29" orientation="longitude" pageId="49" pageNumber="50" precision="925" value="45.483334">45°29′ E</geoCoordinate>
,
<quantity id="87139B370C1BEB4AFBF623541FAA7810" box="[1089,1180,1372,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.134" pageId="49" pageNumber="50" unit="m" value="1134.0">
<elevation id="CBC6D1E10C1BEB4AFBF623541FAA7810" box="[1089,1180,1372,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.134" pageId="49" pageNumber="50" unit="m" value="1134.0">1134 m</elevation>
</quantity>
,
<taxonomicName id="87EB4D510C1BEB4AFB1E23541E3F7811" box="[1193,1289,1372,1398]" class="Malacostraca" family="Trichodactylidae" genus="Valdivia" kingdom="Animalia" order="Decapoda" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="729FEAC00C1BEB4AFB1E23541E3F7811" box="[1193,1289,1372,1398]" italics="true" pageId="49" pageNumber="50">Valdivia</emphasis>
</taxonomicName>
, stn 256, designated by
<bibRefCitation id="247A4B230C1BEB4AFED12388199478FC" author="Hadorn R. &amp; Fraussen K." box="[358,674,1408,1435]" pageId="49" pageNumber="50" pagination="63 - 72" refId="ref34699" refString="Hadorn R. &amp; Fraussen K. 2002. Two new Fusinus from East Africa (Gastropoda: Fasciolariidae). [Dos nuevos Fusinus de Africa oriental (Gastropoda: Fasciolariidae)]. Iberus 20: 63 - 72." type="journal article" year="2002">Hadorn &amp; Fraussen (2002)</bibRefCitation>
(
<collectionCode id="26FAAE170C1BEB4AFD06238919C478FC" box="[689,754,1409,1435]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="49" pageNumber="50" type="Museum">ZMB</collectionCode>
61012).
</materialsCitation>
</paragraph>
<paragraph id="405436D20C1BEB4AFF0A23CB1A8278BA" blockId="49.[189,436,1475,1501]" box="[189,436,1475,1501]" pageId="49" pageNumber="50">
<heading id="1B1C81BE0C1BEB4AFF0A23CB1A8278BA" bold="true" box="[189,436,1475,1501]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="729FEAC00C1BEB4AFF0A23CB1A8278BA" bold="true" box="[189,436,1475,1501]" pageId="49" pageNumber="50">Sequenced material</emphasis>
</heading>
</paragraph>
<paragraph id="405436D20C1BEB4AFF0A23FB19C07B55" blockId="49.[189,1399,1521,1586]" pageId="49" pageNumber="50">
SOUTH
<collectingCountry id="38FC76420C1BEB4AFE9C23FB19367B6A" box="[299,512,1523,1549]" name="Madagascar" pageId="49" pageNumber="50">MADAGASCAR</collectingCountry>
: ATIMO VATAE:
<specimenCount id="56EDFD5B0C1BEB4AFD5023FB182D7B6A" box="[743,795,1523,1549]" pageId="49" pageNumber="50" type="larva">1 lv</specimenCount>
, stn CP3595, South of Pointe Barrow,
<geoCoordinate id="25DF50150C1BEB4AFAB223F91E477B6A" box="[1285,1393,1521,1549]" degrees="25" direction="south" minutes="35" orientation="latitude" pageId="49" pageNumber="50" precision="925" value="-25.583334">25°35′ S</geoCoordinate>
,
<geoCoordinate id="25DF50150C1BEB4AFF0A201D1A137B55" box="[189,293,1557,1586]" degrees="44" direction="east" minutes="15" orientation="longitude" pageId="49" pageNumber="50" precision="925" value="44.25">44°15′ E</geoCoordinate>
,
<quantity id="87139B370C1BEB4AFE85201F1A817B55" box="[306,439,1559,1586]" metricMagnitude="2" metricUnit="m" metricValue="8.655" metricValueMax="9.1" metricValueMin="8.209999999999999" pageId="49" pageNumber="50" unit="m" value="865.5" valueMax="910.0" valueMin="821.0">821–910 m</quantity>
(
<collectionCode id="26FAAE170C1BEB4AFE71201F19147B56" box="[454,546,1559,1585]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="49" pageNumber="50">MNHN</collectionCode>
IM-2009-15087).
</paragraph>
</subSubSection>
<subSubSection id="08F165590C1BEB49FF0A20511F437CF7" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
<paragraph id="405436D20C1BEB4AFF0A20511A787B14" blockId="49.[189,334,1625,1651]" box="[189,334,1625,1651]" pageId="49" pageNumber="50">
<heading id="1B1C81BE0C1BEB4AFF0A20511A787B14" bold="true" box="[189,334,1625,1651]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="729FEAC00C1BEB4AFF0A20511A787B14" bold="true" box="[189,334,1625,1651]" pageId="49" pageNumber="50">Description</emphasis>
</heading>
</paragraph>
<paragraph id="405436D20C1BEB4AFF0A208119AE7A8C" blockId="49.[189,1399,1673,2027]" pageId="49" pageNumber="50">
Shell large for genus, up to
<quantity id="87139B370C1BEB4AFE4B208119787BC3" box="[508,590,1673,1700]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="49" pageNumber="50" unit="mm" value="57.0">57 mm</quantity>
in length, strong, broadly fusiform, gradually constricting to medium long, sinuous, coaxial siphonal canal. Protoconch dome-shaped, paucispiral, of 1.5 convex smooth whorls, greyish, semitransparent, with 3 weak arcuate axial ribs at transition to teleoconch. Protoconch/ teleoconch discontinuity distinct, marked by appearance of teleoconch spiral sculpture. Protoconch diameter
<quantity id="87139B370C1BEB4AFE9B21131AAF7A52" box="[300,409,1819,1845]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="49" pageNumber="50" unit="mm" value="1000.0">1000 µm</quantity>
, exposed height
<quantity id="87139B370C1BEB4AFDD6211319F67A52" box="[609,704,1819,1845]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="49" pageNumber="50" unit="mm" value="700.0">700 µm</quantity>
. Teleoconch with strongly convex whorls. Aperture broad, oval, columella smooth. Spiral sculpture of moderately strong cords, beaded at intersection with axial ribs and much thinner, although distinct, secondary cords, covering interspaces between primary cords; 4–5 primary spiral cords on spire whorls, including penultimate. Axial ribs not aligned along spire, 11– 12 per whorl. Background colour pale flesh to nearly white. Operculum elongate, leaf-shaped, spanning nearly entire aperture. Nucleus terminal.
</paragraph>
<paragraph id="405436D20C18EB49FF0A27031F437CF7" blockId="50.[189,1399,267,401]" pageId="50" pageNumber="51">
Radula (
<bibRefCitation id="247A4B230C18EB49FE96270319607C42" author="Hadorn R. &amp; Fraussen K." box="[289,598,267,294]" pageId="50" pageNumber="51" pagination="63 - 72" refId="ref34699" refString="Hadorn R. &amp; Fraussen K. 2002. Two new Fusinus from East Africa (Gastropoda: Fasciolariidae). [Dos nuevos Fusinus de Africa oriental (Gastropoda: Fasciolariidae)]. Iberus 20: 63 - 72." type="journal article" year="2002">Hadorn &amp; Fraussen 2002</bibRefCitation>
: fig. 21) with medium broad, arcuate, lateral teeth, with attenuated outer anterior corner, producing medium long ‘handle’. Lateral teeth with five slightly recurved cusps, outermost longest, more distant from others. Additional innermost short, but distinct, cusp present. Central tooth small, subrectangular, with three very short cusps, medium longest.
</paragraph>
</subSubSection>
<subSubSection id="08F165590C18EB49FF0A27B018827F65" box="[189,948,440,514]" pageId="50" pageNumber="51" type="distribution">
<paragraph id="405436D20C18EB49FF0A27B01A607CB5" blockId="50.[189,342,440,466]" box="[189,342,440,466]" pageId="50" pageNumber="51">
<heading id="1B1C81BE0C18EB49FF0A27B01A607CB5" bold="true" box="[189,342,440,466]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="729FEAC00C18EB49FF0A27B01A607CB5" bold="true" box="[189,342,440,466]" pageId="50" pageNumber="51">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="405436D20C18EB49FF0A27E018827F65" blockId="50.[189,948,487,514]" box="[189,948,487,514]" pageId="50" pageNumber="51">
East Africa, from
<collectingCountry id="38FC76420C18EB49FE3827EF1AD97F66" box="[399,495,487,513]" name="Somalia" pageId="50" pageNumber="51">Somalia</collectingCountry>
to South
<collectingCountry id="38FC76420C18EB49FDD627EF19C77F66" box="[609,753,487,513]" name="Madagascar" pageId="50" pageNumber="51">Madagascar</collectingCountry>
, in
<quantity id="87139B370C18EB49FCAD27EF18997F65" box="[794,943,487,514]" metricMagnitude="2" metricUnit="m" metricValue="8.17" metricValueMax="11.34" metricValueMin="5.0" pageId="50" pageNumber="51" unit="m" value="817.0" valueMax="1134.0" valueMin="500.0">500–1134 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="08F165590C18EB49FF0A242119F97E67" pageId="50" pageNumber="51" type="discussion">
<paragraph id="405436D20C18EB49FF0A24211A197F24" blockId="50.[189,303,553,579]" box="[189,303,553,579]" pageId="50" pageNumber="51">
<heading id="1B1C81BE0C18EB49FF0A24211A197F24" bold="true" box="[189,303,553,579]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="729FEAC00C18EB49FF0A24211A197F24" bold="true" box="[189,303,553,579]" pageId="50" pageNumber="51">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="405436D20C18EB49FF0A245019F97E67" blockId="50.[189,1399,600,769]" pageId="50" pageNumber="51">
In the COI-based tree
<taxonomicName id="87EB4D510C18EB49FE7B245119807F15" authorityName="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet" authorityYear="2018" baseAuthorityName="Martens" baseAuthorityYear="1901" box="[460,694,601,626]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="retiarius" status="gen. et comb. nov.">
<emphasis id="729FEAC00C18EB49FE7B245119807F15" box="[460,694,601,626]" italics="true" pageId="50" pageNumber="51">Vermeijius retiarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AC57BB0C18EB49FD77245018A97F15" box="[704,927,600,626]" pageId="50" pageNumber="51" rank="species">gen. et comb. nov.</taxonomicNameLabel>
did not cluster with
<taxonomicName id="87EB4D510C18EB49FB2924511E2A7F15" authorityName="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet" authorityYear="2018" baseAuthorityName="Kuroda &amp; Habe" baseAuthorityYear="1961" box="[1182,1308,600,626]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="pallidus" status="gen. et comb. nov.">
<emphasis id="729FEAC00C18EB49FB2924511E2A7F15" box="[1182,1308,600,626]" italics="true" pageId="50" pageNumber="51">V. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AC57BB0C18EB49FA9124501A767FF1" pageId="50" pageNumber="51" rank="species">gen. et comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="87EB4D510C18EB49FECB247419357FF2" authorityName="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet" authorityYear="2018" box="[380,515,636,661]" class="Gastropoda" family="Fasciolariidae" genus="Fusinus" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="virginiae" status="gen. et comb. nov.">
<emphasis id="729FEAC00C18EB49FECB247419357FF2" box="[380,515,636,661]" italics="true" pageId="50" pageNumber="51">V. virginiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AC57BB0C18EB49FDBB247419DC7FF1" box="[524,746,636,662]" pageId="50" pageNumber="51" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, but in the three genes tree it was recovered as a sister group to the clade combining these two species. The relationships are highly supported in the Bayesian analysis, therefore we include it in
<taxonomicName id="87EB4D510C18EB49FDEA24CC19EF7FBA" authority="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet, 2018" authorityName="Kantor &amp; Fedosov &amp; Snyder &amp; Bouchet" authorityYear="2018" box="[605,729,708,733]" class="Gastropoda" family="Fasciolariidae" genus="Vermeijius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neogastropoda" pageId="50" pageNumber="51" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="729FEAC00C18EB49FDEA24CC19EF7FBA" box="[605,729,708,733]" italics="true" pageId="50" pageNumber="51">Vermeijius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AC57BB0C18EB49FD5624CB187B7FBA" box="[737,845,707,733]" pageId="50" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
It differs from the two sequenced species by a more stout shell with paucispiral protoconch.
</paragraph>
</subSubSection>
</treatment>
</document>
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