287 lines
35 KiB
XML
287 lines
35 KiB
XML
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<mods:title id="5DF4AB91CC9AA3D80BE0F7FB16A81F03">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name id="FFBBDF45AB4E8F8E295379F09B87D04B" type="personal">
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<mods:namePart id="8DF2A7B1DD69E10ED57F084D7C3BACFF">Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation id="FC0C75ACB38B382BBEB7F5F798BD4B89">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier id="D2C5833D945265163C08200E430F8AC1" type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8A4D8DAE0B7A6D237038F0173616558C">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier id="A3DEECEFA69882BA196E22B2C32A41FE" type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation id="FF0FF7CEAE92802B5AFA15C7DA32CEA6">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="EF766B5CFEBE3F37C7BB6CD2FF5D28B1">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="90123AEF36DC1B45979EF3CBDB5A1C41">
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<mods:date id="23D0FF1FE010749405382CE19DE4DABF">2022</mods:date>
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<mods:number id="0329EF0925C5EEE45FD56784BE417004">80</mods:number>
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<treatment id="9116DAEFC376544EBEC3CF0CDEAAD952" ID-GBIF-Taxon="199468673" LSID="urn:lsid:plazi:treatment:9116DAEFC376544EBEC3CF0CDEAAD952" httpUri="http://treatment.plazi.org/id/9116DAEFC376544EBEC3CF0CDEAAD952" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
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<subSubSection id="984891EF668D1AA1C95914D24EE98DE1" pageId="0" pageNumber="309" type="nomenclature">
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<paragraph id="9407E7115695B59F3CE28CBC1F61DD87" pageId="0" pageNumber="309">
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3.43.
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<taxonomicName id="58EA461EEF5D0D41621AFC8476483207" ID-CoL="9X65T" LSID="9116DAEF-C376-544E-BEC3-CF0CDEAAD952" authority="(Roewer, 1929)" baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">Metasarcus clavifemur (Roewer, 1929)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="9FF4D8E232E571A76F401098DE40EA05" pageId="0" pageNumber="309" type="description">
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<paragraph id="4680AEDC50242A075389D5F74F97E47F" pageId="0" pageNumber="309">
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<figureCitation id="5102AC64C54B05E619D8EDE47844F9A4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5D</figureCitation>
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<figureCitation id="2A011895FF8617F57B00468FFD4265D7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10E, F</figureCitation>
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<figureCitation id="1D4E52C215C1D8838D921876CA2AF7C2" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">, 19D-F</figureCitation>
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<figureCitation id="92408001D0E4E68A3255828BC675B507" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">, 25C, D</figureCitation>
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<figureCitation id="5D41D292E50F0BF2704EC91D4F5E4FAB" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6010B82F9FB30471BA17B68F86A4A226" pageId="0" pageNumber="309" type="reference_group">
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<paragraph id="553D4759B6F1E7FF92FA15AD7F1FC50A" pageId="0" pageNumber="309">
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<taxonomicName id="D33F09EACF23DC88CBC4B70900ADBA16" authorityName="Roewer" authorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Chacoikeontus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chacoikeontus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">Chacoikeontus clavifemur</taxonomicName>
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Roewer, 1929: 279 (desc.), fig. 45;
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<bibRefCitation id="4F9EF4B3A37852C91A1B66D1149FA113" author="Soares, HEM" journalOrPublisher="Revista Brasileira de Entomologia" pageId="0" pageNumber="309" pagination="1 - 14" refId="B116" refString="Soares, HEM, Soares, BAM, Jim, RLS, 1992. Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae). Revista Brasileira de Entomologia 36 (1): 1 - 14" title="Monografia dos generos de opilioes neotropicos IV. (Opiliones, Gonyleptidae, Prostygninae)." volume="36" year="1992">Soares et al. 1992</bibRefCitation>
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: 3 (cat.);
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<bibRefCitation id="80257F2ACF41A524A79BF13D636F2A8A" DOI="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" author="Kury, AB" journalOrPublisher="Zoo-logical Journal of Linnean Society" pageId="0" pageNumber="309" pagination="143 - 162" refId="B58" refString="Kury, AB, Maury, EA, 1998. A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae). Zoo-logical Journal of Linnean Society 123: 143 - 162, DOI: https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" title="A new genus and five new species of Metasarcinae from Peru (Arachnida, Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.1998.tb01297.x" volume="123" year="1998">Kury and Maury 1998</bibRefCitation>
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: 145 (cit.);
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<bibRefCitation id="6667B760D0569F4B1E406C2CAFF61F34" author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 144 (cat.);
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<bibRefCitation id="09623DFC3DB2F44456CEDBFEF4478EAE" DOI="https://doi.org/10.1111/j.1096-3642.2008.00484.x" author="Yamaguti, HY" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="309" pagination="319 - 362" refId="B127" refString="Yamaguti, HY, Pinto-da-Rocha, R, 2009. Taxonomic review of Bourguyiinae, cladistic analysis, and a new hypothesis of biogeographic relantioships of the Brazilian Atlantic Rainforest (Arachnida: Opiliones, Gonyleptidae). Zoological Journal of the Linnean Society 156: 319 - 362, DOI: https://doi.org/10.1111/j.1096-3642.2008.00484.x" title="Taxonomic review of Bourguyiinae, cladistic analysis, and a new hypothesis of biogeographic relantioships of the Brazilian Atlantic Rainforest (Arachnida: Opiliones, Gonyleptidae)." url="https://doi.org/10.1111/j.1096-3642.2008.00484.x" volume="156" year="2009">Yamaguti and Pinto-da-Rocha 2009</bibRefCitation>
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: 321 (syst.); Kury & Villarreal, 2015: 4-5 (cit.), fig 7 (penis).
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</paragraph>
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<paragraph id="CA25DCFBF7EB9510468FABF3BD04F25A" pageId="0" pageNumber="309">
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<taxonomicName id="4643EE3B2729BD099F304E01F59EE8E1" baseAuthorityName="Roewer" baseAuthorityYear="1929" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus clavifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="clavifemur">Metasarcus clavifemur</taxonomicName>
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:
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<bibRefCitation id="9261601F3C1533BFB929EE82E8E485B1" DOI="https://doi.org/10.1016/j.jcz.2018.12.005" author="Townsend, VR" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="309" pagination="101 - 109" refId="B120" refString="Townsend, VR, Perez-Gonzalez, A, Proud, DN, 2019. Putative mating plugs of harvestmen (Opiliones, Laniatores). Zoologischer Anzeiger 278: 101 - 109, DOI: https://doi.org/10.1016/j.jcz.2018.12.005" title="Putative mating plugs of harvestmen (Opiliones, Laniatores)." url="https://doi.org/10.1016/j.jcz.2018.12.005" volume="278" year="2019">Townsend et al. 2019</bibRefCitation>
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: 102 (cit), 105-106 (biol), fig. 2 (ovipositor).
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</paragraph>
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</subSubSection>
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<subSubSection id="26A110141F3DFFEF1C5BDF95A541D8C5" pageId="0" pageNumber="309" type="redescription">
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<paragraph id="395CBF6E1A9037AC2C579FD4552606EC" pageId="0" pageNumber="309">Redescription.</paragraph>
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<paragraph id="10FD18167EB8ACA29555020354C825A2" pageId="0" pageNumber="309">
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<emphasis id="4D0E791FAADD971C32F6A1BEE39830FB" bold="true" pageId="0" pageNumber="309">
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MALE:
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<emphasis id="0E949DC08D7D757D61B48A4A2B50580B" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis id="59374347A04478F882DF334CE5C442D4" italics="true" pageId="0" pageNumber="309">n</emphasis>
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=5) DSW: 4.6-5.1 (5.0); DSL: 4.3-5.1 (5.1); CL: 1.7-2.0 (1.9). FIVL: 6.6-10.6 (10.6). ChL: 3.1-5.3.
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<emphasis id="581A68E8F64233A9F1B30947058FE605" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
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: (Fig.
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<figureCitation id="C55FD6E84DB5372877C8223B9C7960B7" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">25C</figureCitation>
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) Chelicerae reddish brown with black reticulate pattern.
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<taxonomicName id="292385BCA1B6AA35D2E23A658E5CD13B" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">Pedipalpus</taxonomicName>
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and legs I-III reddish brown. Lateral and posterior margin of DS, free tergites I-III and other segments of legs, dark brown. Anterior margin of DS, carapace and areas I-IV, brown. Small males in a lighter brown tone.
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<emphasis id="7FCFC7094DF32CA05646A1F13EFC63ED" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
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: (Fig.
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<figureCitation id="02D36457EB8735ABA6CDAE68A660B4D6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5D</figureCitation>
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) Kappa-type DSS, with almost straight posterior margin of DS; constriction I not so well marked, constriction II absent, mid-bulge slightly larger than carapace and coda undefined, coalescing with mid-bulge. Ocularium with small granules grouped together close to the eyes. Carapace irregularly granulate, with granules concentrating mainly on the anterior margin of DS, and the median elevation and behind the ocularium. DS with four well defined areas. Area I divided by the scutal grooves I and II; with five pairs of small tubercles. Area II moderately projected toward area I; with seven small tubercles. Area III with two spines, directed posteriorly; seven small tubercles. Area IV with six small tubercles. Lateral margins of DS with granules throughout their length. Posterior margin of DS and free tergites I-III with few granules distributed irregularly.
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<emphasis id="C470E0F5E61CC3B8D4AAEFB2C202EF77" bold="true" italics="true" pageId="0" pageNumber="309">Chelice-rae</emphasis>
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: (Fig.
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<figureCitation id="E2AE69F52C2A700E288B92457D3C6E45" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5D</figureCitation>
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) Segment I with four basal tubercles of different sizes. Segment II with setiferous granules on the front surface; finger with three teeth more apical and a row of tiny denticles. Segment III with three apical teeth, a row of denticles and a large apical tooth, straight apex.
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<taxonomicName id="FA98857CE677963529423AB730E14FAB" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
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<emphasis id="0A481C36C18AF714D30B1FDF7249F1F8" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
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</taxonomicName>
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: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of five spines, the most basal and more apical of which are smallest; granules scattered on prolateral surface. Patella with a proapical spine. Tibia: retrolateral iiIIi, prolateral IiIi. Tarsus: retrolateral iIiIi, prolateral IiIi.
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<emphasis id="53AA43DAE92CE6C1DF843B862AA489CF" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
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: Coxa I with a median row of four setiferous tubercles. Coxae II-IV with sparse granules. Rows of tubercles between coxae II-III and III-IV. Smooth genital area. Free sternites with rows of setiferous granules. Anal operculum granulate.
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<emphasis id="1D0F9701CAED5945849AB6A4BCB027DE" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
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: (Figs
|
||
<figureCitation id="FE2015FC30DDFDBDB618B03A501A3EA8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5D</figureCitation>
|
||
,
|
||
<figureCitation id="7B32B9F582539C095B488A860D217D1D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10E, F</figureCitation>
|
||
) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV unarmed, with granules along all extension. Trochanters I-III smooth and unarmed. Trochanter IV granular with a long retroapical spiniform apophysis and two small dorsoapical tubercles. Femora I-III unarmed and with few sparse granules. Femur IV swollen, with a retrobasal spiniform apophysis; a retrodorsal row of broad, conical tubercles, decreasing in size apically; a row of granules located prolaterally regarding retrodorsal row of tubercles along basal
|
||
<normalizedToken id="48FB983C9D9983DA15E269B2E238FDD0" originalValue="½">1/2</normalizedToken>
|
||
; a prolateral row of tubercles irregularly shaped along basal
|
||
<normalizedToken id="CD6E3FCE9534572AF93DB2E6C60A7FBF" originalValue="½">1/2</normalizedToken>
|
||
; a proventral row of small conical setiferous tubercles and a retroventral row of tiny granules. Patellae I-IV unarmed and with few granules in greater quantity in the patella IV. Tibiae I-III smooth and unarmed. Tibia IV with a retrolateral row of small tubercles. Tarsal segmentation: (
|
||
<emphasis id="6732A0FDAE0D5ACF2E9A5BBD71C8E270" italics="true" pageId="0" pageNumber="309">n</emphasis>
|
||
=5) 7-8 (7), 12-13 (?), 9-10 (10), 10-11 (11).
|
||
<emphasis id="F5A9F3BCED1F76948ABC7FE6AF28DB46" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
|
||
: (Fig.
|
||
<figureCitation id="CE458F0B1ECEEB69CA4B557DE0FBC09A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">19D-F</figureCitation>
|
||
) VP rectangular; distal margin slightly concave. MS C1-C3 apical long and straight; MS A1 sub basal long and straight (smaller than MS C); MS B1 basal short and straight; MS D1 short and conical, laterally placed, near MS C; MS E1 median short and straight, placed more ventrally than MS C and D. Lateral sacs long and apically tapered, with long T3-like microsetae, with short base. Stylus with broad apex, with apical projections. Dorsal process absent. Promontory convex. -
|
||
<emphasis id="91F124F49225D8624128626AF9FF560A" bold="true" pageId="0" pageNumber="309">
|
||
FEMALE:
|
||
<emphasis id="26D7564995C0BE6BB5801A6EF56FE1CD" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
|
||
</emphasis>
|
||
(
|
||
<emphasis id="7E0AD19C1EC971344CAE75A160BDE6BF" italics="true" pageId="0" pageNumber="309">n</emphasis>
|
||
=7) DSW: 4.6-5.1; DSL: 4.0-5.0; CL: 1.5-1.9. FIVL: 8.3-9.6. ChL: 1.7-2.5. (Fig.
|
||
<figureCitation id="1DD9879A96908CF8C9E5B6BFDB4167A3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">25D</figureCitation>
|
||
) Chelicerae not swollen. Trochanter IV unarmed. Femur IV with a small tubercle rather than retroapical apophysis present in males; rows of tubercles and granules absent; longer relative length to that of DS than in male. Color similar to small males, in a lighter brown tone. Lateral and posterior margin of DS and free tergites lighter than larger males. Tarsal segmentation: (
|
||
<emphasis id="DC408E11FFF2CBFAC22D14749240BC76" italics="true" pageId="0" pageNumber="309">n</emphasis>
|
||
=7) 7, 12-14, 10-11, 11-12.
|
||
</paragraph>
|
||
<caption id="207A750127B7E68FB2849242E0200884" doi="10.3897/asp.80.e73829.figure27" httpUri="https://binary.pensoft.net/fig/729901" pageId="0" pageNumber="309" start="Figure 27" startId="F27">
|
||
<paragraph id="1FC735B0ECD0E789C4F6B90B2C352791" pageId="0" pageNumber="309">
|
||
<emphasis id="7BC9CB6D913A55B9A6AA2F3284E98906" bold="true" pageId="0" pageNumber="309">Figure 27.</emphasis>
|
||
Live specimens of
|
||
<taxonomicName id="B8B2EB3A7A53A5ADA58F5314BA5E8851" authorityName="Roewer" authorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Tschaidicancha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tschaidicancha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F6E8771738A87A47395F6781C112666" italics="true" pageId="0" pageNumber="309">Tschaidicancha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="6754232E1C7F27E0A176144BC2378833" bold="true" pageId="0" pageNumber="309">A</emphasis>
|
||
<taxonomicName id="6F262EF25E92B857E1BCC7EA62A3E30D" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. joseochoai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="joseochoai">
|
||
<emphasis id="787EE37C8642CA1E28459254BC27F834" italics="true" pageId="0" pageNumber="309">T. joseochoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B6859434559DCC791E19629F85E4EFA1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
, male;
|
||
<emphasis id="259E083AEC474AAF812B06A816585607" bold="true" pageId="0" pageNumber="309">B</emphasis>
|
||
female;
|
||
<emphasis id="C3397F56FBCB05BAF75970C1766C5A16" bold="true" pageId="0" pageNumber="309">C</emphasis>
|
||
<taxonomicName id="3E013485A733C8BDDCDB5F16B3EB329E" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. chaplini" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="chaplini">
|
||
<emphasis id="3ADF76C75CEBA0730B38ED42481D22D0" italics="true" pageId="0" pageNumber="309">T. chaplini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B4EE5DA75BB68AFDCC3689ED16124CE" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
|
||
, male;
|
||
<emphasis id="B132A29A499A79881A3DAA12EDA5E707" bold="true" pageId="0" pageNumber="309">D</emphasis>
|
||
female;
|
||
<emphasis id="E5CF2023F30DA37E3909A21CFA374114" bold="true" pageId="0" pageNumber="309">E</emphasis>
|
||
<taxonomicName id="ED6914732ECEC1B683EED2FD69360DB3" class="Arachnida" family="Metasarcidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. weyrauchi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="weyrauchi">
|
||
<emphasis id="18AA43FF29CEC4C00ACB2E15394F8025" italics="true" pageId="0" pageNumber="309">T. weyrauchi</emphasis>
|
||
</taxonomicName>
|
||
Roewer, 1957, male;
|
||
<emphasis id="DCC7A18F6CFDA811FCABA7FC669F553C" bold="true" pageId="0" pageNumber="309">F</emphasis>
|
||
: female.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="83FBFC90B718B670F9D1B89258BF94AA" pageId="0" pageNumber="309" type="diagnosis">
|
||
<paragraph id="2DA7C5E2D8B732F1481481AC3DF188C3" pageId="0" pageNumber="309">Diagnosis.</paragraph>
|
||
<paragraph id="4B9965775A3AD4A590D1BC4E71ECAC02" pageId="0" pageNumber="309">
|
||
Differs from other species of the genus by having femur IV swollen and with a long retrobasal apophysis; trochanter IV with long retroapical spiniform apophysis (Fig.
|
||
<figureCitation id="F101C41F4EC4D4FA8EFB4C2ACD3467B7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78D088787C982A420320D44C312A718A" pageId="0" pageNumber="309" type="remarks">
|
||
<paragraph id="A169E2834DD44E75D3440EFFEB119548" pageId="0" pageNumber="309">Remarks.</paragraph>
|
||
<paragraph id="4E145817E281A9D2F2A8EE63AB919AAD" pageId="0" pageNumber="309">See the remarks on section 3.39 for an observation on the recent taxonomic history of the species. Considering that the type material is formed from syntypes, we chose the SMF specimen as the lectotype, in accordance with article 74 of the ICZN. The MNHN material is designated as a paralectotype.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FBA3CE9101D4B3A729E0766377F6D3B" pageId="0" pageNumber="309" type="distribution">
|
||
<paragraph id="6FD380CCB60232704AEC8D0805F8AA53" pageId="0" pageNumber="309">Distribution.</paragraph>
|
||
<paragraph id="B991476991651290FD7F2E28B713401F" pageId="0" pageNumber="309">
|
||
(Fig.
|
||
<figureCitation id="A6B34CA0A6C51D75365F7C47ADBFD604" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
|
||
) BOLIVIA. La Paz. Coroico. La Paz. Yanacachi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E68D713669FE1ADFEE231DF11E834DA6" pageId="0" pageNumber="309" type="materials_examined">
|
||
<paragraph id="EF25EB96096BC880625AF3C19449C9B9" pageId="0" pageNumber="309">Material examined.</paragraph>
|
||
<paragraph id="E97780C4A3B742556CAA8330435A4E29" pageId="0" pageNumber="309">
|
||
<emphasis id="D34BD9C9BB57FCE95E9A6EA782313B51" bold="true" pageId="0" pageNumber="309">
|
||
<emphasis id="22278230EDC3AD678D427E7966E39C4C" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
|
||
:
|
||
<materialsCitation id="85D265842EDC32C4A1B9F815836FD9FA" ID-GBIF-Occurrence="3889641357" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="BOLIVIA" location="La Paz" specimenCount="♂" stateProvince="La Paz" typeStatus="Lectotype">
|
||
<typeStatus id="F7FB1A21D7D17037FEC394F3EA756142">Lectotype</typeStatus>
|
||
<specimenCount id="94488A2E56FDFBDAC9983479ACB81509">♂</specimenCount>
|
||
, '
|
||
<collectingCountry id="21BC5E26B44CA12AD449C9852381B061" name="Bolivia">BOLIVIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="36C200A37B67FCFD3F08C93F4AC8E7E4" country="Bolivia" name="La Paz">La Paz</collectingRegion>
|
||
. Without date and collector data.' (SMF 1006 / SMF 1467/11);
|
||
<emphasis id="4C375C5FCFEE5599255CC2C4D7277EA0" bold="true" pageId="0" pageNumber="309">
|
||
<typeStatus id="F445ED565ACEF7279F079958C689C25F">paralectotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="965C90FE8EE548BDA898E5843B14A87B">♂</specimenCount>
|
||
, 'Without date and collector data.' (MNHN; male not examined)
|
||
</materialsCitation>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="9D5460EDD4928C3AB56EA26850484FC1" bold="true" italics="true" pageId="0" pageNumber="309">Additional material</emphasis>
|
||
:
|
||
<materialsCitation id="FCC3D8C5EB7A3C670A04A12046726004" ID-GBIF-Occurrence="3889641319" collectingDate="2010-11-30" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, A. Saravia" country="BOLIVIA" latitude="-16.215834" location="Road La Paz-Coroico" longLatPrecision="15" longitude="-67.823524" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="La Paz">
|
||
<specimenCount id="081B1F699D3961079E6FBAA6CFF97C37" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="378D441E9CA7C80C8F94228D0B02EC6C" type="female">2 ♀</specimenCount>
|
||
'
|
||
<collectingCountry id="866AFE5107B0D96D7CBEAC239AB59953" name="Bolivia">BOLIVIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="B60AF849C00CB83A68E746320E5DABE3" country="Bolivia" name="La Paz">La Paz</collectingRegion>
|
||
,
|
||
<location id="A6D2D2527DC2B8B442B68A7AEE1A0C26" LSID="urn:lsid:plazi:treatment:9116DAEFC376544EBEC3CF0CDEAAD952:A6D2D2527DC2B8B442B68A7AEE1A0C26" country="BOLIVIA" latitude="-16.215834" longLatPrecision="15" longitude="-67.823524" name="Road La Paz-Coroico" stateProvince="La Paz">Road La Paz-Coroico</location>
|
||
,
|
||
<geoCoordinate id="2D19CC6F440A19CD21E3306F1F6BB6EF" degrees="16" direction="south" minutes="12" orientation="latitude" precision="15" seconds="57" value="-16.215834">16°12′57″S</geoCoordinate>
|
||
<geoCoordinate id="BFECBFEA58E3BE9D20B37F2583D0FD5E" degrees="67" direction="west" minutes="49" orientation="longitude" precision="1" seconds="24.7" value="-67.823524">67°49′24.7″W</geoCoordinate>
|
||
,
|
||
<collectingDate id="9D1EF3779D52092044F3FADE0FCF7761" value="2010-11-30">30/XI/2010</collectingDate>
|
||
,
|
||
<collectorName id="F662FD36FA63A024E66C47D92329365F">R. Pinto-da-Rocha</collectorName>
|
||
,
|
||
<collectorName id="DF5088D63005096DB75AA3679C096C81">A. Benedetti</collectorName>
|
||
,
|
||
<collectorName id="FD20D38C3CA836ED6FAF40DCEF589E75">J. Ochoa</collectorName>
|
||
&
|
||
<collectorName id="94E88ED7945511E3F83B105408EBF958">A. Saravia</collectorName>
|
||
leg. (CBF)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="1310FA098867213477546EA5B265D415" type="male">2 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="17A75CB1A9A5E1DAE1D32DFB9E546635" type="female">2 ♀</specimenCount>
|
||
,
|
||
<normalizedToken id="2206E885C8A7C27A28BF3F6B861B5999" originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(MZSP 36985).
|
||
<materialsCitation id="F30B09C30A51BDC57B3C789235BD6219" ID-GBIF-Occurrence="3889641352" collectingDate="2010-12-12" collectorName="R. Pinto-da-Rocha, A. Benedetti, A. Saravia" country="BOLIVIA" latitude="-16.398472" location="Yanacachi" longLatPrecision="1" longitude="-67.73655" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="La Paz">
|
||
<specimenCount id="70E8C0602F8C430626571D227AAC8FB3" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="18CC2A2A90F9DCDD5AC16AD128F9BF17" type="female">1 ♀</specimenCount>
|
||
'
|
||
<collectingCountry id="FC902F64F9C4F1D90C1EC05541E4B7CB" name="Bolivia">BOLIVIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="A48D283BBB53BAB74DE4BC6B79599340" country="Bolivia" name="La Paz">La Paz</collectingRegion>
|
||
,
|
||
<location id="46DDFD4F42F3875D4CDAC81787F33FA2" LSID="urn:lsid:plazi:treatment:9116DAEFC376544EBEC3CF0CDEAAD952:46DDFD4F42F3875D4CDAC81787F33FA2" country="BOLIVIA" latitude="-16.398472" longLatPrecision="1" longitude="-67.73655" name="Yanacachi" stateProvince="La Paz">Yanacachi</location>
|
||
,
|
||
<geoCoordinate id="B0F7D6C5249066A6DB835D308887F309" degrees="16" direction="south" minutes="23" orientation="latitude" precision="1" seconds="54.5" value="-16.398472">16°23′54.5″S</geoCoordinate>
|
||
<geoCoordinate id="9D26F0B68AF6ED986AC12BEFCC577BA4" degrees="67" direction="west" minutes="44" orientation="longitude" precision="1" seconds="11.6" value="-67.73655">67°44′11.6″W</geoCoordinate>
|
||
,
|
||
<collectingDate id="683923F7CA0C055A8C927985F6510D1B" value="2010-12-12">12/XII/2010</collectingDate>
|
||
,
|
||
<collectorName id="E14F775EFC11E1FCA10505A0BC868447">R. Pinto-da-Rocha</collectorName>
|
||
,
|
||
<collectorName id="B6C2611B92AF347571E05114757065BB">A. Benedetti</collectorName>
|
||
&
|
||
<collectorName id="7C85DE34BC547C334FB9A5B89D33D78C">A. Saravia</collectorName>
|
||
leg. (CBF)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="C9F6BD4328B225E48217F0566D7A4A24" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="78CEC3D2890F41CB5604D75808D4B521" type="female">2 ♀</specimenCount>
|
||
,
|
||
<normalizedToken id="E089555FE1C244CA302292A3B3CDF64A" originalValue="‘ditto’">'ditto'</normalizedToken>
|
||
(MZSP 36986).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |