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<document id="0A750E606A43E659ED6E8E139CB81115" ID-CLB-Dataset="77862" ID-DOI="10.11646/zootaxa.5155.1.6" ID-GBIF-Dataset="dac9d493-3f9f-4432-a7cf-b6987cac23a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6669316" ID-ZooBank="28658B55-F7E0-42FE-B957-F8E0BDC45472" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1655713392415" checkinUser="plazi" docAuthor="Selis, Marco &amp; Femia, Arturo" docDate="2022" docId="FA2E87969E12FFF9FF40568BFEE3FCF7" docLanguage="en" docName="zootaxa.5155.1.6.pdf" docOrigin="Zootaxa 5155 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calligaster williamsi Bequaert 1940" docType="treatment" docVersion="8" lastPageNumber="129" masterDocId="0617FFEE9E17FFFCFFD7561CFFA4FF88" masterDocTitle="The genus Calligaster de Saussure (Hymenoptera: Vespidae: Zethinae) in the Philippine Islands" masterLastPageNumber="132" masterPageNumber="124" pageNumber="129" updateTime="1699339404697" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="88965F45D5F568E7778DD89C01EE0CBB">The genus Calligaster de Saussure (Hymenoptera: Vespidae: Zethinae) in the Philippine Islands</mods:title>
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<mods:namePart id="9E06658ABEC01B62356DC227CCA833A4">Selis, Marco</mods:namePart>
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<mods:namePart id="0E31983F234369EA7240911EF4D6FF1B">Femia, Arturo</mods:namePart>
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<mods:affiliation id="841538F7E804C4CCBF62326BBB74B26F">Via G. Gozzi, 15 - 50141 Firenze, Italy. arfem 93 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6571 - 2741</mods:affiliation>
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<treatment id="FA2E87969E12FFF9FF40568BFEE3FCF7" ID-DOI="http://doi.org/10.5281/zenodo.6684419" ID-GBIF-Taxon="196086733" ID-Zenodo-Dep="6684419" LSID="urn:lsid:plazi:treatment:FA2E87969E12FFF9FF40568BFEE3FCF7" httpUri="http://treatment.plazi.org/id/FA2E87969E12FFF9FF40568BFEE3FCF7" lastPageNumber="129" pageId="5" pageNumber="129">
<subSubSection id="3A9D650B9E12FFF9FF40568BFDF0FF39" box="[151,596,151,178]" pageId="5" pageNumber="129" type="nomenclature">
<paragraph id="723836809E12FFF9FF40568BFDF0FF39" blockId="5.[151,596,151,214]" box="[151,596,151,178]" pageId="5" pageNumber="129">
<heading id="297081EC9E12FFF9FF40568BFDF0FF39" bold="true" box="[151,596,151,178]" fontSize="11" level="1" pageId="5" pageNumber="129" reason="1">
<taxonomicName id="B5874D039E12FFF9FF40568BFDF0FF39" ID-CoL="PQ3M" authority="Bequaert, 1940" authorityName="Bequaert" authorityYear="1940" box="[151,596,151,178]" class="Insecta" family="Eumenidae" genus="Calligaster" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="40F3EA929E12FFF9FF40568BFDF0FF39" bold="true" box="[151,596,151,178]" pageId="5" pageNumber="129">
<emphasis id="40F3EA929E12FFF9FF40568BFE34FF3A" bold="true" box="[151,400,151,178]" italics="true" pageId="5" pageNumber="129">Calligaster williamsi</emphasis>
<bibRefCitation id="16164B719E12FFF9FE40568BFDF0FF39" author="Bequaert, J." box="[407,596,151,177]" pageId="5" pageNumber="129" pagination="124 - 126" refId="ref4132" refString="Bequaert, J. (1940) A new species of Calligaster from the Philippine Islands (Hymenoptera, Vespidae, Subfam. Zethinae). The Pan-Pacific Entomologist, 16, 124 - 126." type="journal article" year="1940">Bequaert, 1940</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="3A9D650B9E12FFF9FF4056A7FED1FF5D" box="[151,373,187,214]" pageId="5" pageNumber="129" type="description">
<paragraph id="723836809E12FFF9FF4056A7FED1FF5D" blockId="5.[151,596,151,214]" box="[151,373,187,214]" pageId="5" pageNumber="129">
(
<figureCitation id="EABC2A059E12FFF9FF4856A7FF4BFF5D" box="[159,239,187,213]" captionStart="FIGURES 12" captionStartId="2.[151,264,1809,1833]" captionTargetBox="[168,1419,630,1769]" captionTargetId="figure-245@2.[154,1433,630,1782]" captionTargetPageId="2" captionText="FIGURES 12. 1, Calligaster ilocana Selis, sp. nov., male paratype, habitus. 2, Calligaster williamsi Bequaert, 1940, male, habitus." figureDoi="http://doi.org/10.5281/zenodo.6669322" httpUri="https://zenodo.org/record/6669322/files/figure.png" pageId="5" pageNumber="129">Figs. 2</figureCitation>
,
<figureCitation id="EABC2A059E12FFF9FF2C56A7FEE6FF5D" box="[251,322,187,214]" captionStart="FIGURES 38" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-17@4.[197,1390,181,1919]" captionTargetPageId="4" captionText="FIGURES 38. 34, 7, Calligaster ilocana Selis, sp. nov. 56, 8, Calligaster williamsi Bequaert, 1940. 3, 5, Head of male in frontal view. 4, 6, Head of female in frontal view. 78, Mesoscutum and scutellum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6669324" httpUri="https://zenodo.org/record/6669324/files/figure.png" pageId="5" pageNumber="129">56, 8</figureCitation>
,
<figureCitation id="EABC2A059E12FFF9FE9856A7FEC8FF5D" box="[335,364,187,213]" captionStart="FIGURES 910" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@6.[225,1363,181,1920]" captionTargetPageId="6" captionText="FIGURES 910. Male genitalia, from left to right: aedeagus in lateral view, aedeagus in ventral view, digitus and volsella. 9, Calligaster ilocana Selis, sp. nov. 10, Calligaster williamsi Bequaert, 1940." figureDoi="http://doi.org/10.5281/zenodo.6669330" httpUri="https://zenodo.org/record/6669330/files/figure.png" pageId="5" pageNumber="129">10</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="529711AE9E12FFF9FF4056E6FBE0FE9A" box="[151,1092,250,274]" pageId="5" pageNumber="129">
<taxonomicName id="B5874D039E12FFF9FF4056E6FEF9FE9A" authorityName="Bingham" box="[151,349,250,274]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="cyanopterus">
<emphasis id="40F3EA929E12FFF9FF4056E6FEF9FE9A" box="[151,349,250,274]" italics="true" pageId="5" pageNumber="129">Zethus cyanopterus</emphasis>
</taxonomicName>
;
<treatmentCitation id="F32610919E12FFF9FEBD56E6FD83FE9A" author="Williams, F. X." box="[362,551,250,274]" page="28" pageId="5" pageNumber="129" year="1919">
<bibRefCitation id="16164B719E12FFF9FEBD56E6FD83FE9A" author="Williams, F. X." box="[362,551,250,274]" pageId="5" pageNumber="129" pagination="19 - 187" refId="ref4546" refString="Williams, F. X. (1919) Philippine wasp studies. Part 2. Description of new species and life history studies. Bulletin of the Experimental Station of the Hawaiian Sugar Plantation Association, 14, 19 - 187." type="journal article" year="1919">Williams 1919: 28</bibRefCitation>
</treatmentCitation>
, 157, figs. 8894 (natural history). Misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="723836809E12FFF9FF405705FA87FEB9" blockId="5.[151,1315,250,305]" box="[151,1315,281,305]" pageId="5" pageNumber="129">
<treatmentCitationGroup id="529711AE9E12FFF9FF405705FA87FEB9" box="[151,1315,281,305]" pageId="5" pageNumber="129">
<treatmentCitation id="F32610919E12FFF9FF405705FD99FEB9" author="Bequaert, J." box="[151,573,281,305]" page="124" pageId="5" pageNumber="129" year="1940">
<taxonomicName id="B5874D039E12FFF9FF405705FD99FEB9" ID-CoL="PQ3M" authority="Bequaert 1940: 124" authorityName="Bequaert" authorityPageNumber="124" authorityYear="1940" box="[151,573,281,305]" class="Insecta" family="Eumenidae" genus="Calligaster" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="40F3EA929E12FFF9FF405705FEC9FEB9" box="[151,365,281,305]" italics="true" pageId="5" pageNumber="129">Calligaster williamsi</emphasis>
<materialsCitation id="C2EF3CDD9E12FFF9FEA45705FD99FEB9" ID-GBIF-Occurrence="3816435309" box="[371,573,281,305]" collectionCode="MCZ" pageId="5" pageNumber="129" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="16164B719E12FFF9FEA45705FD99FEB9" author="Bequaert, J." box="[371,573,281,305]" pageId="5" pageNumber="129" pagination="124 - 126" refId="ref4132" refString="Bequaert, J. (1940) A new species of Calligaster from the Philippine Islands (Hymenoptera, Vespidae, Subfam. Zethinae). The Pan-Pacific Entomologist, 16, 124 - 126." type="journal article" year="1940">Bequaert 1940: 124</bibRefCitation>
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, fig. 1 AD,
<specimenCount id="6481FD099E12FFF9FD175705FD77FEB8" box="[704,723,281,304]" pageId="5" pageNumber="129" type="female"></specimenCount>
,
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—“Los Baños, Luzon, Philippine Is.” (
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MCZ).
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</paragraph>
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<subSubSection id="3A9D650B9E12FFF9FF405745FB42FDA9" pageId="5" pageNumber="129" type="diagnosis">
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<emphasis id="40F3EA929E12FFF9FF405745FEB6FEFB" bold="true" box="[151,274,345,371]" pageId="5" pageNumber="129">Diagnosis.</emphasis>
Similar to previous species but differing as follows: parapsidal furrows shallow and finely punctate, anterior margin of scutellum with an irregular and narrow transverse furrow (
<figureCitation id="EABC2A059E12FFF9FBD45760FBE3FE1F" box="[1027,1095,380,407]" captionStart="FIGURES 38" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-17@4.[197,1390,181,1919]" captionTargetPageId="4" captionText="FIGURES 38. 34, 7, Calligaster ilocana Selis, sp. nov. 56, 8, Calligaster williamsi Bequaert, 1940. 3, 5, Head of male in frontal view. 4, 6, Head of female in frontal view. 78, Mesoscutum and scutellum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6669324" httpUri="https://zenodo.org/record/6669324/files/figure.png" pageId="5" pageNumber="129">Fig. 8</figureCitation>
), scutellum entirely matt with sparse fine punctures, apical emargination of male clypeus semicircular and as deep as wide (
<figureCitation id="EABC2A059E12FFF9FB615783FB5EFE32" box="[1206,1274,415,442]" captionStart="FIGURES 38" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-17@4.[197,1390,181,1919]" captionTargetPageId="4" captionText="FIGURES 38. 34, 7, Calligaster ilocana Selis, sp. nov. 56, 8, Calligaster williamsi Bequaert, 1940. 3, 5, Head of male in frontal view. 4, 6, Head of female in frontal view. 78, Mesoscutum and scutellum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6669324" httpUri="https://zenodo.org/record/6669324/files/figure.png" pageId="5" pageNumber="129">Fig. 5</figureCitation>
), face entirely black in both sexes, female clypeus more sparsely punctured on the disc (
<figureCitation id="EABC2A059E12FFF9FC3057DEFB89FE55" box="[999,1069,450,477]" captionStart="FIGURES 38" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[197,1390,181,1919]" captionTargetId="figure-17@4.[197,1390,181,1919]" captionTargetPageId="4" captionText="FIGURES 38. 34, 7, Calligaster ilocana Selis, sp. nov. 56, 8, Calligaster williamsi Bequaert, 1940. 3, 5, Head of male in frontal view. 4, 6, Head of female in frontal view. 78, Mesoscutum and scutellum in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6669324" httpUri="https://zenodo.org/record/6669324/files/figure.png" pageId="5" pageNumber="129">Fig. 6</figureCitation>
); apical part of aedeagus nearly straight in lateral view, ventral lobes of aedeagus in lateral view subtriangular and pointed and as wide as base of apical part of aedeagus, anteroventral lobe of digitus pointed and forming an acute angle (
<figureCitation id="EABC2A059E12FFF9FB52541BFB7DFDA9" box="[1157,1241,519,545]" captionStart="FIGURES 910" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@6.[225,1363,181,1920]" captionTargetPageId="6" captionText="FIGURES 910. Male genitalia, from left to right: aedeagus in lateral view, aedeagus in ventral view, digitus and volsella. 9, Calligaster ilocana Selis, sp. nov. 10, Calligaster williamsi Bequaert, 1940." figureDoi="http://doi.org/10.5281/zenodo.6669330" httpUri="https://zenodo.org/record/6669330/files/figure.png" pageId="5" pageNumber="129">Fig. 10</figureCitation>
).
</paragraph>
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<emphasis id="40F3EA929E12FFF9FF105435FE17FDCB" bold="true" box="[199,435,553,579]" pageId="5" pageNumber="129">Material examined.</emphasis>
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:
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<collectingDate id="167DE9A89E12FFF9FD1B5436FC9AFDCC" box="[716,830,554,580]" pageId="5" pageNumber="129" value="2018-08">VIII.2018</collectingDate>
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,
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,
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<materialsCitation id="C2EF3CDD9E12FFF9FE4A5500FCD2FCBF" ID-GBIF-Occurrence="3816435303" box="[413,886,796,823]" collectingDate="2017-10" collectionCode="MSVI" location="Hinabangan" pageId="5" pageNumber="129" specimenCount="1" specimenCount-female="1" stateProvince="Samar">
<emphasis id="40F3EA929E12FFF9FE4A5500FE56FCBF" bold="true" box="[413,498,796,823]" pageId="5" pageNumber="129">
<collectingRegion id="B043F8629E12FFF9FE4A5500FE4DFCBD" box="[413,489,796,822]" country="Philippines" name="Samar" pageId="5" pageNumber="129">SAMAR</collectingRegion>
:
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<location id="7758605B9E12FFF9FE2E5500FD27FCBE" LSID="urn:lsid:plazi:treatment:FA2E87969E12FFF9FF40568BFEE3FCF7:7758605B9E12FFF9FE2E5500FD27FCBE" box="[505,643,796,822]" name="Hinabangan" pageId="5" pageNumber="129" stateProvince="Samar">Hinabangan</location>
,
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<collectingDate id="167DE9A89E12FFF9FD585501FD40FCBF" box="[655,740,797,823]" pageId="5" pageNumber="129" value="2017-10">X.2017</collectingDate>
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<location id="7758605B9E12FFF9FC565501FBBAFCBF" LSID="urn:lsid:plazi:treatment:FA2E87969E12FFF9FF40568BFEE3FCF7:7758605B9E12FFF9FC565501FBBAFCBF" box="[897,1054,796,823]" name="Lope de Vega" pageId="5" pageNumber="129" stateProvince="Samar">Lope de Vega</location>
,
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<collectingDate id="167DE9A89E12FFF9FBFE5501FB35FCBF" box="[1065,1169,797,823]" pageId="5" pageNumber="129" value="2017-12">XII.2017</collectingDate>
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</paragraph>
</subSubSection>
<subSubSection id="3A9D650B9E12FFF9FF10555CFEE3FCF7" pageId="5" pageNumber="129" type="distribution">
<paragraph id="723836809E12FFF9FF10555CFEE3FCF7" blockId="5.[151,1437,345,895]" pageId="5" pageNumber="129">
<emphasis id="40F3EA929E12FFF9FF10555CFEC5FCD2" bold="true" box="[199,353,832,858]" pageId="5" pageNumber="129">Distribution.</emphasis>
<collectingCountry id="0A9076109E12FFF9FEB0555CFD90FCD2" box="[359,564,832,858]" name="Philippines" pageId="5" pageNumber="129">Philippine Islands</collectingCountry>
:
<collectingRegion id="B043F8629E12FFF9FDEA555CFD76FCD2" box="[573,722,832,858]" country="Philippines" name="Catanduanes" pageId="5" pageNumber="129">Catanduanes</collectingRegion>
*, Luzon, Mindanao, Mindoro, Negros*,
<collectingRegion id="B043F8629E12FFF9FB41555DFB45FCD3" box="[1174,1249,833,859]" country="Philippines" name="Samar" pageId="5" pageNumber="129">Samar</collectingRegion>
* (
<bibRefCitation id="16164B719E12FFF9FB2E555CFF75FCF7" author="Nugroho, H. &amp; Ubaidillah, R. &amp; Kojima, J." pageId="5" pageNumber="129" pagination="19 - 32" refId="ref4336" refString="Nugroho, H., Ubaidillah, R. &amp; Kojima, J. (2016) Taxonomy of the Indo-Malayan presocial potter wasp of genus Calligaster de Saussure (Hymenoptera, Vespidae, Eumeninae). Journal of Hymenoptera Research, 48, 19 - 32. https: // doi. org / 10.3897 / JHR. 48.7045" type="journal article" year="2016">
Nugroho
<emphasis id="40F3EA929E12FFF9FAB2555DFA39FCD2" box="[1381,1437,832,858]" italics="true" pageId="5" pageNumber="129">et al.</emphasis>
2016
</bibRefCitation>
) (
<figureCitation id="EABC2A059E12FFF9FF3F5578FE9DFCF7" box="[232,313,868,895]" captionStart="FIGURE 11" captionStartId="7.[151,250,1359,1383]" captionTargetBox="[389,1197,181,1335]" captionTargetId="figure-16@7.[389,1197,181,1335]" captionTargetPageId="7" captionText="FIGURE 11. Distributional map illustrating the distribution of Philippines species of Calligaster de Saussure." figureDoi="http://doi.org/10.5281/zenodo.6669332" httpUri="https://zenodo.org/record/6669332/files/figure.png" pageId="5" pageNumber="129">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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