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<document id="C182DB31417769310E9DE364E7723B36" ID-CLB-Dataset="51580" ID-DOI="10.11646/zootaxa.4092.3.2" ID-GBIF-Dataset="dd843b99-1701-4485-b235-1e344c0799db" ID-ISSN="1175-5326" ID-Zenodo-Dep="254902" ID-ZooBank="547625D1-70CC-48A1-9C36-3A0DD41B83A5" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458217128088" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Rahayu, Dwi Listyo" docDate="2016" docId="F56987A9FFA2FFF3FB8FF8B3FE917DED" docLanguage="en" docName="zootaxa.4092.3.2.pdf.imf" docOrigin="Zootaxa 4092 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Selwynia Borradaile 1903" docType="treatment" docVersion="9" lastPageNumber="342" masterDocId="0950FFD1FFA3FFF0FB18FF91FFAE7D15" masterDocTitle="On the genera Selwynia Borradaile, 1903, and Gandoa Kammerer, 2006, with descriptions of two new species from Papua New Guinea and French Polynesia (Crustacea: Decapoda: Brachyura: Aphanodactylidae)" masterLastPageNumber="370" masterPageNumber="339" pageNumber="340" updateTime="1698187634807" updateUser="ExternalLinkService">
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<mods:title id="E043F817EECF8B23000E3F3560C6D115">On the genera Selwynia Borradaile, 1903, and Gandoa Kammerer, 2006, with descriptions of two new species from Papua New Guinea and French Polynesia (Crustacea: Decapoda: Brachyura: Aphanodactylidae)</mods:title>
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<mods:namePart id="D72E5EA0D60248881D508537830E92F6">Ng, Peter K. L.</mods:namePart>
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<mods:date id="99C0920182681EBD69996790554AFC95">2016</mods:date>
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<treatment id="F56987A9FFA2FFF3FB8FF8B3FE917DED" ID-DOI="http://doi.org/10.5281/zenodo.6081356" ID-GBIF-Taxon="117914241" ID-Zenodo-Dep="6081356" LSID="urn:lsid:plazi:treatment:F56987A9FFA2FFF3FB8FF8B3FE917DED" httpUri="http://treatment.plazi.org/id/F56987A9FFA2FFF3FB8FF8B3FE917DED" lastPageId="3" lastPageNumber="342" pageId="1" pageNumber="340">
<subSubSection id="35DA6534FFA2FFF1FB8FF8B3FDAF7A8B" pageId="1" pageNumber="340" type="nomenclature">
<paragraph id="7D7F36BFFFA2FFF1FB8FF8B3FE727A28" blockId="1.[151,476,1826,1853]" box="[151,476,1826,1853]" pageId="1" pageNumber="340">
<heading id="263781D3FFA2FFF1FB8FF8B3FE727A28" bold="true" box="[151,476,1826,1853]" fontSize="11" level="1" pageId="1" pageNumber="340" reason="1">
<taxonomicName id="BAC04D3CFFA2FFF1FB8FF8B3FE727A28" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" box="[151,476,1826,1853]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA2FFF1FB8FF8B3FE727A28" bold="true" box="[151,476,1826,1853]" pageId="1" pageNumber="340">
<emphasis id="4FB4EAADFFA2FFF1FB8FF8B3FEAA7A29" bold="true" box="[151,260,1826,1852]" italics="true" pageId="1" pageNumber="340">Selwynia</emphasis>
Borradaile, 1903
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7D7F36BFFFA2FFF1FB8FF8F8FDAF7A8B" blockId="1.[151,1437,1897,2011]" pageId="1" pageNumber="340">
<taxonomicName id="BAC04D3CFFA2FFF1FB8FF8F8FE777A6A" authority="Borradaile, 1903: 430" authorityName="Borradaile" authorityPageNumber="430" authorityYear="1903" box="[151,473,1897,1919]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA2FFF1FB8FF8F8FF5D7A6A" box="[151,243,1897,1919]" italics="true" pageId="1" pageNumber="340">Selwynia</emphasis>
Borradaile, 1903: 430
</taxonomicName>
[
<typeStatus id="A27B881DFFA2FFF1FAFFF8FBFDBB7A6A" box="[487,533,1898,1919]" pageId="1" pageNumber="340">type</typeStatus>
species:
<taxonomicName id="BAC04D3CFFA2FFF1F96CF8F8FC697A6A" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" box="[628,967,1897,1919]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="4FB4EAADFFA2FFF1F96CF8F8FCBC7A6A" box="[628,786,1897,1919]" italics="true" pageId="1" pageNumber="340">Selwynia laevis</emphasis>
Borradaile, 1903
</taxonomicName>
, by monotypy]; Ng 1997: 98; Ng
<emphasis id="4FB4EAADFFA2FFF1FE3FF8FBFAF57A6A" box="[1319,1371,1897,1919]" italics="true" pageId="1" pageNumber="340">et al.</emphasis>
2008: 144 (list); Ng &amp; Clark 2014: 1.
</paragraph>
</subSubSection>
<subSubSection id="35DA6534FFA2FFF1FB8FF837FB547ACE" pageId="1" pageNumber="340" type="description">
<paragraph id="7D7F36BFFFA2FFF1FB8FF837FB547ACE" blockId="1.[151,1437,1897,2011]" pageId="1" pageNumber="340">
<taxonomicName id="BAC04D3CFFA2FFF1FB8FF837FE407AA9" authority="Tesch, 1918: 283" authorityName="Tesch" authorityPageNumber="283" authorityYear="1918" box="[151,494,1958,1980]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA2FFF1FB8FF837FE947AA9" box="[151,314,1958,1980]" italics="true" pageId="1" pageNumber="340">Aphanodactylus</emphasis>
Tesch, 1918: 283
</taxonomicName>
[
<typeStatus id="A27B881DFFA2FFF1FAE2F836FD867AA9" box="[506,552,1959,1980]" pageId="1" pageNumber="340">type</typeStatus>
species:
<taxonomicName id="BAC04D3CFFA2FFF1F99DF837FBAE7AA9" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[645,1024,1958,1980]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="340" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="4FB4EAADFFA2FFF1F99DF837FCD07AA9" box="[645,894,1958,1980]" italics="true" pageId="1" pageNumber="340">Aphanodactylus sibogae</emphasis>
Tesch, 1918
</taxonomicName>
, by monotypy]; Schmitt
<emphasis id="4FB4EAADFFA2FFF1FFE4F839FA9E7AA9" box="[1276,1328,1958,1981]" italics="true" pageId="1" pageNumber="340">et al.</emphasis>
1973: 127 (in list); Ng
<emphasis id="4FB4EAADFFA2FFF1FA5BF857FED97ACE" box="[323,375,1989,2011]" italics="true" pageId="1" pageNumber="340">et al.</emphasis>
2008: 247; Ng &amp; Naruse 2009: 284; Ahyong &amp; Ng 2009: 34; Castro, 2015: 553 (in list).
</paragraph>
</subSubSection>
<subSubSection id="35DA6534FFA1FFF2FB8FFF06FE417E00" pageId="2" pageNumber="341" type="diagnosis">
<paragraph id="7D7F36BFFFA1FFF2FB8FFF06FE417E00" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="341">
<emphasis id="4FB4EAADFFA1FFF2FB8FFF06FEBF7DA5" bold="true" box="[151,273,151,176]" pageId="2" pageNumber="341">Diagnosis.</emphasis>
Carapace wide, ovate, males 1.4–1.5 times wider than long, females 1.6–1.9 times wider than long; lateral margins cristate or subcristate, but not continuous to external orbital end; surface smooth, glabrous, with few scattered pits. Front narrow, deflexed broadly triangular in frontal view; straight to weakly sinuous in dorsal view. Orbit transverse; margins entire, unarmed; infraorbital margin mesially terminating in acute angle; supraorbital margin demarcated from antennular fossa by even curve; infraorbital and supraorbital margins not extending laterally beyond cornea or continuing laterally or posteriorly beyond orbit. Eyes short, mobile, fully. Antennules articulating transversely to slightly obliquely. Epistome short, medially sunken. Third maxilliped ischium longer than merus, both subquadrate. Thoracic sternites 1, 2 completely fused, broadly angular anteriorly; sternites 2/3 demarcated by shallow groove; sternites 3/4 fused, laterally unarmed; sternites 4–8 demarcated by narrow grooves, those demarcating sternites 4–7 incomplete medially; without longitudinal groove. Cheliped merus and carpus unarmed, with sparse plumose setae. P2–5 similar in shape, P3 longest; meri about twice as long as high, as long as or slightly longer than respective carpi and propodi combined; with plumose setae on flexor and extensor margins; meri with or without teeth along flexor margin; P5 markedly shorter than preceding legs; P2–5 with short spines on distoflexor angle of propodus opposing dactylus; dactylus very short, claw-like. Male abdomen simple, triangular; telson and all somites free; widest at somite 1; thoracic sternite 8 exposed when abdomen closed. Female abdomen with telson, all somites freely articulating, widest at somite 4. Vulva ovate, maximum width, may exceed half width of sternite 6. G1 simple, slender, broadly curved. G2 small, about 1/3 length of G1, slender, apex blunt. (Modified from Ahyong &amp; Ng 2009: 36).
</paragraph>
</subSubSection>
<subSubSection id="35DA6534FFA1FFF2FBDFFC8EFBF07AC8" pageId="2" pageNumber="341" type="discussion">
<paragraph id="7D7F36BFFFA1FFF2FBDFFC8EFE8379D0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="341">
<emphasis id="4FB4EAADFFA1FFF2FBDFFC8EFE9A7E2D" bold="true" box="[199,308,799,824]" pageId="2" pageNumber="341">Remarks</emphasis>
.
<taxonomicName id="BAC04D3CFFA1FFF2FA5DFCB1FE047E2D" box="[325,426,800,824]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2FA5DFCB1FE047E2D" box="[325,426,800,824]" italics="true" pageId="2" pageNumber="341">Selwynia</emphasis>
</taxonomicName>
is the most species-rich genus, with six described species, with another two described here. As discussed by Ng &amp; Naruse (2009), the various species of
<taxonomicName id="BAC04D3CFFA1FFF2F89DFCD5FC447E49" box="[901,1002,836,860]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2F89DFCD5FC447E49" box="[901,1002,836,860]" italics="true" pageId="2" pageNumber="341">Selwynia</emphasis>
</taxonomicName>
(as
<taxonomicName id="BAC04D3CFFA1FFF2FF03FCD5FB617E49" box="[1051,1231,836,860]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2FF03FCD5FB617E49" box="[1051,1231,836,860]" italics="true" pageId="2" pageNumber="341">Aphanodactylus</emphasis>
</taxonomicName>
) closely resemble each other in general appearance. The shape of the G1 is also quite similar in most species. Ng
<emphasis id="4FB4EAADFFA1FFF2FFD8FCF8FB597E95" box="[1216,1271,872,896]" italics="true" pageId="2" pageNumber="341">et al.</emphasis>
(2010) argued that most of characters used by Konishi &amp; Noda (1999) to separate the species of
<taxonomicName id="BAC04D3CFFA1FFF2FF5EFC1DFB057EB1" box="[1094,1195,908,932]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2FF5EFC1DFB057EB1" box="[1094,1195,908,932]" italics="true" pageId="2" pageNumber="341">Selwynia</emphasis>
</taxonomicName>
(as
<taxonomicName id="BAC04D3CFFA1FFF2FFC6FC1DFA3C7EB1" box="[1246,1426,908,932]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2FFC6FC1DFA3C7EB1" box="[1246,1426,908,932]" italics="true" pageId="2" pageNumber="341">Aphanodactylus</emphasis>
</taxonomicName>
) such as width to length ratio of the carapace, depth and extent of dorsal punctae on the carapace, shape of the frontal margin, and the number of antennular segments were unreliable, although the ventral armature of the merus of P2–P5 was useful at least to separate
<taxonomicName id="BAC04D3CFFA1FFF2F947FC69FCAD7905" box="[607,771,1016,1040]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2F947FC69FCAD7905" box="[607,771,1016,1040]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="BAC04D3CFFA1FFF2F824FC69FB6A7905" authority="Konishi &amp; Noda, 1999" authorityName="Konishi &amp; Noda" authorityYear="1999" box="[828,1220,1015,1040]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="lomiae">
<emphasis id="4FB4EAADFFA1FFF2F824FC69FC0B7905" box="[828,933,1016,1040]" italics="true" pageId="2" pageNumber="341">S. lomiae</emphasis>
(Konishi &amp; Noda, 1999)
</taxonomicName>
, but not
<taxonomicName id="BAC04D3CFFA1FFF2FE30FC69FE9B7921" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="4FB4EAADFFA1FFF2FE30FC69FA357905" box="[1320,1435,1016,1040]" italics="true" pageId="2" pageNumber="341">S. sibogae</emphasis>
(Tesch, 1918)
</taxonomicName>
. Instead, the species of
<taxonomicName id="BAC04D3CFFA1FFF2F95CFB8DFD077921" box="[580,681,1052,1076]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="4FB4EAADFFA1FFF2F95CFB8DFD077921" box="[580,681,1052,1076]" italics="true" pageId="2" pageNumber="341">Selwynia</emphasis>
</taxonomicName>
can be distinguished by the shape of the carapace, ambulatory leg proportions and armature, structure of the third maxilliped, and subtle differences in the shape of the G1. Unfortunately, no single species is represented by a large sample size of various size-groups, so variation within a species is not well understood. For now, the differences observed between taxa are regarded as significant at the species level.
</paragraph>
<paragraph id="7D7F36BFFFA1FFF2FBDFFB40FBF07AC8" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="341">
For females, most species have teeth on the flexor margins of P2–P4, with denticles of varying numbers. In species like
<taxonomicName id="BAC04D3CFFA1FFF2FA39FB65FE3A7819" box="[289,404,1268,1292]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="4FB4EAADFFA1FFF2FA39FB65FE3A7819" box="[289,404,1268,1292]" italics="true" pageId="2" pageNumber="341">S. sibogae</emphasis>
</taxonomicName>
,
<taxonomicName id="BAC04D3CFFA1FFF2FABBFB65FDBA7819" box="[419,532,1268,1292]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="loimiae">
<emphasis id="4FB4EAADFFA1FFF2FABBFB65FDBA7819" box="[419,532,1268,1292]" italics="true" pageId="2" pageNumber="341">S. loimiae</emphasis>
</taxonomicName>
,
<emphasis id="4FB4EAADFFA1FFF2F93BFB65FD0C7819" box="[547,674,1268,1292]" italics="true" pageId="2" pageNumber="341">
<taxonomicName id="BAC04D3CFFA1FFF2F93BFB65FD307819" box="[547,670,1268,1292]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="panglao">S. panglao</taxonomicName>
,
</emphasis>
and
<taxonomicName id="BAC04D3CFFA1FFF2F9C2FB65FCD37819" box="[730,893,1268,1292]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2F9C2FB65FCD37819" box="[730,893,1268,1292]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
, there are two distinct teeth on each merus with varying numbers of denticles (e.g.,
<figureCitation id="E5FB2A3AFFA1FFF2F939FA86FDC37825" box="[545,621,1303,1328]" captionStart="FIGURE 4" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[170,1419,204,1881]" captionTargetId="figure@8.[151,1436,193,1896]" captionTargetPageId="8" captionText="FIGURE 4. Selwynia sibogae (Tesch, 1918). A – E, male (7.7 × 5.3 mm) (ZRC 2010.0298), Lombok, Indonesia; F – K, ovigerous female lectotype (10.9 × 6.1 mm). A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, left P 3; H, left P 4; I, left P 5; J, right P 3; K, right P 4. Scales: A – E = 1.0 mm; F = 2.0 mm." httpUri="https://zenodo.org/record/254906/files/figure.png" pageId="2" pageNumber="341">Figs. 4</figureCitation>
G, H, J, K, 5C; Konishi &amp; Noda 1999: fig. 1e; Ng
<emphasis id="4FB4EAADFFA1FFF2FF80FA88FB7E7825" box="[1176,1232,1304,1328]" italics="true" pageId="2" pageNumber="341">et al.</emphasis>
2010: fig. 1D; Ng &amp; Naruse 2009: fig. 1a). In
<taxonomicName id="BAC04D3CFFA1FFF2FACCFAADFDE27841" box="[468,588,1340,1364]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2FACCFAADFDE27841" box="[468,588,1340,1364]" italics="true" pageId="2" pageNumber="341">S. madang</emphasis>
</taxonomicName>
<emphasis id="4FB4EAADFFA1FFF2F94CFAADFD3A7840" bold="true" box="[596,660,1340,1365]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="548757D6FFA1FFF2F94CFAADFD3A7840" box="[596,660,1340,1365]" pageId="2" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="BAC04D3CFFA1FFF2F9CCFAADFCFD7841" box="[724,851,1340,1364]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2F9CCFAADFCFD7841" box="[724,851,1340,1364]" italics="true" pageId="2" pageNumber="341">S. punctata</emphasis>
</taxonomicName>
<emphasis id="4FB4EAADFFA1FFF2F843FAADFC0C7840" bold="true" box="[859,930,1340,1365]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="548757D6FFA1FFF2F843FAADFC327840" box="[859,924,1340,1365]" pageId="2" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
there is one distinct tooth, and several or no denticles on each merus (e.g.,
<figureCitation id="E5FB2A3AFFA1FFF2FAF1FACEFDEC786D" box="[489,578,1375,1400]" captionStart="FIGURE 12" captionStartId="18.[151,250,1789,1811]" captionTargetBox="[176,1425,198,1750]" captionTargetId="figure@18.[151,1437,193,1767]" captionTargetPageId="18" captionText="FIGURE 12. Selwynia madang n. sp. A – E, male paratype (8.2 × 5.4 mm) (MNHN IU- 2013 - 1428 b), Papua New Guinea, F – H ovigerous female holotype (12.1 × 7.5 mm) (MNHN IU- 2013 - 1428 a), Papua New Guinea. A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, right P 4; H, right P 5. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254914/files/figure.png" pageId="2" pageNumber="341">Figs. 12</figureCitation>
G, 16H; Ng &amp; Naruse 2009: table 1; Ahyong &amp; Ng 2009: fig. 2E). As for the P5, females generally have an unarmed flexor margin, although the right merus of one specimen of a female
<taxonomicName id="BAC04D3CFFA1FFF2FE9FFA15FF5F78D5" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2FE9FFA15FF5F78D5" italics="true" pageId="2" pageNumber="341">S. madang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="548757D6FFA1FFF2FBE1FA36FE9778D5" box="[249,313,1447,1472]" pageId="2" pageNumber="341" rank="species">
<emphasis id="4FB4EAADFFA1FFF2FBE1FA36FE9C78D5" bold="true" box="[249,306,1447,1472]" pageId="2" pageNumber="341">n. sp</emphasis>
.
</taxonomicNameLabel>
(the
<typeStatus id="A27B881DFFA1FFF2FA6DFA36FE7778D5" box="[373,473,1447,1472]" pageId="2" pageNumber="341" type="holotype">holotype</typeStatus>
) has a small denticle (
<figureCitation id="E5FB2A3AFFA1FFF2F9C0FA36FC8478D5" box="[728,810,1447,1472]" captionStart="FIGURE 12" captionStartId="18.[151,250,1789,1811]" captionTargetBox="[176,1425,198,1750]" captionTargetId="figure@18.[151,1437,193,1767]" captionTargetPageId="18" captionText="FIGURE 12. Selwynia madang n. sp. A – E, male paratype (8.2 × 5.4 mm) (MNHN IU- 2013 - 1428 b), Papua New Guinea, F – H ovigerous female holotype (12.1 × 7.5 mm) (MNHN IU- 2013 - 1428 a), Papua New Guinea. A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, right P 4; H, right P 5. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254914/files/figure.png" pageId="2" pageNumber="341">Fig. 12</figureCitation>
H). There also seems to be variation in
<taxonomicName id="BAC04D3CFFA1FFF2FFF2FA39FAF578D5" box="[1258,1371,1448,1472]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="loimiae">
<emphasis id="4FB4EAADFFA1FFF2FFF2FA39FAF578D5" box="[1258,1371,1448,1472]" italics="true" pageId="2" pageNumber="341">S. loimiae</emphasis>
</taxonomicName>
, with the margin unarmed or with one prominent tooth and denticles (cf. Ng
<emphasis id="4FB4EAADFFA1FFF2F8F1FA5CFB8878F1" box="[1001,1062,1484,1508]" italics="true" pageId="2" pageNumber="341">et al.</emphasis>
2010).
<taxonomicName id="BAC04D3CFFA1FFF2FF9DFA5DFAD378F1" box="[1157,1405,1484,1508]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2FF9DFA5DFAD378F1" box="[1157,1405,1484,1508]" italics="true" pageId="2" pageNumber="341">Selwynia edmondsoni</emphasis>
</taxonomicName>
is distinctive among its congeners in that the flexor margin of the P5 merus has a large proximal tooth with several denticles (
<figureCitation id="E5FB2A3AFFA1FFF2FA12F985FEE37B39" box="[266,333,1556,1581]" captionStart="FIGURE 7" captionStartId="12.[151,250,1782,1804]" captionTargetBox="[226,1383,213,1738]" captionTargetId="figure@12.[212,1392,192,1745]" captionTargetPageId="12" captionText="FIGURE 7. Selwynia edmondsoni (Rathbun, 1932). A – G, male (10.8 × 8.1 mm) (ZRC 2000.0542); H, I, female (16.5 × 10 mm) (ZRC 2000.0542); both O‘ahu, Hawai‘i. A, left third maxilliped; B, male abdomen; C, left P 4 (inner view); D, left P 4 (outer view); E, right P 5; F, G, left G 1; H, right P 5; I, female abdomen. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254909/files/figure.png" pageId="2" pageNumber="341">Fig. 7</figureCitation>
E). For males, the P2–P4 flexor margin of the merus has 2–4 teeth and several or no denticles in
<taxonomicName id="BAC04D3CFFA1FFF2FE9FF985FF407B45" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="4FB4EAADFFA1FFF2FE9FF985FF407B45" italics="true" pageId="2" pageNumber="341">S. sibogae</emphasis>
</taxonomicName>
and
<taxonomicName id="BAC04D3CFFA1FFF2FA31F9A9FE0D7B45" box="[297,419,1592,1616]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="panglao">
<emphasis id="4FB4EAADFFA1FFF2FA31F9A9FE0D7B45" box="[297,419,1592,1616]" italics="true" pageId="2" pageNumber="341">S. panglao</emphasis>
</taxonomicName>
, with one tooth and low denticles in
<taxonomicName id="BAC04D3CFFA1FFF2F84DF9A9FC7B7B45" box="[853,981,1592,1616]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2F84DF9A9FC7B7B45" box="[853,981,1592,1616]" italics="true" pageId="2" pageNumber="341">S. punctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="548757D6FFA1FFF2F8C7F9A6FB8E7B5A" box="[991,1056,1591,1616]" pageId="2" pageNumber="341" rank="species">
<emphasis id="4FB4EAADFFA1FFF2F8C7F9A6FBB77B45" bold="true" box="[991,1049,1591,1616]" pageId="2" pageNumber="341">n. sp</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="BAC04D3CFFA1FFF2FF43F9A9FAAF7B45" box="[1115,1281,1592,1616]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2FF43F9A9FAAF7B45" box="[1115,1281,1592,1616]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
, only several denticles in
<taxonomicName id="BAC04D3CFFA1FFF2FA38F9CDFE387B61" box="[288,406,1628,1652]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2FA38F9CDFE387B61" box="[288,406,1628,1652]" italics="true" pageId="2" pageNumber="341">S. madang</emphasis>
</taxonomicName>
<emphasis id="4FB4EAADFFA1FFF2FA86F9CDFE487B60" bold="true" box="[414,486,1628,1653]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="548757D6FFA1FFF2FA86F9CDFE4E7B60" box="[414,480,1628,1653]" pageId="2" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
almost unarmed or with one denticle in
<taxonomicName id="BAC04D3CFFA1FFF2F8A8F9CDFBA07B61" box="[944,1038,1628,1652]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="4FB4EAADFFA1FFF2F8A8F9CDFBA07B61" box="[944,1038,1628,1652]" italics="true" pageId="2" pageNumber="341">S. laevis</emphasis>
</taxonomicName>
(
<figureCitation id="E5FB2A3AFFA1FFF2FF06F9CDFBC37B61" box="[1054,1133,1628,1653]" captionStart="FIGURE 4" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[170,1419,204,1881]" captionTargetId="figure@8.[151,1436,193,1896]" captionTargetPageId="8" captionText="FIGURE 4. Selwynia sibogae (Tesch, 1918). A – E, male (7.7 × 5.3 mm) (ZRC 2010.0298), Lombok, Indonesia; F – K, ovigerous female lectotype (10.9 × 6.1 mm). A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, left P 3; H, left P 4; I, left P 5; J, right P 3; K, right P 4. Scales: A – E = 1.0 mm; F = 2.0 mm." httpUri="https://zenodo.org/record/254906/files/figure.png" pageId="2" pageNumber="341">Figs. 4</figureCitation>
C, 7C, D, 12C, 16C; Ng &amp; Naruse 2009: fig. 3d; Ng &amp; Clark 2014: fig. 2B–G). The flexor margin of the P5 merus in most of the species is unarmed, with the exception of
<taxonomicName id="BAC04D3CFFA1FFF2F917F935FD187BA9" box="[527,694,1700,1724]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2F917F935FD187BA9" box="[527,694,1700,1724]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="BAC04D3CFFA1FFF2F9D0F935FCEC7BA9" box="[712,834,1700,1724]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2F9D0F935FCEC7BA9" box="[712,834,1700,1724]" italics="true" pageId="2" pageNumber="341">S. madang</emphasis>
</taxonomicName>
<emphasis id="4FB4EAADFFA1FFF2F855F935FC397BA8" bold="true" box="[845,919,1700,1725]" pageId="2" pageNumber="341">
<taxonomicNameLabel id="548757D6FFA1FFF2F855F935FC3F7BA8" box="[845,913,1700,1725]" pageId="2" pageNumber="341" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
and
<taxonomicName id="BAC04D3CFFA1FFF2F8CDF935FBF97BA9" box="[981,1111,1700,1724]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2F8CDF935FBF97BA9" box="[981,1111,1700,1724]" italics="true" pageId="2" pageNumber="341">S. punctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="548757D6FFA1FFF2FF7AF935FB0A7BA9" box="[1122,1188,1700,1725]" pageId="2" pageNumber="341" rank="species">
<emphasis id="4FB4EAADFFA1FFF2FF7AF935FB317BA8" bold="true" box="[1122,1183,1700,1725]" pageId="2" pageNumber="341">n. sp</emphasis>
.
</taxonomicNameLabel>
, which has only one distinct tooth and/or denticles; being largest in
<taxonomicName id="BAC04D3CFFA1FFF2F9A5F959FCCD7BF5" box="[701,867,1736,1760]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2F9A5F959FCCD7BF5" box="[701,867,1736,1760]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
(
<figureCitation id="E5FB2A3AFFA1FFF2F86EF956FC697BCA" box="[886,967,1735,1760]" captionStart="FIGURE 7" captionStartId="12.[151,250,1782,1804]" captionTargetBox="[226,1383,213,1738]" captionTargetId="figure@12.[212,1392,192,1745]" captionTargetPageId="12" captionText="FIGURE 7. Selwynia edmondsoni (Rathbun, 1932). A – G, male (10.8 × 8.1 mm) (ZRC 2000.0542); H, I, female (16.5 × 10 mm) (ZRC 2000.0542); both O‘ahu, Hawai‘i. A, left third maxilliped; B, male abdomen; C, left P 4 (inner view); D, left P 4 (outer view); E, right P 5; F, G, left G 1; H, right P 5; I, female abdomen. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254909/files/figure.png" pageId="2" pageNumber="341">Figs. 7</figureCitation>
E, 9A, 16D).
<taxonomicName id="BAC04D3CFFA1FFF2FF70F959FA9C7BF5" box="[1128,1330,1736,1760]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2FF70F959FA9C7BF5" box="[1128,1330,1736,1760]" italics="true" pageId="2" pageNumber="341">Selwynia madang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="548757D6FFA1FFF2FE24F956FAD17BCA" box="[1340,1407,1735,1760]" pageId="2" pageNumber="341" rank="species">
<emphasis id="4FB4EAADFFA1FFF2FE24F956FAD97BF5" bold="true" box="[1340,1399,1735,1760]" pageId="2" pageNumber="341">n. sp</emphasis>
.
</taxonomicNameLabel>
is unusual in that the basis-ischium of the female P5 also has a distinct small or large denticle on the ventral margin (
<figureCitation id="E5FB2A3AFFA1FFF2FB87F89EFF5A7A3D" box="[159,244,1807,1832]" captionStart="FIGURE 12" captionStartId="18.[151,250,1789,1811]" captionTargetBox="[176,1425,198,1750]" captionTargetId="figure@18.[151,1437,193,1767]" captionTargetPageId="18" captionText="FIGURE 12. Selwynia madang n. sp. A – E, male paratype (8.2 × 5.4 mm) (MNHN IU- 2013 - 1428 b), Papua New Guinea, F – H ovigerous female holotype (12.1 × 7.5 mm) (MNHN IU- 2013 - 1428 a), Papua New Guinea. A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, right P 4; H, right P 5. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254914/files/figure.png" pageId="2" pageNumber="341">Fig. 12</figureCitation>
H). An unusual character present only in male
<taxonomicName id="BAC04D3CFFA1FFF2F805F881FC6A7A3D" box="[797,964,1808,1832]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2F805F881FC6A7A3D" box="[797,964,1808,1832]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
is that the anterior margin of the flexor surface is armed with three teeth (
<figureCitation id="E5FB2A3AFFA1FFF2F90DF8A5FDF77A59" box="[533,601,1844,1869]" captionStart="FIGURE 7" captionStartId="12.[151,250,1782,1804]" captionTargetBox="[226,1383,213,1738]" captionTargetId="figure@12.[212,1392,192,1745]" captionTargetPageId="12" captionText="FIGURE 7. Selwynia edmondsoni (Rathbun, 1932). A – G, male (10.8 × 8.1 mm) (ZRC 2000.0542); H, I, female (16.5 × 10 mm) (ZRC 2000.0542); both O‘ahu, Hawai‘i. A, left third maxilliped; B, male abdomen; C, left P 4 (inner view); D, left P 4 (outer view); E, right P 5; F, G, left G 1; H, right P 5; I, female abdomen. Scales = 1.0 mm." httpUri="https://zenodo.org/record/254909/files/figure.png" pageId="2" pageNumber="341">Fig. 7</figureCitation>
C); in all congeners, even in females of
<taxonomicName id="BAC04D3CFFA1FFF2FF02F8A5FB107A59" box="[1050,1214,1844,1868]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="edmondsoni">
<emphasis id="4FB4EAADFFA1FFF2FF02F8A5FB107A59" box="[1050,1214,1844,1868]" italics="true" pageId="2" pageNumber="341">S. edmondsoni</emphasis>
</taxonomicName>
, this part is smooth (e.g.,
<figureCitation id="E5FB2A3AFFA1FFF2FBCFF8C6FE8A7A65" box="[215,292,1879,1904]" captionStart="FIGURE 4" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[170,1419,204,1881]" captionTargetId="figure@8.[151,1436,193,1896]" captionTargetPageId="8" captionText="FIGURE 4. Selwynia sibogae (Tesch, 1918). A – E, male (7.7 × 5.3 mm) (ZRC 2010.0298), Lombok, Indonesia; F – K, ovigerous female lectotype (10.9 × 6.1 mm). A, left third maxilliped; B, male abdomen; C, right P 4; D, E, left G 1; F, female abdomen; G, left P 3; H, left P 4; I, left P 5; J, right P 3; K, right P 4. Scales: A – E = 1.0 mm; F = 2.0 mm." httpUri="https://zenodo.org/record/254906/files/figure.png" pageId="2" pageNumber="341">Figs. 4</figureCitation>
H, 12C, 16H; Ng &amp; Naruse 2009: fig. 3d; Ng &amp; Clark 2014: fig. 2). The extensor margin of the merus of the P2–P5, it is usually unarmed, although the surface is granular and uneven. In
<taxonomicName id="BAC04D3CFFA1FFF2FF4FF8EDFB627A81" box="[1111,1228,1916,1940]" class="Malacostraca" family="Aphanodactylidae" genus="Aphanodactylus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="4FB4EAADFFA1FFF2FF4FF8EDFB627A81" box="[1111,1228,1916,1940]" italics="true" pageId="2" pageNumber="341">S. sibogae</emphasis>
</taxonomicName>
and
<taxonomicName id="BAC04D3CFFA1FFF2FE1DF8EDFAD07A81" box="[1285,1406,1916,1940]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="madang" status="sp. nov.">
<emphasis id="4FB4EAADFFA1FFF2FE1DF8EDFAD07A81" box="[1285,1406,1916,1940]" italics="true" pageId="2" pageNumber="341">S. madang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="548757D6FFA1FFF2FE9DF8EDFF197AAD" pageId="2" pageNumber="341" rank="species">
<emphasis id="4FB4EAADFFA1FFF2FE9DF8EDFF1F7AAC" bold="true" pageId="2" pageNumber="341">n. sp</emphasis>
.
</taxonomicNameLabel>
, the proximal part of this margin can be uneven but it is not dentate. The only species in which the extensor margin of P2 and P3 is distinctly dentiform is
<taxonomicName id="BAC04D3CFFA1FFF2F984F855FCBD7AC9" box="[668,787,1988,2012]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="341" phylum="Arthropoda" rank="species" species="panglao">
<emphasis id="4FB4EAADFFA1FFF2F984F855FCBD7AC9" box="[668,787,1988,2012]" italics="true" pageId="2" pageNumber="341">S. panglao</emphasis>
</taxonomicName>
(see Ng &amp; Naruse 2009: 3d).
</paragraph>
</subSubSection>
<subSubSection id="35DA6534FFA0FFF3FBDFFF06FE917DED" pageId="3" pageNumber="342" type="materials_examined">
<paragraph id="7D7F36BFFFA0FFF3FBDFFF06FE917DED" blockId="3.[151,1436,151,248]" pageId="3" pageNumber="342">
<emphasis id="4FB4EAADFFA0FFF3FBDFFF06FE7E7DA5" bold="true" box="[199,464,151,176]" pageId="3" pageNumber="342">Comparative material</emphasis>
.
<taxonomicName id="BAC04D3CFFA0FFF3FAC7FF09FCFB7DA5" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" box="[479,853,151,176]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="4FB4EAADFFA0FFF3FAC7FF09FD237DA5" box="[479,653,152,176]" italics="true" pageId="3" pageNumber="342">Selwynia laevis</emphasis>
Borradaile, 1903
</taxonomicName>
: see Ng &amp; Clark (2014);
<taxonomicName id="BAC04D3CFFA0FFF3FF6AFF09FEE97DC1" authority="Konishi &amp; Noda, 1999" authorityName="Konishi &amp; Noda" authorityYear="1999" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="loimiae">
<emphasis id="4FB4EAADFFA0FFF3FF6AFF09FA9C7DA5" box="[1138,1330,152,176]" italics="true" pageId="3" pageNumber="342">Selwynia loimiae</emphasis>
(Konishi &amp; Noda, 1999)
</taxonomicName>
: see Ahyong &amp; Ng (2009); Ng
<emphasis id="4FB4EAADFFA0FFF3F9AEFF2CFD5F7DC1" box="[694,753,188,212]" italics="true" pageId="3" pageNumber="342">et al.</emphasis>
(2010);
<taxonomicName id="BAC04D3CFFA0FFF3F84DFF2DFB8E7DC1" box="[853,1056,188,212]" class="Malacostraca" family="Pilumnidae" genus="Selwynia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="342" phylum="Arthropoda" rank="species" species="panglao">
<emphasis id="4FB4EAADFFA0FFF3F84DFF2DFB8E7DC1" box="[853,1056,188,212]" italics="true" pageId="3" pageNumber="342">Selwynia panglao</emphasis>
</taxonomicName>
(Ng &amp; Naruse, 2009): see Ng &amp; Naruse (2009).
</paragraph>
</subSubSection>
</treatment>
</document>
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