331 lines
39 KiB
XML
331 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1195.112112" ID-Pensoft-Pub="1313-2970-1195-157" ID-Pensoft-UUID="E26F0ADE4B4C5BCB943E2FCC63E349C4" ID-ZooBank="7CFF082DDA924ECF8E7C0A9972F6CD08" ModsDocID="1313-2970-1195-157" checkinTime="1710460428610" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Lwin, Ngwe, Pall-Gergely, Barna, Chanabun, Ratmanee, Pholyotha, Arthit, Prasankok, Pongpun, Seesamut, Teerapong, Siriwut, Warut, Srisonchai, Ruttapon, Sutcharit, Chirasak & Panha, Somsak" docDate="2024" docId="F528EC79C66755AC9FE6834FBD02544A" docLanguage="en" docName="ZooKeys 1195: 157-197" docOrigin="ZooKeys 1195" docPubDate="2024-03-14" docSource="http://dx.doi.org/10.3897/zookeys.1195.112112" docTitle="Gyliotrachela aunglini Tongkerd & Panha 2024, sp. nov." docType="treatment" docUuid="76E8570A-172A-4E2C-BB8A-7A1D898CD149" docUuidSource="ZooBank" docVersion="1" id="E26F0ADE4B4C5BCB943E2FCC63E349C4" lastPageNumber="157" masterDocId="E26F0ADE4B4C5BCB943E2FCC63E349C4" masterDocTitle="Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species" masterLastPageNumber="197" masterPageNumber="157" pageNumber="157" updateTime="1710460428610" updateUser="pensoft">
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<mods:titleInfo id="17C4E04F17271468EAB6AC4F3082FC19">
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<mods:title id="4E9108458CF2607A80577E688A3316BF">Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species</mods:title>
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<mods:namePart id="934C11C3B971645CF42411DAB23A1582">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="88B73F6CD3C78FC73991ECA4A25481F3">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="E5F6858D65A1DDFAEBDD9B53C6CB42FB">Lwin, Ngwe</mods:namePart>
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<mods:affiliation id="DD27FCB25F442ED31485B30B2F57006E">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="9188DA16419A8ABBBF382AED452F7B9B">Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation id="6FFCF15621B615B70D47CBC3A0900E78">Plant Protection Institute, Centre for Agricultural Research, ELKH, Herman Otto ut 15, 1022 Budapest, Hungary</mods:affiliation>
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<mods:namePart id="39D248D9C8533F44B3E326B2CFE012FD">Chanabun, Ratmanee</mods:namePart>
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<mods:affiliation id="25CB283AF6217590ACBBC912C2D57489">Program in Animal Science, Faculty of Agricultural Technology, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
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<mods:namePart id="47EC3B0858DEDAC88CD8A9A8023D4746">Pholyotha, Arthit</mods:namePart>
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<mods:affiliation id="6B4356717F2E82E89215D8FB56CAF8D5">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="706C50D731AA12E423F9F1AB90B0B3B9">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="6B2204683514BCD16D0516AF427244D2">Biodiversity and Utilization Research Unit, Center of Excellence in Modern Agriculture, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
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<mods:namePart id="A9F61B596CC7463996559518F6140127">Seesamut, Teerapong</mods:namePart>
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<mods:affiliation id="959F3697BD8D4FBB7B8240D929016DB1">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="F73AAFEC983354AF9607174EF966C36A">Siriwut, Warut</mods:namePart>
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<mods:affiliation id="9C2B7CD3E8C852F3DE392BB673F8CECC">Department of Biology, Faculty of Science, Rangsit University, Pathum Thani 12000, Thailand</mods:affiliation>
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<mods:namePart id="A7517C41C522BC7BB4C2B2FB96AFB2C2">Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation id="449D8F2BEE65DFE356A88601187E2504">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="EF7D01F129547AC7B9FC7149163CB032">Sutcharit, Chirasak</mods:namePart>
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<mods:nameIdentifier id="ACC1088181D77C93D9C4F254907C3071" type="ORCID">https://orcid.org/0000-0001-7670-9540</mods:nameIdentifier>
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<mods:affiliation id="B6BEDF5EC56F41FD41CEC7BDA4011A15">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="B86761CCC31FF6D7FBA327BB0F370BF3" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:namePart id="87092FAFA292E7B003087BA603FABF35">Panha, Somsak</mods:namePart>
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<mods:nameIdentifier id="273BC5DEB262BE5AE29061C45B1AAEE0" type="ORCID">https://orcid.org/0000-0002-4431-2458</mods:nameIdentifier>
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<mods:affiliation id="7C989BF609D07360A47D5253DAB09E49">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<mods:nameIdentifier id="29810D093CAAC1F4074EE33DC1FB16E2" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="02FB3173662AB2124EB3853A75B633A9">ZooKeys</mods:title>
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<mods:part id="91E1BBFCDBD9D049CC984B7651CD26A9">
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<mods:date id="102DCD13FDBFBB026D45572657C1AC4B">2024</mods:date>
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<mods:number id="796999771F150351655FBA1FF0249472">2024-03-14</mods:number>
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<mods:detail id="EC83F9D59AA9DD717A7253FF378A5653" type="volume">
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<mods:number id="FDDA006449300647918519C4C383BD44">1195</mods:number>
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<mods:start id="5B9DFF89AA68B81FC8438935BF9F0B8B">157</mods:start>
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<treatment id="F528EC79C66755AC9FE6834FBD02544A" LSID="urn:lsid:zoobank.org:act:76E8570A-172A-4E2C-BB8A-7A1D898CD149" httpUri="http://treatment.plazi.org/id/F528EC79C66755AC9FE6834FBD02544A" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection id="8D4568F11490F888F761865641437D02" pageId="0" pageNumber="157" type="nomenclature">
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<paragraph id="B4DF86C1D4091172750BB917A382865D" pageId="0" pageNumber="157">
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<taxonomicName id="9E9B8B2EAB7122B8F8D08EBA03FF38A0" LSID="https://zoobank.org/76E8570A-172A-4E2C-BB8A-7A1D898CD149" authority="Tongkerd & Panha" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela aunglini" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="aunglini" status="sp. nov.">Gyliotrachela aunglini Tongkerd & Panha</taxonomicName>
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<taxonomicNameLabel id="B7E94E1CFC83211B706139689C0A4236" pageId="0" pageNumber="157">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3180E42ABF56350804CC5C5198A86E1" pageId="0" pageNumber="157" type="description">
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<paragraph id="3DA93C203C891274ACA4A62B88A516CF" pageId="0" pageNumber="157">
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<figureCitation id="084FF45FA185137AD599BDD2F72F6783" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Gyliotrachela aunglini sp. nov. from Hpa-An, Kayin State A, B holotype CUMZ 14383, (A) shell and (B) protoconch C, D paratypes CUMZ 14412 from the type locality. Abbreviations: a: angular lamella, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure12" httpUri="https://binary.pensoft.net/fig/1002061" pageId="0" pageNumber="157">Figs 12</figureCitation>
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<figureCitation id="DC98CBF9DB7F87021DE817B9B697C212" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">, 13P</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="82589CE6320C01428C057C7149904C48" pageId="0" pageNumber="157" type="type locality">
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<paragraph id="867663640DCEC10EDE1155D6A13F13B6" pageId="0" pageNumber="157">Type locality.</paragraph>
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<paragraph id="BEA2CB34B4047DF40684B1075171CB3F" pageId="0" pageNumber="157">
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Kaw Gon Cave, Hpa-An, Kayin State, Myanmar (locality code PA5;
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<geoCoordinate id="900F68D19F1898341B4194A77D1A4732" degrees="16" direction="north" minutes="49" orientation="latitude" precision="1" seconds="22.2" value="16.822834">16°49'22.2"N</geoCoordinate>
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,
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<geoCoordinate id="9F11B3A7BFFF6C4AFB5D67BAA7E2404A" degrees="97" direction="east" minutes="35" orientation="longitude" precision="1" seconds="08.9" value="97.58581">97°35'08.9"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="6A2F1FF3F814876C62DA77A5A6E2A554" pageId="0" pageNumber="157" type="type material">
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<paragraph id="CB25A954F05402FBC311428D7879EC54" pageId="0" pageNumber="157">Type material.</paragraph>
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<paragraph id="FB5EE4E576F88F9DC53D578311818C6A" pageId="0" pageNumber="157">
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<emphasis id="BCD2A027CBA48047912B63E6878FA52A" bold="true" italics="true" pageId="0" pageNumber="157">Holotype</emphasis>
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CUMZ 14383 (height 1.8 mm, width 1.6 mm; Fig.
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<figureCitation id="5C6D3D01A6CA8F4EF7EB04CF5B733338" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Gyliotrachela aunglini sp. nov. from Hpa-An, Kayin State A, B holotype CUMZ 14383, (A) shell and (B) protoconch C, D paratypes CUMZ 14412 from the type locality. Abbreviations: a: angular lamella, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure12" httpUri="https://binary.pensoft.net/fig/1002061" pageId="0" pageNumber="157">12A, B</figureCitation>
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).
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<emphasis id="4100E00AE3ECA4B1D7D5A6891EF0E002" bold="true" italics="true" pageId="0" pageNumber="157">Paratypes</emphasis>
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CUMZ 14384 (2 shells); CUMZ 14411 (16 shells; measured); CUMZ 14412 (2 shell; Figs
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<figureCitation id="3A43CFA5EE727937670C1D57AA570CE6" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Gyliotrachela aunglini sp. nov. from Hpa-An, Kayin State A, B holotype CUMZ 14383, (A) shell and (B) protoconch C, D paratypes CUMZ 14412 from the type locality. Abbreviations: a: angular lamella, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure12" httpUri="https://binary.pensoft.net/fig/1002061" pageId="0" pageNumber="157">12C, D</figureCitation>
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,
|
|
<figureCitation id="F5F3262CD29B7BB775654ECDA0FA05CF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">13P</figureCitation>
|
|
), NHMUK 20230593 (2 shells), SMF 373021 (2 shells) all from the type locality.
|
|
</paragraph>
|
|
<caption id="4FD924C0F04790E266CB881D47E31DD5" doi="10.3897/zookeys.1195.112112.figure12" httpUri="https://binary.pensoft.net/fig/1002061" pageId="0" pageNumber="157" start="Figure 12" startId="F12">
|
|
<paragraph id="A101268AF29AF8F87E87286D998A468D" pageId="0" pageNumber="157">
|
|
<emphasis id="7859933BEFC4D35DFFAB5D6BEFC63A78" bold="true" pageId="0" pageNumber="157">Figure 12.</emphasis>
|
|
<taxonomicName id="6F70C22C470F8BA386ED484700208C1A" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela aunglini" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="aunglini">
|
|
<emphasis id="D61573B55E479F03725F9B77B9F3C638" italics="true" pageId="0" pageNumber="157">Gyliotrachela aunglini</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. from Hpa-An, Kayin State
|
|
<emphasis id="754028F3354B1BBCF6D05EED3DEB6E08" bold="true" pageId="0" pageNumber="157">A, B</emphasis>
|
|
holotype CUMZ 14383, (
|
|
<emphasis id="CB8BE2233CDB9C3484455EA18453F146" bold="true" pageId="0" pageNumber="157">A</emphasis>
|
|
) shell and (
|
|
<emphasis id="66A1F0C53D1F271D46BBC52F9C69CD53" bold="true" pageId="0" pageNumber="157">B</emphasis>
|
|
) protoconch
|
|
<emphasis id="6706429A4656D970D714C912D60386ED" bold="true" pageId="0" pageNumber="157">C, D</emphasis>
|
|
paratypes CUMZ 14412 from the type locality. Abbreviations:
|
|
<emphasis id="F24F3C02390BE3CD73C157F0049E6A06" bold="true" pageId="0" pageNumber="157">a</emphasis>
|
|
: angular lamella,
|
|
<emphasis id="7D5A91A9CF37405DAC9DFF2E0B5DFD3B" bold="true" pageId="0" pageNumber="157">c</emphasis>
|
|
: columellar lamella,
|
|
<emphasis id="35C0AB7EB0F77D84264E1119FAFCE6E2" bold="true" pageId="0" pageNumber="157">lpl</emphasis>
|
|
: lower palatal plica,
|
|
<emphasis id="9A562CB79953BA7868914DB562B9D0E7" bold="true" pageId="0" pageNumber="157">p</emphasis>
|
|
: parietal lamella,
|
|
<emphasis id="B3E12514F2BEB521E0458258CC7E7EF3" bold="true" pageId="0" pageNumber="157">upl</emphasis>
|
|
: upper palatal plica.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="A84C1F8ADE3192DFDB1B077A5BDC78B4" pageId="0" pageNumber="157" type="diagnosis">
|
|
<paragraph id="9406BB0D4B13E02D482EC0832E956032" pageId="0" pageNumber="157">Diagnosis.</paragraph>
|
|
<paragraph id="AE52BA32A1501D2075FB456C6CD54A80" pageId="0" pageNumber="157">Shell ovate-conical with shouldered and flat-sided last whorl, aperture not detached from penultimate whorl; shell surface irregularly wrinkled, colour pale reddish brown; aperture roundly triangular, with five or six apertural dentitions (angular, parietal, upper palatal, lower palatal and columellar).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BEB47EC982570F46DB752A7FEE44374B" pageId="0" pageNumber="157" type="etymology">
|
|
<paragraph id="D2CBB6B950D5FB73372CEE9AF89B1200" pageId="0" pageNumber="157">Etymology.</paragraph>
|
|
<paragraph id="F53497B4826ADC65F4C3F4A8E3717D91" pageId="0" pageNumber="157">
|
|
The specific name
|
|
<emphasis id="5F4E679539B6A55DD62B11435C540A1F" italics="true" pageId="0" pageNumber="157">aunglini</emphasis>
|
|
is named after Mr. Aung Lin, the FFI coordinator, who took care of the survey team and arranged the limestone survey trip in Myanmar between 2015 and 2016.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="410237794A6CA2125367A1BC9D715A04" pageId="0" pageNumber="157" type="description">
|
|
<paragraph id="C2DAA7E194550FA700C4D5E1285C2A00" pageId="0" pageNumber="157">Description.</paragraph>
|
|
<paragraph id="5810447A95701C7290D2D2537DBD268D" pageId="0" pageNumber="157">
|
|
Shell broadly ovate-conical, pale reddish brown in colour, moderate spire, and
|
|
<normalizedToken id="E0C1FA36F044ECC2D22B410A9755F863" originalValue="4½">41/2</normalizedToken>
|
|
-5 widely convex whorls. Shell height 1.6-1.8 mm and shell width 1.6-1.8 mm. Apex blunt; protoconch with
|
|
<normalizedToken id="3D528D31C5FF487D3B228E37DFCD663E" originalValue="1¾">13/4</normalizedToken>
|
|
whorls, wrinkled, with roundish pits and weak spiral ridges. Teleoconch with narrowly spaced radial growth lines, parallel with very strong, irregular wrinkles; suture well impressed. Last whorl bluntly shouldered and flat-sided with shallow groove below shoulder. Tuba absent; peristome adnate and widely expanded; lip whitish to yellowish. Aperture roundly triangular and with five dentitions. Parietal lamella large, strong, blunt and located somewhat deeper inside aperture than angular lamella. Angular lamella strong and reaching peristome. Upper palatal plica strong, tall and approximately same size as parietal lamella; lower palatal plica strong and lower than upper palatal plica. Columellar lamella strong, distinct and pointing slightly upwards. Umbilicus widely perforate, ~ 1/3 of shell width, rounded, shallow, and surrounded by curved periumbilical keel.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CF6386512E8E96CCB47F9540D9F2F6A2" pageId="0" pageNumber="157" type="distribution">
|
|
<paragraph id="D2D7D38DD926B541312B13CD35E3EA53" pageId="0" pageNumber="157">Distribution.</paragraph>
|
|
<paragraph id="AFAEE3F9F8398CBF0196DA91AAA44A2C" pageId="0" pageNumber="157">This new species is currently known only from the type locality in Kayin State. This snail lives on the limestone walls and under leaf litter.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5142C9FC817BFBAF3BF397D3DDA45A34" pageId="0" pageNumber="157" type="differential diagnosis">
|
|
<paragraph id="52887FC8F7260B15C48D5A8C4A6EC1DC" pageId="0" pageNumber="157">Differential diagnosis.</paragraph>
|
|
<paragraph id="EB6C40CC8E26091A89C3FD4E1653306B" pageId="0" pageNumber="157">
|
|
<taxonomicName id="8B2D8242F731D3B5341293EDF055EC51" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela aunglini" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="aunglini">
|
|
<emphasis id="484927D36783EA9C234F288B9983C395" italics="true" pageId="0" pageNumber="157">Gyliotrachela aunglini</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="3AC8B58CCBF282787DE27DCC8E110BB4" lsidName="G. bensonianum" pageId="0" pageNumber="157" rank="species" species="bensonianum">
|
|
<emphasis id="0F1258B1FEDA0799AACD4E05F85CF6E7" italics="true" pageId="0" pageNumber="157">G. bensonianum</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="09780829683DA96A5259ADF525301174" lsidName="G. tianxingqiaoensis" pageId="0" pageNumber="157" rank="species" species="tianxingqiaoensis">
|
|
<emphasis id="8A7091D4FFB86D72985E50386FB5C568" italics="true" pageId="0" pageNumber="157">G. tianxingqiaoensis</emphasis>
|
|
</taxonomicName>
|
|
from Myanmar in having high spire, peristome adnate to preceding whorl, shouldered last whorl, with very weak spiral striae on protoconch, and only five apertural dentitions (parietal, angular, upper- and lower- palatal, and columellar). In contrast, the other two species possess a depressed spire, a short and slightly ascending tuba, and an angular last whorl. Additionally,
|
|
<taxonomicName id="F3EA3946B32225A0D12DB837B232192A" lsidName="G. bensonianum" pageId="0" pageNumber="157" rank="species" species="bensonianum">
|
|
<emphasis id="CA656310A090AF1FACF60115B5FCAAFF" italics="true" pageId="0" pageNumber="157">G. bensonianum</emphasis>
|
|
</taxonomicName>
|
|
has a spirally striated teleoconch, and it has a basal plica and infraparietal lamella;
|
|
<taxonomicName id="9DF1B2C4FAA540EF784CE6A184C1E23D" lsidName="G. tianxingqiaoensis" pageId="0" pageNumber="157" rank="species" species="tianxingqiaoensis">
|
|
<emphasis id="C326C05E39F95FBD946B15F625510132" italics="true" pageId="0" pageNumber="157">G. tianxingqiaoensis</emphasis>
|
|
</taxonomicName>
|
|
possesses numerous small accessory plicae and lamellae (
|
|
<bibRefCitation id="FCB7226E50D7226DB4AF7F4DFE53EB2A" DOI="https://doi.org/10.1127/arch.moll/151/131-139" author="Gojsina, V" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="157" pagination="131 - 139" refId="B24" refString="Gojsina, V, Chen, Z-U, Pall-Gergely, B, 2022. Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar. Archiv fuer Molluskenkunde 151 (2): 131 - 139, DOI: https://doi.org/10.1127/arch.moll/151/131-139" title="Notes on Hypselostomatidae (Gastropoda: Eupulmonata: Pupilloidea) from Myanmar." url="https://doi.org/10.1127/arch.moll/151/131-139" volume="151" year="2022">
|
|
<normalizedToken id="EDD78EF57039731EE143019567B04897" originalValue="Gojšina">Gojsina</normalizedToken>
|
|
et al. 2022
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="9B596A02592B1C13E054156F51C17A1C" pageId="0" pageNumber="157">
|
|
This new species can be distinguished from
|
|
<taxonomicName id="A4628C885D692E9C26F07719FA7F5CEF" lsidName="G. saraburiensis" pageId="0" pageNumber="157" rank="species" species="saraburiensis">
|
|
<emphasis id="AF6C14E185239C04C164184281B4AC72" italics="true" pageId="0" pageNumber="157">G. saraburiensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="9B564E977FDA7ECDC6628570DC749264" lsidName="G. muangon" pageId="0" pageNumber="157" rank="species" species="muangon">
|
|
<emphasis id="97AD5ED7CD6FE7D83A9DB396262597BA" italics="true" pageId="0" pageNumber="157">G. muangon</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="89721A01A2684B241E125E7A96992832" lsidName="G. cultura" pageId="0" pageNumber="157" rank="species" species="cultura">
|
|
<emphasis id="F44D47469BA7D4F21C01649EA1114216" italics="true" pageId="0" pageNumber="157">G. cultura</emphasis>
|
|
</taxonomicName>
|
|
(all known from Thailand) by having no tuba (i.e., peristome adnate to preceding whorl), possessing a shouldered last whorl, strong wrinkles on the shell surface and bearing only five to six major dentitions. In contrast, the three species from Thailand tend to have a short and slightly ascending tuba, an angular to weakly angular last whorl, spiral striae on the teleoconch, and numerous accessory plicae and lamellae inside the aperture (
|
|
<bibRefCitation id="43AE2E4C6D7C6F8613F92AB7E07435E6" author="Burch, JB" journalOrPublisher="D. dissertation. Evolutionary Ecology Group, University of Antwerp, Antwerp" pageId="0" pageNumber="157" refId="B13" refString="Burch, JB, Panha, S, Tongkerd, P, 2003. New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana 13[2002](29/30): 129-187." title="New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana 13 [2002] (29 / 30): 129 - 187." year="2003">Burch et al. 2003</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="085BAB58FD1E8A37A37F6D9D6CD2AAA7" author="Panha, S" journalOrPublisher="The Natural History Journal of Chulalongkorn University" pageId="0" pageNumber="157" pagination="57 - 82" refId="B65" refString="Panha, S, Tongkerd, P, Sucharit, C, Burch, JB, 2004. New pupillid species from Thailand (Pulmonata: Pupillidae). The Natural History Journal of Chulalongkorn University 4 (2): 57 - 82" title="New pupillid species from Thailand (Pulmonata: Pupillidae)." volume="4" year="2004">Panha et al. 2004</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="8E40A0CFC0B77E6B4F06D1FF72E3BD86" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B63" refString="Panha, S, Burch, JB, 2005. An introduction to the microsnails of Thailand. Malacological Review 37/38(2004-2005): 1-155. [Date of publication follow Jirapatrasilp et al. (2023)]" title="An introduction to the microsnails of Thailand. Malacological Review 37 / 38 (2004 - 2005): 1 - 155. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2005">Panha and Burch 2005</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="8202B102EDB9E23A7AF7F6AC7CCFD830" author="Tanmuangpak, K" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="157" pagination="407 - 416" refId="B85" refString="Tanmuangpak, K, Dumrongrojwattana, P, 2022. Gyliotrachela cultura, a new species of terrestrial microsnail (Gastropoda: Eupulmonata: Vertiginidae) from Thailand. The Raffles Bulletin of Zoology 70: 407 - 416" title="Gyliotrachela cultura, a new species of terrestrial microsnail (Gastropoda: Eupulmonata: Vertiginidae) from Thailand." volume="70" year="2022">Tanmuangpak and Dumrongrojwattana 2022</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="321471ABF18AD8A2916EB7574A36A097" pageId="0" pageNumber="157">
|
|
Several
|
|
<taxonomicName id="136057C9B08000B9D60605CD11F9E176" authorityName="le Tomlin" authorityYear="1930" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
|
<emphasis id="896A90C2AFA4707974DEA68B7A4C31A4" italics="true" pageId="0" pageNumber="157">Gyliotrachela</emphasis>
|
|
</taxonomicName>
|
|
species from southern Thailand are similar to the new species by having very short tuba, and few apertural dentitions.
|
|
<taxonomicName id="1201384A862C83E14114E6BB9B00CC61" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela aunglini" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="aunglini">
|
|
<emphasis id="CCC28ADF01044F97CE851B4366725022" italics="true" pageId="0" pageNumber="157">Gyliotrachela aunglini</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="2CE98F347B552883B4B3F7D8E243F62C" lsidName="G. transitans" pageId="0" pageNumber="157" rank="species" species="transitans">
|
|
<emphasis id="4A109E32B65B9453C056F4C176C29991" italics="true" pageId="0" pageNumber="157">G. transitans</emphasis>
|
|
</taxonomicName>
|
|
(von
|
|
<normalizedToken id="D9E0A8AE6E674318864320687384E8D6" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
, 1894) in the short and slightly descending tuba, the angular last whorl, and the apertural dentition with supracolumellar lamella and a basal plica;
|
|
<taxonomicName id="90ABB24F771E2638EBEEB3321E49942F" lsidName="G. tarutao" pageId="0" pageNumber="157" rank="species" species="tarutao">
|
|
<emphasis id="E689FAC920E922CC6BA89EC67A88A87D" italics="true" pageId="0" pageNumber="157">G. tarutao</emphasis>
|
|
</taxonomicName>
|
|
(Panha & Burch, 2002) has an elevated spire, an angular last whorl, a very short tuba, and the apertural dentition is characterised by a subcolumellar lamella, an infraparietal plica and a basal plica (
|
|
<bibRefCitation id="57020BB9159EC7581EA9E12D203AB794" author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="157" pagination="146 - 156" publicationUrl="https://biodiversitylibrary.org/page/35990426" refId="B97" refString="von Moellendorff, OF, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156, https://biodiversitylibrary.org/page/35990426" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." url="https://biodiversitylibrary.org/page/35990426" volume="62" year="1894">
|
|
von
|
|
<normalizedToken id="377EA1AF3F18A8FDE68B3CB7C79BB6F6" originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
1894
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="FAE82F45CAC9A740699BC633E8B6FC2E" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B61" refString="Panha, S, Burch, JB, 2002b. The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12[2001](28): 77-94. [Date of publication follow Jirapatrasilp et al. (2023)]" title="The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12 [2001] (28): 77 - 94. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2002 b">Panha and Burch 2002b</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="4366D9CE878BDFE435FB2AC62A5719AE" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B63" refString="Panha, S, Burch, JB, 2005. An introduction to the microsnails of Thailand. Malacological Review 37/38(2004-2005): 1-155. [Date of publication follow Jirapatrasilp et al. (2023)]" title="An introduction to the microsnails of Thailand. Malacological Review 37 / 38 (2004 - 2005): 1 - 155. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2005">2005</bibRefCitation>
|
|
). The new species differs from
|
|
<taxonomicName id="4FB33A1A9CA30C170FACA1E723A8B21B" lsidName="G. adela" pageId="0" pageNumber="157" rank="species" species="adela">
|
|
<emphasis id="2FA88B2C90B4D924DE26813B49E7FF86" italics="true" pageId="0" pageNumber="157">G. adela</emphasis>
|
|
</taxonomicName>
|
|
Thompson & Upatham, 1997 in the depressed spire, the short and descending tuba, with prominent spiral striae on the teleoconch, and apertural dentition with a basal plica (
|
|
<bibRefCitation id="5FDE250F6D4D962D0CFB0475C4438805" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida Museum of Natural History" pageId="0" pageNumber="157" pagination="221 - 245" publicationUrl="http://ufdc.ufl.edu/UF00095785/00001" refId="B87" refString="Thompson, FG, Upatham, S, 1997. Vertiginid land snails from Thailand (Gastropoda, Pulmonata, Pupilloidea). Bulletin of the Florida Museum of Natural History 39 (7): 221 - 245, http://ufdc.ufl.edu/UF00095785/00001" title="Vertiginid land snails from Thailand (Gastropoda, Pulmonata, Pupilloidea)." url="http://ufdc.ufl.edu/UF00095785/00001" volume="39" year="1997">Thompson and Upatham 1997</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="A05840157C0AFF53E0DD172A994BBE6A" pageId="0" pageNumber="157">
|
|
Further comparison can be made with species from Peninsular Malaysia that have a very short tuba and few apertural dentitions.
|
|
<taxonomicName id="DBEB2D04BD705F309A17872921A1C857" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Gyliotrachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gyliotrachela aunglini" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="aunglini">
|
|
<emphasis id="93F720A1DFAE0F0DB190CEF1C6AE1BCD" italics="true" pageId="0" pageNumber="157">Gyliotrachela aunglini</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
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<taxonomicName id="597EFB8534D7B48B528BDB9D4898FF7A" lsidName="G. emergens" pageId="0" pageNumber="157" rank="species" species="emergens">
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<emphasis id="FDB71E890304C7FE7765F9FC517A6CD4" italics="true" pageId="0" pageNumber="157">G. emergens</emphasis>
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</taxonomicName>
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van Benthem Jutting, 1950 and
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<taxonomicName id="F1075C0CE5CB0CC5F4D5B64FB0EB33BA" lsidName="G. modesta" pageId="0" pageNumber="157" rank="species" species="modesta">
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<emphasis id="0D8202F5654AF4DE0BB8D72C5542072A" italics="true" pageId="0" pageNumber="157">G. modesta</emphasis>
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</taxonomicName>
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van Benthem Jutting, 1950 in having an elevated spire, short and slightly descending tuba and an angular or weakly angular last whorl. In addition, the apertural dentition of the former species includes a basal plica, and supra- and sub- columellar lamellae, while the latter species has a supracolumellar lamella and a weak infrapalatal plica. Additionally, this new species differs from
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<taxonomicName id="35FA95255CB21EEB03326B2B0AEE0BF2" lsidName="G. troglodytes" pageId="0" pageNumber="157" rank="species" species="troglodytes">
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<emphasis id="BDC036A77FD622679A1EBE12A3E113FE" italics="true" pageId="0" pageNumber="157">G. troglodytes</emphasis>
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</taxonomicName>
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van Benthem Jutting, 1950 in the depressed spire, the very short and descending tuba, the angular last whorl, the apertural dentition with infraparietal and weak angular lamellae, and the strongly wrinkled shell surface (
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<bibRefCitation id="8D0D002D9DBF248467A5AA03CED912A5" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="157" pagination="5 - 47" refId="B94" refString="van Benthem Jutting, WSS, 1950. The Malayan species of Boysidia, Paraboysidia, Hypselostoma and Gyliotrachela (Gastropoda, Pulmonata, Vertiginidae) with a catalogue of all the species hitherto described. Bulletin of the Raffles Museum 21: 5 - 47" title="The Malayan species of Boysidia, Paraboysidia, Hypselostoma and Gyliotrachela (Gastropoda, Pulmonata, Vertiginidae) with a catalogue of all the species hitherto described." volume="21" year="1950">van Benthem Jutting 1950</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="719C52A382B7584C365924A7EF4B04C0" pageId="0" pageNumber="157" type="remarks">
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<paragraph id="50132943F08C5C6ADF1C1C25BDBBF00D" pageId="0" pageNumber="157">Remarks.</paragraph>
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<paragraph id="54A6340B1238B6367399D96F18DABB80" pageId="0" pageNumber="157">The material examined here shows minor variability in terms of shell size, but the key morphological characters, shell sculpture and apertural dentition, are reliable.</paragraph>
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</subSubSection>
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</treatment>
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</document> |