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<mods:title id="DBCFDBB4283C5E266AEEEC50D3F662CA">Review of Baeolidia, the largest genus of Aeolidiidae (Mollusca: Nudibranchia), with the description of five new species</mods:title>
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<mods:namePart id="84F8A82F8316004762EC95E7D511EA44">Carmona, Leila</mods:namePart>
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<mods:namePart id="CA94C2267492B61A296E98B3E1ED05EA">Gosliner, Terrence M.</mods:namePart>
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<mods:namePart id="D3D24AD4BEA59BC4E6CF5F4490570F5E">Cervera, Juan Lucas</mods:namePart>
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<paragraph id="65204B47A030FF8EFF1BFC18E09FFC7D" blockId="12.[151,746,943,970]" box="[151,746,943,970]" pageId="12" pageNumber="489">
<heading id="3E68FC2BA030FF8EFF1BFC18E09FFC7D" bold="true" box="[151,746,943,970]" fontSize="11" level="1" pageId="12" pageNumber="489" reason="1">
<taxonomicName id="A29F30C4A030FF8EFF1BFC18E09FFC7D" ID-CoL="KCLJ" authority="Er. Marcus &amp; Ev. Marcus, 1960" authorityName="Er. Marcus &amp; Ev. Marcus" authorityYear="1960" box="[151,746,943,970]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="species" species="dela">
<emphasis id="57EB9755A030FF8EFF1BFC18E09FFC7D" bold="true" box="[151,746,943,970]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF1BFC18E330FC7E" bold="true" box="[151,325,943,969]" italics="true" pageId="12" pageNumber="489">Baeolidia dela</emphasis>
(
<bibRefCitation id="010E36B6A030FF8EFED8FC07E097FC7D" author="Er" box="[340,738,944,970]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article" year="1960">Er. Marcus &amp; Ev. Marcus, 1960</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="2D8518CCA030FF8EFF1BFC41E155FBBB" box="[151,800,1014,1036]" pageId="12" pageNumber="489" type="reference_group">
<paragraph id="65204B47A030FF8EFF1BFC41E155FBBB" blockId="12.[151,800,1014,1036]" box="[151,800,1014,1036]" pageId="12" pageNumber="489">
<treatmentCitationGroup id="458F6C69A030FF8EFF1BFC41E155FBBB" box="[151,800,1014,1036]" pageId="12" pageNumber="489">
<taxonomicName id="A29F30C4A030FF8EFF1BFC41E0E2FBBB" ID-CoL="LKTX" authority="Er. Marcus &amp; Ev. Marcus, 1960: 924" authorityName="Er. Marcus &amp; Ev. Marcus" authorityPageNumber="924" authorityYear="1960" box="[151,663,1014,1036]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="species" species="dela">
<emphasis id="57EB9755A030FF8EFF1BFC41E36FFBBB" box="[151,282,1014,1036]" italics="true" pageId="12" pageNumber="489">Berghia dela</emphasis>
<treatmentCitation id="E43E6D56A030FF8EFEACFC40E0E2FBBB" author="Er" box="[288,663,1014,1036]" page="924" pageId="12" pageNumber="489" year="1960">
<bibRefCitation id="010E36B6A030FF8EFEACFC40E010FBBB" author="Er" box="[288,613,1014,1036]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article" year="1960">Er. Marcus &amp; Ev. Marcus, 1960</bibRefCitation>
: 924
</treatmentCitation>
</taxonomicName>
, Figs. 8386.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2D8518CCA030FF8EFF1BFB8EE01FFAB1" pageId="12" pageNumber="489" type="materials_examined">
<paragraph id="65204B47A030FF8EFF1BFB8EE0E4FBE5" blockId="12.[151,1436,1081,1718]" box="[151,657,1081,1106]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF1BFB8EE340FBE5" bold="true" box="[151,309,1081,1106]" pageId="12" pageNumber="489">
<typeStatus id="BA24F5E5A030FF8EFF1BFB8EE2A7FBE5" box="[151,210,1081,1106]" pageId="12" pageNumber="489">Type</typeStatus>
locality.
</emphasis>
Addu-Atoll,
<collectingCountry id="1D880BD7A030FF8EFE47FB8EE046FBE5" box="[459,563,1081,1106]" name="Maldives" pageId="12" pageNumber="489">Maldives</collectingCountry>
Islands.
</paragraph>
<paragraph id="65204B47A030FF8EFF4BFBEAE127FBC1" blockId="12.[151,1436,1081,1718]" box="[199,850,1117,1142]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF4BFBEAE300FBC1" bold="true" box="[199,373,1117,1142]" pageId="12" pageNumber="489">
<typeStatus id="BA24F5E5A030FF8EFF4BFBEAE377FBC1" box="[199,258,1117,1142]" pageId="12" pageNumber="489">Type</typeStatus>
material.
</emphasis>
To our knowledge no
<typeStatus id="BA24F5E5A030FF8EFDFDFBE9E0D4FBC1" box="[625,673,1118,1142]" pageId="12" pageNumber="489">type</typeStatus>
material exists.
</paragraph>
<paragraph id="65204B47A030FF8EFF4BFB36E34EFB09" blockId="12.[151,1436,1081,1718]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF4BFB36E073FB2D" bold="true" box="[199,518,1153,1178]" pageId="12" pageNumber="489">Geographical distribution.</emphasis>
So far, this species is only known from the
<collectingCountry id="1D880BD7A030FF8EFB8EFB36E61FFB2D" box="[1026,1130,1153,1178]" name="Maldives" pageId="12" pageNumber="489">Maldives</collectingCountry>
Islands (
<bibRefCitation id="010E36B6A030FF8EFB5FFB35E35AFB09" author="Er" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article" year="1960">Er. Marcus &amp; Ev. Marcus 1960</bibRefCitation>
).
</paragraph>
<paragraph id="65204B47A030FF8EFF4BFB7EE01FFAB1" blockId="12.[151,1436,1081,1718]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF4BFB7EE00BFB55" bold="true" box="[199,638,1225,1250]" pageId="12" pageNumber="489">External and internal morphology.</emphasis>
Described and figured by
<bibRefCitation id="010E36B6A030FF8EFC41FB7DE712FB55" box="[973,1383,1225,1250]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article">Er. Marcus &amp; Ev. Marcus (1960)</bibRefCitation>
. No specimens available for the present study.
</paragraph>
</subSubSection>
<subSubSection id="2D8518CCA030FF8EFF4BFAA6E033F901" pageId="12" pageNumber="489" type="discussion">
<paragraph id="65204B47A030FF8EFF4BFAA6E033F901" blockId="12.[151,1436,1081,1718]" pageId="12" pageNumber="489">
<emphasis id="57EB9755A030FF8EFF4BFAA6E34EFA9D" bold="true" box="[199,315,1297,1322]" pageId="12" pageNumber="489">Remarks.</emphasis>
Ernest &amp; Eveline
<bibRefCitation id="010E36B6A030FF8EFD98FAA5E0C9FA9D" author="Marcus" box="[532,700,1297,1322]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article" year="1960">Marcus (1960)</bibRefCitation>
described
<taxonomicName id="A29F30C4A030FF8EFCB7FAA6E1BBFA9D" box="[827,974,1297,1322]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="species" species="dela">
<emphasis id="57EB9755A030FF8EFCB7FAA6E1BBFA9D" box="[827,974,1297,1322]" italics="true" pageId="12" pageNumber="489">Berghia dela</emphasis>
</taxonomicName>
, based on a single specimen. Since this specimen was already preserved,
<bibRefCitation id="010E36B6A030FF8EFD94FA81E1E0FAF9" box="[536,917,1333,1358]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article">Er. Marcus &amp; Ev. Marcus (1960)</bibRefCitation>
could not provide any information about the colouration of the living animal. However, they described and drew the papillate rhinophores, the jaws, the radular tooth and the penis (
<bibRefCitation id="010E36B6A030FF8EFEF6FAC9E0ACFA21" author="Er" box="[378,729,1405,1430]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article" year="1960">Er. Marcus &amp; Ev. Marcus 1960</bibRefCitation>
, figs. 8486). The cerata were depicted as cylindrical and were arranged in arches plus rows. Recently,
<bibRefCitation id="010E36B6A030FF8EFDE7FA15E10FFA0D" box="[619,890,1441,1466]" pageId="12" pageNumber="489" refString="Carmona, L., Pola, M, Gosliner, T. M., Cervera, J. L. (2014 b) The Atlantic-Mediterranean genus Berghia (Mollusca, Nudibranchia, Aeolidiidae). Taxonomic review and phylogenetic analysis. Journal of Molluscan Studies. [in press]" type="book">
Carmona
<emphasis id="57EB9755A030FF8EFD50FA15E16DFA0D" box="[732,792,1441,1466]" italics="true" pageId="12" pageNumber="489">et al.</emphasis>
(2014b)
</bibRefCitation>
stated the main characteristics of the aeolidiid genus
<taxonomicName id="A29F30C4A030FF8EFF6EFA72E34EFA69" box="[226,315,1477,1502]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A030FF8EFF6EFA72E34EFA69" box="[226,315,1477,1502]" italics="true" pageId="12" pageNumber="489">Berghia</emphasis>
</taxonomicName>
. One of the most important is the papillate morphology of the rhinophores, which can be easily distinguished from the rhinophorial ornamentation of
<taxonomicName id="A29F30C4A030FF8EFD73FA5EE119F9B5" box="[767,876,1513,1538]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A030FF8EFD73FA5EE119F9B5" box="[767,876,1513,1538]" italics="true" pageId="12" pageNumber="489">Baeolidia</emphasis>
</taxonomicName>
(see general Discussion). Additionally, based on these authors investigations,
<taxonomicName id="A29F30C4A030FF8EFE68F9BAE048F991" box="[484,573,1549,1574]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A030FF8EFE68F9BAE048F991" box="[484,573,1549,1574]" italics="true" pageId="12" pageNumber="489">Berghia</emphasis>
</taxonomicName>
is restricted to the Atlantic and Mediterranean. Therefore, according to these statements,
<taxonomicName id="A29F30C4A030FF8EFE91F986E3C5F9FD" box="[285,432,1585,1610]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="species" species="dela">
<emphasis id="57EB9755A030FF8EFE91F986E3C5F9FD" box="[285,432,1585,1610]" italics="true" pageId="12" pageNumber="489">Berghia dela</emphasis>
</taxonomicName>
should be transferred to
<taxonomicName id="A29F30C4A030FF8EFD58F986E134F9FD" box="[724,833,1585,1610]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A030FF8EFD58F986E134F9FD" box="[724,833,1585,1610]" italics="true" pageId="12" pageNumber="489">Baeolidia</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="010E36B6A030FF8EFC49F985E734F9FD" box="[965,1345,1585,1610]" pageId="12" pageNumber="489" refString="Marcus, Er. &amp; Marcus, Ev. (1960) Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 12, 873 - 934." type="journal article">Er. Marcus &amp; Ev. Marcus (1960)</bibRefCitation>
did not give any information about colouration, which is usually critical to recognize and separate species within
<taxonomicName id="A29F30C4A030FF8EFF1BF9CEE36FF925" box="[151,282,1657,1682]" class="Gastropoda" family="Aeolidiidae" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="489" phylum="Mollusca" rank="family">Aeolidiidae</taxonomicName>
(personal observation), the cylindrical cerata could be the distinctive feature that may help to identify this species if it is ever reported again.
</paragraph>
</subSubSection>
</treatment>
</document>