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<document id="D31485FEAA3F99001CB967EAC9EE61CA" ID-DOI="10.11646/zootaxa.5486.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="13210082" ID-ZooBank="3181DB14-02DC-470C-9BE3-C7B291BBFA3F" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1722788564901" checkinUser="plazi" docAuthor="Roca-Cusachs, Marcos, Rider, David A., Kment, Petr &amp; Jung, Sunghoon" docDate="2024" docId="E6426A70B914FFD585C51864FD15FE57" docLanguage="en" docName="zootaxa.5486.2.3.5.pdf" docOrigin="Zootaxa 5486 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vitruvius insignis Distant 1901" docType="treatment" docVersion="3" lastPageNumber="426" masterDocId="1A7B1208B917FFD285521C53FFDAFFFA" masterDocTitle="Revision of Vitruvius (Hemiptera: Heteroptera: Pentatomidae) with the description of a new species from Cambodia and Thailand" masterLastPageNumber="434" masterPageNumber="419" pageNumber="422" updateTime="1726250890409" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:title id="9F78BA94B98C7D20A2834B471BBF675D">Revision of Vitruvius (Hemiptera: Heteroptera: Pentatomidae) with the description of a new species from Cambodia and Thailand</mods:title>
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<mods:namePart id="360073DC2D9AFF85F467305D9E184221">Roca-Cusachs, Marcos</mods:namePart>
<mods:affiliation id="592C2B51CECB9D60DA4EA51CF855E79A">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and Life Sciences, Chungnam National University, Daejeon, Korea &amp; IRBio. Institut de Recerca a la Biodiversitat, Universitat de Barcelona, Barcelona, Spain &amp; Evolution &amp; Ecology Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia</mods:affiliation>
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<mods:namePart id="95E752E7032B1E22C0D0A6166F5A7CF1">Rider, David A.</mods:namePart>
<mods:affiliation id="4C622B43BBC08604C1061A3364BC4961">Department of Entomology, North Dakota State University, Fargo, North Dakota, USA</mods:affiliation>
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<mods:namePart id="5C7F15C244CD13F7B3C54DF3F21FA5A6">Kment, Petr</mods:namePart>
<mods:affiliation id="796AEBB160EC5F85BF37DDABE07E742C">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic</mods:affiliation>
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<mods:namePart id="2827E7E6452F31316326DBCE02F22BB2">Jung, Sunghoon</mods:namePart>
<mods:affiliation id="D6286070AB6257DDB417C2448466C4E4">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and Life Sciences, Chungnam National University, Daejeon, Korea &amp; Department of Smart Agriculture Systems, College of Agriculture and Life Sciences, Chungnam National University, Daejeon, Korea * Corresponding authors</mods:affiliation>
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<heading id="351C6C0AB914FFD185C51864FDC2FBAA" bold="true" box="[151,536,1078,1105]" fontSize="11" level="1" pageId="3" pageNumber="422" reason="1">
<taxonomicName id="A9EBA0E5B914FFD185C51864FDC2FBAA" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[151,536,1078,1105]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B914FFD185C51864FDC2FBAA" bold="true" box="[151,536,1078,1105]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185C51864FEB6FBAB" bold="true" box="[151,364,1079,1105]" italics="true" pageId="3" pageNumber="422">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B914FFD184211865FDC2FBAA" author="Distant, W. L." box="[371,536,1078,1104]" pageId="3" pageNumber="422" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="26F188EDB914FFD185C51808FE39FB8F" box="[151,483,1115,1141]" pageId="3" pageNumber="422" type="description">
<paragraph id="6E54DB66B914FFD185C51808FE39FB8F" blockId="3.[151,536,1078,1141]" box="[151,483,1115,1141]" pageId="3" pageNumber="422">
(
<figureCitation id="F6D0C7E3B914FFD185CD1808FED0FB8F" box="[159,266,1115,1141]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="3" pageNumber="422">Figures 1</figureCitation>
,
<figureCitation id="F6D0C7E3B914FFD1844A1808FE01FB8F" box="[280,475,1115,1141]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">2A, 2B, 2C &amp; 2E</figureCitation>
)
</paragraph>
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<treatmentCitation id="EF4AFD77B914FFD185C518C9FDD3FB48" author="Distant, W. L." box="[151,521,1178,1202]" page="112" pageId="3" pageNumber="422" year="1901">
<taxonomicName id="A9EBA0E5B914FFD185C518C9FDD3FB48" authority="Distant, 1901: 112" authorityName="Distant" authorityPageNumber="112" authorityYear="1901" box="[151,521,1178,1202]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B914FFD185C518C9FE92FB4B" box="[151,328,1178,1201]" italics="true" pageId="3" pageNumber="422">Vitruvius insignis</emphasis>
<materialsCitation id="DE83D13BB914FFD1841C18C9FDD3FB48" box="[334,521,1178,1202]" collectionCode="MCSN" country="Myanmar" location="Rangoon'" pageId="3" pageNumber="422" specimenCount="1" typeStatus="syntype">
<bibRefCitation id="0A7AA697B914FFD1841C18C9FDD3FB48" author="Distant, W. L." box="[334,521,1178,1202]" pageId="3" pageNumber="422" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901: 112</bibRefCitation>
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(original description).
<typeStatus id="B15065C4B914FFD187A018C9FC96FB48" box="[754,844,1178,1202]" pageId="3" pageNumber="422" type="syntype">Syntype</typeStatus>
(s):
<collectingCountry id="16FC9BF6B914FFD1862318C9FC67FB48" box="[881,957,1178,1202]" name="Myanmar" pageId="3" pageNumber="422">Burma</collectingCountry>
,
<location id="6B348DBDB914FFD1869518C9FBF3FB48" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB914FFD1869518C9FBF3FB48" box="[967,1065,1178,1202]" country="Myanmar" name="Rangoon'" pageId="3" pageNumber="422">Rangoon</location>
(MCSN).
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</paragraph>
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<taxonomicName id="A9EBA0E5B914FFD185C518EAFE90FB2A" authorityName="Distant" authorityYear="1901" box="[151,330,1209,1232]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B914FFD185C518EAFE90FB2A" box="[151,330,1209,1232]" italics="true" pageId="3" pageNumber="422">Vitruvius insignis</emphasis>
</taxonomicName>
:
<treatmentCitation id="EF4AFD77B914FFD1840B18EBFDFFFB2B" author="Distant, W. L." box="[345,549,1208,1233]" page="278" pageId="3" pageNumber="422" year="1902">
<bibRefCitation id="0A7AA697B914FFD1840B18EBFE29FB2B" author="Distant, W. L." box="[345,499,1208,1233]" pageId="3" pageNumber="422" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant (1902)</bibRefCitation>
: 278
</treatmentCitation>
279 (redescription, figures);
<treatmentCitation id="EF4AFD77B914FFD1860818EBFBECFB2A" author="Bergroth, E." box="[858,1078,1208,1233]" page="186" pageId="3" pageNumber="422" year="1908">
<bibRefCitation id="0A7AA697B914FFD1860818EBFBDEFB2B" author="Bergroth, E." box="[858,1028,1208,1233]" pageId="3" pageNumber="422" pagination="131 - 200" refId="ref7616" refString="Bergroth, E. (1908) Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique, 15 (10), 131 - 200." type="journal article" year="1908">Bergroth (1908)</bibRefCitation>
: 186
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1819218EBFA46FB2A" author="Kirkaldy, G. W." box="[1216,1436,1208,1233]" page="353" pageId="3" pageNumber="422" year="1909">
<bibRefCitation id="0A7AA697B914FFD1819218EBFAB0FB2B" author="Kirkaldy, G. W." box="[1216,1386,1208,1233]" pageId="3" pageNumber="422" refId="ref8800" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, (xl) + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 353
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1841B1884FDCFFB15" author="Distant, W. L." box="[329,533,1239,1263]" page="164" pageId="3" pageNumber="422" year="1921">
<bibRefCitation id="0A7AA697B914FFD1841B1884FE39FB15" author="Distant, W. L." box="[329,483,1239,1263]" pageId="3" pageNumber="422" pagination="164 - 169" refId="ref7952" refString="Distant, W. L. (1921) The Heteroptera of Indo-China. Entomologist, London, 54, 164 - 169." type="journal article" year="1921">Distant (1921)</bibRefCitation>
: 164
</treatmentCitation>
(listed from Indo-China, no exact locality);
<treatmentCitation id="EF4AFD77B914FFD186BB1884FB0EFB15" author="Hoffmann, W. E." box="[1001,1236,1239,1263]" page="144" pageId="3" pageNumber="422" year="1931">
<bibRefCitation id="0A7AA697B914FFD186BB1884FB7BFB15" author="Hoffmann, W. E." box="[1001,1185,1239,1263]" pageId="3" pageNumber="422" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann (1931)</bibRefCitation>
: 144
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD1840618A5FD99FAF4" author="Hoffmann, W. E." box="[340,579,1270,1294]" page="10" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1840618A5FDC1FAF4" author="Hoffmann, W. E." box="[340,539,1270,1294]" pageId="3" pageNumber="422" pagination="6 - 11" refId="ref8297" refString="Hoffmann, W. E. (1932 a) A list of the Pentatomidae, Plataspidae, and Coreidae (order Hemiptera) of China, Korea, and Indo-China. Journal of the Pan-Pacific Research Institution, 7, 6 - 11." type="journal article" year="1932">Hoffmann (1932a)</bibRefCitation>
: 10
</treatmentCitation>
(checklist);
<treatmentCitation id="EF4AFD77B914FFD1879C18A5FC16FAF4" author="Hoffmann, W. E." box="[718,972,1270,1294]" page="139" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1879C18A5FC4DFAF4" author="Hoffmann, W. E." box="[718,919,1270,1294]" pageId="3" pageNumber="422" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">Hoffmann (1932b)</bibRefCitation>
: 139
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD1806718A5FE86FAD6" author="Hoffmann, W. E." page="1016" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1806718A5FEC0FAD7" author="Hoffmann, W. E." pageId="3" pageNumber="422" pagination="1010 - 1027" refId="ref8379" refString="Hoffmann, W. E. (1932 c) Notes on the bionomics of some oriental Pentatomidae (Hemiptera). Archivo Zoologico Italiano, 16, 1010 - 1027." type="journal article" year="1932">Hoffmann (1932c)</bibRefCitation>
: 1016
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD187931947FC1AFAD7" author="Hoffmann, W. E." box="[705,960,1300,1325]" page="470" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD187931947FC53FAD6" author="Hoffmann, W. E." box="[705,905,1300,1324]" pageId="3" pageNumber="422" refId="ref8411" refString="Hoffmann, W. E. (1932 d) Brief note on Vitruvius insignis Distant. Lingnan Science Journal, 11 (3), 470, pl. 8." type="journal volume" year="1932">Hoffmann (1932d)</bibRefCitation>
: 470
</treatmentCitation>
, pl. 8: figs 14 (larval morphology, habitus drawings of adult and larva, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD186641960FC05FAB1" author="Cheo, M. - T." box="[822,991,1331,1355]" page="31" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD186641960FC61FAB1" author="Cheo, M. - T." box="[822,955,1331,1355]" pageId="3" pageNumber="422" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo (1935)</bibRefCitation>
: 31
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD180671960FE90FA90" author="Hoffmann, W. E." page="132" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD180671960FEC0FA90" author="Hoffmann, W. E." pageId="3" pageNumber="422" pagination="1 - 294" refId="ref8443" refString="Hoffmann, W. E. (1935 a) An abridged catalogue of certain Scutelleroidea (Plataspidae, Scutelleridae, and Pentatomidae) of China, Chosen, Indo-China, and Taiwan. Lingnan University Science Bulletin, 7, 1 - 294, 2 maps." type="book chapter" year="1935">Hoffmann (1935a)</bibRefCitation>
: 132
</treatmentCitation>
, 177 (catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD1872F1901FC84FA90" author="Hoffmann, W. E." box="[637,862,1362,1386]" page="61" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD1872F1901FCE4FA90" author="Hoffmann, W. E." box="[637,830,1362,1386]" pageId="3" pageNumber="422" pagination="57 - 62" refId="ref8491" refString="Hoffmann, W. E. (1935 b) Notes on eight pentatomids not hitherto recorded from Formosa. Transactions of the Natural History Society of Formosa, 25 (136 - 139), 57 - 62." type="journal article" year="1935">Hoffmann (1935b)</bibRefCitation>
: 61
</treatmentCitation>
62 (catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD181DB1901FAFDFA90" author="Wu, C. F." box="[1161,1319,1362,1386]" page="225" pageId="3" pageNumber="422" year="1933">
<bibRefCitation id="0A7AA697B914FFD181DB1901FB23FA90" author="Wu, C. F." box="[1161,1273,1362,1386]" pageId="3" pageNumber="422" pagination="203 - 231" refId="ref10735" refString="Wu, C. F. (1933) A preliminary check list of Hemiptera Heteroptera recorded from Kwangtung Province, South China. Lingnan Science Journal, Supplement, 12, 203 - 231." type="journal article" year="1933">Wu (1933)</bibRefCitation>
: 225
</treatmentCitation>
(catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD184031922FE25FA72" author="Tang, T. - H." box="[337,511,1392,1417]" page="366" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD184031922FE0AFA72" author="Tang, T. - H." box="[337,464,1392,1417]" pageId="3" pageNumber="422" pagination="275 - 382" refId="ref10478" refString="Tang, T. - H. (1935) Family Pentatomidae. In: Wu, C. - F. F., Catalogus insectorum sinensium. (Catalogue of Chinese Insects). Vol. II. Fan Memorial Institute of Biology, Peiping, pp. 275 - 382. [Book pagination: 634 pp.]" type="book chapter" year="1935">Tang (1935)</bibRefCitation>
: 366
</treatmentCitation>
(catalogue, distribution); Blöte (1945): 299 (distribution);
<treatmentCitation id="EF4AFD77B914FFD1811D1922FAB7FA72" author="Kumar, R. &amp; Ghauri, M. S. K." box="[1103,1389,1392,1417]" page="25" pageId="3" pageNumber="422" year="1970">
<bibRefCitation id="0A7AA697B914FFD1811D1922FA92FA73" author="Kumar, R. &amp; Ghauri, M. S. K." box="[1103,1352,1392,1417]" pageId="3" pageNumber="422" pagination="1 - 32" refId="ref9189" refString="Kumar, R. &amp; Ghauri, M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae). Deutsche Entomologische Zeitschrift, 17 (1 - 3), 1 - 32. https: // doi. org / 10.1002 / mmnd. 4810170102" type="journal article" year="1970">Kumar &amp; Ghauri (1970)</bibRefCitation>
: 25
</treatmentCitation>
: figs 96100, 26, 28: fig. 117 (description and drawings of male and female genitalia);
<treatmentCitation id="EF4AFD77B914FFD186AC19DCFAD6FA5D" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[1022,1292,1423,1447]" page="66" pageId="3" pageNumber="422" year="1977">
<bibRefCitation id="0A7AA697B914FFD186AC19DCFB3DFA5D" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[1022,1255,1423,1447]" pageId="3" pageNumber="422" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng (1977)</bibRefCitation>
: 66
</treatmentCitation>
, pl. 7: fig. 110 (diagnosis, habitus photograph, distribution);
<treatmentCitation id="EF4AFD77B914FFD187C119FDFC9EFA3C" author="Chen, Z. - Y." box="[659,836,1454,1478]" page="133" pageId="3" pageNumber="422" year="1982">
<bibRefCitation id="0A7AA697B914FFD187C119FDFCCCFA3C" author="Chen, Z. - Y." box="[659,790,1454,1478]" pageId="3" pageNumber="422" refId="ref7694" refString="Chen, Z. - Y. (1982) On the Pentatomidae of Dinghushan, Guangdong Province. Acta Scientiarum Naturalium Universitatis Sunyatseni, 1981 (4), 133 - 135. [in Chinese, English summary]" type="book" year="1982">Chen (1982)</bibRefCitation>
: 133
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1868E19FDFB05FA3C" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[988,1247,1454,1478]" page="113" pageId="3" pageNumber="422" year="1982">
<bibRefCitation id="0A7AA697B914FFD1868E19FDFB68FA3C" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[988,1202,1454,1478]" pageId="3" pageNumber="422" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou (1982)</bibRefCitation>
: 113
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD18406199FFDD0FA1F" author="Chen, Q. - H." box="[340,522,1484,1509]" page="117" pageId="3" pageNumber="422" year="1985">
<bibRefCitation id="0A7AA697B914FFD18406199FFE03FA1E" author="Chen, Q. - H." box="[340,473,1484,1508]" pageId="3" pageNumber="422" refId="ref7647" refString="Chen, Q. - H. (1985) The catalogue and geographical distribution of bugs (Hemiptera: Pentatomoidea) in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 11 (1), 115 - 127. [in Chinese, English summary]" type="book" year="1985">Chen (1985)</bibRefCitation>
: 117
</treatmentCitation>
(distribution); Zhang (1985): 4950, pl. XLVI: fig. 155 (redescription, habitus drawing, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD1875E19B8FD26F9F9" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[524,764,1515,1539]" page="10" pageId="3" pageNumber="422" year="1985">
<bibRefCitation id="0A7AA697B914FFD1875E19B8FD02F9F9" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[524,728,1515,1539]" pageId="3" pageNumber="422" refId="ref10917" refString="Zhang, S. - M., Lin, Y. - J. &amp; Chen, Z. - Q. (1985) Faunal analysis of the pentatomid bugs in Jian-Feng-Ling, Hainan. Acta Agriculturae Universitatis Jiangxiensis, 1985 (2), 9 - 12. [in Chinese, English summary]" type="book" year="1985">
Zhang
<emphasis id="5C9F0774B914FFD1870719BFFD50F9F9" box="[597,650,1515,1539]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(1985)
</bibRefCitation>
: 10
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD186CB19B8FB0BF9F9" author="Schaefer, C. W. &amp; Ahmad, I." box="[921,1233,1515,1539]" page="27" pageId="3" pageNumber="422" year="1987">
<bibRefCitation id="0A7AA697B914FFD186CB19B8FB77F9F9" author="Schaefer, C. W. &amp; Ahmad, I." box="[921,1197,1515,1539]" pageId="3" pageNumber="422" pagination="21 - 34" refId="ref10436" refString="Schaefer, C. W. &amp; Ahmad, I. (1987) The food plants of four pentatomoid families (Hemiptera: Acanthosomatidae, Tessaratomidae, Urostylidae, and Dinidoridae). Phytophaga, 1, 21 - 34." type="journal article" year="1987">Schaefer &amp; Ahmad (1987)</bibRefCitation>
: 27
</treatmentCitation>
(plant association);
<treatmentCitation id="EF4AFD77B914FFD185951A59FE04F9D8" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[199,478,1546,1570]" page="290" pageId="3" pageNumber="422" year="1988">
<bibRefCitation id="0A7AA697B914FFD185951A59FE74F9D8" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[199,430,1546,1570]" pageId="3" pageNumber="422" refId="ref10813" refString="Zhang, S. - M. &amp; Chen, Q. - H. (1988) Faunal structure of the pentatomid bugs in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 14 (3), 287 - 291. [in Chinese, English summary]" type="book" year="1988">Zhang &amp; Chen (1988)</bibRefCitation>
: 290
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1872A1A59FCC5F9D8" author="Chen, Z. - Y." box="[632,799,1546,1570]" page="59" pageId="3" pageNumber="422" year="1989">
<bibRefCitation id="0A7AA697B914FFD1872A1A59FD26F9D8" author="Chen, Z. - Y." box="[632,764,1546,1570]" pageId="3" pageNumber="422" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">Chen (1989)</bibRefCitation>
: 59
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD181201A59FAD4F9D8" author="Hua, L. - Z." box="[1138,1294,1546,1570]" page="45" pageId="3" pageNumber="422" year="1989">
<bibRefCitation id="0A7AA697B914FFD181201A59FB31F9D8" author="Hua, L. - Z." box="[1138,1259,1546,1570]" pageId="3" pageNumber="422" refId="ref8652" refString="Hua, L. - Z. (1989) List of insect specimens of Zhongshan (Sun Yatsen) University. Taxonomy Division, Institute of Entomology, Zhongshan (Sun Yatsen) University, Ghungzhou, ii + 158 pp. [in Chinese, English summary]" type="book" year="1989">Hua (1989)</bibRefCitation>
: 45
</treatmentCitation>
(species list);
<treatmentCitation id="EF4AFD77B914FFD185951A7BFE6AF9BB" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[199,432,1576,1601]" page="2" pageId="3" pageNumber="422" year="1990">
<bibRefCitation id="0A7AA697B914FFD185951A7BFE43F9BB" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[199,409,1576,1601]" pageId="3" pageNumber="422" refId="ref10862" refString="Zhang, S. - M. &amp; Lin, Y. - J. (1990) The newly recorded pentatomid bugs in Xishuangbana [sic!] and their faunal structures. Acta Agriculturae Universitatis Jiangxiensis, 12 (1), 287 - 291. [in Chinese, English summary]" type="book" year="1990">Zhang &amp; Lin (1990)</bibRefCitation>
: 2
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1871B1A7BFD20F9BB" author="Zheng, L. - Y." box="[585,762,1576,1601]" page="92" pageId="3" pageNumber="422" year="1994">
<bibRefCitation id="0A7AA697B914FFD1871B1A7BFD0DF9BB" author="Zheng, L. - Y." box="[585,727,1576,1601]" pageId="3" pageNumber="422" pagination="91 - 96" refId="ref11035" refString="Zheng, L. - Y. (1994) Heteropteran insects (Hemiptera) feeding on bamboos in China. Annals of the Entomological Society of America, 87 (1), 91 - 96. https: // doi. org / 10.1093 / aesa / 87.1.91" type="journal article" year="1994">Zheng (1994)</bibRefCitation>
: 92
</treatmentCitation>
(plant association);
<treatmentCitation id="EF4AFD77B914FFD1869A1A7AFBAAF9BB" author="Hua, L. - Z." box="[968,1136,1577,1601]" page="170" pageId="3" pageNumber="422" year="2000">
<bibRefCitation id="0A7AA697B914FFD1869A1A7AFB9AF9BB" author="Hua, L. - Z." box="[968,1088,1577,1601]" pageId="3" pageNumber="422" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua (2000)</bibRefCitation>
: 170
</treatmentCitation>
(catalogue, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD184031A14FDE7F9A5" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[337,573,1607,1631]" page="146" pageId="3" pageNumber="422" year="2002">
<bibRefCitation id="0A7AA697B914FFD184031A14FDD7F9A5" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[337,525,1607,1631]" pageId="3" pageNumber="422" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B914FFD184DD1A1BFE61F9A5" box="[399,443,1607,1631]" italics="true" pageId="3" pageNumber="422">et al</emphasis>
. (2002)
</bibRefCitation>
: 146
</treatmentCitation>
(checklist, distribution);
<treatmentCitation id="EF4AFD77B914FFD186691A14FB96F9A5" author="Zheng, L. - Y. &amp; Chen, C." box="[827,1100,1607,1631]" page="256" pageId="3" pageNumber="422" year="2002">
<bibRefCitation id="0A7AA697B914FFD186691A14FBC6F9A5" author="Zheng, L. - Y. &amp; Chen, C." box="[827,1052,1607,1631]" pageId="3" pageNumber="422" pagination="256 - 261" refId="ref11085" refString="Zheng, L. - Y. &amp; Chen, C. (2002) Hemiptera: Pentatomidae. In Huang, F. - S. (Ed.), Forest Insects of Hainan. Science Press, Beijing, pp. 256 - 261. [Book pagination: 1064 pp., in Chinese, English summary]" type="book chapter" year="2002">Zheng &amp; Chen (2002)</bibRefCitation>
: 256
</treatmentCitation>
(identification key);
<treatmentCitation id="EF4AFD77B914FFD180721A14FE9AF984" author="Liu, G. - Q. &amp; Wang, H. - J." page="189" pageId="3" pageNumber="422" year="2004">
<bibRefCitation id="0A7AA697B914FFD180721A14FED4F984" author="Liu, G. - Q. &amp; Wang, H. - J." pageId="3" pageNumber="422" pagination="173 - 189" refId="ref9251" refString="Liu, G. - Q. &amp; Wang, H. - J. (2004) Hemiptera: Scutelleridae, Tessaratomidae, Dinidoridae and Pentatomidae. In: Yang, X. - K. (Ed.), Insects from Mt. Shiwandashan area of Guangxi. China Forestry Publishing House, Beijing, pp. 173 - 189. [in Chinese, English summary]" type="book chapter" year="2004">Liu &amp; Wang (2004)</bibRefCitation>
: 189
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1848E1A35FD4EF984" author="Rider, D. A." box="[476,660,1638,1662]" page="376" pageId="3" pageNumber="422" year="2006">
<bibRefCitation id="0A7AA697B914FFD1848E1A35FDB9F984" author="Rider, D. A." box="[476,611,1638,1662]" pageId="3" pageNumber="422" refId="ref9388" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402. [Book pagination: xiii + 550 pp.]" type="book" year="2006">Rider (2006)</bibRefCitation>
: 376
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1864F1A35FC1EF984" author="Fan, Z. - H." box="[797,964,1638,1662]" page="521" pageId="3" pageNumber="422" year="2011">
<bibRefCitation id="0A7AA697B914FFD1864F1A35FC48F984" author="Fan, Z. - H." box="[797,914,1638,1662]" pageId="3" pageNumber="422" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan (2011)</bibRefCitation>
: 521
</treatmentCitation>
, 522: fig. 2.275, 523 (redescription, drawings of male and female genitalia, genitalia), 609: figs 578579, 610 (habitus photographs);
<treatmentCitation id="EF4AFD77B914FFD181151AD7FAE0F967" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1095,1338,1668,1693]" page="110" pageId="3" pageNumber="422" year="2018">
<bibRefCitation id="0A7AA697B914FFD181151AD7FAD3F967" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1095,1289,1668,1693]" pageId="3" pageNumber="422" pagination="25 - 201" refId="ref9452" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of the Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): Biology, higher systematics, semiochemistry, and management. CRC Press, Boca Raton, Florida, pp. 25 - 201. [Book pagination: xix + 819 pp.] https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider
<emphasis id="5C9F0774B914FFD181D51AD6FB61F966" box="[1159,1211,1668,1692]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(2018)
</bibRefCitation>
: 110
</treatmentCitation>
, 193: fig. 2.24H (habitus photograph).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6E54DB66B914FFD185C51A91FC77F920" blockId="3.[151,1437,1178,1754]" box="[151,941,1730,1754]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185C51A91FF25F923" box="[151,255,1730,1753]" italics="true" pageId="3" pageNumber="422">Vitruvinus</emphasis>
[
<emphasis id="5C9F0774B914FFD1845C1A91FEE9F923" box="[270,307,1730,1753]" italics="true" pageId="3" pageNumber="422">sic!</emphasis>
]
<emphasis id="5C9F0774B914FFD184131A91FE4AF923" box="[321,400,1730,1753]" italics="true" pageId="3" pageNumber="422">insignis</emphasis>
:
<bibRefCitation id="0A7AA697B914FFD184CF1A91FD9BF920" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[413,577,1730,1754]" pageId="3" pageNumber="422" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B914FFD184931A90FE2EF920" box="[449,500,1730,1754]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(2011)
</bibRefCitation>
: 59 (plant association, distribution).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB914FFD485C51B5BFCA4FD7F" lastPageId="6" lastPageNumber="425" pageId="3" pageNumber="422" type="materials_examined">
<paragraph id="6E54DB66B914FFD185C51B5BFB90F848" blockId="3.[151,1437,1800,1971]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185C51B5BFE9CF8D8" bold="true" box="[151,326,1800,1826]" pageId="3" pageNumber="422">Type material.</emphasis>
<materialsCitation id="DE83D13BB914FFD1841F1B5AFB90F848" collectingDate="1888-12" collectorName="M. Roca-Cuschachs" pageId="3" pageNumber="422" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="B15065C4B914FFD1841F1B5AFE67F8D9" box="[333,445,1801,1827]" pageId="3" pageNumber="422" type="lectotype">Lectotype</typeStatus>
(here designated):
<specimenCount id="78ED10EFB914FFD187C51B5AFD76F8D9" box="[663,684,1801,1827]" count="1" pageId="3" pageNumber="422" type="female"></specimenCount>
(
<figureCitation id="F6D0C7E3B914FFD187EE1B5BFCCCF8D8" box="[700,790,1800,1826]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">Fig. 2A</figureCitation>
), “g,
<date id="1A55FDA6B914FFD186061B5AFC67F8D8" box="[852,957,1800,1827]" pageId="3" pageNumber="422" value="1888-12">
<collectingDate id="0A11044EB914FFD186061B5AFC67F8D8" box="[852,957,1800,1827]" pageId="3" pageNumber="422" value="1888-12">XII 1888</collectingDate>
</date>
” [darkend, mostly illegible, interpreted by comparison with other locality labels from Fea] // “Typus” [printed in orange] //
<taxonomicName id="A9EBA0E5B914FFD181481B7FFB58F8BC" authorityName="Distant" authorityYear="1901" box="[1050,1154,1836,1862]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Vitruvius</taxonomicName>
/ insignis / Dist. / Typus! [handwritten in black ink, orange label] // “
<taxonomicName id="A9EBA0E5B914FFD187C51B03FD25F890" authorityName="Distant" authorityYear="1901" box="[663,767,1872,1898]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Vitruvius</taxonomicName>
/ insignis / Dist.” [handwritten in black ink, white label, Distants handwritting] //
<taxonomicName id="A9EBA0E5B914FFD184E51B27FD80F874" authority="Dist." authorityName="Distant" authorityYear="1901" box="[439,602,1908,1934]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">“insignis Dist.</taxonomicName>
” [white handwritten label in black ink] // “
<typeStatus id="B15065C4B914FFD181651B27FB36F874" box="[1079,1260,1908,1934]" pageId="3" pageNumber="422" type="lectotype">LECTOTYPUS</typeStatus>
/
<taxonomicName id="A9EBA0E5B914FFD180501B27FA54F874" authorityName="Distant" authorityYear="1901" box="[1282,1422,1908,1934]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="5C9F0774B914FFD180501B27FA54F874" box="[1282,1422,1908,1934]" italics="true" pageId="3" pageNumber="422">VITRUVIUS</emphasis>
</taxonomicName>
/
<emphasis id="5C9F0774B914FFD185C51BCBFED3F848" box="[151,265,1944,1970]" italics="true" pageId="3" pageNumber="422">INSIGNIS</emphasis>
/
<bibRefCitation id="0A7AA697B914FFD1844D1BCBFE6FF848" author="Distant, W. L." box="[287,437,1944,1970]" pageId="3" pageNumber="422" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
/ des.
<collectorName id="C31EBEB0B914FFD184AD1BCAFD3AF848" box="[511,736,1944,1971]" pageId="3" pageNumber="422">M. Roca-Cuschachs</collectorName>
<emphasis id="5C9F0774B914FFD187B51BCAFCC5F848" box="[743,799,1944,1970]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
2024” [printed, red label].
</materialsCitation>
</paragraph>
<caption id="3A948BEEB913FFD685CA1B4DFDFBF810" ID-DOI="http://doi.org/10.5281/zenodo.13210084" ID-Zenodo-Dep="13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="4" pageNumber="423" startId="4.[152,255,1822,1846]" targetBox="[203,1344,199,1769]" targetPageId="4" targetType="figure">
<paragraph id="6E54DB66B913FFD685CA1B4DFDFBF810" blockId="4.[151,1437,1822,2026]" pageId="4" pageNumber="423">
<emphasis id="5C9F0774B913FFD685CA1B4DFEC6F8CC" bold="true" box="[152,284,1822,1846]" pageId="4" pageNumber="423">FIGURE 1.</emphasis>
<taxonomicName id="A9EBA0E5B913FFD684741B4CFDAEF8CC" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[294,628,1822,1846]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B913FFD684741B4CFE00F8CC" box="[294,474,1823,1846]" italics="true" pageId="4" pageNumber="423">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B913FFD684B61B4DFDAEF8CC" author="Distant, W. L." box="[484,628,1822,1846]" pageId="4" pageNumber="423" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs.
</paragraph>
</caption>
<caption id="3A948BEEB912FFD785CA1B4CFB26F811" ID-DOI="http://doi.org/10.5281/zenodo.13210086" ID-Zenodo-Dep="13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="5" pageNumber="424" startId="5.[152,255,1823,1847]" targetBox="[151,1436,185,1799]" targetPageId="5" targetType="figure">
<paragraph id="6E54DB66B912FFD785CA1B4CFB26F811" blockId="5.[151,1436,1823,2028]" pageId="5" pageNumber="424">
<emphasis id="5C9F0774B912FFD785CA1B4CFEC3F8CD" bold="true" box="[152,281,1823,1847]" pageId="5" pageNumber="424">FIGURE 2.</emphasis>
<taxonomicName id="A9EBA0E5B912FFD784721B73FDBEF8CD" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[288,612,1823,1847]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B912FFD784721B73FE0BF8CD" box="[288,465,1824,1847]" italics="true" pageId="5" pageNumber="424">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B912FFD7848A1B4CFDBEF8CD" author="Distant, W. L." box="[472,612,1823,1847]" pageId="5" pageNumber="424" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—
<taxonomicName id="A9EBA0E5B912FFD7879F1B17FCB4F8A1" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[717,878,1859,1883]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B912FFD7879F1B17FCB4F8A1" box="[717,878,1859,1883]" italics="true" pageId="5" pageNumber="424">Vitruvius dobby</emphasis>
</taxonomicName>
<emphasis id="5C9F0774B912FFD786271B10FC1CF8A1" bold="true" box="[885,966,1859,1883]" pageId="5" pageNumber="424">
<taxonomicNameLabel id="47ACBA0FB912FFD786271B10FC1CF8A1" box="[885,966,1859,1883]" pageId="5" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female genital plates; E—
<taxonomicName id="A9EBA0E5B912FFD7818C1B17FA90F8A1" authorityName="Distant" authorityYear="1901" box="[1246,1354,1860,1883]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B912FFD7818C1B17FB37F8A1" box="[1246,1261,1860,1883]" italics="true" pageId="5" pageNumber="424">V</emphasis>
.
<emphasis id="5C9F0774B912FFD781A91B17FA90F8A1" box="[1275,1354,1860,1883]" italics="true" pageId="5" pageNumber="424">insignis</emphasis>
</taxonomicName>
, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs.
</paragraph>
</caption>
<paragraph id="6E54DB66B911FFD485951CC4FED0FE2B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
It is impossible to tell from the original description how many specimens
<bibRefCitation id="0A7AA697B911FFD486A51CC4FB42FF4B" author="Distant, W. L." box="[1015,1176,151,177]" pageId="6" pageNumber="425" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant (1901)</bibRefCitation>
had when he described this species (though provided measurements suggests probably only single one) so any type specimens must be regarded as
<typeStatus id="B15065C4B911FFD484721C8CFE58FF03" box="[288,386,223,249]" pageId="6" pageNumber="425" type="syntype">syntypes</typeStatus>
(cf.
<bibRefCitation id="0A7AA697B911FFD484EA1C8CFDE7FF03" author="ICZN &amp; International Comission on Zoological Nomenclature" box="[440,573,223,249]" pageId="6" pageNumber="425" refId="ref8754" refString="ICZN (International Comission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp. https: // doi. org / 10.5962 / bhl. title. 50608" type="book" year="1999">ICZN 1999</bibRefCitation>
: Recommendation 73F;
<bibRefCitation id="0A7AA697B911FFD486031C8CFC0EFF03" author="Rider, D. A." box="[849,980,223,249]" pageId="6" pageNumber="425" refId="ref9388" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402. [Book pagination: xiii + 550 pp.]" type="book" year="2006">Rider 2006</bibRefCitation>
).
<specimenCount id="78ED10EFB911FFD486B51C8CFBB0FF03" box="[999,1130,223,249]" count="1" pageId="6" pageNumber="425" type="female">One female</specimenCount>
<typeStatus id="B15065C4B911FFD481231C8CFB12FF03" box="[1137,1224,223,249]" pageId="6" pageNumber="425" type="syntype">syntype</typeStatus>
was located in the Museo Civico di Storia Naturale di Genova (
<collectionCode id="08FA43A3B911FFD487D71D50FD00FEE7" box="[645,730,259,285]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="6" pageNumber="425">MCSN</collectionCode>
), and has been examined by means of high quality images made available by Maria Tavano (
<figureCitation id="F6D0C7E3B911FFD484871D74FDEAFEBB" box="[469,560,295,321]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">Fig. 2A</figureCitation>
). That, along with the illustration provided by
<bibRefCitation id="0A7AA697B911FFD481131D74FB3DFEBB" author="Distant, W. L." box="[1089,1255,295,321]" pageId="6" pageNumber="425" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant (1902)</bibRefCitation>
, which is also a female specimen, leaves little doubt about the identity of this species. However, for the purpose of nomenclatural stability, we herein designate this specimen as the
<typeStatus id="B15065C4B911FFD487871D3CFCE5FE73" box="[725,831,367,393]" pageId="6" pageNumber="425" type="lectotype">lectotype</typeStatus>
. The specimen has with five labels preserved with it. In the original description (
<bibRefCitation id="0A7AA697B911FFD4849F1DC0FDB9FE57" author="Distant, W. L." box="[461,611,403,429]" pageId="6" pageNumber="425" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant 1901</bibRefCitation>
), the type locality was given as “
<collectingCountry id="16FC9BF6B911FFD4868F1DC0FBF4FE57" box="[989,1070,403,429]" name="Myanmar" pageId="6" pageNumber="425">Burma</collectingCountry>
,
<collectingRegion id="AC2F1584B911FFD481691DC0FB7FFE57" box="[1083,1189,403,429]" country="Myanmar" name="Yangon" pageId="6" pageNumber="425">Rangoon</collectingRegion>
”, currently
<collectingRegion id="AC2F1584B911FFD4807A1DC0FAA4FE57" box="[1320,1406,403,429]" country="Myanmar" name="Yangon" pageId="6" pageNumber="425">Yangon</collectingRegion>
in
<collectingCountry id="16FC9BF6B911FFD485C51DE4FEDCFE2B" box="[151,262,439,465]" name="Myanmar" pageId="6" pageNumber="425">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="6E54DB66B911FFD485951D89FCA4FD7F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951D89FDE7FE0E" bold="true" box="[199,573,474,500]" pageId="6" pageNumber="425">Additional material examined.</emphasis>
<materialsCitation id="DE83D13BB911FFD4871A1D89FE3AFDE3" collectingDate="1966-03-02" collectionCode="DARC" collectorName="Bishop Museum" country="Laos" location="Sayaboury" municipality="Sayabouri prov." pageId="6" pageNumber="425" specimenCount="1" specimenCount-male="1">
<emphasis id="5C9F0774B911FFD4871A1D89FD44FE0E" bold="true" box="[584,670,474,500]" pageId="6" pageNumber="425">
<collectingCountry id="16FC9BF6B911FFD4871A1D89FD42FE0E" box="[584,664,474,500]" name="Laos" pageId="6" pageNumber="425">LAOS</collectingCountry>
:
</emphasis>
<collectingMunicipality id="8E30411CB911FFD487F81D88FCB8FE0F" box="[682,866,475,501]" pageId="6" pageNumber="425">Sayabouri prov.</collectingMunicipality>
,
<location id="6B348DBDB911FFD486201D88FC31FE0F" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD486201D88FC31FE0F" box="[882,1003,475,501]" country="Laos" municipality="Sayabouri prov." name="Sayaboury" pageId="6" pageNumber="425">Sayaboury</location>
,
<date id="1A55FDA6B911FFD486A81D88FB79FE0F" box="[1018,1187,475,501]" pageId="6" pageNumber="425" value="1966-03-02">
<collectingDate id="0A11044EB911FFD486A81D88FB79FE0F" box="[1018,1187,475,501]" pageId="6" pageNumber="425" value="1966-03-02">2 March 1966</collectingDate>
</date>
, native collector (
<specimenCount id="78ED10EFB911FFD480281D88FA47FE0F" box="[1402,1437,475,501]" count="1" pageId="6" pageNumber="425" type="male">1♂</specimenCount>
<collectorName id="C31EBEB0B911FFD485C51DACFE8CFDE3" box="[151,342,511,537]" pageId="6" pageNumber="425">Bishop Museum</collectorName>
, in
<collectionCode id="08FA43A3B911FFD484D71DACFE02FDE3" box="[389,472,511,537]" pageId="6" pageNumber="425">DARC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="DE83D13BB911FFD484BC1DADFE23FDC7" collectingDate="1963-09-21" collectionCode="DARC" country="Thailand" location="Khao" pageId="6" pageNumber="425" specimenCount="1" specimenCount-female="1">
<emphasis id="5C9F0774B911FFD484BC1DADFD54FDE2" bold="true" box="[494,654,510,536]" pageId="6" pageNumber="425">
<collectingCountry id="16FC9BF6B911FFD484BC1DADFD53FDE2" box="[494,649,510,536]" name="Thailand" pageId="6" pageNumber="425">THAILAND</collectingCountry>
:
</emphasis>
<location id="6B348DBDB911FFD487CB1DACFD0CFDE3" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD487CB1DACFD0CFDE3" box="[665,726,511,537]" country="Thailand" name="Khao" pageId="6" pageNumber="425">Khao</location>
yai,
<date id="1A55FDA6B911FFD486461DACFC22FDE3" box="[788,1016,511,537]" pageId="6" pageNumber="425" value="1963-09-21">
<collectingDate id="0A11044EB911FFD486461DACFC22FDE3" box="[788,1016,511,537]" pageId="6" pageNumber="425" value="1963-09-21">21 September 1963</collectingDate>
</date>
,
<taxonomicName id="A9EBA0E5B911FFD481551DACFB17FDE2" authorityName="Distant" authorityYear="1901" box="[1031,1229,511,536]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B911FFD481551DACFB17FDE2" box="[1031,1229,511,536]" italics="true" pageId="6" pageNumber="425">Vitruvius insignis</emphasis>
</taxonomicName>
[in a big, folded, handwritten label] (
<specimenCount id="78ED10EFB911FFD484241E70FE4DFDC7" box="[374,407,547,573]" count="1" pageId="6" pageNumber="425" type="female">1♀</specimenCount>
<collectionCode id="08FA43A3B911FFD484CC1E70FE2BFDC7" box="[414,497,547,573]" pageId="6" pageNumber="425">DARC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB911FFD487561E70FD0EFD9B" collectingDate="1997-08-14" collectionCode="MRCB" collectorName="Dr. W. Ullrich &amp; Dr. Wolfgang Ullrich" country="Thailand" county="Um Phang" location="Schreb." municipality="Mae Kok river" pageId="6" pageNumber="425" specimenCount="1" specimenCount-male="1">
<collectingCounty id="8735A3EAB911FFD487561E70FDA4FDC7" box="[516,638,547,573]" pageId="6" pageNumber="425">Um Phang</collectingCounty>
,
<collectingMunicipality id="8E30411CB911FFD487DB1E70FCEAFDC7" box="[649,816,547,573]" pageId="6" pageNumber="425">Mae Kok river</collectingMunicipality>
,
<collectorName id="C31EBEB0B911FFD486691E70FC03FDC7" box="[827,985,547,573]" pageId="6" pageNumber="425">Dr. W. Ullrich</collectorName>
leg.,
<taxonomicName id="A9EBA0E5B911FFD4814A1E70FB1AFDC7" box="[1048,1216,547,573]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="5C9F0774B911FFD4814A1E70FB5AFDC7" box="[1048,1152,547,573]" italics="true" pageId="6" pageNumber="425">Bambusa</emphasis>
spec.
</taxonomicName>
<location id="6B348DBDB911FFD481951E70FAC5FDC7" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD481951E70FAC5FDC7" box="[1223,1311,547,573]" country="Thailand" county="Um Phang" municipality="Mae Kok river" name="Schreb." pageId="6" pageNumber="425">Schreb.</location>
,
<date id="1A55FDA6B911FFD480781E70FF08FD9B" pageId="6" pageNumber="425" value="1997-08-14">
<collectingDate id="0A11044EB911FFD480781E70FF08FD9B" pageId="6" pageNumber="425" value="1997-08-14">14 August 1997</collectingDate>
</date>
,
<collectorName id="C31EBEB0B911FFD4858E1E14FE12FD9B" box="[220,456,583,609]" pageId="6" pageNumber="425">Dr. Wolfgang Ullrich</collectorName>
collection (
<specimenCount id="78ED10EFB911FFD4871F1E14FDAAFD9B" box="[589,624,583,609]" count="1" pageId="6" pageNumber="425" type="male">1♂</specimenCount>
<collectionCode id="08FA43A3B911FFD487241E14FD16FD9B" box="[630,716,583,609]" pageId="6" pageNumber="425">MRCB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB911FFD4878C1E14FCA0FD7F" collectionCode="BMNH" collectorName="Siam &amp; Pres. By" country="Thailand" location="Prince" pageId="6" pageNumber="425" specimenCount="1" specimenCount-female="1" stateProvince="Chumphon">
<collectorName id="C31EBEB0B911FFD4878C1E14FCC0FD9B" box="[734,794,583,609]" pageId="6" pageNumber="425">Siam</collectorName>
,
<collectorName id="C31EBEB0B911FFD486771E14FC59FD9B" box="[805,899,583,609]" pageId="6" pageNumber="425">Pres. By</collectorName>
H.R.H.
<location id="6B348DBDB911FFD4868D1E14FBFDFD9B" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD4868D1E14FBFDFD9B" box="[991,1063,583,609]" country="Thailand" name="Prince" pageId="6" pageNumber="425" stateProvince="Chumphon">Prince</location>
of
<collectingRegion id="AC2F1584B911FFD481191E14FB1DFD9B" box="[1099,1223,583,609]" country="Thailand" name="Chumphon" pageId="6" pageNumber="425">Chumphon</collectingRegion>
(unprecise locality and no date of collection); [in a small, printed label] (
<specimenCount id="78ED10EFB911FFD487A31E38FCC8FD7F" box="[753,786,619,645]" count="1" pageId="6" pageNumber="425" type="female">1♀</specimenCount>
<collectionCode id="08FA43A3B911FFD4864B1E38FCA8FD7F" box="[793,882,619,645]" pageId="6" pageNumber="425">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951EDDFDB7FC7B" pageId="6" pageNumber="425" type="diagnosis">
<paragraph id="6E54DB66B911FFD485951EDDFDB7FC7B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951EDDFE98FD52" bold="true" box="[199,322,654,680]" pageId="6" pageNumber="425">Diagnosis.</emphasis>
Very similar to
<taxonomicName id="A9EBA0E5B911FFD484AB1EDCFD83FD53" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[505,601,655,681]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="dobby">
<emphasis id="5C9F0774B911FFD484AB1EDCFD83FD53" box="[505,601,655,681]" italics="true" pageId="6" pageNumber="425">V. dobby</emphasis>
</taxonomicName>
described herein, differing externally in having colouration generally less contrasted, somewhat larger head, the pronotal humeri slightly more protruding laterad, with the apical angles slightly wider, and the anterolateral margins of pronotum are nearly in right angle to the body axis (
<figureCitation id="F6D0C7E3B911FFD481AD1E84FA87FD0B" box="[1279,1373,727,753]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Figs 1A</figureCitation>
,
<figureCitation id="F6D0C7E3B911FFD480381E84FA4AFD0B" box="[1386,1424,727,753]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">2A</figureCitation>
). The genital capsule (
<figureCitation id="F6D0C7E3B911FFD484D21EA8FE0FFCEF" box="[384,469,763,789]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1B</figureCitation>
) is wider apically (two times wider than base over dorsal rim insertion), posterolateral angles extremely broad at apices, with rhomboid outline in ventral view, somewhat spatulate, flat. Posterior rim ventrally broadly U-shaped, deeply concave, bearing a tuft of dense setae projecting ventro-medially. Posterior rim dorsally broadly U-shaped, with flat base.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951FD9FE85F9D7" pageId="6" pageNumber="425" type="description">
<paragraph id="6E54DB66B911FFD485951FD9FCCBFB87" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951FD9FDF6FC5F" bold="true" box="[199,556,906,933]" pageId="6" pageNumber="425">
Redescription.
<emphasis id="5C9F0774B911FFD4842F1FD9FDFFFC5F" bold="true" box="[381,549,906,933]" italics="true" pageId="6" pageNumber="425">Male genitalia</emphasis>
.
</emphasis>
Genital capsule (
<figureCitation id="F6D0C7E3B911FFD487A61FD8FC90FC5F" box="[756,842,907,933]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1B</figureCitation>
) apically two times wider than base over dorsal rim insertion, posterolateral angles very broad at apices, rhomboid in outline, somewhat spatulate, flat. Ventral posterior rim broadly U-shaped, deeply concave, bearing a tuft of dense setae projecting ventro-medially. Posterior rim dorsally broadly U-shaped, with flat base. Proctiger robust, quadrangular in shape. Phallus (
<figureCitation id="F6D0C7E3B911FFD481F71FA4FB20FBEB" box="[1189,1274,1015,1041]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1C</figureCitation>
) with exterior pair of conjunctival processes semi-triangular, broad at base. Internal conjunctival processes with hook-like apical portion shallowly sclerotized. Parameres (
<figureCitation id="F6D0C7E3B911FFD4873D186CFD1CFBA3" box="[623,710,1087,1113]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1D</figureCitation>
) with finger-like process apically, projecting caudad, apical lobe clearly much wider at base, somewhat triangular, robust.
</paragraph>
<paragraph id="6E54DB66B911FFD4859518D5FC58FACB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD4859518D5FE55FB5B" bold="true" box="[199,399,1158,1185]" italics="true" pageId="6" pageNumber="425">Female genitalia</emphasis>
(
<figureCitation id="F6D0C7E3B911FFD484F618D4FDD8FB5B" box="[420,514,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">Fig. 2C</figureCitation>
). Valvifers VIII greatly reduced, partially covered by abdominal segment VII, semicircular, with mesial margins contacting each other. Valvifers IX fused, elongate trapezoid, mesially with projection on each posteromesial angle. Laterotergites IX rectangular, elongate, slightly surpassing apices of laterotergites VIII, quadrate at apices. Sternite X quadrate. Laterotergites VIII somewhat triangular with rounded angles, sinuous at base. Internal genitalia as in generic description.
</paragraph>
<paragraph id="6E54DB66B911FFD485951968FB4BFA3B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951968FEABFAAF" bold="true" box="[199,369,1339,1365]" italics="true" pageId="6" pageNumber="425">Measurements</emphasis>
(
<specimenCount id="78ED10EFB911FFD484D61968FE41FAAF" box="[388,411,1339,1365]" count="1" pageId="6" pageNumber="425" type="male"></specimenCount>
, n = 1 /
<specimenCount id="78ED10EFB911FFD487571968FDC0FAAF" box="[517,538,1339,1365]" count="1" pageId="6" pageNumber="425" type="female"></specimenCount>
, n = 1). Total body length: 18.30/23.86; abdominal width at hemelytral base: 10.72/13.94; maximum abdominal width: 11.70/14.40; head length: 2.04/2.40; width of head (including eyes): 4.10/4.94; anterior head width: 2.86/3.44; interocular distance: 2.42/3.02; pronotal length: 3.74/4.34; maximum pronotal width: 12.38/15.32; scutellar length: 8.02/11.22; scutellar width at base: 7.00/8.74
</paragraph>
<paragraph id="6E54DB66B911FFD485951998FD0EFA1F" blockId="6.[151,1437,151,2013]" box="[199,724,1483,1509]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951998FECDFA1F" bold="true" box="[199,279,1483,1509]" pageId="6" pageNumber="425">Larva.</emphasis>
For description see
<bibRefCitation id="0A7AA697B911FFD484A81998FD0AFA1F" author="Hoffmann, W. E." box="[506,720,1483,1509]" pageId="6" pageNumber="425" refId="ref8411" refString="Hoffmann, W. E. (1932 d) Brief note on Vitruvius insignis Distant. Lingnan Science Journal, 11 (3), 470, pl. 8." type="journal volume" year="1932">Hoffmann (1932d)</bibRefCitation>
.
</paragraph>
<paragraph id="6E54DB66B911FFD4859519BDFE85F9D7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD4859519BDFEE5F9F2" bold="true" box="[199,319,1518,1544]" pageId="6" pageNumber="425">Variation.</emphasis>
There is a considerable size variation between individuals of the two sexes; females are considerably larger than males.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951A64FEF8F98F" pageId="6" pageNumber="425" type="etymology">
<paragraph id="6E54DB66B911FFD485951A64FEF8F98F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951A64FE94F9AB" bold="true" box="[199,334,1591,1617]" pageId="6" pageNumber="425">Etymology.</emphasis>
The species epithet is the Latin adjective
<taxonomicName id="A9EBA0E5B911FFD486741A64FCA7F9AA" authorityName="Distant" authorityYear="1901" box="[806,893,1591,1616]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B911FFD486741A64FCA7F9AA" box="[806,893,1591,1616]" italics="true" pageId="6" pageNumber="425">insignis</emphasis>
</taxonomicName>
, meaning e.g. conspicuous, eminent, famous, or outstanding.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951A2DFD72F8B7" pageId="6" pageNumber="425" type="biology_ecology">
<paragraph id="6E54DB66B911FFD485951A2DFD72F8B7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951A2DFEFDF962" bold="true" box="[199,295,1662,1688]" pageId="6" pageNumber="425">Biology.</emphasis>
<bibRefCitation id="0A7AA697B911FFD484631A2CFE2BF963" author="Hoffmann, W. E." box="[305,497,1663,1689]" pageId="6" pageNumber="425" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann (1931</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487521A2CFD96F963" author="Hoffmann, W. E." box="[512,588,1663,1689]" pageId="6" pageNumber="425" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">1932b</bibRefCitation>
,c,d) was the first to record this species feeding and breeding on terminal branches of bamboo in
<collectingRegion id="AC2F1584B911FFD484CC1AF0FE35F947" box="[414,495,1699,1725]" country="China" name="Hainan" pageId="6" pageNumber="425">Hainan</collectingRegion>
Island, being classified as its minor pest. It was recorded from unidentified bamboo also by
<bibRefCitation id="0A7AA697B911FFD485BE1A94FEA0F91B" author="Cheo, M. - T." box="[236,378,1734,1761]" pageId="6" pageNumber="425" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo (1935)</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484D71A94FDB7F91B" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[389,621,1735,1761]" pageId="6" pageNumber="425" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou (1982)</bibRefCitation>
, Zhang (1985),
<bibRefCitation id="0A7AA697B911FFD4864F1A94FC73F91B" author="Chen, Z. - Y." box="[797,937,1735,1761]" pageId="6" pageNumber="425" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">Chen (1989)</bibRefCitation>
and
<bibRefCitation id="0A7AA697B911FFD4868F1A94FB57F91B" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[989,1165,1735,1761]" pageId="6" pageNumber="425" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B911FFD481561A94FBE1F91B" box="[1028,1083,1735,1761]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
(2011)
</bibRefCitation>
. Additionally, according to the label data from one of the examined specimens, this species has been found on a species of
<taxonomicName id="A9EBA0E5B911FFD481901AB8FAF0F8FF" box="[1218,1322,1771,1797]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="genus">
<emphasis id="5C9F0774B911FFD481901AB8FAF0F8FF" box="[1218,1322,1771,1797]" italics="true" pageId="6" pageNumber="425">Bambusa</emphasis>
</taxonomicName>
(
<taxonomicName id="A9EBA0E5B911FFD480651AB8FA43F8FF" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1335,1433,1771,1797]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
:
<taxonomicName id="A9EBA0E5B911FFD485C51B5CFE9BF8D3" authorityName="Luerssen" authorityYear="1893" box="[151,321,1807,1833]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="subFamily" subFamily="Bambusoideae">Bambusoideae</taxonomicName>
). It was also recorded from
<taxonomicName id="A9EBA0E5B911FFD487281B5CFCEDF8D3" authorityName="Blanco" authorityYear="1837" box="[634,823,1807,1833]" class="Insecta" family="Tachinidae" genus="Melia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="azaderach">
<emphasis id="5C9F0774B911FFD487281B5CFCEDF8D3" box="[634,823,1807,1833]" italics="true" pageId="6" pageNumber="425">Melia azaderach</emphasis>
</taxonomicName>
(
<taxonomicName id="A9EBA0E5B911FFD486171B5CFC66F8D3" box="[837,956,1807,1833]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
) without any details (
<bibRefCitation id="0A7AA697B911FFD481E31B5CFAF8F8D3" author="Hua, L. - Z." box="[1201,1314,1807,1833]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
), however, this record represents probably a sitting record.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD585951B05FC42FF2F" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425" type="distribution">
<paragraph id="6E54DB66B911FFD585951B05FC42FF2F" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,150,429]" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951B05FEBBF88A" bold="true" box="[199,353,1878,1904]" pageId="6" pageNumber="425">Distribution.</emphasis>
<collectingCountry id="16FC9BF6B911FFD484341B04FE69F88B" box="[358,435,1879,1905]" name="China" pageId="6" pageNumber="425">China</collectingCountry>
!:
<collectingRegion id="AC2F1584B911FFD484ED1B04FD98F88B" box="[447,578,1879,1905]" country="China" name="Guangdong" pageId="6" pageNumber="425">Guangdong</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD4871D1B04FD16F88B" author="Cheo, M. - T." box="[591,716,1878,1905]" pageId="6" pageNumber="425" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo 1935</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD487841B04FC4CF88B" author="Hoffmann, W. E." box="[726,918,1878,1905]" pageId="6" pageNumber="425" pagination="1 - 294" refId="ref8443" refString="Hoffmann, W. E. (1935 a) An abridged catalogue of certain Scutelleroidea (Plataspidae, Scutelleridae, and Pentatomidae) of China, Chosen, Indo-China, and Taiwan. Lingnan University Science Bulletin, 7, 1 - 294, 2 maps." type="book chapter" year="1935">Hoffmann 1935a</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD486F21B04FBC1F88B" author="Chen, Z. - Y." box="[928,1051,1879,1905]" pageId="6" pageNumber="425" refId="ref7694" refString="Chen, Z. - Y. (1982) On the Pentatomidae of Dinghushan, Guangdong Province. Acta Scientiarum Naturalium Universitatis Sunyatseni, 1981 (4), 133 - 135. [in Chinese, English summary]" type="book" year="1982">Chen 1982</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD481771B04FBBAF88B" author="Chen, Z. - Y." box="[1061,1120,1879,1905]" pageId="6" pageNumber="425" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">1989</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD481381B04FB00F88B" author="Hua, L. - Z." box="[1130,1242,1879,1905]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481BE1B04FA96F88B" box="[1260,1356,1879,1905]" country="China" name="Guangxi" pageId="6" pageNumber="425">Guangxi</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480091B04FEE5F86F" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." pageId="6" pageNumber="425" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484191B28FDCCF86F" author="Zhang, Y. - Q. &amp; You, Q. - J. &amp; Pu, T. - S. &amp; Lin, R. - Z." box="[331,534,1915,1941]" pageId="6" pageNumber="425" refId="ref10975" refString="Zhang, Y. - Q., You, Q. - J., Pu, T. - S. &amp; Lin, R. - Z. (Eds.) (1994) Insect Catalogue of Guangxi. Guangxi Science and Technology Press, Nanning, 8 + 438 pp. [in Chinese]" type="book" year="1994">
Zhang
<emphasis id="5C9F0774B911FFD484C81B28FE09F86F" box="[410,467,1915,1941]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
1994
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487731B28FD49F86F" author="Hua, L. - Z." box="[545,659,1915,1941]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487CD1B28FCBAF86F" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[671,864,1915,1941]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD487B71B28FCC4F86F" box="[741,798,1915,1941]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4863E1B28FBE1F86F" author="Liu, G. - Q. &amp; Wang, H. - J." box="[876,1083,1915,1941]" pageId="6" pageNumber="425" pagination="173 - 189" refId="ref9251" refString="Liu, G. - Q. &amp; Wang, H. - J. (2004) Hemiptera: Scutelleridae, Tessaratomidae, Dinidoridae and Pentatomidae. In: Yang, X. - K. (Ed.), Insects from Mt. Shiwandashan area of Guangxi. China Forestry Publishing House, Beijing, pp. 173 - 189. [in Chinese, English summary]" type="book chapter" year="2004">Liu &amp; Wang 2004</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD481151B28FB6EF86F" author="Fan, Z. - H." box="[1095,1204,1915,1941]" pageId="6" pageNumber="425" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan 2011</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD4819A1B28FAC3F86F" box="[1224,1305,1915,1941]" country="China" name="Hainan" pageId="6" pageNumber="425">Hainan</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480781B28FF0BF843" author="Hoffmann, W. E." pageId="6" pageNumber="425" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann 1931</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4858D1BCCFEF1F843" author="Hoffmann, W. E." box="[223,299,1951,1977]" pageId="6" pageNumber="425" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">1932b</bibRefCitation>
,c,d, 1935a;
<bibRefCitation id="0A7AA697B911FFD484E11BCCFD72F843" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[435,680,1951,1977]" pageId="6" pageNumber="425" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD487E71BCCFC5EF843" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[693,900,1950,1977]" pageId="6" pageNumber="425" refId="ref10917" refString="Zhang, S. - M., Lin, Y. - J. &amp; Chen, Z. - Q. (1985) Faunal analysis of the pentatomid bugs in Jian-Feng-Ling, Hainan. Acta Agriculturae Universitatis Jiangxiensis, 1985 (2), 9 - 12. [in Chinese, English summary]" type="book" year="1985">
Zhang
<emphasis id="5C9F0774B911FFD486541BCCFC9AF843" box="[774,832,1951,1977]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
1985
</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD486C01BCCFBDFF843" author="Hua, L. - Z." box="[914,1029,1951,1977]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD481401BCCFB02F843" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[1042,1240,1951,1977]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD4810B1BCCFB4EF843" box="[1113,1172,1951,1977]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481BF1BCCFA99F843" box="[1261,1347,1951,1977]" country="China" name="Yunnan" pageId="6" pageNumber="425">Yunnan</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480011BCCFEFDF827" author="Zheng, L. - Y. &amp; Zou, H. - G." pageId="6" pageNumber="425" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou 1982</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484661B90FDD4F827" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[308,526,1987,2013]" pageId="6" pageNumber="425" refId="ref10862" refString="Zhang, S. - M. &amp; Lin, Y. - J. (1990) The newly recorded pentatomid bugs in Xishuangbana [sic!] and their faunal structures. Acta Agriculturae Universitatis Jiangxiensis, 12 (1), 287 - 291. [in Chinese, English summary]" type="book" year="1990">Zhang &amp; Lin 1990</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487491B90FD54F827" author="Hua, L. - Z." box="[539,654,1987,2013]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487C91B90FCBAF827" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[667,864,1987,2013]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD487B01B90FCC6F827" box="[738,796,1987,2013]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4863F1B90FC00F827" author="Fan, Z. - H." box="[877,986,1987,2013]" pageId="6" pageNumber="425" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan 2011</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD486B51B90FB54F827" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[999,1166,1987,2013]" pageId="6" pageNumber="425" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B911FFD481421B90FB90F827" box="[1040,1098,1987,2013]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2011
</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481F11B90FADDF827" box="[1187,1287,1987,2013]" country="China" name="Zhejiang" pageId="6" pageNumber="425">Zhejiang</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480451B90FA42F827" author="Chen, Q. - H." box="[1303,1432,1986,2013]" pageId="6" pageNumber="425" refId="ref7647" refString="Chen, Q. - H. (1985) The catalogue and geographical distribution of bugs (Hemiptera: Pentatomoidea) in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 11 (1), 115 - 127. [in Chinese, English summary]" type="book" year="1985">Chen 1985</bibRefCitation>
,
<bibRefCitation id="0A7AA697B910FFD585C51CC4FEA5FF4B" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[151,383,151,177]" pageId="7" pageNumber="426" refId="ref10813" refString="Zhang, S. - M. &amp; Chen, Q. - H. (1988) Faunal structure of the pentatomid bugs in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 14 (3), 287 - 291. [in Chinese, English summary]" type="book" year="1988">Zhang &amp; Chen 1988</bibRefCitation>
);?
<collectingCountry id="16FC9BF6B910FFD584F41CC4FDCDFF4B" box="[422,535,151,177]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
:
<collectingRegion id="AC2F1584B910FFD587731CC4FD89FF4B" box="[545,595,151,177]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="426">Java</collectingRegion>
(Blöte 1945);
<collectingCountry id="16FC9BF6B910FFD587A71CC4FCEBFF4B" box="[757,817,151,177]" name="Laos" pageId="7" pageNumber="426">Laos</collectingCountry>
! (new record),
<collectingCountry id="16FC9BF6B910FFD5868A1CC4FB91FF4B" box="[984,1099,151,177]" name="Myanmar" pageId="7" pageNumber="426">Myanmar</collectingCountry>
! (
<bibRefCitation id="0A7AA697B910FFD5810E1CC4FB2AFF4B" author="Distant, W. L." box="[1116,1264,151,177]" pageId="7" pageNumber="426" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant 1901</bibRefCitation>
,
<bibRefCitation id="0A7AA697B910FFD581A91CC4FAEDFF4B" author="Distant, W. L." box="[1275,1335,151,177]" pageId="7" pageNumber="426" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">1902</bibRefCitation>
);
<collectingCountry id="16FC9BF6B910FFD580181CC4FA46FF4B" box="[1354,1436,151,177]" name="Taiwan" pageId="7" pageNumber="426">Taiwan</collectingCountry>
(
<bibRefCitation id="0A7AA697B910FFD585F31CE8FEBEFF2F" author="Hoffmann, W. E." box="[161,356,186,213]" pageId="7" pageNumber="426" pagination="57 - 62" refId="ref8491" refString="Hoffmann, W. E. (1935 b) Notes on eight pentatomids not hitherto recorded from Formosa. Transactions of the Natural History Society of Formosa, 25 (136 - 139), 57 - 62." type="journal article" year="1935">Hoffmann 1935b</bibRefCitation>
);
<collectingCountry id="16FC9BF6B910FFD5842A1CE8FE38FF2F" box="[376,482,187,213]" name="Thailand" pageId="7" pageNumber="426">Thailand</collectingCountry>
! (new record);?
<collectingCountry id="16FC9BF6B910FFD587CF1CE8FD26FF2F" box="[669,764,187,213]" name="Vietnam" pageId="7" pageNumber="426">Vietnam</collectingCountry>
(Blöte 1945).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB910FFD585951C8CFD15FE57" pageId="7" pageNumber="426" type="discussion">
<paragraph id="6E54DB66B910FFD585951C8CFD15FE57" blockId="7.[151,1436,150,429]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585951C8CFEE1FF03" bold="true" box="[199,315,223,249]" pageId="7" pageNumber="426">Remarks.</emphasis>
<bibRefCitation id="0A7AA697B910FFD584111C8CFE64FF03" author="Fan, Z. - H." box="[323,446,223,249]" pageId="7" pageNumber="426" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan (2011)</bibRefCitation>
provided a redescription and illustrations of the male and female genitalia of a Chinese specimens identified as
<taxonomicName id="A9EBA0E5B910FFD584FB1D50FDC7FEE6" authorityName="Distant" authorityYear="1901" box="[425,541,259,284]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B910FFD584FB1D50FDC7FEE6" box="[425,541,259,284]" italics="true" pageId="7" pageNumber="426">V. insignis</emphasis>
</taxonomicName>
. Her illustrations of the genitalia correspond with the species we recognize as
<taxonomicName id="A9EBA0E5B910FFD585C51D74FED6FEBA" authorityName="Distant" authorityYear="1901" box="[151,268,295,320]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B910FFD585C51D74FED6FEBA" box="[151,268,295,320]" italics="true" pageId="7" pageNumber="426">V. insignis</emphasis>
</taxonomicName>
; therefore, we can verify her identifications from
<collectingRegion id="AC2F1584B910FFD586001D74FC68FEBB" box="[850,946,295,321]" country="China" name="Guangxi" pageId="7" pageNumber="426">Guangxi</collectingRegion>
and
<collectingRegion id="AC2F1584B910FFD586BF1D74FB99FEBB" box="[1005,1091,295,321]" country="China" name="Yunnan" pageId="7" pageNumber="426">Yunnan</collectingRegion>
(
<collectingCountry id="16FC9BF6B910FFD581071D74FB44FEBB" box="[1109,1182,295,321]" name="China" pageId="7" pageNumber="426">China</collectingCountry>
). We were not able to verify the records from
<collectingCountry id="16FC9BF6B910FFD584CD1D18FE24FE9F" box="[415,510,331,357]" name="Vietnam" pageId="7" pageNumber="426">Vietnam</collectingCountry>
and
<collectingCountry id="16FC9BF6B910FFD587661D18FD7FFE9F" box="[564,677,331,357]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
:
<collectingRegion id="AC2F1584B910FFD587FD1D18FD3BFE9F" box="[687,737,331,357]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="426">Java</collectingRegion>
by Blöte (1945). The genus is present in
<collectingCountry id="16FC9BF6B910FFD581FC1D18FAC1FE9F" box="[1198,1307,331,357]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
throughout pictures uploaded to iNaturalist, but many of them probably belong to an undescribed species, therefore the precise distribution of this species requires further studies.
</paragraph>
</subSubSection>
</treatment>
</document>