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<document id="D15E344A7804A43CA39D7DAF731C03DB" ID-DOI="10.25226/bboc.v142i1.2022.a2" ID-ISSN="2513-9894" ID-Zenodo-Dep="11558820" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718072209933" checkinUser="felipe" docAuthor="Gouraud, Christophe" docDate="2022" docId="0A4987B3C23AFF89C4B081A515F24E24" docLanguage="en" docName="BullBritOrnitholClub.142.1.5-9.pdf" docOrigin="Bulletin of the British Ornithologists’ Club 142 (1)" docSource="https://bioone.org/journals/bulletin-of-the-british-ornithologists-club/volume-142/issue-1/bboc.v142i1.2022.a2/Note-on-the-nomenclature-of-Myrmothera-guttata-Vieillot-1824-Passeriformes/10.25226/bboc.v142i1.2022.a2.full" docStyle="DocumentStyle:EFA7F293C51FD4806C5290561C16F223.2:BullBritOrnitholClub.2017-.journal_article" docStyleId="EFA7F293C51FD4806C5290561C16F223" docStyleName="BullBritOrnitholClub.2017-.journal_article" docStyleVersion="2" docTitle="Myrmothera tessellata Vieillot 1822" docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="F670FFCBC23BFF8DC53A841F14634F7D" masterDocTitle="Note on the nomenclature of Myrmothera guttata Vieillot, 1824 (Passeriformes, Thamnophilidae)" masterLastPageNumber="9" masterPageNumber="5" pageNumber="5" updateTime="1726251417691" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="B4BF14D2ABC42CC4F14074DB118FE97C">Note on the nomenclature of Myrmothera guttata Vieillot, 1824 (Passeriformes, Thamnophilidae)</mods:title>
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<mods:namePart id="015E91A336C8EC3E2F8BCDD256A85B88">Gouraud, Christophe</mods:namePart>
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<subSubSection id="CAFA652EC23AFF8CC4B081A517F14AA5" box="[394,914,1466,1496]" pageId="1" pageNumber="5" type="nomenclature">
<paragraph id="825F36A5C23AFF8CC4B081A517F14AA5" blockId="1.[394,914,1466,1496]" box="[394,914,1466,1496]" pageId="1" pageNumber="5">
<heading id="D91781C9C23AFF8CC4B081A517F14AA5" box="[394,914,1466,1496]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="5" reason="3">
<taxonomicName id="45E04D26C23AFF8CC4B081A517F14AA5" authority="Vieillot, 1822" authorityName="Vieillot" authorityYear="1822" box="[394,914,1466,1496]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="1" pageNumber="5" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C23AFF8CC4B081A516B14AAA" bold="true" box="[394,722,1466,1495]" italics="true" pageId="1" pageNumber="5">Myrmothera tessellata</emphasis>
Vieillot, 1822
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</heading>
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<subSubSection id="CAFA652EC23AFF8CC5FA81F115FA495B" pageId="1" pageNumber="5" type="description">
<paragraph id="825F36A5C23AFF8CC5FA81F115FA495B" blockId="1.[144,1164,1518,1702]" pageId="1" pageNumber="5">
<heading id="D91781C9C23AFF8CC5FA81F115FA495B" fontSize="10" level="2" pageId="1" pageNumber="5" reason="2">
<emphasis id="B094EAB7C23AFF8CC5FA81F11535497B" bold="true" box="[192,342,1518,1542]" italics="true" pageId="1" pageNumber="5">Full reference</emphasis>
.
<emphasis id="B094EAB7C23AFF8CC46781F115ED497B" box="[349,398,1518,1542]" italics="true" pageId="1" pageNumber="5">
<emphasis id="B094EAB7C23AFF8CC46781F115154978" bold="true" box="[349,374,1518,1541]" italics="true" pageId="1" pageNumber="5">—</emphasis>
M
</emphasis>
[
<emphasis id="B094EAB7C23AFF8CC4AD81F11662497B" box="[407,513,1518,1542]" italics="true" pageId="1" pageNumber="5">yrmothera</emphasis>
]
<taxonomicName id="45E04D26C23AFF8CC72A81F116A8497B" authority="Vieillot" authorityName="Vieillot" authorityYear="1822" box="[528,715,1518,1542]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="1" pageNumber="7" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C23AFF8CC72A81F11611497B" box="[528,626,1518,1542]" italics="true" pageId="1" pageNumber="5">Tessellata</emphasis>
Vieillot
</taxonomicName>
<emphasis id="B094EAB7C23AFF8CC7E881F01684497B" box="[722,743,1519,1542]" italics="true" pageId="1" pageNumber="5">in</emphasis>
Bonnaterre &amp;
<bibCitation id="57CA3EC9C23AFF8CC6B781F115F5495B" journal="Vieillot" page="683" pageId="1" pageNumber="5" score="1.97023359760351E-6" volume="1822" year="1822">
Vieillot, 1822: 683–684, livraison 91 (
<date id="F65E1065C23AFF8CC41B821115ED495B" box="[289,398,1550,1574]" pageId="1" pageNumber="5" value="1822-07">July 1822</date>
)
</bibCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CAFA652EC23AFF8EC5FA823016034BD6" lastPageId="3" lastPageNumber="7" pageId="1" pageNumber="5" type="reference_group">
<paragraph id="825F36A5C23AFF8EC5FA823015944C96" blockId="1.[144,1164,1518,1702]" lastBlockId="3.[144,1165,756,1644]" lastPageId="3" lastPageNumber="7" pageId="1" pageNumber="5">
<taxonomicName id="45E04D26C23AFF8CC5FA82301522493B" authorityName="Vieillot" authorityYear="1822" box="[192,321,1582,1606]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="1" pageNumber="5" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C23AFF8CC5FA82301522493B" box="[192,321,1582,1606]" italics="true" pageId="1" pageNumber="5">M. tessellata</emphasis>
</taxonomicName>
was introduced by Vieillot in
<quantity id="45189B40C23AFF8CC7A982311681493B" box="[659,738,1582,1606]" metricMagnitude="1" metricUnit="m" metricValue="4.62788" pageId="1" pageNumber="5" unit="in" value="1822.0">1822 in</quantity>
the
<emphasis id="B094EAB7C23AFF8CC62882311063493B" box="[786,1024,1582,1606]" italics="true" pageId="1" pageNumber="5">Tableau encyclopédique</emphasis>
. The section
<emphasis id="B094EAB7C23AFF8CC5AA82511570491B" box="[144,275,1614,1638]" italics="true" pageId="1" pageNumber="5">Ornithologie</emphasis>
in which this taxon was described comprises eight livraisons, the dates of which were detailed by
<bibRefCitation id="E6714B54C23AFF8CC49D827116EB49FB" author="Dickinson, E. C." box="[423,648,1646,1670]" pageId="1" pageNumber="5" refId="ref1673" refString="Dickinson, E. C. 2011. Bonnaterre, L'Abbe &amp; L. P. Vieillot (1790 - 1823). Tableau encyclopedique et methodique des trois regnes de la nature. 2 eme edition. Ornithologie. P. 78 in Dickinson, E. C., Overstreet, L. K., Dowsett, R. J. &amp; Bruce, M. D. (eds.) Priority! The dating of scientific names in ornithology. Aves Press, Northampton." type="journal volume" year="2011">Dickinson (2011: 78</bibRefCitation>
, and references therein). Livraison 91, which contains pp. 529‒848 and concerns us here, appeared in
<date id="F65E1065C23AFF8CC622829117E749DB" box="[792,900,1678,1702]" pageId="1" pageNumber="5" value="1822-07">July 1822</date>
. The bird described by Vieillot is a good match for a female Rufous-bellied Antwren
<taxonomicName id="45E04D26C238FF8EC66A86EB10EF4C71" ID-CoL="3Q7BY" authority="(Vieillot, 1824)" baseAuthorityName="Vieillot" baseAuthorityYear="1824" box="[848,1164,756,780]" class="Aves" family="Thamnophilidae" genus="Isleria" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC66A86EB17824C76" box="[848,993,756,780]" italics="true" pageId="3" pageNumber="7">Isleria guttata</emphasis>
(Vieillot, 1824)
</taxonomicName>
whose description, of a male, appeared two years later in Vieillot &amp; Oudart’s
<emphasis id="B094EAB7C238FF8EC13A870B17564C36" italics="true" pageId="3" pageNumber="7">La galerie des oiseaux, dédiée à son Altesse Royale Madame, Duchesse du Berri</emphasis>
(1820–26, hereafter
<emphasis id="B094EAB7C238FF8EC120872B14BF4C16" italics="true" pageId="3" pageNumber="7">Galerie des oiseaux</emphasis>
) under the name
<taxonomicName id="45E04D26C238FF8EC493874B161F4C16" authorityName="Vieillot" authorityYear="1824" box="[425,636,852,876]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC493874B161F4C16" box="[425,636,852,876]" italics="true" pageId="3" pageNumber="7">Myrmothera guttata</emphasis>
</taxonomicName>
. More precisely, the text appeared on the first page of livraison 44 (p. 251) and the plate (155; see
<figureCitation id="1ADB2A20C238FF8EC7E5876B177F4CF1" box="[735,796,884,908]" captionStart="Figure 1" captionStartId="2.[144,205,1631,1650]" captionTargetBox="[144,1145,129,1615]" captionTargetId="figure-101@2.[144,1164,126,1617]" captionTargetPageId="2" captionText="Figure 1. Plate 155 of the Galerie des oiseaux: ‘Le Fourmilier Moucheté (Myrmothera guttata)’ (© Ernst Mayr Library, Museum of Comparative Zoology, Harvard University, Cambridge, via Biodiversity Heritage Library: https://www.biodiversitylibrary.org)" figureDoi="http://doi.org/10.5281/zenodo.11558822" httpUri="https://zenodo.org/record/11558822/files/figure.png" pageId="3" pageNumber="7">Fig. 1</figureCitation>
) was the last of livraison 45 (A. Lebossé
<emphasis id="B094EAB7C238FF8EC5CB878B154F4CD1" box="[241,300,916,940]" italics="true" pageId="3" pageNumber="7">in litt</emphasis>
. 2020). Both livraisons were dated 1824 by
<bibRefCitation id="E6714B54C238FF8EC61B878B10EB4CD6" author="Lebosse, A. &amp; Dickinson, E. C." box="[801,1160,915,940]" pageId="3" pageNumber="7" pagination="25 - 58" refId="ref1958" refString="Lebosse, A. &amp; Dickinson, E. C. 2015. Fresh information relevant to the make-up of the livraisons of the &quot; Galerie des Oiseaux &quot; by Vieillot (1748 - 1831) &amp; Oudart (1796 - 1860). Zool. Bibliogr. 3: 25 - 58." type="journal article" year="2015">Lebossé &amp; Dickinson (2015: 51)</bibRefCitation>
. Thus,
<taxonomicName id="45E04D26C238FF8EC5EE87AB16314CB1" authority="Vieillot, 1822" authorityName="Vieillot" authorityYear="1822" box="[212,594,948,972]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C238FF8EC5EE87AB15DA4CB1" box="[212,441,948,972]" italics="true" pageId="3" pageNumber="7">Myrmothera tessellata</emphasis>
Vieillot, 1822
</taxonomicName>
, is a senior synonym of
<taxonomicName id="45E04D26C238FF8EC66487AB14A64C91" authority="Vieillot, 1824" authorityName="Vieillot" authorityYear="1824" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC66487AB104D4CB6" box="[862,1070,948,972]" italics="true" pageId="3" pageNumber="7">Myrmothera guttata</emphasis>
Vieillot, 1824
</taxonomicName>
, the name currently in use.
</paragraph>
<caption id="D69F662DC239FF8FC5AA8240165849DE" ID-DOI="http://doi.org/10.5281/zenodo.11558822" ID-Zenodo-Dep="11558822" httpUri="https://zenodo.org/record/11558822/files/figure.png" pageId="2" pageNumber="6" startId="2.[144,205,1631,1650]" targetBox="[144,1145,129,1615]" targetPageId="2" targetType="figure">
<paragraph id="825F36A5C239FF8FC5AA8240165849DE" blockId="2.[144,1164,1630,1699]" pageId="2" pageNumber="6">
Figure 1. Plate 155 of the
<emphasis id="B094EAB7C239FF8FC4B282401653490C" box="[392,560,1631,1650]" italics="true" pageId="2" pageNumber="6">Galerie des oiseaux</emphasis>
: ‘Le Fourmilier Moucheté (
<taxonomicName id="45E04D26C239FF8FC6018240178F490C" authorityName="Vieillot" authorityYear="1824" box="[827,1004,1631,1650]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="2" pageNumber="6" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C239FF8FC6018240178F490C" box="[827,1004,1631,1650]" italics="true" pageId="2" pageNumber="6">Myrmothera guttata</emphasis>
</taxonomicName>
)’ (© Ernst Mayr Library, Museum of Comparative Zoology, Harvard University, Cambridge, via Biodiversity Heritage Library: https://www.biodiversitylibrary.org)
</paragraph>
</caption>
<caption id="D69F662DC238FF8EC5AA866B168F4DAF" ID-DOI="http://doi.org/10.5281/zenodo.11558824" ID-Zenodo-Dep="11558824" httpUri="https://zenodo.org/record/11558824/files/figure.png" pageId="3" pageNumber="7" startId="3.[144,205,628,647]" targetBox="[144,1165,129,613]" targetPageId="3" targetType="figure">
<paragraph id="825F36A5C238FF8EC5AA866B168F4DAF" blockId="3.[144,1164,628,722]" pageId="3" pageNumber="7">
Figure 2A: Specimen MLC.2011.0.1526, holotype of
<taxonomicName id="45E04D26C238FF8EC74A866B17CC4DF5" authority="Vieillot, 1822" authorityName="Vieillot" authorityYear="1822" box="[624,943,628,648]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C238FF8EC74A866B174C4DFA" box="[624,815,628,647]" italics="true" pageId="3" pageNumber="7">Myrmothera tessellata</emphasis>
Vieillot, 1822
</taxonomicName>
. B: the original pedestal inscriptions read: ‘
<taxonomicName id="45E04D26C238FF8EC47E869215DD4DDD" authorityName="Vieillot" authorityYear="1816" box="[324,446,653,672]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="genus">Myrmothera</taxonomicName>
/ tessellata V[ieillot] / fourmilier / marqueté Vieill[ot] / Encycl[opédie] / Cayenne / M. Bécoeur’. The red label to indicate type status will be added as soon as possible (© Christophe Gouraud / Musée George Sand et de la Vallée Noire, La Châtre)
</paragraph>
</caption>
<paragraph id="825F36A5C238FF8EC5FA87EC16034BD6" blockId="3.[144,1165,756,1644]" pageId="3" pageNumber="7">
<emphasis id="B094EAB7C238FF8EC5FA87EC15A94B76" bold="true" box="[192,458,1011,1035]" italics="true" pageId="3" pageNumber="7">Reversal of precedence</emphasis>
.—Both conditions of Art. 23.9.1 are met and therefore the younger name (
<taxonomicName id="45E04D26C238FF8EC473800B167D4B56" authorityName="Vieillot" authorityYear="1824" box="[329,542,1044,1068]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC473800B167D4B56" box="[329,542,1044,1068]" italics="true" pageId="3" pageNumber="7">Myrmothera guttata</emphasis>
</taxonomicName>
) is valid. The conditions of Art. 23.9.1.2 are met (see list of works in Appendix). To my knowledge, the condition in Art. 23.9.1.1 applies. Thus, the younger name has precedence. Applying Art. 23.9.2, the younger but valid name
<taxonomicName id="45E04D26C238FF8EC5AA806C16694BF1" authority="Vieillot, 1824" authorityName="Vieillot" authorityYear="1824" box="[144,522,1139,1164]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC5AA806C15054BF6" box="[144,358,1139,1163]" italics="true" pageId="3" pageNumber="7">Myrmothera guttata</emphasis>
Vieillot, 1824
</taxonomicName>
, is a
<emphasis id="B094EAB7C238FF8EC76A806B17694BF6" box="[592,778,1140,1163]" italics="true" pageId="3" pageNumber="7">nomen protectum</emphasis>
and the older name
<taxonomicName id="45E04D26C238FF8EC131806C15E44BD1" authority="Vieillot, 1822" authorityName="Vieillot" authorityYear="1822" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C238FF8EC131806C148E4BD6" italics="true" pageId="3" pageNumber="7">Myrmothera tessellata</emphasis>
Vieillot, 1822
</taxonomicName>
, is a
<emphasis id="B094EAB7C238FF8EC485808B16394BD6" box="[447,602,1171,1195]" italics="true" pageId="3" pageNumber="7">nomen oblitum</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="CAFA652EC238FF89C5FA80AB15604FC4" lastPageId="4" lastPageNumber="8" pageId="3" pageNumber="7" type="description">
<paragraph id="825F36A5C238FF89C5FA80AB15604FC4" blockId="3.[144,1165,756,1644]" lastBlockId="4.[144,1165,129,345]" lastPageId="4" lastPageNumber="8" pageId="3" pageNumber="7">
<emphasis id="B094EAB7C238FF8EC5FA80AB15614BB6" bold="true" box="[192,258,1204,1227]" italics="true" pageId="3" pageNumber="7">Types</emphasis>
.—Although the
<emphasis id="B094EAB7C238FF8EC4F280AC16F54BB6" box="[456,662,1203,1227]" italics="true" pageId="3" pageNumber="7">Galerie des oiseaux</emphasis>
aimed to describe and figure at least one species of each genus in the Muséum national d’Histoire naturelle, Paris (
<collectionCode id="E4F1AE60C238FF8EC11680CB10E64B91" box="[1068,1157,1236,1260]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="7">MNHN</collectionCode>
) collections, Vieillot’s protologue for
<taxonomicName id="45E04D26C238FF8EC70980EB16C74A76" authorityName="Vieillot" authorityYear="1824" box="[563,676,1268,1291]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="7" phylum="Chordata" rank="species" species="guttata">
<emphasis id="B094EAB7C238FF8EC70980EB16C74A76" box="[563,676,1268,1291]" italics="true" pageId="3" pageNumber="7">M. guttata</emphasis>
</taxonomicName>
is based on a bird from Cayenne (
<collectingCountry id="FAF77635C238FF8EC10480EC14804A56" name="French Guiana" pageId="3" pageNumber="7">French Guiana</collectingCountry>
) originally owned by the Comte de Riocour, most of whose collection was acquired by Adolphe Boucard in 1889 (
<bibRefCitation id="E6714B54C238FF8EC4E3812C16D34A31" author="Renshaw, G." box="[473,688,1331,1356]" pageId="3" pageNumber="7" pagination="418 - 422" refId="ref2032" refString="Renshaw, G. 1905. The Reunion Starling. The Zoologist (4) 9: 418 - 422." type="journal article" year="1905">Renshaw 1905: 422</bibRefCitation>
). The Boucard collection was subsequently dispersed among several museums, especially
<collectionCode id="E4F1AE60C238FF8EC788814B176A4A16" box="[690,777,1364,1387]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="7">MNHN</collectionCode>
(25,000 specimens), the National Museum of Natural History, Smithsonian Institution, Washington DC (10,000 duplicates) and those in Madrid and Lisbon
<superScript id="75959BEDC238FF8EC735818E16744AE2" attach="left" box="[527,535,1425,1439]" fontSize="6" pageId="3" pageNumber="7">1</superScript>
, which acquired 8,000 birds (
<bibRefCitation id="E6714B54C238FF8EC64F818B14DB4AB6" author="Mearns, B. &amp; Mearns, R." pageId="3" pageNumber="7" refId="ref2012" refString="Mearns, B. &amp; Mearns, R. 1998. The bird collectors. Academic Press, London." type="book" year="1998">Mearns &amp; Mearns 1998: 286</bibRefCitation>
, 302). The
<typeStatus id="5D5B8807C238FF8EC40D81AB15F94AB1" box="[311,410,1460,1484]" pageId="3" pageNumber="7" type="holotype">holotype</typeStatus>
could not be found at
<collectionCode id="E4F1AE60C238FF8EC79381AB17634AB6" box="[681,768,1460,1483]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="7">MNHN</collectionCode>
(P. Boussès
<emphasis id="B094EAB7C238FF8EC6AE81AB17AC4AB1" box="[916,975,1460,1484]" italics="true" pageId="3" pageNumber="7">in litt</emphasis>
. 2021). It is also not in Washington (not listed by
<bibRefCitation id="E6714B54C238FF8EC73E81CB16FC4A96" author="Deignan, H. G." box="[516,671,1491,1516]" pageId="3" pageNumber="7" pagination="1 - 718" refId="ref1641" refString="Deignan, H. G. 1961. Type specimens of birds in the United States National Museum. Bull. US Natl. Mus. 221: 1 - 718." type="journal article" year="1961">Deignan 1961</bibRefCitation>
; confirmed by C. Milensky
<emphasis id="B094EAB7C238FF8EC6E281CB10724A91" box="[984,1041,1492,1516]" italics="true" pageId="3" pageNumber="7">in litt</emphasis>
. 2021) and Philadelphia (N. H. Rice
<emphasis id="B094EAB7C238FF8EC49981EB15B84971" box="[419,475,1524,1548]" italics="true" pageId="3" pageNumber="7">in litt</emphasis>
. 2021), where some specimens ended up. The Natural History Museum, Tring, also possesses some Boucard specimens, but the type was not listed by
<bibRefCitation id="E6714B54C238FF8EC5AA822B15A64931" author="Warren, R. L. M. &amp; Harrison, C. J. O." box="[144,453,1588,1612]" pageId="3" pageNumber="7" refId="ref2162" refString="Warren, R. L. M. &amp; Harrison, C. J. O. 1971. Type-specimens of birds in the British Museum (Natural History), vol. 2. Trustees of the Brit. Mus. (Nat. Hist.), London." type="book" year="1971">Warren &amp; Harrison (1971)</bibRefCitation>
or in the online
<collectionCode id="E4F1AE60C238FF8EC7A1822B17604936" box="[667,771,1588,1611]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="7" type="Museum">NHMUK</collectionCode>
database (https://data.nhm.ac.uk/ search). Finally, it has not been found in the Museo Nacional de Ciencias Naturales, Madrid (J. Barreiro Rodríguez
<emphasis id="B094EAB7C23FFF89C4AE849D15AD4FE4" box="[404,462,129,153]" italics="true" pageId="4" pageNumber="8">in litt</emphasis>
. 2021). In summary the whereabouts of the
<typeStatus id="5D5B8807C23FFF89C6F7849E179C4FE4" box="[973,1023,129,153]" pageId="4" pageNumber="8">type</typeStatus>
specimen is unknown.
</paragraph>
</subSubSection>
<paragraph id="825F36A5C238FF8EC5B5828B17E549D4" blockId="3.[143,902,1684,1706]" box="[143,902,1684,1706]" pageId="3" pageNumber="7">
<superScript id="75959BEDC238FF8EC5B5828B14F649DD" attach="left" box="[143,149,1684,1696]" fontSize="5" pageId="3" pageNumber="7">1</superScript>
The entire collection was destroyed by fire in
<date id="F65E1065C238FF8EC76B828916DC49D4" box="[593,703,1686,1705]" pageId="3" pageNumber="7" value="1978-03">March 1978</date>
(
<bibRefCitation id="E6714B54C238FF8EC7F08289171849D4" author="Roselaar, C. S." box="[714,891,1686,1705]" pageId="3" pageNumber="7" refId="ref2053" refString="Roselaar, C. S. 2003. An inventory of major European bird collections. Pp. 253 - 337 in Collar, N. J., Fisher, C. &amp; Feare, C. J. (eds.) Why museums matter: avian archives in an age of extinction. Bull. Brit. Orn. Cl. Suppl. 123 A." type="journal volume" year="2003">Roselaar 2003: 275</bibRefCitation>
).
</paragraph>
<subSubSection id="CAFA652EC23FFF89C5FA84DE15F24E24" pageId="4" pageNumber="8" type="materials_examined">
<paragraph id="825F36A5C23FFF89C5FA84DE15F24E24" blockId="4.[144,1165,129,345]" pageId="4" pageNumber="8">
<materialsCitation id="32883CF8C23FFF89C5FA84DE15EE4E24" collectionCode="MLC" pageId="4" pageNumber="8" specimenCount="1" typeStatus="holotype">
With respect to
<taxonomicName id="45E04D26C23FFF89C44784DE16054FA4" authorityName="Vieillot" authorityYear="1822" box="[381,614,193,217]" class="Aves" family="Grallariidae" genus="Myrmothera" kingdom="Animalia" order="Passeriformes" pageId="4" pageNumber="8" phylum="Chordata" rank="species" species="tessellata">
<emphasis id="B094EAB7C23FFF89C44784DE16054FA4" box="[381,614,193,217]" italics="true" pageId="4" pageNumber="8">Myrmothera tessellata</emphasis>
</taxonomicName>
, Vieillot’s description clearly indicates that the individual described was in the Baillon Collection and came from Cayenne. The
<typeStatus id="5D5B8807C23FFF89C11384FE10EF4F84" box="[1065,1164,225,249]" pageId="4" pageNumber="8" type="holotype">holotype</typeStatus>
has been found (
<collectionCode id="E4F1AE60C23FFF89C45B851E15F94E64" box="[353,410,257,281]" pageId="4" pageNumber="8">MLC</collectionCode>
.2011.0.1526;
<figureCitation id="1ADB2A20C23FFF89C70F851E16E44E64" box="[565,647,257,281]" captionStart="Figure 2" captionStartId="3.[144,205,628,647]" captionTargetBox="[144,1165,129,613]" captionTargetId="figure-531@3.[144,697,126,615]" captionTargetPageId="3" captionText="Figure 2A: Specimen MLC.2011.0.1526, holotype of Myrmothera tessellata Vieillot, 1822. B: the original pedestal inscriptions read: ‘Myrmothera / tessellata V[ieillot] / fourmilier / marqueté Vieill[ot] / Encycl[opédie] / Cayenne / M. Bécoeur’. The red label to indicate type status will be added as soon as possible (© Christophe Gouraud / Musée George Sand et de la Vallée Noire, La Châtre)" figureDoi="http://doi.org/10.5281/zenodo.11558824" httpUri="https://zenodo.org/record/11558824/files/figure.png" pageId="4" pageNumber="8">Fig. 2a</figureCitation>
) and it matches Vieillot’s protologue. The pedestal inscriptions also state that the bird is from Cayenne (
<figureCitation id="1ADB2A20C23FFF89C66C853E17C74E44" box="[854,932,289,313]" captionStart="Figure 2" captionStartId="3.[144,205,628,647]" captionTargetBox="[144,1165,129,613]" captionTargetId="figure-531@3.[144,697,126,615]" captionTargetPageId="3" captionText="Figure 2A: Specimen MLC.2011.0.1526, holotype of Myrmothera tessellata Vieillot, 1822. B: the original pedestal inscriptions read: ‘Myrmothera / tessellata V[ieillot] / fourmilier / marqueté Vieill[ot] / Encycl[opédie] / Cayenne / M. Bécoeur’. The red label to indicate type status will be added as soon as possible (© Christophe Gouraud / Musée George Sand et de la Vallée Noire, La Châtre)" figureDoi="http://doi.org/10.5281/zenodo.11558824" httpUri="https://zenodo.org/record/11558824/files/figure.png" pageId="4" pageNumber="8">Fig. 2b</figureCitation>
) in accordance with the given type locality
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

434
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<heading id="351C6C0AB910FFD585C51DACFC5AFDE2" bold="true" box="[151,896,510,537]" fontSize="11" level="1" pageId="7" pageNumber="426" reason="1">
<emphasis id="5C9F0774B910FFD585C51DACFC5AFDE2" bold="true" box="[151,896,510,537]" pageId="7" pageNumber="426">
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<emphasis id="5C9F0774B910FFD585C51DACFE8FFDE3" bold="true" box="[151,341,510,537]" italics="true" pageId="7" pageNumber="426">Vitruvius dobby</emphasis>
Roca-Cusachs, Rider &amp; Kment
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,
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<paragraph id="6E54DB66B910FFD585C51E70FE76FDC7" blockId="7.[151,896,510,573]" box="[151,428,546,573]" pageId="7" pageNumber="426">
(
<figureCitation id="F6D0C7E3B910FFD585CD1E70FEC4FDC7" box="[159,286,547,573]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="7" pageNumber="426">Figures 2C</figureCitation>
,
<figureCitation id="F6D0C7E3B910FFD584781E70FEE3FDC7" box="[298,313,547,573]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="7" pageNumber="426">3</figureCitation>
,
<figureCitation id="F6D0C7E3B910FFD584141E70FE8CFDC7" box="[326,342,547,573]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="7" pageNumber="426">4</figureCitation>
,
<figureCitation id="F6D0C7E3B910FFD584301E71FEAAFDC7" box="[354,368,546,573]" captionStart="FIGURE 5" captionStartId="10.[152,255,1553,1577]" captionTargetBox="[151,1435,181,1529]" captionTargetId="figure-37@10.[151,1435,181,1529]" captionTargetPageId="10" captionText="FIGURE 5. Vitruvius dobby sp. nov., structures of abdomen: A—spiracle and trichobothria on ventrite VI (magnification 85×); B—detail of trichobothria on ventrite VI (500×); C–F—male genital capsule (C—caudal view, 55×; D—lateral view, 60×; E—dorsal view, 37×; F—ventral view, 42×). Abbreviations: pa = paramere; pla = posterolateral angle of genital capsule; sp = spiracle; tr = trichobothrium; x = proctiger (segment X). Scale bars: A, D—0.5 mm, B—0.1 mm; C, E–F—1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210094" httpUri="https://zenodo.org/record/13210094/files/figure.png" pageId="7" pageNumber="426">5</figureCitation>
&amp;
<figureCitation id="F6D0C7E3B910FFD584C61E70FE79FDC7" box="[404,419,547,573]" captionStart="FIGURE 6" captionStartId="11.[152,255,949,973]" captionTargetBox="[156,1436,192,921]" captionTargetId="figure-24@11.[151,1436,181,925]" captionTargetPageId="11" captionText="FIGURE 6. Vitruvius dobby sp. nov., variability of colouration.A—female, Cambodia, Kampong Speu, Chambok (11°21′25″N 104°07′09″); B–C—females, Cambodia, Preah Vihear, BeTreed Adventures (13°29′44″N 104°42′36″). Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210096" httpUri="https://zenodo.org/record/13210096/files/figure.png" pageId="7" pageNumber="426">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB910FFD585C51E38FA4FFBEB" pageId="7" pageNumber="426" type="materials_examined">
<paragraph id="6E54DB66B910FFD585C51E38FA4FFBEB" blockId="7.[151,1437,618,1905]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585C51E38FE92FD7E" bold="true" box="[151,328,618,645]" pageId="7" pageNumber="426">Type material.</emphasis>
<materialsCitation id="DE83D13BB910FFD584001E38FC5FFD37" collectingDate="2006-08-07" collectingDateMax="2006-08-14" collectingDateMin="2006-08-07" collectionCode="USNM" collectorName="N. Hongyothi &amp; L. Janteab" country="Thailand" latitude="16.749384" location="Tham Pra Laad Forest" longLatPrecision="1" longitude="101.46388" municipality="Tham Pra Laad Forest Unit" pageId="7" pageNumber="426" specimenCount="1" specimenCount-male="1" stateProvince="Phetchabun" typeStatus="holotype">
<typeStatus id="B15065C4B910FFD584001E38FE65FD7F" box="[338,447,619,645]" pageId="7" pageNumber="426" type="holotype">Holotype</typeStatus>
:
<specimenCount id="78ED10EFB910FFD5849F1E38FE3EFD7F" box="[461,484,619,645]" count="1" pageId="7" pageNumber="426" type="male">♂</specimenCount>
, “
<emphasis id="5C9F0774B910FFD584AD1E39FDB6FD7E" bold="true" box="[511,620,618,644]" pageId="7" pageNumber="426">
<collectingCountry id="16FC9BF6B910FFD584AD1E39FDB6FD7E" box="[511,620,618,644]" name="Thailand" pageId="7" pageNumber="426">Thailand</collectingCountry>
</emphasis>
.
<collectingRegion id="AC2F1584B910FFD5872C1E38FCDBFD7F" box="[638,769,619,645]" country="Thailand" name="Phetchabun" pageId="7" pageNumber="426">Phetchabun</collectingRegion>
Nam Nao NP.,
<collectingMunicipality id="8E30411CB910FFD586EF1E38FADAFD7F" box="[957,1280,619,645]" pageId="7" pageNumber="426">
<location id="6B348DBDB910FFD586EF1E38FB1AFD7F" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD586EF1E38FB1AFD7F" box="[957,1216,619,645]" country="Thailand" latitude="16.749384" longLatPrecision="1" longitude="101.46388" municipality="Tham Pra Laad Forest Unit" name="Tham Pra Laad Forest" pageId="7" pageNumber="426" stateProvince="Phetchabun">Tham Pra Laad Forest</location>
Unit
</collectingMunicipality>
,
<geoCoordinate id="0BDFBDA1B910FFD5805C1E3FFA46FD7D" box="[1294,1436,620,647]" degrees="16" direction="north" minutes="44.963" orientation="latitude" pageId="7" pageNumber="426" precision="1" value="16.749384">16°44.963′N</geoCoordinate>
<geoCoordinate id="0BDFBDA1B910FFD585C51EC3FEE9FD51" box="[151,307,656,683]" degrees="101" direction="east" minutes="27.833" orientation="longitude" pageId="7" pageNumber="426" precision="1" value="101.46388">101°27.833′E</geoCoordinate>
,
<date id="1A55FDA6B910FFD5846C1EDCFE3FFD53" box="[318,485,655,681]" pageId="7" pageNumber="426" value="2006-08-07" valueMax="2006-08-14" valueMin="2006-08-07">
<collectingDate id="0A11044EB910FFD5846C1EDCFE3FFD53" box="[318,485,655,681]" pageId="7" pageNumber="426" value="2006-08-07" valueMax="2006-08-14" valueMin="2006-08-07">
7–14.viii.
<quantity id="A9137683B910FFD584F51EDCFE3FFD53" box="[423,485,655,681]" metricMagnitude="6" metricUnit="m" metricValue="2.006711" pageId="7" pageNumber="426" unit="m" value="2006711.0">2006</quantity>
</collectingDate>
</date>
, 711 m,
<collectorName id="C31EBEB0B910FFD587101EDCFD06FD53" box="[578,732,655,681]" pageId="7" pageNumber="426">N. Hongyothi</collectorName>
&amp;
<collectorName id="C31EBEB0B910FFD587AD1EDCFCABFD53" box="[767,881,655,681]" pageId="7" pageNumber="426">L. Janteab</collectorName>
leg.,
<collectingMethod id="B7AAA371B910FFD586FD1EDCFBE0FD53" box="[943,1082,655,681]" pageId="7" pageNumber="426">Malaise trap</collectingMethod>
T424, DAR_0545” //
<taxonomicName id="A9EBA0E5B910FFD580601EDCFD90FD37" authority="Roca-Cusachs, Rider &amp; Kment" authorityName="Roca-Cusachs, Rider &amp; Kment" authorityYear="2024" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="dobby">
<emphasis id="5C9F0774B910FFD580601EDCFD90FD37" bold="true" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD580601EDCFF01FD37" bold="true" italics="true" pageId="7" pageNumber="426">Vitruvius dobby</emphasis>
Roca-Cusachs, Rider &amp; Kment
</emphasis>
</taxonomicName>
[printed, red label] (
<collectionCode id="08FA43A3B910FFD586751EE0FCA7FD37" box="[807,893,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="426" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="DE83D13BB910FFD586DF1EE0FD15FD0B" collectingDate="2006-08-07" collectingDateMax="2006-08-14" collectingDateMin="2006-08-07" collectionCode="MRCB" collectorName="Same" country="Thailand" latitude="16.749384" location="Tham Pra Laad Forest" longLatPrecision="1" longitude="101.46388" municipality="Tham Pra Laad Forest Unit" pageId="7" pageNumber="426" specimenCode="MRCUCE0073" specimenCount="1" specimenCount-male="1" stateProvince="Phetchabun" typeStatus="paratype">
<typeStatus id="B15065C4B910FFD586DF1EE0FC24FD37" box="[909,1022,691,717]" pageId="7" pageNumber="426" type="paratype">Paratypes</typeStatus>
:
<emphasis id="5C9F0774B910FFD581571EE1FB7CFD36" bold="true" box="[1029,1190,690,716]" pageId="7" pageNumber="426">
<collectingCountry id="16FC9BF6B910FFD581571EE1FB7AFD36" box="[1029,1184,690,716]" name="Thailand" pageId="7" pageNumber="426">THAILAND</collectingCountry>
:
</emphasis>
<collectorName id="C31EBEB0B910FFD581F81EE0FB32FD37" box="[1194,1256,691,717]" pageId="7" pageNumber="426">Same</collectorName>
data as holotype except, DAR_0544 (
<specimenCount id="78ED10EFB910FFD584D01E84FE7CFD0B" box="[386,422,727,753]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
,
<collectionCode id="08FA43A3B910FFD584E11E84FDDDFD0B" box="[435,519,727,753]" pageId="7" pageNumber="426">MRCB</collectionCode>
;
<specimenCode id="3E4D731DB910FFD587411E84FD1DFD0B" box="[531,711,727,753]" collectionCode="MRCB" pageId="7" pageNumber="426">MRCUCE0073</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB910FFD587891E84FF28FCC3" collectionCode="DARC" collectorName="S. Chachumnan &amp; S. Singtong" country="Thailand" elevation="2007168" latitude="16.659817" location="Khao Kho" longLatPrecision="1" longitude="101.13641" pageId="7" pageNumber="426" specimenCount="1" specimenCount-male="1" stateProvince="Phetchabun" typeStatus="paratype">
<collectingRegion id="AC2F1584B910FFD587891E84FCBBFD0B" box="[731,865,727,753]" country="Thailand" name="Phetchabun" pageId="7" pageNumber="426">Phetchabun</collectingRegion>
,
<location id="6B348DBDB910FFD5863E1E84FC3AFD0B" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD5863E1E84FC3AFD0B" box="[876,992,727,753]" country="Thailand" latitude="16.659817" longLatPrecision="1" longitude="101.13641" name="Khao Kho" pageId="7" pageNumber="426" stateProvince="Phetchabun">Khao Kho</location>
NP.,
<geoCoordinate id="0BDFBDA1B910FFD5814D1E8BFB75FD09" box="[1055,1199,728,755]" degrees="16" direction="north" minutes="39.589" orientation="latitude" pageId="7" pageNumber="426" precision="1" value="16.659817">16°39.589′N</geoCoordinate>
,
<geoCoordinate id="0BDFBDA1B910FFD581EE1E8BFA82FD09" box="[1212,1368,728,755]" degrees="101" direction="east" minutes="08.185" orientation="longitude" pageId="7" pageNumber="426" precision="1" value="101.13641">101°08.185′E</geoCoordinate>
,
<date id="1A55FDA6B910FFD580361E84FEEEFCEF" pageId="7" pageNumber="426" value="2007-01-08" valueMax="2007-01-18" valueMin="2007-01-08">
8–18 January
<quantity id="A9137683B910FFD585AB1EA8FEEEFCEF" box="[249,308,763,789]" metricMagnitude="6" metricUnit="m" metricValue="2.007168" pageId="7" pageNumber="426" unit="m" value="2007168.0">
<elevation id="E5C63C55B910FFD585AB1EA8FEEEFCEF" box="[249,308,763,789]" metricMagnitude="6" metricUnit="m" metricValue="2.007168" pageId="7" pageNumber="426" unit="m" value="2007168.0">2007</elevation>
</quantity>
</date>
, 168 m, mixed deciduous,
<collectingMethod id="B7AAA371B910FFD587221EA8FD0AFCEF" box="[624,720,763,789]" pageId="7" pageNumber="426">Pan trap</collectingMethod>
T1386,
<collectorName id="C31EBEB0B910FFD586661EA8FC33FCEF" box="[820,1001,763,789]" pageId="7" pageNumber="426">S. Chachumnan</collectorName>
&amp;
<collectorName id="C31EBEB0B910FFD581411EA8FB43FCEF" box="[1043,1177,763,789]" pageId="7" pageNumber="426">S. Singtong</collectorName>
leg., DAR_0532 (
<specimenCount id="78ED10EFB910FFD580211EA8FA4DFCEF" box="[1395,1431,763,789]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
,
<collectionCode id="08FA43A3B910FFD585C51F4CFF30FCC3" box="[151,234,799,825]" pageId="7" pageNumber="426">DARC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB910FFD585AF1F4CFD07FCA7" collectionCode="DARC" collectorName="L. Sapsiri" country="Thailand" latitude="15.620167" location="Entrance of Huay Pok Substation" longLatPrecision="1" longitude="105.6153" municipality="Pha Taem" pageId="7" pageNumber="426" specimenCount="1" specimenCount-male="1" stateProvince="Ubon Ratchathani" typeStatus="paratype">
<collectingRegion id="AC2F1584B910FFD585AF1F4CFE11FCC3" box="[253,459,799,825]" country="Thailand" name="Ubon Ratchathani" pageId="7" pageNumber="426">Ubon Ratchathani</collectingRegion>
,
<collectingMunicipality id="8E30411CB910FFD584871F4CFD99FCC3" box="[469,579,799,825]" pageId="7" pageNumber="426">Pha Taem</collectingMunicipality>
NP.,
<geoCoordinate id="0BDFBDA1B910FFD5872D1F73FCCBFCC1" box="[639,785,800,827]" degrees="15" direction="north" minutes="37.210" orientation="latitude" pageId="7" pageNumber="426" precision="1" value="15.620167">15°37.210′N</geoCoordinate>
,
<geoCoordinate id="0BDFBDA1B910FFD586491F73FC6DFCC1" box="[795,951,800,827]" degrees="105" direction="east" minutes="36.918" orientation="longitude" pageId="7" pageNumber="426" precision="1" value="105.6153">105°36.918′E</geoCoordinate>
,
<location id="6B348DBDB910FFD586901F4CFAE3FCC3" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD586901F4CFAE3FCC3" box="[962,1337,799,825]" country="Thailand" latitude="15.620167" longLatPrecision="1" longitude="105.6153" municipality="Pha Taem" name="Entrance of Huay Pok Substation" pageId="7" pageNumber="426" stateProvince="Ubon Ratchathani">Entrance of Huay Pok Substation</location>
,
<collectingMethod id="B7AAA371B910FFD580161F4CFF19FCA7" pageId="7" pageNumber="426">Malaise trap</collectingMethod>
T2172,
<collectorName id="C31EBEB0B910FFD5844F1F10FE52FCA7" box="[285,392,835,861]" pageId="7" pageNumber="426">L. Sapsiri</collectorName>
leg., DAR_0529 (
<specimenCount id="78ED10EFB910FFD587011F10FDADFCA7" box="[595,631,835,861]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
,
<collectionCode id="08FA43A3B910FFD587D01F10FD0FFCA7" box="[642,725,835,861]" pageId="7" pageNumber="426">DARC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB910FFD587B41F10FEBBFC7B" collectingDate="2018-10-28" collectionCode="MRCB" collectorName="Chiangmai &amp; V. Ustinov" country="Thailand" location="Chiangdao" pageId="7" pageNumber="426" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectorName id="C31EBEB0B910FFD587B41F10FCBEFCA7" box="[742,868,835,861]" pageId="7" pageNumber="426">Chiangmai</collectorName>
,
<location id="6B348DBDB910FFD5863F1F10FC3CFCA7" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD5863F1F10FC3CFCA7" box="[877,998,835,861]" country="Thailand" name="Chiangdao" pageId="7" pageNumber="426">Chiangdao</location>
hill resort,
<date id="1A55FDA6B910FFD581301F10FAC4FCA7" box="[1122,1310,835,861]" pageId="7" pageNumber="426" value="2018-10-28">
<collectingDate id="0A11044EB910FFD581301F10FAC4FCA7" box="[1122,1310,835,861]" pageId="7" pageNumber="426" value="2018-10-28">28 October 2018</collectingDate>
</date>
,
<collectorName id="C31EBEB0B910FFD580751F10FA47FCA7" box="[1319,1437,835,861]" pageId="7" pageNumber="426">V. Ustinov</collectorName>
leg. (
<specimenCount id="78ED10EFB910FFD585861F34FF2FFC7B" box="[212,245,871,897]" count="1" pageId="7" pageNumber="426" type="female">1♀</specimenCount>
,
<collectionCode id="08FA43A3B910FFD584511F34FE83FC7B" box="[259,345,871,897]" pageId="7" pageNumber="426">MRCB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="DE83D13BB910FFD5843E1F35FCB7FC5F" collectingDate="2015-05-04" collectingDateMax="2015-05-08" collectingDateMin="2015-05-04" collectionCode="ISNB, NMPC" collectorName="Day &amp; J. Constant &amp; V. Sougnez" country="Cambodia" latitude="11.356945" location="Chambok" pageId="7" pageNumber="426" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kampong Speu" typeStatus="paratype">
<emphasis id="5C9F0774B910FFD5843E1F35FDCEFC7A" bold="true" box="[364,532,870,896]" pageId="7" pageNumber="426">
<collectingCountry id="16FC9BF6B910FFD5843E1F35FDD5FC7A" box="[364,527,870,896]" name="Cambodia" pageId="7" pageNumber="426">CAMBODIA</collectingCountry>
:
</emphasis>
<collectingRegion id="AC2F1584B910FFD5874E1F34FD17FC7B" box="[540,717,871,897]" country="Cambodia" name="Kampong Speu" pageId="7" pageNumber="426">Kampong Speu</collectingRegion>
,
<location id="6B348DBDB910FFD5878A1F34FC9FFC7B" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD5878A1F34FC9FFC7B" box="[728,837,871,897]" country="Cambodia" latitude="11.356945" name="Chambok" pageId="7" pageNumber="426" stateProvince="Kampong Speu">Chambok</location>
11°21′25″N, 104°7′9″,
<collectorName id="C31EBEB0B910FFD581001F34FB5BFC7B" box="[1106,1153,871,897]" pageId="7" pageNumber="426">Day</collectorName>
coll.,
<date id="1A55FDA6B910FFD5819A1F35FA97FC7B" box="[1224,1357,870,897]" pageId="7" pageNumber="426" value="2015-05-04" valueMax="2015-05-08" valueMin="2015-05-04">
<collectingDate id="0A11044EB910FFD5819A1F35FA97FC7B" box="[1224,1357,870,897]" pageId="7" pageNumber="426" value="2015-05-04" valueMax="2015-05-08" valueMin="2015-05-04">4.–8.v.2015</collectingDate>
</date>
, leg.
<collectorName id="C31EBEB0B910FFD580D81F34FF21FC5F" pageId="7" pageNumber="426">J. Constant</collectorName>
&amp;
<collectorName id="C31EBEB0B910FFD5844E1FD8FE47FC5F" box="[284,413,907,933]" pageId="7" pageNumber="426">V. Sougnez</collectorName>
, I.G. 33.022 (
<specimenCount id="78ED10EFB910FFD587651FD8FD80FC5F" box="[567,602,907,933]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
<specimenCount id="78ED10EFB910FFD587321FD8FD59FC5F" box="[608,643,907,933]" count="1" pageId="7" pageNumber="426" type="female">1♀</specimenCount>
,
<collectionCode id="08FA43A3B910FFD587C21FD8FD08FC5F" box="[656,722,907,933]" pageId="7" pageNumber="426">ISNB</collectionCode>
,
<specimenCount id="78ED10EFB910FFD5878F1FD8FCDBFC5F" box="[733,769,907,933]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
,
<collectionCode id="08FA43A3B910FFD5865F1FD8FCB8FC5F" box="[781,866,907,933]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="426">NMPC</collectionCode>
);
</materialsCitation>
<materialsCitation id="DE83D13BB910FFD586261FD8FC4AFC33" collectingDate="2015-05-04" collectingDateMax="2015-05-08" collectingDateMin="2015-05-04" collectingMethod="pitfall trap" collectionCode="ISNB" collectorName="J. Constant &amp; V. Sougnez" country="Cambodia" latitude="11.356945" location="Kampong Speu" pageId="7" pageNumber="426" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Kampong Speu" typeStatus="paratype">
<collectingRegion id="AC2F1584B910FFD586261FD8FBF9FC5F" box="[884,1059,907,933]" country="Cambodia" name="Kampong Speu" pageId="7" pageNumber="426">Kampong Speu</collectingRegion>
, Chambok 11°21′25″N, 104°7′9″,
<collectingMethod id="B7AAA371B910FFD585C51FFCFECAFC33" box="[151,272,943,969]" pageId="7" pageNumber="426">pitfall trap</collectingMethod>
, leg.
<collectorName id="C31EBEB0B910FFD5841C1FFCFE16FC33" box="[334,460,943,969]" pageId="7" pageNumber="426">J. Constant</collectorName>
&amp;
<collectorName id="C31EBEB0B910FFD584A31FFCFDA9FC33" box="[497,627,943,969]" pageId="7" pageNumber="426">V. Sougnez</collectorName>
, I.G. 33.022 (
<specimenCount id="78ED10EFB910FFD586411FFCFCEDFC33" box="[787,823,943,969]" count="1" pageId="7" pageNumber="426" type="male">1♂</specimenCount>
,
<collectionCode id="08FA43A3B910FFD586161FFCFC52FC33" box="[836,904,943,969]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="426">ISNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB910FFD586CF1FFCFCF7FC17" collectingDate="2005-05-11" collectingDateMax="2005-05-17" collectingDateMin="2005-05-11" collectionCode="ISNB" collectorName="K. Smets &amp; I. Var" country="Cambodia" location="Wildlife Sanct. Forest" municipality="Phnom Samkos" pageId="7" pageNumber="426" specimenCount="1" specimenCount-female="1" stateProvince="Pursat Prov." typeStatus="paratype">
<collectingRegion id="AC2F1584B910FFD586CF1FFCFBF2FC33" box="[925,1064,943,969]" country="Cambodia" name="Pursat" pageId="7" pageNumber="426">Pursat Prov.</collectingRegion>
,
<collectingMunicipality id="8E30411CB910FFD581661FFCFB3DFC33" box="[1076,1255,943,969]" pageId="7" pageNumber="426">Phnom Samkos</collectingMunicipality>
,
<location id="6B348DBDB910FFD581A11FFCFF3AFC17" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD581A11FFCFF3AFC17" country="Cambodia" municipality="Phnom Samkos" name="Wildlife Sanct. Forest" pageId="7" pageNumber="426" stateProvince="Pursat Prov.">Wildlife Sanct. Forest</location>
,
<date id="1A55FDA6B910FFD585BF1F81FE54FC17" box="[237,398,978,1005]" pageId="7" pageNumber="426" value="2005-05-11" valueMax="2005-05-17" valueMin="2005-05-11">
<collectingDate id="0A11044EB910FFD585BF1F81FE54FC17" box="[237,398,978,1005]" pageId="7" pageNumber="426" value="2005-05-11" valueMax="2005-05-17" valueMin="2005-05-11">11.–17.v.2005</collectingDate>
</date>
, leg.
<collectorName id="C31EBEB0B910FFD5849F1F80FDEFFC17" box="[461,565,979,1005]" pageId="7" pageNumber="426">K. Smets</collectorName>
&amp;
<collectorName id="C31EBEB0B910FFD5870F1F80FD46FC17" box="[605,668,979,1005]" pageId="7" pageNumber="426">I. Var</collectorName>
(
<specimenCount id="78ED10EFB910FFD587E31F80FD08FC17" box="[689,722,979,1005]" count="1" pageId="7" pageNumber="426" type="female">1♀</specimenCount>
,
<collectionCode id="08FA43A3B910FFD587B31F80FCFFFC17" box="[737,805,979,1005]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="426">ISNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB910FFD586691F80FA48FBEB" collectingDate="2017-10-16" collectingDateMax="2017-10-21" collectingDateMin="2017-10-16" collectionCode="ISNB, NMPC" collectorName="J. Constant &amp; Vermeersch" country="Cambodia" latitude="13.495556" location="BeTreed Adventures" longLatPrecision="21" longitude="104.71" municipality="Praeh Vihear Prov." pageId="7" pageNumber="426" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="8E30411CB910FFD586691F80FBC9FC17" box="[827,1043,979,1005]" pageId="7" pageNumber="426">Praeh Vihear Prov.</collectingMunicipality>
,
<location id="6B348DBDB910FFD581721F80FAD0FC17" LSID="urn:lsid:plazi:treatment:E6426A70B910FFD985C51DACFDBCFB33:6B348DBDB910FFD581721F80FAD0FC17" box="[1056,1290,979,1005]" country="Cambodia" latitude="13.495556" longLatPrecision="21" longitude="104.71" municipality="Praeh Vihear Prov." name="BeTreed Adventures" pageId="7" pageNumber="426">BeTreed Adventures</location>
,
<geoCoordinate id="0BDFBDA1B910FFD580451F87FA47FC15" box="[1303,1437,980,1007]" degrees="13" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="426" precision="15" seconds="44" value="13.495556">13°29′44″N</geoCoordinate>
<geoCoordinate id="0BDFBDA1B910FFD585C51FABFEF0FBE9" box="[151,298,1016,1043]" degrees="104" direction="east" minutes="42" orientation="longitude" pageId="7" pageNumber="426" precision="15" seconds="36" value="104.71">104°42′36″E</geoCoordinate>
, GTI Project,
<date id="1A55FDA6B910FFD5849A1FA4FDB6FBEB" box="[456,620,1015,1041]" pageId="7" pageNumber="426" value="2017-10-16" valueMax="2017-10-21" valueMin="2017-10-16">
<collectingDate id="0A11044EB910FFD5849A1FA4FDB6FBEB" box="[456,620,1015,1041]" pageId="7" pageNumber="426" value="2017-10-16" valueMax="2017-10-21" valueMin="2017-10-16">16.–21.x.2017</collectingDate>
</date>
, leg.
<collectorName id="C31EBEB0B910FFD587F51FA4FCFEFBEB" box="[679,804,1015,1041]" pageId="7" pageNumber="426">J. Constant</collectorName>
&amp; X.
<collectorName id="C31EBEB0B910FFD5863B1FA4FC2AFBEB" box="[873,1008,1015,1041]" pageId="7" pageNumber="426">Vermeersch</collectorName>
, I.G. 33.551 (
<specimenCount id="78ED10EFB910FFD581DC1FA4FB75FBEB" box="[1166,1199,1015,1041]" count="1" pageId="7" pageNumber="426" type="female">1♀</specimenCount>
,
<collectionCode id="08FA43A3B910FFD581EE1FA4FB25FBEB" box="[1212,1279,1015,1041]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="7" pageNumber="426">ISNB</collectionCode>
;
<specimenCount id="78ED10EFB910FFD580591FA4FAF4FBEB" box="[1291,1326,1015,1041]" count="1" pageId="7" pageNumber="426" type="female">1♀</specimenCount>
<collectionCode id="08FA43A3B910FFD580671FA4FA50FBEB" box="[1333,1418,1015,1041]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="426">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB910FFD585951849FC5FFB13" pageId="7" pageNumber="426" type="diagnosis">
<paragraph id="6E54DB66B910FFD585951849FC5FFB13" blockId="7.[151,1437,618,1905]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585951849FE98FBCE" bold="true" box="[199,322,1050,1076]" pageId="7" pageNumber="426">Diagnosis.</emphasis>
Very similar to
<taxonomicName id="A9EBA0E5B910FFD587521848FDACFBCE" authorityName="Distant" authorityYear="1901" box="[512,630,1051,1076]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B910FFD587521848FDACFBCE" box="[512,630,1051,1076]" italics="true" pageId="7" pageNumber="426">V. insignis</emphasis>
</taxonomicName>
, somewhat smaller head, pronotal humeri less protruding, with apical angles slightly more rounded, and by the anterolateral borders of the pronotum more at an obtuse angle (declivent) to longitudinal body axis (
<figureCitation id="F6D0C7E3B910FFD584941830FDF8FB87" box="[454,546,1123,1149]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="7" pageNumber="426">Fig. 2A</figureCitation>
). The genital capsule (
<figureCitation id="F6D0C7E3B910FFD586741830FCA4FB87" box="[806,894,1123,1149]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="7" pageNumber="426">Fig. 2B</figureCitation>
) is narrower (clearly less than two times wider than base over dorsal rim insertion), posterolateral angles acute, projecting caudally, bearing conspicuous tuft of long setae apically, ventral posterior rim laterally sinuous, medially U-shaped, shallow, without tuft of dense setae projecting ventro-medially. Dorsal posterior rim broadly V-shaped.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB910FFD8859518A1FB32F86B" lastPageId="10" lastPageNumber="429" pageId="7" pageNumber="426" type="description">
<paragraph id="6E54DB66B910FFD5859518A1FF36F9F3" blockId="7.[151,1437,618,1905]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD5859518A1FDCAFAF7" bold="true" box="[199,528,1266,1293]" pageId="7" pageNumber="426">
Description.
<emphasis id="5C9F0774B910FFD5843318A1FDCAFAF7" bold="true" box="[353,528,1266,1293]" italics="true" pageId="7" pageNumber="426">Male genitalia.</emphasis>
</emphasis>
Genital capsule (
<figureCitation id="F6D0C7E3B910FFD5878518A0FCF4FAF7" box="[727,814,1267,1293]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="7" pageNumber="426">Fig. 3B</figureCitation>
) with apex clearly less than two times wider than base over dorsal rim insertion, posterolateral angles acute, projecting caudally, bearing conspicuous tuft of long setae apically; ventral posterior rim laterally sinuous, medially U-shaped, shallow, without tuft of dense setae projecting ventro-medially; dorsal posterior rim broadly V-shaped. Proctiger elongate, with apical portion narrower than base. Phallus (
<figureCitation id="F6D0C7E3B910FFD585A819D0FE8AFA67" box="[250,336,1411,1437]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="7" pageNumber="426">Fig. 3C</figureCitation>
) with exterior pair of conjunctival processes elongate, finger-like, of similar diameter throughout entire length. Internal conjunctival processes with hook-like apical portion strongly sclerotized. Parameres (
<figureCitation id="F6D0C7E3B910FFD5803D19F4FF60FA1F" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="7" pageNumber="426">Fig. 3D</figureCitation>
) with finger-like process located laterad, slightly curved caudally, apical lobe clearly much broader at apex than at base.
</paragraph>
<paragraph id="6E54DB66B910FFD585951A41FD3BF947" blockId="7.[151,1437,618,1905]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585951A41FE55F9D7" bold="true" box="[199,399,1554,1581]" italics="true" pageId="7" pageNumber="426">Female genitalia</emphasis>
(
<figureCitation id="F6D0C7E3B910FFD584F61A40FDD9F9D7" box="[420,515,1555,1581]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="7" pageNumber="426">Fig. 2D</figureCitation>
). Valvifers VIII greatly reduced, partially covered by abdominal segment VII, semicircular, with mesial margins contiguous. Valvifers IX fused, elongate trapezoid, mesially with projection on each posterolateral angle. Laterotergites IX rectangular, elongate, not surpassing apices of laterotergites VIII, rounded at apices. Sternite X quadrate. Laterotergites VIII somewhat triangular with rounded angles, flat at base. Internal genitalia as in generic description (
<figureCitation id="F6D0C7E3B910FFD5872F1AF0FD0EF947" box="[637,724,1699,1725]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="7" pageNumber="426">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="6E54DB66B910FFD585951A94FBE8F88B" blockId="7.[151,1437,618,1905]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585951A94FEABF91B" bold="true" box="[199,369,1735,1761]" italics="true" pageId="7" pageNumber="426">Measurements</emphasis>
(
<specimenCount id="78ED10EFB910FFD5842D1A94FE4CF91B" box="[383,406,1735,1761]" count="1" pageId="7" pageNumber="426" type="male">♂</specimenCount>
, n = 7 /
<specimenCount id="78ED10EFB910FFD584B51A94FE26F91B" box="[487,508,1735,1761]" count="1" pageId="7" pageNumber="426" type="female">♀</specimenCount>
, n = 4). Total body length: 13.05–18.36/14.02–17.40; abdominal width at hemelytral base: 7.16–9.78/7.45–9.90; maximum abdominal width: 6.65–11.2/7.45–11.60; head length: 1.72–2.32/1.81–1.9; width of head (including eyes): 2.94–3.94/2.94–3.60; anterior head width: 1.96–2.82/2.11–2.50; interocular distance: 1.76–2.46/1.81–2.20; pronotal length: 3.20–3.38/3.33–3.90; maximum pronotal width: 8.04–11.12/8.14–10.40; scutellar length: 5.39–7.8/5.69–7.20; scutellar width at base: 4.31–6.14/4.41–5.29.
</paragraph>
<caption id="3A948BEEB91FFFDA85CA1B4DFEEAF810" ID-DOI="http://doi.org/10.5281/zenodo.13210088" ID-Zenodo-Dep="13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="8" pageNumber="427" startId="8.[152,255,1822,1846]" targetBox="[213,1350,191,1794]" targetPageId="8" targetType="figure">
<paragraph id="6E54DB66B91FFFDA85CA1B4DFEEAF810" blockId="8.[151,1436,1822,2026]" pageId="8" pageNumber="427">
<emphasis id="5C9F0774B91FFFDA85CA1B4DFECDF8CC" bold="true" box="[152,279,1822,1846]" pageId="8" pageNumber="427">FIGURE 3.</emphasis>
<taxonomicName id="A9EBA0E5B91FFFDA844E1B4CFE60F8CC" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[284,442,1822,1846]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="427" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B91FFFDA844E1B4CFE60F8CC" box="[284,442,1822,1846]" italics="true" pageId="8" pageNumber="427">Vitruvius dobby</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47ACBA0FB91FFFDA84ED1B4DFDD7F8CF" box="[447,525,1822,1845]" pageId="8" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs.
</paragraph>
</caption>
<caption id="3A948BEEB91EFFDB85CA1B13FAA9F812" ID-DOI="http://doi.org/10.5281/zenodo.13210090" ID-Zenodo-Dep="13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="9" pageNumber="428" startId="9.[152,255,1856,1880]" targetBox="[205,1382,181,1832]" targetPageId="9" targetType="figure">
<paragraph id="6E54DB66B91EFFDB85CA1B13FAA9F812" blockId="9.[151,1437,1856,2025]" pageId="9" pageNumber="428">
<emphasis id="5C9F0774B91EFFDB85CA1B13FEC0F8A2" bold="true" box="[152,282,1856,1880]" pageId="9" pageNumber="428">FIGURE 4.</emphasis>
<taxonomicName id="A9EBA0E5B91EFFDB84711B12FE1EF8A3" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[291,452,1857,1881]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="428" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B91EFFDB84711B12FE1EF8A3" box="[291,452,1857,1881]" italics="true" pageId="9" pageNumber="428">Vitruvius dobby</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47ACBA0FB91EFFDB849E1B12FDFBF8A2" box="[460,545,1857,1881]" pageId="9" pageNumber="428" rank="species">
<emphasis id="5C9F0774B91EFFDB849E1B12FDC0F8A3" bold="true" box="[460,538,1857,1881]" pageId="9" pageNumber="428">sp. nov</emphasis>
.
</taxonomicNameLabel>
, structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment.
</paragraph>
</caption>
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<paragraph id="6E54DB66B91DFFD885CA1A42FF39F943" blockId="10.[151,1437,1553,1721]" pageId="10" pageNumber="429">
<emphasis id="5C9F0774B91DFFD885CA1A42FEC1F9D3" bold="true" box="[152,283,1553,1577]" pageId="10" pageNumber="429">FIGURE 5.</emphasis>
<taxonomicName id="A9EBA0E5B91DFFD884771A41FE1DF9D3" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[293,455,1553,1577]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="429" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B91DFFD884771A41FE1DF9D3" box="[293,455,1553,1577]" italics="true" pageId="10" pageNumber="429">Vitruvius dobby</emphasis>
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<emphasis id="5C9F0774B91DFFD884831A42FDC5F9D3" bold="true" box="[465,543,1553,1577]" pageId="10" pageNumber="429">sp. nov</emphasis>
.
</taxonomicNameLabel>
, structures of abdomen: A—spiracle and trichobothria on ventrite VI (magnification 85×); B—detail of trichobothria on ventrite VI (500×); C–F—male genital capsule (C—caudal view, 55×; D—lateral view, 60×; E—dorsal view, 37×; F—ventral view, 42×). Abbreviations: pa = paramere; pla = posterolateral angle of genital capsule; sp = spiracle; tr = trichobothrium; x = proctiger (segment X). Scale bars: A, D—0.5 mm, B—0.1 mm; C, E–F—1 mm. Orig. Petr Kment.
</paragraph>
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<paragraph id="6E54DB66B91DFFD885951B58FB32F86B" blockId="10.[151,1436,1803,1938]" pageId="10" pageNumber="429">
<emphasis id="5C9F0774B91DFFD885951B58FEE5F8DF" bold="true" box="[199,319,1803,1829]" pageId="10" pageNumber="429">Variation.</emphasis>
There is a considerable size variation between individuals of the two sexes; females are considerably larger than males (see Measurements). The colouration of the specimens vary from straw yellow specimens with only inconspicuous minute dark brown markings to specimens with prominent dark brown colouration on pronotum, scutellum, clavus and endocorium forming vaguely delimited longitudinal stripes (see
<figureCitation id="F6D0C7E3B91DFFD881331B24FB7CF868" box="[1121,1190,1911,1938]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="10" pageNumber="429">Figs 3</figureCitation>
&amp;
<figureCitation id="F6D0C7E3B91DFFD881981B24FB00F86B" box="[1226,1242,1911,1937]" captionStart="FIGURE 6" captionStartId="11.[152,255,949,973]" captionTargetBox="[156,1436,192,921]" captionTargetId="figure-24@11.[151,1436,181,925]" captionTargetPageId="11" captionText="FIGURE 6. Vitruvius dobby sp. nov., variability of colouration.A—female, Cambodia, Kampong Speu, Chambok (11°21′25″N 104°07′09″); B–C—females, Cambodia, Preah Vihear, BeTreed Adventures (13°29′44″N 104°42′36″). Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210096" httpUri="https://zenodo.org/record/13210096/files/figure.png" pageId="10" pageNumber="429">6</figureCitation>
).
</paragraph>
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<paragraph id="6E54DB66B91CFFD985CA1FE6FAB9FC0B" blockId="11.[151,1436,949,1011]" pageId="11" pageNumber="430">
<emphasis id="5C9F0774B91CFFD985CA1FE6FECDFC37" bold="true" box="[152,279,949,973]" pageId="11" pageNumber="430">FIGURE 6.</emphasis>
<taxonomicName id="A9EBA0E5B91CFFD9844E1FE5FE60FC37" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[284,442,949,973]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="430" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B91CFFD9844E1FE5FE60FC37" box="[284,442,949,973]" italics="true" pageId="11" pageNumber="430">Vitruvius dobby</emphasis>
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<emphasis id="5C9F0774B91CFFD984ED1FE6FDD3FC37" bold="true" box="[447,521,949,973]" pageId="11" pageNumber="430">sp. nov</emphasis>
.
</taxonomicNameLabel>
, variability of colouration. A—female, Cambodia, Kampong Speu, Chambok (11°21′25″N 104°07′09″); B–C—females, Cambodia, Preah Vihear, BeTreed Adventures (13°29′44″N 104°42′36″). Orig. Petr Kment.
</paragraph>
</caption>
<subSubSection id="26F188EDB91CFFD985951810FC98FB5F" pageId="11" pageNumber="430" type="etymology">
<paragraph id="6E54DB66B91CFFD985951810FC98FB5F" blockId="11.[151,1436,1091,1225]" pageId="11" pageNumber="430">
<emphasis id="5C9F0774B91CFFD985951810FE94FBA7" bold="true" box="[199,334,1091,1117]" pageId="11" pageNumber="430">Etymology.</emphasis>
The specific name refers to Dobby the house-elf from Harry Potter’s novelistic (and later cinematographic) series, by J. K. Rowling, who was a loving character with which many members of the first author’s generation have grown up with. Noun in apposition.
</paragraph>
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<subSubSection id="26F188EDB91CFFD9859518FCFDBCFB33" box="[199,614,1199,1225]" pageId="11" pageNumber="430" type="distribution">
<paragraph id="6E54DB66B91CFFD9859518FCFDBCFB33" blockId="11.[151,1436,1091,1225]" box="[199,614,1199,1225]" pageId="11" pageNumber="430">
<emphasis id="5C9F0774B91CFFD9859518FCFEBBFB33" bold="true" box="[199,353,1199,1225]" pageId="11" pageNumber="430">Distribution.</emphasis>
<collectingCountry id="16FC9BF6B91CFFD9843518FCFE39FB33" box="[359,483,1199,1225]" name="Cambodia" pageId="11" pageNumber="430">Cambodia</collectingCountry>
!,
<collectingCountry id="16FC9BF6B91CFFD984A318FCFD84FB33" box="[497,606,1199,1225]" name="Thailand" pageId="11" pageNumber="430">Thailand</collectingCountry>
!.
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735
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<emphasis id="5C9F0774B914FFD185C51864FEB6FBAB" bold="true" box="[151,364,1079,1105]" italics="true" pageId="3" pageNumber="422">Vitruvius insignis</emphasis>
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(
<figureCitation id="F6D0C7E3B914FFD185CD1808FED0FB8F" box="[159,266,1115,1141]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="3" pageNumber="422">Figures 1</figureCitation>
,
<figureCitation id="F6D0C7E3B914FFD1844A1808FE01FB8F" box="[280,475,1115,1141]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">2A, 2B, 2C &amp; 2E</figureCitation>
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(original description).
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(s): ‘
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,
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(MCSN).
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</paragraph>
<paragraph id="6E54DB66B914FFD185C518EAFE3DF941" blockId="3.[151,1437,1178,1754]" pageId="3" pageNumber="422">
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<taxonomicName id="A9EBA0E5B914FFD185C518EAFE90FB2A" authorityName="Distant" authorityYear="1901" box="[151,330,1209,1232]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B914FFD185C518EAFE90FB2A" box="[151,330,1209,1232]" italics="true" pageId="3" pageNumber="422">Vitruvius insignis</emphasis>
</taxonomicName>
:
<treatmentCitation id="EF4AFD77B914FFD1840B18EBFDFFFB2B" author="Distant, W. L." box="[345,549,1208,1233]" page="278" pageId="3" pageNumber="422" year="1902">
<bibRefCitation id="0A7AA697B914FFD1840B18EBFE29FB2B" author="Distant, W. L." box="[345,499,1208,1233]" pageId="3" pageNumber="422" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant (1902)</bibRefCitation>
: 278
</treatmentCitation>
–279 (redescription, figures);
<treatmentCitation id="EF4AFD77B914FFD1860818EBFBECFB2A" author="Bergroth, E." box="[858,1078,1208,1233]" page="186" pageId="3" pageNumber="422" year="1908">
<bibRefCitation id="0A7AA697B914FFD1860818EBFBDEFB2B" author="Bergroth, E." box="[858,1028,1208,1233]" pageId="3" pageNumber="422" pagination="131 - 200" refId="ref7616" refString="Bergroth, E. (1908) Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique, 15 (10), 131 - 200." type="journal article" year="1908">Bergroth (1908)</bibRefCitation>
: 186
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1819218EBFA46FB2A" author="Kirkaldy, G. W." box="[1216,1436,1208,1233]" page="353" pageId="3" pageNumber="422" year="1909">
<bibRefCitation id="0A7AA697B914FFD1819218EBFAB0FB2B" author="Kirkaldy, G. W." box="[1216,1386,1208,1233]" pageId="3" pageNumber="422" refId="ref8800" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, (xl) + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 353
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1841B1884FDCFFB15" author="Distant, W. L." box="[329,533,1239,1263]" page="164" pageId="3" pageNumber="422" year="1921">
<bibRefCitation id="0A7AA697B914FFD1841B1884FE39FB15" author="Distant, W. L." box="[329,483,1239,1263]" pageId="3" pageNumber="422" pagination="164 - 169" refId="ref7952" refString="Distant, W. L. (1921) The Heteroptera of Indo-China. Entomologist, London, 54, 164 - 169." type="journal article" year="1921">Distant (1921)</bibRefCitation>
: 164
</treatmentCitation>
(listed from Indo-China, no exact locality);
<treatmentCitation id="EF4AFD77B914FFD186BB1884FB0EFB15" author="Hoffmann, W. E." box="[1001,1236,1239,1263]" page="144" pageId="3" pageNumber="422" year="1931">
<bibRefCitation id="0A7AA697B914FFD186BB1884FB7BFB15" author="Hoffmann, W. E." box="[1001,1185,1239,1263]" pageId="3" pageNumber="422" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann (1931)</bibRefCitation>
: 144
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD1840618A5FD99FAF4" author="Hoffmann, W. E." box="[340,579,1270,1294]" page="10" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1840618A5FDC1FAF4" author="Hoffmann, W. E." box="[340,539,1270,1294]" pageId="3" pageNumber="422" pagination="6 - 11" refId="ref8297" refString="Hoffmann, W. E. (1932 a) A list of the Pentatomidae, Plataspidae, and Coreidae (order Hemiptera) of China, Korea, and Indo-China. Journal of the Pan-Pacific Research Institution, 7, 6 - 11." type="journal article" year="1932">Hoffmann (1932a)</bibRefCitation>
: 10
</treatmentCitation>
(checklist);
<treatmentCitation id="EF4AFD77B914FFD1879C18A5FC16FAF4" author="Hoffmann, W. E." box="[718,972,1270,1294]" page="139" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1879C18A5FC4DFAF4" author="Hoffmann, W. E." box="[718,919,1270,1294]" pageId="3" pageNumber="422" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">Hoffmann (1932b)</bibRefCitation>
: 139
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD1806718A5FE86FAD6" author="Hoffmann, W. E." page="1016" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD1806718A5FEC0FAD7" author="Hoffmann, W. E." pageId="3" pageNumber="422" pagination="1010 - 1027" refId="ref8379" refString="Hoffmann, W. E. (1932 c) Notes on the bionomics of some oriental Pentatomidae (Hemiptera). Archivo Zoologico Italiano, 16, 1010 - 1027." type="journal article" year="1932">Hoffmann (1932c)</bibRefCitation>
: 1016
</treatmentCitation>
(distribution, plant association);
<treatmentCitation id="EF4AFD77B914FFD187931947FC1AFAD7" author="Hoffmann, W. E." box="[705,960,1300,1325]" page="470" pageId="3" pageNumber="422" year="1932">
<bibRefCitation id="0A7AA697B914FFD187931947FC53FAD6" author="Hoffmann, W. E." box="[705,905,1300,1324]" pageId="3" pageNumber="422" refId="ref8411" refString="Hoffmann, W. E. (1932 d) Brief note on Vitruvius insignis Distant. Lingnan Science Journal, 11 (3), 470, pl. 8." type="journal volume" year="1932">Hoffmann (1932d)</bibRefCitation>
: 470
</treatmentCitation>
, pl. 8: figs 1–4 (larval morphology, habitus drawings of adult and larva, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD186641960FC05FAB1" author="Cheo, M. - T." box="[822,991,1331,1355]" page="31" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD186641960FC61FAB1" author="Cheo, M. - T." box="[822,955,1331,1355]" pageId="3" pageNumber="422" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo (1935)</bibRefCitation>
: 31
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD180671960FE90FA90" author="Hoffmann, W. E." page="132" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD180671960FEC0FA90" author="Hoffmann, W. E." pageId="3" pageNumber="422" pagination="1 - 294" refId="ref8443" refString="Hoffmann, W. E. (1935 a) An abridged catalogue of certain Scutelleroidea (Plataspidae, Scutelleridae, and Pentatomidae) of China, Chosen, Indo-China, and Taiwan. Lingnan University Science Bulletin, 7, 1 - 294, 2 maps." type="book chapter" year="1935">Hoffmann (1935a)</bibRefCitation>
: 132
</treatmentCitation>
, 177 (catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD1872F1901FC84FA90" author="Hoffmann, W. E." box="[637,862,1362,1386]" page="61" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD1872F1901FCE4FA90" author="Hoffmann, W. E." box="[637,830,1362,1386]" pageId="3" pageNumber="422" pagination="57 - 62" refId="ref8491" refString="Hoffmann, W. E. (1935 b) Notes on eight pentatomids not hitherto recorded from Formosa. Transactions of the Natural History Society of Formosa, 25 (136 - 139), 57 - 62." type="journal article" year="1935">Hoffmann (1935b)</bibRefCitation>
: 61
</treatmentCitation>
–62 (catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD181DB1901FAFDFA90" author="Wu, C. F." box="[1161,1319,1362,1386]" page="225" pageId="3" pageNumber="422" year="1933">
<bibRefCitation id="0A7AA697B914FFD181DB1901FB23FA90" author="Wu, C. F." box="[1161,1273,1362,1386]" pageId="3" pageNumber="422" pagination="203 - 231" refId="ref10735" refString="Wu, C. F. (1933) A preliminary check list of Hemiptera Heteroptera recorded from Kwangtung Province, South China. Lingnan Science Journal, Supplement, 12, 203 - 231." type="journal article" year="1933">Wu (1933)</bibRefCitation>
: 225
</treatmentCitation>
(catalogue, distribution);
<treatmentCitation id="EF4AFD77B914FFD184031922FE25FA72" author="Tang, T. - H." box="[337,511,1392,1417]" page="366" pageId="3" pageNumber="422" year="1935">
<bibRefCitation id="0A7AA697B914FFD184031922FE0AFA72" author="Tang, T. - H." box="[337,464,1392,1417]" pageId="3" pageNumber="422" pagination="275 - 382" refId="ref10478" refString="Tang, T. - H. (1935) Family Pentatomidae. In: Wu, C. - F. F., Catalogus insectorum sinensium. (Catalogue of Chinese Insects). Vol. II. Fan Memorial Institute of Biology, Peiping, pp. 275 - 382. [Book pagination: 634 pp.]" type="book chapter" year="1935">Tang (1935)</bibRefCitation>
: 366
</treatmentCitation>
(catalogue, distribution); Blöte (1945): 299 (distribution);
<treatmentCitation id="EF4AFD77B914FFD1811D1922FAB7FA72" author="Kumar, R. &amp; Ghauri, M. S. K." box="[1103,1389,1392,1417]" page="25" pageId="3" pageNumber="422" year="1970">
<bibRefCitation id="0A7AA697B914FFD1811D1922FA92FA73" author="Kumar, R. &amp; Ghauri, M. S. K." box="[1103,1352,1392,1417]" pageId="3" pageNumber="422" pagination="1 - 32" refId="ref9189" refString="Kumar, R. &amp; Ghauri, M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae). Deutsche Entomologische Zeitschrift, 17 (1 - 3), 1 - 32. https: // doi. org / 10.1002 / mmnd. 4810170102" type="journal article" year="1970">Kumar &amp; Ghauri (1970)</bibRefCitation>
: 25
</treatmentCitation>
: figs 96–100, 26, 28: fig. 117 (description and drawings of male and female genitalia);
<treatmentCitation id="EF4AFD77B914FFD186AC19DCFAD6FA5D" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[1022,1292,1423,1447]" page="66" pageId="3" pageNumber="422" year="1977">
<bibRefCitation id="0A7AA697B914FFD186AC19DCFB3DFA5D" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[1022,1255,1423,1447]" pageId="3" pageNumber="422" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng (1977)</bibRefCitation>
: 66
</treatmentCitation>
, pl. 7: fig. 110 (diagnosis, habitus photograph, distribution);
<treatmentCitation id="EF4AFD77B914FFD187C119FDFC9EFA3C" author="Chen, Z. - Y." box="[659,836,1454,1478]" page="133" pageId="3" pageNumber="422" year="1982">
<bibRefCitation id="0A7AA697B914FFD187C119FDFCCCFA3C" author="Chen, Z. - Y." box="[659,790,1454,1478]" pageId="3" pageNumber="422" refId="ref7694" refString="Chen, Z. - Y. (1982) On the Pentatomidae of Dinghushan, Guangdong Province. Acta Scientiarum Naturalium Universitatis Sunyatseni, 1981 (4), 133 - 135. [in Chinese, English summary]" type="book" year="1982">Chen (1982)</bibRefCitation>
: 133
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1868E19FDFB05FA3C" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[988,1247,1454,1478]" page="113" pageId="3" pageNumber="422" year="1982">
<bibRefCitation id="0A7AA697B914FFD1868E19FDFB68FA3C" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[988,1202,1454,1478]" pageId="3" pageNumber="422" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou (1982)</bibRefCitation>
: 113
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD18406199FFDD0FA1F" author="Chen, Q. - H." box="[340,522,1484,1509]" page="117" pageId="3" pageNumber="422" year="1985">
<bibRefCitation id="0A7AA697B914FFD18406199FFE03FA1E" author="Chen, Q. - H." box="[340,473,1484,1508]" pageId="3" pageNumber="422" refId="ref7647" refString="Chen, Q. - H. (1985) The catalogue and geographical distribution of bugs (Hemiptera: Pentatomoidea) in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 11 (1), 115 - 127. [in Chinese, English summary]" type="book" year="1985">Chen (1985)</bibRefCitation>
: 117
</treatmentCitation>
(distribution); Zhang (1985): 49–50, pl. XLVI: fig. 155 (redescription, habitus drawing, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD1875E19B8FD26F9F9" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[524,764,1515,1539]" page="10" pageId="3" pageNumber="422" year="1985">
<bibRefCitation id="0A7AA697B914FFD1875E19B8FD02F9F9" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[524,728,1515,1539]" pageId="3" pageNumber="422" refId="ref10917" refString="Zhang, S. - M., Lin, Y. - J. &amp; Chen, Z. - Q. (1985) Faunal analysis of the pentatomid bugs in Jian-Feng-Ling, Hainan. Acta Agriculturae Universitatis Jiangxiensis, 1985 (2), 9 - 12. [in Chinese, English summary]" type="book" year="1985">
Zhang
<emphasis id="5C9F0774B914FFD1870719BFFD50F9F9" box="[597,650,1515,1539]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(1985)
</bibRefCitation>
: 10
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD186CB19B8FB0BF9F9" author="Schaefer, C. W. &amp; Ahmad, I." box="[921,1233,1515,1539]" page="27" pageId="3" pageNumber="422" year="1987">
<bibRefCitation id="0A7AA697B914FFD186CB19B8FB77F9F9" author="Schaefer, C. W. &amp; Ahmad, I." box="[921,1197,1515,1539]" pageId="3" pageNumber="422" pagination="21 - 34" refId="ref10436" refString="Schaefer, C. W. &amp; Ahmad, I. (1987) The food plants of four pentatomoid families (Hemiptera: Acanthosomatidae, Tessaratomidae, Urostylidae, and Dinidoridae). Phytophaga, 1, 21 - 34." type="journal article" year="1987">Schaefer &amp; Ahmad (1987)</bibRefCitation>
: 27
</treatmentCitation>
(plant association);
<treatmentCitation id="EF4AFD77B914FFD185951A59FE04F9D8" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[199,478,1546,1570]" page="290" pageId="3" pageNumber="422" year="1988">
<bibRefCitation id="0A7AA697B914FFD185951A59FE74F9D8" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[199,430,1546,1570]" pageId="3" pageNumber="422" refId="ref10813" refString="Zhang, S. - M. &amp; Chen, Q. - H. (1988) Faunal structure of the pentatomid bugs in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 14 (3), 287 - 291. [in Chinese, English summary]" type="book" year="1988">Zhang &amp; Chen (1988)</bibRefCitation>
: 290
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1872A1A59FCC5F9D8" author="Chen, Z. - Y." box="[632,799,1546,1570]" page="59" pageId="3" pageNumber="422" year="1989">
<bibRefCitation id="0A7AA697B914FFD1872A1A59FD26F9D8" author="Chen, Z. - Y." box="[632,764,1546,1570]" pageId="3" pageNumber="422" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">Chen (1989)</bibRefCitation>
: 59
</treatmentCitation>
(plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD181201A59FAD4F9D8" author="Hua, L. - Z." box="[1138,1294,1546,1570]" page="45" pageId="3" pageNumber="422" year="1989">
<bibRefCitation id="0A7AA697B914FFD181201A59FB31F9D8" author="Hua, L. - Z." box="[1138,1259,1546,1570]" pageId="3" pageNumber="422" refId="ref8652" refString="Hua, L. - Z. (1989) List of insect specimens of Zhongshan (Sun Yatsen) University. Taxonomy Division, Institute of Entomology, Zhongshan (Sun Yatsen) University, Ghungzhou, ii + 158 pp. [in Chinese, English summary]" type="book" year="1989">Hua (1989)</bibRefCitation>
: 45
</treatmentCitation>
(species list);
<treatmentCitation id="EF4AFD77B914FFD185951A7BFE6AF9BB" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[199,432,1576,1601]" page="2" pageId="3" pageNumber="422" year="1990">
<bibRefCitation id="0A7AA697B914FFD185951A7BFE43F9BB" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[199,409,1576,1601]" pageId="3" pageNumber="422" refId="ref10862" refString="Zhang, S. - M. &amp; Lin, Y. - J. (1990) The newly recorded pentatomid bugs in Xishuangbana [sic!] and their faunal structures. Acta Agriculturae Universitatis Jiangxiensis, 12 (1), 287 - 291. [in Chinese, English summary]" type="book" year="1990">Zhang &amp; Lin (1990)</bibRefCitation>
: 2
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1871B1A7BFD20F9BB" author="Zheng, L. - Y." box="[585,762,1576,1601]" page="92" pageId="3" pageNumber="422" year="1994">
<bibRefCitation id="0A7AA697B914FFD1871B1A7BFD0DF9BB" author="Zheng, L. - Y." box="[585,727,1576,1601]" pageId="3" pageNumber="422" pagination="91 - 96" refId="ref11035" refString="Zheng, L. - Y. (1994) Heteropteran insects (Hemiptera) feeding on bamboos in China. Annals of the Entomological Society of America, 87 (1), 91 - 96. https: // doi. org / 10.1093 / aesa / 87.1.91" type="journal article" year="1994">Zheng (1994)</bibRefCitation>
: 92
</treatmentCitation>
(plant association);
<treatmentCitation id="EF4AFD77B914FFD1869A1A7AFBAAF9BB" author="Hua, L. - Z." box="[968,1136,1577,1601]" page="170" pageId="3" pageNumber="422" year="2000">
<bibRefCitation id="0A7AA697B914FFD1869A1A7AFB9AF9BB" author="Hua, L. - Z." box="[968,1088,1577,1601]" pageId="3" pageNumber="422" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua (2000)</bibRefCitation>
: 170
</treatmentCitation>
(catalogue, plant association, distribution);
<treatmentCitation id="EF4AFD77B914FFD184031A14FDE7F9A5" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[337,573,1607,1631]" page="146" pageId="3" pageNumber="422" year="2002">
<bibRefCitation id="0A7AA697B914FFD184031A14FDD7F9A5" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[337,525,1607,1631]" pageId="3" pageNumber="422" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B914FFD184DD1A1BFE61F9A5" box="[399,443,1607,1631]" italics="true" pageId="3" pageNumber="422">et al</emphasis>
. (2002)
</bibRefCitation>
: 146
</treatmentCitation>
(checklist, distribution);
<treatmentCitation id="EF4AFD77B914FFD186691A14FB96F9A5" author="Zheng, L. - Y. &amp; Chen, C." box="[827,1100,1607,1631]" page="256" pageId="3" pageNumber="422" year="2002">
<bibRefCitation id="0A7AA697B914FFD186691A14FBC6F9A5" author="Zheng, L. - Y. &amp; Chen, C." box="[827,1052,1607,1631]" pageId="3" pageNumber="422" pagination="256 - 261" refId="ref11085" refString="Zheng, L. - Y. &amp; Chen, C. (2002) Hemiptera: Pentatomidae. In Huang, F. - S. (Ed.), Forest Insects of Hainan. Science Press, Beijing, pp. 256 - 261. [Book pagination: 1064 pp., in Chinese, English summary]" type="book chapter" year="2002">Zheng &amp; Chen (2002)</bibRefCitation>
: 256
</treatmentCitation>
(identification key);
<treatmentCitation id="EF4AFD77B914FFD180721A14FE9AF984" author="Liu, G. - Q. &amp; Wang, H. - J." page="189" pageId="3" pageNumber="422" year="2004">
<bibRefCitation id="0A7AA697B914FFD180721A14FED4F984" author="Liu, G. - Q. &amp; Wang, H. - J." pageId="3" pageNumber="422" pagination="173 - 189" refId="ref9251" refString="Liu, G. - Q. &amp; Wang, H. - J. (2004) Hemiptera: Scutelleridae, Tessaratomidae, Dinidoridae and Pentatomidae. In: Yang, X. - K. (Ed.), Insects from Mt. Shiwandashan area of Guangxi. China Forestry Publishing House, Beijing, pp. 173 - 189. [in Chinese, English summary]" type="book chapter" year="2004">Liu &amp; Wang (2004)</bibRefCitation>
: 189
</treatmentCitation>
(distribution);
<treatmentCitation id="EF4AFD77B914FFD1848E1A35FD4EF984" author="Rider, D. A." box="[476,660,1638,1662]" page="376" pageId="3" pageNumber="422" year="2006">
<bibRefCitation id="0A7AA697B914FFD1848E1A35FDB9F984" author="Rider, D. A." box="[476,611,1638,1662]" pageId="3" pageNumber="422" refId="ref9388" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402. [Book pagination: xiii + 550 pp.]" type="book" year="2006">Rider (2006)</bibRefCitation>
: 376
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B914FFD1864F1A35FC1EF984" author="Fan, Z. - H." box="[797,964,1638,1662]" page="521" pageId="3" pageNumber="422" year="2011">
<bibRefCitation id="0A7AA697B914FFD1864F1A35FC48F984" author="Fan, Z. - H." box="[797,914,1638,1662]" pageId="3" pageNumber="422" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan (2011)</bibRefCitation>
: 521
</treatmentCitation>
, 522: fig. 2.275, 523 (redescription, drawings of male and female genitalia, genitalia), 609: figs 578–579, 610 (habitus photographs);
<treatmentCitation id="EF4AFD77B914FFD181151AD7FAE0F967" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1095,1338,1668,1693]" page="110" pageId="3" pageNumber="422" year="2018">
<bibRefCitation id="0A7AA697B914FFD181151AD7FAD3F967" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1095,1289,1668,1693]" pageId="3" pageNumber="422" pagination="25 - 201" refId="ref9452" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of the Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): Biology, higher systematics, semiochemistry, and management. CRC Press, Boca Raton, Florida, pp. 25 - 201. [Book pagination: xix + 819 pp.] https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider
<emphasis id="5C9F0774B914FFD181D51AD6FB61F966" box="[1159,1211,1668,1692]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(2018)
</bibRefCitation>
: 110
</treatmentCitation>
, 193: fig. 2.24H (habitus photograph).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6E54DB66B914FFD185C51A91FC77F920" blockId="3.[151,1437,1178,1754]" box="[151,941,1730,1754]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185C51A91FF25F923" box="[151,255,1730,1753]" italics="true" pageId="3" pageNumber="422">Vitruvinus</emphasis>
[
<emphasis id="5C9F0774B914FFD1845C1A91FEE9F923" box="[270,307,1730,1753]" italics="true" pageId="3" pageNumber="422">sic!</emphasis>
]
<emphasis id="5C9F0774B914FFD184131A91FE4AF923" box="[321,400,1730,1753]" italics="true" pageId="3" pageNumber="422">insignis</emphasis>
:
<bibRefCitation id="0A7AA697B914FFD184CF1A91FD9BF920" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[413,577,1730,1754]" pageId="3" pageNumber="422" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B914FFD184931A90FE2EF920" box="[449,500,1730,1754]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
(2011)
</bibRefCitation>
: 59 (plant association, distribution).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB914FFD485C51B5BFCA4FD7F" lastPageId="6" lastPageNumber="425" pageId="3" pageNumber="422" type="materials_examined">
<paragraph id="6E54DB66B914FFD185C51B5BFB90F848" blockId="3.[151,1437,1800,1971]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185C51B5BFE9CF8D8" bold="true" box="[151,326,1800,1826]" pageId="3" pageNumber="422">Type material.</emphasis>
<materialsCitation id="DE83D13BB914FFD1841F1B5AFB90F848" collectingDate="1888-12" collectorName="M. Roca-Cuschachs" pageId="3" pageNumber="422" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="B15065C4B914FFD1841F1B5AFE67F8D9" box="[333,445,1801,1827]" pageId="3" pageNumber="422" type="lectotype">Lectotype</typeStatus>
(here designated):
<specimenCount id="78ED10EFB914FFD187C51B5AFD76F8D9" box="[663,684,1801,1827]" count="1" pageId="3" pageNumber="422" type="female">♀</specimenCount>
(
<figureCitation id="F6D0C7E3B914FFD187EE1B5BFCCCF8D8" box="[700,790,1800,1826]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">Fig. 2A</figureCitation>
), “g,
<date id="1A55FDA6B914FFD186061B5AFC67F8D8" box="[852,957,1800,1827]" pageId="3" pageNumber="422" value="1888-12">
<collectingDate id="0A11044EB914FFD186061B5AFC67F8D8" box="[852,957,1800,1827]" pageId="3" pageNumber="422" value="1888-12">XII 1888</collectingDate>
</date>
” [darkend, mostly illegible, interpreted by comparison with other locality labels from Fea] // “Typus” [printed in orange] //
<taxonomicName id="A9EBA0E5B914FFD181481B7FFB58F8BC" authorityName="Distant" authorityYear="1901" box="[1050,1154,1836,1862]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Vitruvius</taxonomicName>
/ insignis / Dist. / Typus! [handwritten in black ink, orange label] // “
<taxonomicName id="A9EBA0E5B914FFD187C51B03FD25F890" authorityName="Distant" authorityYear="1901" box="[663,767,1872,1898]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Vitruvius</taxonomicName>
/ insignis / Dist.” [handwritten in black ink, white label, Distant’s handwritting] //
<taxonomicName id="A9EBA0E5B914FFD184E51B27FD80F874" authority="Dist." authorityName="Distant" authorityYear="1901" box="[439,602,1908,1934]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="insignis">“insignis Dist.</taxonomicName>
” [white handwritten label in black ink] // “
<typeStatus id="B15065C4B914FFD181651B27FB36F874" box="[1079,1260,1908,1934]" pageId="3" pageNumber="422" type="lectotype">LECTOTYPUS</typeStatus>
/
<taxonomicName id="A9EBA0E5B914FFD180501B27FA54F874" authorityName="Distant" authorityYear="1901" box="[1282,1422,1908,1934]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="5C9F0774B914FFD180501B27FA54F874" box="[1282,1422,1908,1934]" italics="true" pageId="3" pageNumber="422">VITRUVIUS</emphasis>
</taxonomicName>
/
<emphasis id="5C9F0774B914FFD185C51BCBFED3F848" box="[151,265,1944,1970]" italics="true" pageId="3" pageNumber="422">INSIGNIS</emphasis>
/
<bibRefCitation id="0A7AA697B914FFD1844D1BCBFE6FF848" author="Distant, W. L." box="[287,437,1944,1970]" pageId="3" pageNumber="422" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
/ des.
<collectorName id="C31EBEB0B914FFD184AD1BCAFD3AF848" box="[511,736,1944,1971]" pageId="3" pageNumber="422">M. Roca-Cuschachs</collectorName>
<emphasis id="5C9F0774B914FFD187B51BCAFCC5F848" box="[743,799,1944,1970]" italics="true" pageId="3" pageNumber="422">et al.</emphasis>
2024” [printed, red label].
</materialsCitation>
</paragraph>
<caption id="3A948BEEB913FFD685CA1B4DFDFBF810" ID-DOI="http://doi.org/10.5281/zenodo.13210084" ID-Zenodo-Dep="13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="4" pageNumber="423" startId="4.[152,255,1822,1846]" targetBox="[203,1344,199,1769]" targetPageId="4" targetType="figure">
<paragraph id="6E54DB66B913FFD685CA1B4DFDFBF810" blockId="4.[151,1437,1822,2026]" pageId="4" pageNumber="423">
<emphasis id="5C9F0774B913FFD685CA1B4DFEC6F8CC" bold="true" box="[152,284,1822,1846]" pageId="4" pageNumber="423">FIGURE 1.</emphasis>
<taxonomicName id="A9EBA0E5B913FFD684741B4CFDAEF8CC" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[294,628,1822,1846]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="423" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B913FFD684741B4CFE00F8CC" box="[294,474,1823,1846]" italics="true" pageId="4" pageNumber="423">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B913FFD684B61B4DFDAEF8CC" author="Distant, W. L." box="[484,628,1822,1846]" pageId="4" pageNumber="423" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs.
</paragraph>
</caption>
<caption id="3A948BEEB912FFD785CA1B4CFB26F811" ID-DOI="http://doi.org/10.5281/zenodo.13210086" ID-Zenodo-Dep="13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="5" pageNumber="424" startId="5.[152,255,1823,1847]" targetBox="[151,1436,185,1799]" targetPageId="5" targetType="figure">
<paragraph id="6E54DB66B912FFD785CA1B4CFB26F811" blockId="5.[151,1436,1823,2028]" pageId="5" pageNumber="424">
<emphasis id="5C9F0774B912FFD785CA1B4CFEC3F8CD" bold="true" box="[152,281,1823,1847]" pageId="5" pageNumber="424">FIGURE 2.</emphasis>
<taxonomicName id="A9EBA0E5B912FFD784721B73FDBEF8CD" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[288,612,1823,1847]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B912FFD784721B73FE0BF8CD" box="[288,465,1824,1847]" italics="true" pageId="5" pageNumber="424">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B912FFD7848A1B4CFDBEF8CD" author="Distant, W. L." box="[472,612,1823,1847]" pageId="5" pageNumber="424" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—
<taxonomicName id="A9EBA0E5B912FFD7879F1B17FCB4F8A1" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[717,878,1859,1883]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="dobby" status="sp. nov.">
<emphasis id="5C9F0774B912FFD7879F1B17FCB4F8A1" box="[717,878,1859,1883]" italics="true" pageId="5" pageNumber="424">Vitruvius dobby</emphasis>
</taxonomicName>
<emphasis id="5C9F0774B912FFD786271B10FC1CF8A1" bold="true" box="[885,966,1859,1883]" pageId="5" pageNumber="424">
<taxonomicNameLabel id="47ACBA0FB912FFD786271B10FC1CF8A1" box="[885,966,1859,1883]" pageId="5" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female genital plates; E—
<taxonomicName id="A9EBA0E5B912FFD7818C1B17FA90F8A1" authorityName="Distant" authorityYear="1901" box="[1246,1354,1860,1883]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="424" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B912FFD7818C1B17FB37F8A1" box="[1246,1261,1860,1883]" italics="true" pageId="5" pageNumber="424">V</emphasis>
.
<emphasis id="5C9F0774B912FFD781A91B17FA90F8A1" box="[1275,1354,1860,1883]" italics="true" pageId="5" pageNumber="424">insignis</emphasis>
</taxonomicName>
, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs.
</paragraph>
</caption>
<paragraph id="6E54DB66B911FFD485951CC4FED0FE2B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
It is impossible to tell from the original description how many specimens
<bibRefCitation id="0A7AA697B911FFD486A51CC4FB42FF4B" author="Distant, W. L." box="[1015,1176,151,177]" pageId="6" pageNumber="425" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant (1901)</bibRefCitation>
had when he described this species (though provided measurements suggests probably only single one) so any type specimens must be regarded as
<typeStatus id="B15065C4B911FFD484721C8CFE58FF03" box="[288,386,223,249]" pageId="6" pageNumber="425" type="syntype">syntypes</typeStatus>
(cf.
<bibRefCitation id="0A7AA697B911FFD484EA1C8CFDE7FF03" author="ICZN &amp; International Comission on Zoological Nomenclature" box="[440,573,223,249]" pageId="6" pageNumber="425" refId="ref8754" refString="ICZN (International Comission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp. https: // doi. org / 10.5962 / bhl. title. 50608" type="book" year="1999">ICZN 1999</bibRefCitation>
: Recommendation 73F;
<bibRefCitation id="0A7AA697B911FFD486031C8CFC0EFF03" author="Rider, D. A." box="[849,980,223,249]" pageId="6" pageNumber="425" refId="ref9388" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402. [Book pagination: xiii + 550 pp.]" type="book" year="2006">Rider 2006</bibRefCitation>
).
<specimenCount id="78ED10EFB911FFD486B51C8CFBB0FF03" box="[999,1130,223,249]" count="1" pageId="6" pageNumber="425" type="female">One female</specimenCount>
<typeStatus id="B15065C4B911FFD481231C8CFB12FF03" box="[1137,1224,223,249]" pageId="6" pageNumber="425" type="syntype">syntype</typeStatus>
was located in the Museo Civico di Storia Naturale di Genova (
<collectionCode id="08FA43A3B911FFD487D71D50FD00FEE7" box="[645,730,259,285]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="6" pageNumber="425">MCSN</collectionCode>
), and has been examined by means of high quality images made available by Maria Tavano (
<figureCitation id="F6D0C7E3B911FFD484871D74FDEAFEBB" box="[469,560,295,321]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">Fig. 2A</figureCitation>
). That, along with the illustration provided by
<bibRefCitation id="0A7AA697B911FFD481131D74FB3DFEBB" author="Distant, W. L." box="[1089,1255,295,321]" pageId="6" pageNumber="425" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant (1902)</bibRefCitation>
, which is also a female specimen, leaves little doubt about the identity of this species. However, for the purpose of nomenclatural stability, we herein designate this specimen as the
<typeStatus id="B15065C4B911FFD487871D3CFCE5FE73" box="[725,831,367,393]" pageId="6" pageNumber="425" type="lectotype">lectotype</typeStatus>
. The specimen has with five labels preserved with it. In the original description (
<bibRefCitation id="0A7AA697B911FFD4849F1DC0FDB9FE57" author="Distant, W. L." box="[461,611,403,429]" pageId="6" pageNumber="425" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant 1901</bibRefCitation>
), the type locality was given as “
<collectingCountry id="16FC9BF6B911FFD4868F1DC0FBF4FE57" box="[989,1070,403,429]" name="Myanmar" pageId="6" pageNumber="425">Burma</collectingCountry>
,
<collectingRegion id="AC2F1584B911FFD481691DC0FB7FFE57" box="[1083,1189,403,429]" country="Myanmar" name="Yangon" pageId="6" pageNumber="425">Rangoon</collectingRegion>
”, currently
<collectingRegion id="AC2F1584B911FFD4807A1DC0FAA4FE57" box="[1320,1406,403,429]" country="Myanmar" name="Yangon" pageId="6" pageNumber="425">Yangon</collectingRegion>
in
<collectingCountry id="16FC9BF6B911FFD485C51DE4FEDCFE2B" box="[151,262,439,465]" name="Myanmar" pageId="6" pageNumber="425">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="6E54DB66B911FFD485951D89FCA4FD7F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951D89FDE7FE0E" bold="true" box="[199,573,474,500]" pageId="6" pageNumber="425">Additional material examined.</emphasis>
<materialsCitation id="DE83D13BB911FFD4871A1D89FE3AFDE3" collectingDate="1966-03-02" collectionCode="DARC" collectorName="Bishop Museum" country="Laos" location="Sayaboury" municipality="Sayabouri prov." pageId="6" pageNumber="425" specimenCount="1" specimenCount-male="1">
<emphasis id="5C9F0774B911FFD4871A1D89FD44FE0E" bold="true" box="[584,670,474,500]" pageId="6" pageNumber="425">
<collectingCountry id="16FC9BF6B911FFD4871A1D89FD42FE0E" box="[584,664,474,500]" name="Laos" pageId="6" pageNumber="425">LAOS</collectingCountry>
:
</emphasis>
<collectingMunicipality id="8E30411CB911FFD487F81D88FCB8FE0F" box="[682,866,475,501]" pageId="6" pageNumber="425">Sayabouri prov.</collectingMunicipality>
,
<location id="6B348DBDB911FFD486201D88FC31FE0F" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD486201D88FC31FE0F" box="[882,1003,475,501]" country="Laos" municipality="Sayabouri prov." name="Sayaboury" pageId="6" pageNumber="425">Sayaboury</location>
,
<date id="1A55FDA6B911FFD486A81D88FB79FE0F" box="[1018,1187,475,501]" pageId="6" pageNumber="425" value="1966-03-02">
<collectingDate id="0A11044EB911FFD486A81D88FB79FE0F" box="[1018,1187,475,501]" pageId="6" pageNumber="425" value="1966-03-02">2 March 1966</collectingDate>
</date>
, native collector (
<specimenCount id="78ED10EFB911FFD480281D88FA47FE0F" box="[1402,1437,475,501]" count="1" pageId="6" pageNumber="425" type="male">1♂</specimenCount>
<collectorName id="C31EBEB0B911FFD485C51DACFE8CFDE3" box="[151,342,511,537]" pageId="6" pageNumber="425">Bishop Museum</collectorName>
, in
<collectionCode id="08FA43A3B911FFD484D71DACFE02FDE3" box="[389,472,511,537]" pageId="6" pageNumber="425">DARC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="DE83D13BB911FFD484BC1DADFE23FDC7" collectingDate="1963-09-21" collectionCode="DARC" country="Thailand" location="Khao" pageId="6" pageNumber="425" specimenCount="1" specimenCount-female="1">
<emphasis id="5C9F0774B911FFD484BC1DADFD54FDE2" bold="true" box="[494,654,510,536]" pageId="6" pageNumber="425">
<collectingCountry id="16FC9BF6B911FFD484BC1DADFD53FDE2" box="[494,649,510,536]" name="Thailand" pageId="6" pageNumber="425">THAILAND</collectingCountry>
:
</emphasis>
<location id="6B348DBDB911FFD487CB1DACFD0CFDE3" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD487CB1DACFD0CFDE3" box="[665,726,511,537]" country="Thailand" name="Khao" pageId="6" pageNumber="425">Khao</location>
yai,
<date id="1A55FDA6B911FFD486461DACFC22FDE3" box="[788,1016,511,537]" pageId="6" pageNumber="425" value="1963-09-21">
<collectingDate id="0A11044EB911FFD486461DACFC22FDE3" box="[788,1016,511,537]" pageId="6" pageNumber="425" value="1963-09-21">21 September 1963</collectingDate>
</date>
,
<taxonomicName id="A9EBA0E5B911FFD481551DACFB17FDE2" authorityName="Distant" authorityYear="1901" box="[1031,1229,511,536]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B911FFD481551DACFB17FDE2" box="[1031,1229,511,536]" italics="true" pageId="6" pageNumber="425">Vitruvius insignis</emphasis>
</taxonomicName>
[in a big, folded, handwritten label] (
<specimenCount id="78ED10EFB911FFD484241E70FE4DFDC7" box="[374,407,547,573]" count="1" pageId="6" pageNumber="425" type="female">1♀</specimenCount>
<collectionCode id="08FA43A3B911FFD484CC1E70FE2BFDC7" box="[414,497,547,573]" pageId="6" pageNumber="425">DARC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB911FFD487561E70FD0EFD9B" collectingDate="1997-08-14" collectionCode="MRCB" collectorName="Dr. W. Ullrich &amp; Dr. Wolfgang Ullrich" country="Thailand" county="Um Phang" location="Schreb." municipality="Mae Kok river" pageId="6" pageNumber="425" specimenCount="1" specimenCount-male="1">
<collectingCounty id="8735A3EAB911FFD487561E70FDA4FDC7" box="[516,638,547,573]" pageId="6" pageNumber="425">Um Phang</collectingCounty>
,
<collectingMunicipality id="8E30411CB911FFD487DB1E70FCEAFDC7" box="[649,816,547,573]" pageId="6" pageNumber="425">Mae Kok river</collectingMunicipality>
,
<collectorName id="C31EBEB0B911FFD486691E70FC03FDC7" box="[827,985,547,573]" pageId="6" pageNumber="425">Dr. W. Ullrich</collectorName>
leg.,
<taxonomicName id="A9EBA0E5B911FFD4814A1E70FB1AFDC7" box="[1048,1216,547,573]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="5C9F0774B911FFD4814A1E70FB5AFDC7" box="[1048,1152,547,573]" italics="true" pageId="6" pageNumber="425">Bambusa</emphasis>
spec.
</taxonomicName>
<location id="6B348DBDB911FFD481951E70FAC5FDC7" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD481951E70FAC5FDC7" box="[1223,1311,547,573]" country="Thailand" county="Um Phang" municipality="Mae Kok river" name="Schreb." pageId="6" pageNumber="425">Schreb.</location>
,
<date id="1A55FDA6B911FFD480781E70FF08FD9B" pageId="6" pageNumber="425" value="1997-08-14">
<collectingDate id="0A11044EB911FFD480781E70FF08FD9B" pageId="6" pageNumber="425" value="1997-08-14">14 August 1997</collectingDate>
</date>
,
<collectorName id="C31EBEB0B911FFD4858E1E14FE12FD9B" box="[220,456,583,609]" pageId="6" pageNumber="425">Dr. Wolfgang Ullrich</collectorName>
collection (
<specimenCount id="78ED10EFB911FFD4871F1E14FDAAFD9B" box="[589,624,583,609]" count="1" pageId="6" pageNumber="425" type="male">1♂</specimenCount>
<collectionCode id="08FA43A3B911FFD487241E14FD16FD9B" box="[630,716,583,609]" pageId="6" pageNumber="425">MRCB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DE83D13BB911FFD4878C1E14FCA0FD7F" collectionCode="BMNH" collectorName="Siam &amp; Pres. By" country="Thailand" location="Prince" pageId="6" pageNumber="425" specimenCount="1" specimenCount-female="1" stateProvince="Chumphon">
<collectorName id="C31EBEB0B911FFD4878C1E14FCC0FD9B" box="[734,794,583,609]" pageId="6" pageNumber="425">Siam</collectorName>
,
<collectorName id="C31EBEB0B911FFD486771E14FC59FD9B" box="[805,899,583,609]" pageId="6" pageNumber="425">Pres. By</collectorName>
H.R.H.
<location id="6B348DBDB911FFD4868D1E14FBFDFD9B" LSID="urn:lsid:plazi:treatment:E6426A70B914FFD585C51864FD15FE57:6B348DBDB911FFD4868D1E14FBFDFD9B" box="[991,1063,583,609]" country="Thailand" name="Prince" pageId="6" pageNumber="425" stateProvince="Chumphon">Prince</location>
of
<collectingRegion id="AC2F1584B911FFD481191E14FB1DFD9B" box="[1099,1223,583,609]" country="Thailand" name="Chumphon" pageId="6" pageNumber="425">Chumphon</collectingRegion>
(unprecise locality and no date of collection); [in a small, printed label] (
<specimenCount id="78ED10EFB911FFD487A31E38FCC8FD7F" box="[753,786,619,645]" count="1" pageId="6" pageNumber="425" type="female">1♀</specimenCount>
<collectionCode id="08FA43A3B911FFD4864B1E38FCA8FD7F" box="[793,882,619,645]" pageId="6" pageNumber="425">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951EDDFDB7FC7B" pageId="6" pageNumber="425" type="diagnosis">
<paragraph id="6E54DB66B911FFD485951EDDFDB7FC7B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951EDDFE98FD52" bold="true" box="[199,322,654,680]" pageId="6" pageNumber="425">Diagnosis.</emphasis>
Very similar to
<taxonomicName id="A9EBA0E5B911FFD484AB1EDCFD83FD53" authorityName="Roca-Cusachs &amp; Rider &amp; Kment &amp; Jung" authorityYear="2024" box="[505,601,655,681]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="dobby">
<emphasis id="5C9F0774B911FFD484AB1EDCFD83FD53" box="[505,601,655,681]" italics="true" pageId="6" pageNumber="425">V. dobby</emphasis>
</taxonomicName>
described herein, differing externally in having colouration generally less contrasted, somewhat larger head, the pronotal humeri slightly more protruding laterad, with the apical angles slightly wider, and the anterolateral margins of pronotum are nearly in right angle to the body axis (
<figureCitation id="F6D0C7E3B911FFD481AD1E84FA87FD0B" box="[1279,1373,727,753]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Figs 1A</figureCitation>
,
<figureCitation id="F6D0C7E3B911FFD480381E84FA4AFD0B" box="[1386,1424,727,753]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">2A</figureCitation>
). The genital capsule (
<figureCitation id="F6D0C7E3B911FFD484D21EA8FE0FFCEF" box="[384,469,763,789]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1B</figureCitation>
) is wider apically (two times wider than base over dorsal rim insertion), posterolateral angles extremely broad at apices, with rhomboid outline in ventral view, somewhat spatulate, flat. Posterior rim ventrally broadly U-shaped, deeply concave, bearing a tuft of dense setae projecting ventro-medially. Posterior rim dorsally broadly U-shaped, with flat base.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951FD9FE85F9D7" pageId="6" pageNumber="425" type="description">
<paragraph id="6E54DB66B911FFD485951FD9FCCBFB87" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951FD9FDF6FC5F" bold="true" box="[199,556,906,933]" pageId="6" pageNumber="425">
Redescription.
<emphasis id="5C9F0774B911FFD4842F1FD9FDFFFC5F" bold="true" box="[381,549,906,933]" italics="true" pageId="6" pageNumber="425">Male genitalia</emphasis>
.
</emphasis>
Genital capsule (
<figureCitation id="F6D0C7E3B911FFD487A61FD8FC90FC5F" box="[756,842,907,933]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1B</figureCitation>
) apically two times wider than base over dorsal rim insertion, posterolateral angles very broad at apices, rhomboid in outline, somewhat spatulate, flat. Ventral posterior rim broadly U-shaped, deeply concave, bearing a tuft of dense setae projecting ventro-medially. Posterior rim dorsally broadly U-shaped, with flat base. Proctiger robust, quadrangular in shape. Phallus (
<figureCitation id="F6D0C7E3B911FFD481F71FA4FB20FBEB" box="[1189,1274,1015,1041]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1C</figureCitation>
) with exterior pair of conjunctival processes semi-triangular, broad at base. Internal conjunctival processes with hook-like apical portion shallowly sclerotized. Parameres (
<figureCitation id="F6D0C7E3B911FFD4873D186CFD1CFBA3" box="[623,710,1087,1113]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="6" pageNumber="425">Fig. 1D</figureCitation>
) with finger-like process apically, projecting caudad, apical lobe clearly much wider at base, somewhat triangular, robust.
</paragraph>
<paragraph id="6E54DB66B911FFD4859518D5FC58FACB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD4859518D5FE55FB5B" bold="true" box="[199,399,1158,1185]" italics="true" pageId="6" pageNumber="425">Female genitalia</emphasis>
(
<figureCitation id="F6D0C7E3B911FFD484F618D4FDD8FB5B" box="[420,514,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="6" pageNumber="425">Fig. 2C</figureCitation>
). Valvifers VIII greatly reduced, partially covered by abdominal segment VII, semicircular, with mesial margins contacting each other. Valvifers IX fused, elongate trapezoid, mesially with projection on each posteromesial angle. Laterotergites IX rectangular, elongate, slightly surpassing apices of laterotergites VIII, quadrate at apices. Sternite X quadrate. Laterotergites VIII somewhat triangular with rounded angles, sinuous at base. Internal genitalia as in generic description.
</paragraph>
<paragraph id="6E54DB66B911FFD485951968FB4BFA3B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951968FEABFAAF" bold="true" box="[199,369,1339,1365]" italics="true" pageId="6" pageNumber="425">Measurements</emphasis>
(
<specimenCount id="78ED10EFB911FFD484D61968FE41FAAF" box="[388,411,1339,1365]" count="1" pageId="6" pageNumber="425" type="male">♂</specimenCount>
, n = 1 /
<specimenCount id="78ED10EFB911FFD487571968FDC0FAAF" box="[517,538,1339,1365]" count="1" pageId="6" pageNumber="425" type="female">♀</specimenCount>
, n = 1). Total body length: 18.30/23.86; abdominal width at hemelytral base: 10.72/13.94; maximum abdominal width: 11.70/14.40; head length: 2.04–/2.40; width of head (including eyes): 4.10/4.94; anterior head width: 2.86/3.44; interocular distance: 2.42/3.02; pronotal length: 3.74/4.34; maximum pronotal width: 12.38/15.32; scutellar length: 8.02/11.22; scutellar width at base: 7.00/8.74
</paragraph>
<paragraph id="6E54DB66B911FFD485951998FD0EFA1F" blockId="6.[151,1437,151,2013]" box="[199,724,1483,1509]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951998FECDFA1F" bold="true" box="[199,279,1483,1509]" pageId="6" pageNumber="425">Larva.</emphasis>
For description see
<bibRefCitation id="0A7AA697B911FFD484A81998FD0AFA1F" author="Hoffmann, W. E." box="[506,720,1483,1509]" pageId="6" pageNumber="425" refId="ref8411" refString="Hoffmann, W. E. (1932 d) Brief note on Vitruvius insignis Distant. Lingnan Science Journal, 11 (3), 470, pl. 8." type="journal volume" year="1932">Hoffmann (1932d)</bibRefCitation>
.
</paragraph>
<paragraph id="6E54DB66B911FFD4859519BDFE85F9D7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD4859519BDFEE5F9F2" bold="true" box="[199,319,1518,1544]" pageId="6" pageNumber="425">Variation.</emphasis>
There is a considerable size variation between individuals of the two sexes; females are considerably larger than males.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951A64FEF8F98F" pageId="6" pageNumber="425" type="etymology">
<paragraph id="6E54DB66B911FFD485951A64FEF8F98F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951A64FE94F9AB" bold="true" box="[199,334,1591,1617]" pageId="6" pageNumber="425">Etymology.</emphasis>
The species epithet is the Latin adjective
<taxonomicName id="A9EBA0E5B911FFD486741A64FCA7F9AA" authorityName="Distant" authorityYear="1901" box="[806,893,1591,1616]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B911FFD486741A64FCA7F9AA" box="[806,893,1591,1616]" italics="true" pageId="6" pageNumber="425">insignis</emphasis>
</taxonomicName>
, meaning e.g. conspicuous, eminent, famous, or outstanding.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD485951A2DFD72F8B7" pageId="6" pageNumber="425" type="biology_ecology">
<paragraph id="6E54DB66B911FFD485951A2DFD72F8B7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951A2DFEFDF962" bold="true" box="[199,295,1662,1688]" pageId="6" pageNumber="425">Biology.</emphasis>
<bibRefCitation id="0A7AA697B911FFD484631A2CFE2BF963" author="Hoffmann, W. E." box="[305,497,1663,1689]" pageId="6" pageNumber="425" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann (1931</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487521A2CFD96F963" author="Hoffmann, W. E." box="[512,588,1663,1689]" pageId="6" pageNumber="425" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">1932b</bibRefCitation>
,c,d) was the first to record this species feeding and breeding on terminal branches of bamboo in
<collectingRegion id="AC2F1584B911FFD484CC1AF0FE35F947" box="[414,495,1699,1725]" country="China" name="Hainan" pageId="6" pageNumber="425">Hainan</collectingRegion>
Island, being classified as its minor pest. It was recorded from unidentified bamboo also by
<bibRefCitation id="0A7AA697B911FFD485BE1A94FEA0F91B" author="Cheo, M. - T." box="[236,378,1734,1761]" pageId="6" pageNumber="425" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo (1935)</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484D71A94FDB7F91B" author="Zheng, L. - Y. &amp; Zou, H. - G." box="[389,621,1735,1761]" pageId="6" pageNumber="425" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou (1982)</bibRefCitation>
, Zhang (1985),
<bibRefCitation id="0A7AA697B911FFD4864F1A94FC73F91B" author="Chen, Z. - Y." box="[797,937,1735,1761]" pageId="6" pageNumber="425" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">Chen (1989)</bibRefCitation>
and
<bibRefCitation id="0A7AA697B911FFD4868F1A94FB57F91B" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[989,1165,1735,1761]" pageId="6" pageNumber="425" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B911FFD481561A94FBE1F91B" box="[1028,1083,1735,1761]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
(2011)
</bibRefCitation>
. Additionally, according to the label data from one of the examined specimens, this species has been found on a species of
<taxonomicName id="A9EBA0E5B911FFD481901AB8FAF0F8FF" box="[1218,1322,1771,1797]" class="Liliopsida" family="Poaceae" genus="Bambusa" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="genus">
<emphasis id="5C9F0774B911FFD481901AB8FAF0F8FF" box="[1218,1322,1771,1797]" italics="true" pageId="6" pageNumber="425">Bambusa</emphasis>
</taxonomicName>
(
<taxonomicName id="A9EBA0E5B911FFD480651AB8FA43F8FF" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1335,1433,1771,1797]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
:
<taxonomicName id="A9EBA0E5B911FFD485C51B5CFE9BF8D3" authorityName="Luerssen" authorityYear="1893" box="[151,321,1807,1833]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="subFamily" subFamily="Bambusoideae">Bambusoideae</taxonomicName>
). It was also recorded from
<taxonomicName id="A9EBA0E5B911FFD487281B5CFCEDF8D3" authorityName="Blanco" authorityYear="1837" box="[634,823,1807,1833]" class="Insecta" family="Tachinidae" genus="Melia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="425" phylum="Arthropoda" rank="species" species="azaderach">
<emphasis id="5C9F0774B911FFD487281B5CFCEDF8D3" box="[634,823,1807,1833]" italics="true" pageId="6" pageNumber="425">Melia azaderach</emphasis>
</taxonomicName>
(
<taxonomicName id="A9EBA0E5B911FFD486171B5CFC66F8D3" box="[837,956,1807,1833]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="425" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
) without any details (
<bibRefCitation id="0A7AA697B911FFD481E31B5CFAF8F8D3" author="Hua, L. - Z." box="[1201,1314,1807,1833]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
), however, this record represents probably a sitting record.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB911FFD585951B05FC42FF2F" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425" type="distribution">
<paragraph id="6E54DB66B911FFD585951B05FC42FF2F" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,150,429]" lastPageId="7" lastPageNumber="426" pageId="6" pageNumber="425">
<emphasis id="5C9F0774B911FFD485951B05FEBBF88A" bold="true" box="[199,353,1878,1904]" pageId="6" pageNumber="425">Distribution.</emphasis>
<collectingCountry id="16FC9BF6B911FFD484341B04FE69F88B" box="[358,435,1879,1905]" name="China" pageId="6" pageNumber="425">China</collectingCountry>
!:
<collectingRegion id="AC2F1584B911FFD484ED1B04FD98F88B" box="[447,578,1879,1905]" country="China" name="Guangdong" pageId="6" pageNumber="425">Guangdong</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD4871D1B04FD16F88B" author="Cheo, M. - T." box="[591,716,1878,1905]" pageId="6" pageNumber="425" pagination="5 - 37" refId="ref7774" refString="Cheo, M. - T. (1935) A preliminary list of the insects and arachnids injurious to economic plants in China. Peking Natural History Bulletin, 10 (1), 5 - 37." type="journal article" year="1935">Cheo 1935</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD487841B04FC4CF88B" author="Hoffmann, W. E." box="[726,918,1878,1905]" pageId="6" pageNumber="425" pagination="1 - 294" refId="ref8443" refString="Hoffmann, W. E. (1935 a) An abridged catalogue of certain Scutelleroidea (Plataspidae, Scutelleridae, and Pentatomidae) of China, Chosen, Indo-China, and Taiwan. Lingnan University Science Bulletin, 7, 1 - 294, 2 maps." type="book chapter" year="1935">Hoffmann 1935a</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD486F21B04FBC1F88B" author="Chen, Z. - Y." box="[928,1051,1879,1905]" pageId="6" pageNumber="425" refId="ref7694" refString="Chen, Z. - Y. (1982) On the Pentatomidae of Dinghushan, Guangdong Province. Acta Scientiarum Naturalium Universitatis Sunyatseni, 1981 (4), 133 - 135. [in Chinese, English summary]" type="book" year="1982">Chen 1982</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD481771B04FBBAF88B" author="Chen, Z. - Y." box="[1061,1120,1879,1905]" pageId="6" pageNumber="425" refId="ref7735" refString="Chen, Z. - Y. (1989) The Hemiptera on bamboos from Guangdong Province. Journal of Bamboo Research, 8 (3), 58 - 60. [in Chinese, English summary]" type="book" year="1989">1989</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD481381B04FB00F88B" author="Hua, L. - Z." box="[1130,1242,1879,1905]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481BE1B04FA96F88B" box="[1260,1356,1879,1905]" country="China" name="Guangxi" pageId="6" pageNumber="425">Guangxi</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480091B04FEE5F86F" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." pageId="6" pageNumber="425" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484191B28FDCCF86F" author="Zhang, Y. - Q. &amp; You, Q. - J. &amp; Pu, T. - S. &amp; Lin, R. - Z." box="[331,534,1915,1941]" pageId="6" pageNumber="425" refId="ref10975" refString="Zhang, Y. - Q., You, Q. - J., Pu, T. - S. &amp; Lin, R. - Z. (Eds.) (1994) Insect Catalogue of Guangxi. Guangxi Science and Technology Press, Nanning, 8 + 438 pp. [in Chinese]" type="book" year="1994">
Zhang
<emphasis id="5C9F0774B911FFD484C81B28FE09F86F" box="[410,467,1915,1941]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
1994
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487731B28FD49F86F" author="Hua, L. - Z." box="[545,659,1915,1941]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487CD1B28FCBAF86F" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[671,864,1915,1941]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD487B71B28FCC4F86F" box="[741,798,1915,1941]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4863E1B28FBE1F86F" author="Liu, G. - Q. &amp; Wang, H. - J." box="[876,1083,1915,1941]" pageId="6" pageNumber="425" pagination="173 - 189" refId="ref9251" refString="Liu, G. - Q. &amp; Wang, H. - J. (2004) Hemiptera: Scutelleridae, Tessaratomidae, Dinidoridae and Pentatomidae. In: Yang, X. - K. (Ed.), Insects from Mt. Shiwandashan area of Guangxi. China Forestry Publishing House, Beijing, pp. 173 - 189. [in Chinese, English summary]" type="book chapter" year="2004">Liu &amp; Wang 2004</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD481151B28FB6EF86F" author="Fan, Z. - H." box="[1095,1204,1915,1941]" pageId="6" pageNumber="425" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan 2011</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD4819A1B28FAC3F86F" box="[1224,1305,1915,1941]" country="China" name="Hainan" pageId="6" pageNumber="425">Hainan</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480781B28FF0BF843" author="Hoffmann, W. E." pageId="6" pageNumber="425" pagination="138 - 151" refId="ref8251" refString="Hoffmann, W. E. (1931) Notes on Hemiptera and Homoptera at Canton, Kwangtung Province, Southern China 1924 - 1929. United States Department of Agriculture, Insect Pest Survey Bulletin, 11 (3), 138 - 151." type="journal article" year="1931">Hoffmann 1931</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4858D1BCCFEF1F843" author="Hoffmann, W. E." box="[223,299,1951,1977]" pageId="6" pageNumber="425" pagination="139 - 141" refId="ref8342" refString="Hoffmann, W. E. (1932 b) On pentatomids collected by the Lingnan University fifth Hainan Island expedition, 1929. Lingnan Science Journal, 11 (1), 139 - 141." type="journal article" year="1932">1932b</bibRefCitation>
,c,d, 1935a;
<bibRefCitation id="0A7AA697B911FFD484E11BCCFD72F843" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[435,680,1951,1977]" pageId="6" pageNumber="425" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD487E71BCCFC5EF843" author="Zhang, S. - M. &amp; Lin, Y. - J. &amp; Chen, Z. - Q." box="[693,900,1950,1977]" pageId="6" pageNumber="425" refId="ref10917" refString="Zhang, S. - M., Lin, Y. - J. &amp; Chen, Z. - Q. (1985) Faunal analysis of the pentatomid bugs in Jian-Feng-Ling, Hainan. Acta Agriculturae Universitatis Jiangxiensis, 1985 (2), 9 - 12. [in Chinese, English summary]" type="book" year="1985">
Zhang
<emphasis id="5C9F0774B911FFD486541BCCFC9AF843" box="[774,832,1951,1977]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
1985
</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD486C01BCCFBDFF843" author="Hua, L. - Z." box="[914,1029,1951,1977]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
;
<bibRefCitation id="0A7AA697B911FFD481401BCCFB02F843" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[1042,1240,1951,1977]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD4810B1BCCFB4EF843" box="[1113,1172,1951,1977]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481BF1BCCFA99F843" box="[1261,1347,1951,1977]" country="China" name="Yunnan" pageId="6" pageNumber="425">Yunnan</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480011BCCFEFDF827" author="Zheng, L. - Y. &amp; Zou, H. - G." pageId="6" pageNumber="425" pagination="113 - 120" refId="ref11147" refString="Zheng, L. - Y. &amp; Zou, H. - G. (1982) Records of heteropterous insects on bamboo from Yunnan. Zoological Research, 3 (Supplement), 113 - 120. [in Chinese with English summary]" type="journal article" year="1982">Zheng &amp; Zou 1982</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD484661B90FDD4F827" author="Zhang, S. - M. &amp; Lin, Y. - J." box="[308,526,1987,2013]" pageId="6" pageNumber="425" refId="ref10862" refString="Zhang, S. - M. &amp; Lin, Y. - J. (1990) The newly recorded pentatomid bugs in Xishuangbana [sic!] and their faunal structures. Acta Agriculturae Universitatis Jiangxiensis, 12 (1), 287 - 291. [in Chinese, English summary]" type="book" year="1990">Zhang &amp; Lin 1990</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487491B90FD54F827" author="Hua, L. - Z." box="[539,654,1987,2013]" pageId="6" pageNumber="425" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD487C91B90FCBAF827" author="Rider, D. A. &amp; Zheng, L. Y. &amp; Kerzhner, I. M." box="[667,864,1987,2013]" pageId="6" pageNumber="425" pagination="135 - 153" refId="ref9565" refString="Rider, D. A., Zheng, L. Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135 - 153. https: // doi. org / 10.31610 / zsr / 2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="5C9F0774B911FFD487B01B90FCC6F827" box="[738,796,1987,2013]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD4863F1B90FC00F827" author="Fan, Z. - H." box="[877,986,1987,2013]" pageId="6" pageNumber="425" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan 2011</bibRefCitation>
,
<bibRefCitation id="0A7AA697B911FFD486B51B90FB54F827" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[999,1166,1987,2013]" pageId="6" pageNumber="425" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B911FFD481421B90FB90F827" box="[1040,1098,1987,2013]" italics="true" pageId="6" pageNumber="425">et al.</emphasis>
2011
</bibRefCitation>
),
<collectingRegion id="AC2F1584B911FFD481F11B90FADDF827" box="[1187,1287,1987,2013]" country="China" name="Zhejiang" pageId="6" pageNumber="425">Zhejiang</collectingRegion>
(
<bibRefCitation id="0A7AA697B911FFD480451B90FA42F827" author="Chen, Q. - H." box="[1303,1432,1986,2013]" pageId="6" pageNumber="425" refId="ref7647" refString="Chen, Q. - H. (1985) The catalogue and geographical distribution of bugs (Hemiptera: Pentatomoidea) in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 11 (1), 115 - 127. [in Chinese, English summary]" type="book" year="1985">Chen 1985</bibRefCitation>
,
<bibRefCitation id="0A7AA697B910FFD585C51CC4FEA5FF4B" author="Zhang, S. - M. &amp; Chen, Q. - H." box="[151,383,151,177]" pageId="7" pageNumber="426" refId="ref10813" refString="Zhang, S. - M. &amp; Chen, Q. - H. (1988) Faunal structure of the pentatomid bugs in Zhejiang Province. Acta Agriculturae Universitatis Zhejiangensis, 14 (3), 287 - 291. [in Chinese, English summary]" type="book" year="1988">Zhang &amp; Chen 1988</bibRefCitation>
);?
<collectingCountry id="16FC9BF6B910FFD584F41CC4FDCDFF4B" box="[422,535,151,177]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
:
<collectingRegion id="AC2F1584B910FFD587731CC4FD89FF4B" box="[545,595,151,177]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="426">Java</collectingRegion>
(Blöte 1945);
<collectingCountry id="16FC9BF6B910FFD587A71CC4FCEBFF4B" box="[757,817,151,177]" name="Laos" pageId="7" pageNumber="426">Laos</collectingCountry>
! (new record),
<collectingCountry id="16FC9BF6B910FFD5868A1CC4FB91FF4B" box="[984,1099,151,177]" name="Myanmar" pageId="7" pageNumber="426">Myanmar</collectingCountry>
! (
<bibRefCitation id="0A7AA697B910FFD5810E1CC4FB2AFF4B" author="Distant, W. L." box="[1116,1264,151,177]" pageId="7" pageNumber="426" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant 1901</bibRefCitation>
,
<bibRefCitation id="0A7AA697B910FFD581A91CC4FAEDFF4B" author="Distant, W. L." box="[1275,1335,151,177]" pageId="7" pageNumber="426" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">1902</bibRefCitation>
);
<collectingCountry id="16FC9BF6B910FFD580181CC4FA46FF4B" box="[1354,1436,151,177]" name="Taiwan" pageId="7" pageNumber="426">Taiwan</collectingCountry>
(
<bibRefCitation id="0A7AA697B910FFD585F31CE8FEBEFF2F" author="Hoffmann, W. E." box="[161,356,186,213]" pageId="7" pageNumber="426" pagination="57 - 62" refId="ref8491" refString="Hoffmann, W. E. (1935 b) Notes on eight pentatomids not hitherto recorded from Formosa. Transactions of the Natural History Society of Formosa, 25 (136 - 139), 57 - 62." type="journal article" year="1935">Hoffmann 1935b</bibRefCitation>
);
<collectingCountry id="16FC9BF6B910FFD5842A1CE8FE38FF2F" box="[376,482,187,213]" name="Thailand" pageId="7" pageNumber="426">Thailand</collectingCountry>
! (new record);?
<collectingCountry id="16FC9BF6B910FFD587CF1CE8FD26FF2F" box="[669,764,187,213]" name="Vietnam" pageId="7" pageNumber="426">Vietnam</collectingCountry>
(Blöte 1945).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB910FFD585951C8CFD15FE57" pageId="7" pageNumber="426" type="discussion">
<paragraph id="6E54DB66B910FFD585951C8CFD15FE57" blockId="7.[151,1436,150,429]" pageId="7" pageNumber="426">
<emphasis id="5C9F0774B910FFD585951C8CFEE1FF03" bold="true" box="[199,315,223,249]" pageId="7" pageNumber="426">Remarks.</emphasis>
<bibRefCitation id="0A7AA697B910FFD584111C8CFE64FF03" author="Fan, Z. - H." box="[323,446,223,249]" pageId="7" pageNumber="426" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan (2011)</bibRefCitation>
provided a redescription and illustrations of the male and female genitalia of a Chinese specimens identified as
<taxonomicName id="A9EBA0E5B910FFD584FB1D50FDC7FEE6" authorityName="Distant" authorityYear="1901" box="[425,541,259,284]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B910FFD584FB1D50FDC7FEE6" box="[425,541,259,284]" italics="true" pageId="7" pageNumber="426">V. insignis</emphasis>
</taxonomicName>
. Her illustrations of the genitalia correspond with the species we recognize as
<taxonomicName id="A9EBA0E5B910FFD585C51D74FED6FEBA" authorityName="Distant" authorityYear="1901" box="[151,268,295,320]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="426" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5C9F0774B910FFD585C51D74FED6FEBA" box="[151,268,295,320]" italics="true" pageId="7" pageNumber="426">V. insignis</emphasis>
</taxonomicName>
; therefore, we can verify her identifications from
<collectingRegion id="AC2F1584B910FFD586001D74FC68FEBB" box="[850,946,295,321]" country="China" name="Guangxi" pageId="7" pageNumber="426">Guangxi</collectingRegion>
and
<collectingRegion id="AC2F1584B910FFD586BF1D74FB99FEBB" box="[1005,1091,295,321]" country="China" name="Yunnan" pageId="7" pageNumber="426">Yunnan</collectingRegion>
(
<collectingCountry id="16FC9BF6B910FFD581071D74FB44FEBB" box="[1109,1182,295,321]" name="China" pageId="7" pageNumber="426">China</collectingCountry>
). We were not able to verify the records from
<collectingCountry id="16FC9BF6B910FFD584CD1D18FE24FE9F" box="[415,510,331,357]" name="Vietnam" pageId="7" pageNumber="426">Vietnam</collectingCountry>
and
<collectingCountry id="16FC9BF6B910FFD587661D18FD7FFE9F" box="[564,677,331,357]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
:
<collectingRegion id="AC2F1584B910FFD587FD1D18FD3BFE9F" box="[687,737,331,357]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="426">Java</collectingRegion>
by Blöte (1945). The genus is present in
<collectingCountry id="16FC9BF6B910FFD581FC1D18FAC1FE9F" box="[1198,1307,331,357]" name="Indonesia" pageId="7" pageNumber="426">Indonesia</collectingCountry>
throughout pictures uploaded to iNaturalist, but many of them probably belong to an undescribed species, therefore the precise distribution of this species requires further studies.
</paragraph>
</subSubSection>
</treatment>
</document>

303
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<emphasis id="5C9F0774B915FFD085C51C8CFDD0FF03" bold="true" box="[151,522,223,249]" pageId="2" pageNumber="421">
Genus
<taxonomicName id="A9EBA0E5B915FFD085BC1C8CFDD0FF03" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[238,522,223,249]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="5C9F0774B915FFD085BC1C8CFE87FF03" bold="true" box="[238,349,223,249]" italics="true" pageId="2" pageNumber="421">Vitruvius</emphasis>
<bibRefCitation id="0A7AA697B915FFD084371C8CFDD0FF03" author="Distant, W. L." box="[357,522,223,249]" pageId="2" pageNumber="421" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<treatmentCitation id="EF4AFD77B915FFD085C51D77FE33FEC6" author="Distant, W. L." box="[151,489,292,316]" page="112" pageId="2" pageNumber="421" year="1901">
<taxonomicName id="A9EBA0E5B915FFD085C51D77FE33FEC6" authority="Distant, 1901: 111 - 112" authorityName="Distant" authorityPageNumber="111 - 112" authorityYear="1901" box="[151,489,292,316]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="5C9F0774B915FFD085C51D77FF28FEC1" box="[151,242,292,315]" italics="true" pageId="2" pageNumber="421">Vitruvius</emphasis>
<bibRefCitation id="0A7AA697B915FFD085A81D77FE33FEC6" author="Distant, W. L." box="[250,489,292,316]" pageId="2" pageNumber="421" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901: 111–112</bibRefCitation>
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(original description, systematic placement).
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species:
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<emphasis id="5C9F0774B915FFD081001D77FADEFEC1" box="[1106,1284,292,315]" italics="true" pageId="2" pageNumber="421">Vitruvius insignis</emphasis>
<bibRefCitation id="0A7AA697B915FFD0805E1D77FA42FEC6" author="Distant, W. L." box="[1292,1432,292,316]" pageId="2" pageNumber="421" pagination="99 - 114" refId="ref7813" refString="Distant, W. L. (1901) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB915FFD085C51D31FB81FDE8" pageId="2" pageNumber="421" type="reference_group">
<paragraph id="6E54DB66B915FFD085C51D31FCD5FE0E" blockId="2.[151,1437,292,530]" pageId="2" pageNumber="421">
<treatmentCitationGroup id="4EFBFC48B915FFD085C51D31FCD5FE0E" pageId="2" pageNumber="421">
<taxonomicName id="A9EBA0E5B915FFD085C51D31FF28FE83" authorityName="Distant" authorityYear="1901" box="[151,242,354,377]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="5C9F0774B915FFD085C51D31FF28FE83" box="[151,242,354,377]" italics="true" pageId="2" pageNumber="421">Vitruvius</emphasis>
</taxonomicName>
:
<treatmentCitation id="EF4AFD77B915FFD084521D32FE13FE83" author="Distant, W. L." box="[256,457,353,377]" page="278" pageId="2" pageNumber="421" year="1902">
<bibRefCitation id="0A7AA697B915FFD084521D32FE43FE83" author="Distant, W. L." box="[256,409,353,377]" pageId="2" pageNumber="421" pagination="1 - 438" refId="ref7882" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant (1902)</bibRefCitation>
: 278
</treatmentCitation>
(redescription);
<treatmentCitation id="EF4AFD77B915FFD087261D32FC96FE83" author="Bergroth, E." box="[628,844,353,377]" page="186" pageId="2" pageNumber="421" year="1908">
<bibRefCitation id="0A7AA697B915FFD087261D32FCC6FE83" author="Bergroth, E." box="[628,796,353,377]" pageId="2" pageNumber="421" pagination="131 - 200" refId="ref7616" refString="Bergroth, E. (1908) Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique, 15 (10), 131 - 200." type="journal article" year="1908">Bergroth (1908)</bibRefCitation>
: 186
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B915FFD086861D32FB77FE83" author="Kirkaldy, G. W." box="[980,1197,353,377]" page="353" pageId="2" pageNumber="421" year="1909">
<bibRefCitation id="0A7AA697B915FFD086861D32FBA7FE83" author="Kirkaldy, G. W." box="[980,1149,353,377]" pageId="2" pageNumber="421" refId="ref8800" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, (xl) + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 353
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B915FFD080671D32FE9BFE62" author="Hoffmann, W. E." page="10" pageId="2" pageNumber="421" year="1932">
<bibRefCitation id="0A7AA697B915FFD080671D32FEC0FE62" author="Hoffmann, W. E." pageId="2" pageNumber="421" pagination="6 - 11" refId="ref8297" refString="Hoffmann, W. E. (1932 a) A list of the Pentatomidae, Plataspidae, and Coreidae (order Hemiptera) of China, Korea, and Indo-China. Journal of the Pan-Pacific Research Institution, 7, 6 - 11." type="journal article" year="1932">Hoffmann (1932a)</bibRefCitation>
: 10
</treatmentCitation>
(checklist);
<treatmentCitation id="EF4AFD77B915FFD084981DD3FD24FE6D" author="Kumar, R. &amp; Ghauri, M. S. K." box="[458,766,383,408]" page="15" pageId="2" pageNumber="421" year="1970">
<bibRefCitation id="0A7AA697B915FFD084981DD3FD0EFE62" author="Kumar, R. &amp; Ghauri, M. S. K." box="[458,724,384,408]" pageId="2" pageNumber="421" pagination="1 - 32" refId="ref9189" refString="Kumar, R. &amp; Ghauri, M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae). Deutsche Entomologische Zeitschrift, 17 (1 - 3), 1 - 32. https: // doi. org / 10.1002 / mmnd. 4810170102" type="journal article" year="1970">Kumar &amp; Ghauri (1970)</bibRefCitation>
: 15
</treatmentCitation>
, 29 (systematic placement, transferred from Tessaratomidae to Pentatomidae);
<treatmentCitation id="EF4AFD77B915FFD084381DCDFD58FE4C" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[362,642,414,438]" page="63" pageId="2" pageNumber="421" year="1977">
<bibRefCitation id="0A7AA697B915FFD084381DCDFD87FE4C" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[362,605,414,438]" pageId="2" pageNumber="421" pagination="5 - 25" refId="ref8531" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297 - 300, pls. 5 - 25. [Book pagination: iii + 330 pp., 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng (1977)</bibRefCitation>
: 63
</treatmentCitation>
(key to genera);
<treatmentCitation id="EF4AFD77B915FFD0866E1DCCFC33FE4C" author="Hua, L. - Z." box="[828,1001,414,438]" page="170" pageId="2" pageNumber="421" year="2000">
<bibRefCitation id="0A7AA697B915FFD0866E1DCCFC6CFE4C" author="Hua, L. - Z." box="[828,950,414,438]" pageId="2" pageNumber="421" pagination="162 - 216" refId="ref8702" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Zhongshan University Press, pp. 162 - 216. [Book pagination: 448 pp.]" type="book chapter" year="2000">Hua (2000)</bibRefCitation>
: 170
</treatmentCitation>
(placed in Tessaratomidae);
<treatmentCitation id="EF4AFD77B915FFD0804B1DCDFEE4FE2F" author="Liu, G. - Q. &amp; Wang, H. - J." page="176" pageId="2" pageNumber="421" year="2004">
<bibRefCitation id="0A7AA697B915FFD0804B1DCDFED4FE2F" author="Liu, G. - Q. &amp; Wang, H. - J." pageId="2" pageNumber="421" pagination="173 - 189" refId="ref9251" refString="Liu, G. - Q. &amp; Wang, H. - J. (2004) Hemiptera: Scutelleridae, Tessaratomidae, Dinidoridae and Pentatomidae. In: Yang, X. - K. (Ed.), Insects from Mt. Shiwandashan area of Guangxi. China Forestry Publishing House, Beijing, pp. 173 - 189. [in Chinese, English summary]" type="book chapter" year="2004">Liu &amp; Wang (2004)</bibRefCitation>
: 176
</treatmentCitation>
(key to genera);
<treatmentCitation id="EF4AFD77B915FFD084B91DEEFD7BFE2F" author="Rider, D. A." box="[491,673,445,469]" page="376" pageId="2" pageNumber="421" year="2006">
<bibRefCitation id="0A7AA697B915FFD084B91DEEFDABFE2F" author="Rider, D. A." box="[491,625,445,469]" pageId="2" pageNumber="421" refId="ref9388" refString="Rider, D. A. (2006) Family Pentatomidae. In: Aukema, B. &amp; Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402. [Book pagination: xiii + 550 pp.]" type="book" year="2006">Rider (2006)</bibRefCitation>
: 376
</treatmentCitation>
(catalogue);
<treatmentCitation id="EF4AFD77B915FFD086751DEEFC66FE2F" author="Fan, Z. - H." box="[807,956,445,469]" page="16" pageId="2" pageNumber="421" year="2011">
<bibRefCitation id="0A7AA697B915FFD086751DEEFC40FE2F" author="Fan, Z. - H." box="[807,922,445,469]" pageId="2" pageNumber="421" refId="ref8054" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Nankai University, Tianjin, 631 pp. [unpublished dissertation; in Chinese.]" type="book" year="2011">Fan (2011)</bibRefCitation>
: 16
</treatmentCitation>
(key to genera), 521 (generic diagnosis);
<treatmentCitation id="EF4AFD77B915FFD080361DEEFEACFE0E" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." page="110" pageId="2" pageNumber="421" year="2018">
<bibRefCitation id="0A7AA697B915FFD080361DEEFE92FE0E" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." pageId="2" pageNumber="421" pagination="25 - 201" refId="ref9452" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of the Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): Biology, higher systematics, semiochemistry, and management. CRC Press, Boca Raton, Florida, pp. 25 - 201. [Book pagination: xix + 819 pp.] https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider
<emphasis id="5C9F0774B915FFD085951D8FFF20FE0E" box="[199,250,476,500]" italics="true" pageId="2" pageNumber="421">et al.</emphasis>
(2018)
</bibRefCitation>
: 110
</treatmentCitation>
–111 (morphology, systematic position).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6E54DB66B915FFD085C51DA8FB81FDE8" blockId="2.[151,1437,292,530]" box="[151,1115,506,530]" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD085C51DA8FF25FDE8" box="[151,255,507,530]" italics="true" pageId="2" pageNumber="421">Vitruvinus</emphasis>
(incorrect subsequent spelling):
<bibRefCitation id="0A7AA697B915FFD087191DA8FD35FDE8" author="He, Q. - J. &amp; Yi, Ch. - H. &amp; Yang, S." box="[587,751,506,530]" pageId="2" pageNumber="421" pagination="55 - 65" refId="ref8199" refString="He, Q. - J., Yi, Ch. - H. &amp; Yang, S. (2011) Faunal analysis of Pentatomomorpha (Hemiptera: Heteroptera) in bamboo forest of Yunnan. Chinese Agricultural Science Bulletin, 27 (22), 55 - 65." type="journal article" year="2011">
He
<emphasis id="5C9F0774B915FFD0873D1DA8FD78FDE8" box="[623,674,506,530]" italics="true" pageId="2" pageNumber="421">et al.</emphasis>
(2011)
</bibRefCitation>
: 59 (plant association, distribution).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB915FFD185C51E13FF1DFD0B" lastPageId="3" lastPageNumber="422" pageId="2" pageNumber="421" type="description">
<paragraph id="6E54DB66B915FFD085C51E13FC8AFD58" blockId="2.[151,1437,576,2042]" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD085C51E13FE9FFDA0" bold="true" box="[151,325,576,602]" pageId="2" pageNumber="421">Redescription.</emphasis>
Body strongly deltoid, widest across humeral angles, slightly flattened dorsoventrally, compact in appearance. General colouration yellowish to brownish, with darker punctures, sometimes forming vague longitudinal bands dorsally, ventrally ochraceous, pale brown.
</paragraph>
<paragraph id="6E54DB66B915FFD085951EFFFAC5FBD5" blockId="2.[151,1437,576,2042]" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD085951EFFFEDFFD3D" bold="true" box="[199,261,684,711]" italics="true" pageId="2" pageNumber="421">Head</emphasis>
(
<figureCitation id="F6D0C7E3B915FFD0844B1EFFFE41FD3D" box="[281,411,684,711]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Figs 4A–C</figureCitation>
) flat, with semi-circular outline. Mandibular plates extending beyond apex of clypeus, convergent, clearly enclosing clypeus (
<figureCitation id="F6D0C7E3B915FFD0870C1E83FD3DFD11" box="[606,743,720,747]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Figs 4A, C</figureCitation>
); mandibular plates broad, shallowly concave laterally and clearly covering antenniferous tubercles (
<figureCitation id="F6D0C7E3B915FFD087361EA7FD67FCF5" box="[612,701,756,783]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4A</figureCitation>
). Lateral margins of mandibular plates slightly deflected upwards. Anteocular tooth absent, however, anteocular region slightly prominent. Eyes globose, protruding from head outline. Ocelli located just behind imaginary line between posterior margin of eyes (
<figureCitation id="F6D0C7E3B915FFD086A01F6FFB94FCAD" box="[1010,1102,828,855]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4A</figureCitation>
). Bucculae short and low, not covering labrum and labiomere 1, shallowly concave in anterior half, convex in posterior half, its anterolateral angle obtusangulate (nearly rectangular), continually narrowing posteriorly (
<figureCitation id="F6D0C7E3B915FFD086B81FD7FB98FC65" box="[1002,1090,900,927]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4B</figureCitation>
). Labium reaching mesocoxae, labiomere 1 reaching posterior margin of head (
<figureCitation id="F6D0C7E3B915FFD087931FFBFC9EFC39" box="[705,836,936,963]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Figs 4B, C</figureCitation>
), labiomere 2 longest, labiomere 3 somewhat wider apically, labiomere 4 with apex black. Antennae pentamerous, with scape stout, not surpassing apex of head; length of antennomeres: III and IV (subequal)&gt; IIa and IIb (subequal)&gt; I, basipedicellite (IIa) and distipedicellite (IIb) about 2.5× times as long as scape, basiflagellum (III) and distiflagellum (IV) about 3× as long as scape.
</paragraph>
<paragraph id="6E54DB66B915FFD08595186BFC56F995" blockId="2.[151,1437,576,2042]" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD08595186BFEF9FBA9" bold="true" box="[199,291,1080,1107]" italics="true" pageId="2" pageNumber="421">Thorax.</emphasis>
Pronotum trapezoidal, clearly broader than long, anterior margin behind head distinctly concave, Ushaped (
<figureCitation id="F6D0C7E3B915FFD085A7180FFE89FB8D" box="[245,339,1116,1143]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">Figs 1A</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD08432180EFE5EFB8D" box="[352,388,1117,1143]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="2" pageNumber="421">2A</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD084C3180EFE6DFB8D" box="[401,439,1117,1143]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="2" pageNumber="421">3A</figureCitation>
). Anterior angles transversally projected, clearly visible but small, anterolateral margins deflected, anterior margins straight, flattened, uncalloused, with thin black line. Humeri protruding anterolaterad, reaching beyond hemelytral bases. Ventral surface of pronotum evenly punctured, punctures concolourous with surface.Scutellum triangular,elongate;apex narrow,rounded apically; anterolateral angles of scutellum concolourous, slightly foveate (
<figureCitation id="F6D0C7E3B915FFD0840518BFFE6CFAFD" box="[343,438,1260,1287]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">Figs 1A</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD0849618BEFE33FAFD" box="[452,489,1261,1287]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="2" pageNumber="421">3A</figureCitation>
). Corium slightly convex, laterally lined in black, membrane hyaline, veins clearly marked, concolourous with membrane, with no black or coloured patterns. Apex of membrane somewhat obtuse, flat apically (broadly U-shaped). Coria covering most of connexiva, basal segments partially exposed. Prosternum extremely reduced, neither grooved or ridged (
<figureCitation id="F6D0C7E3B915FFD087EC190BFCC4FA89" box="[702,798,1368,1395]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4D</figureCitation>
). Mesosternum with a slightly elevated medial carina; metasternum somewhat elevated medially, very slightly gibbous (
<figureCitation id="F6D0C7E3B915FFD086DB192FFC3CFA6D" box="[905,998,1404,1431]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4D</figureCitation>
). Ostioles large, oval-shaped, opening ventrally (
<figureCitation id="F6D0C7E3B915FFD0845F19F3FEBCFA41" box="[269,358,1440,1467]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4F</figureCitation>
); peritreme well developed but short, spout-shaped, only slightly elevated above surrounding pleuron, spatulate apically, its peritremal surface oriented ventrally (
<figureCitation id="F6D0C7E3B915FFD086F11997FBDBFA25" box="[931,1025,1476,1503]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="2" pageNumber="421">Figs 2B</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD0815C1996FB95FA25" box="[1038,1103,1477,1503]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">4E–G</figureCitation>
). Metapleural evaporatorium large, situated in anterior and central portions of metapleuron, with well developed, sublateral ridge near to its lateral margin (
<figureCitation id="F6D0C7E3B915FFD085AB1A5FFE89F9DD" box="[249,339,1548,1575]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4E</figureCitation>
), distinctly gyrificated (
<figureCitation id="F6D0C7E3B915FFD0873B1A5FFD18F9DD" box="[617,706,1548,1575]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4F</figureCitation>
). Mesopleural evaporatorium extending in posterior and lateral areas of mesopleuron. Legs moderately robust, bearing short, blackish, spine-like setae. Tibiae strongly sulcate, tarsi trimerous, tarsomere II with a protrusion basally near tarsomere III.
</paragraph>
<paragraph id="6E54DB66B915FFD085951A2BFD57F891" blockId="2.[151,1437,576,2042]" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD085951A2BFEE6F969" bold="true" box="[199,316,1656,1683]" italics="true" pageId="2" pageNumber="421">Abdomen.</emphasis>
Connexiva concolourous with dorsal ground colouration, darker in connexival outline. Ventrolateral sutures between tergites not prominent, last abdominal segment (VII) with posterolateral extension slightly angled, not acute (
<figureCitation id="F6D0C7E3B915FFD0845F1A93FEADF921" box="[269,375,1728,1755]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">Figs 1A1</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD084D61A92FE77F921" box="[388,429,1729,1755]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="2" pageNumber="421">3A</figureCitation>
<figureCitation id="F6D0C7E3B915FFD084FF1A92FE62F921" box="[429,440,1729,1755]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">1</figureCitation>
). Ventral surface of abdomen evenly punctured except central longitudinal line, which is impunctate (
<figureCitation id="F6D0C7E3B915FFD084731AB7FE50F905" box="[289,394,1764,1791]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">Figs 1A1</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD084C71AB6FE64F905" box="[405,446,1765,1791]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="2" pageNumber="421">3A</figureCitation>
<figureCitation id="F6D0C7E3B915FFD084EC1AB6FE13F905" box="[446,457,1765,1791]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">1</figureCitation>
). Ventral abdominal base wide, with a dorsoventrally flattened tubercle slightly covering and embraced by sternal plates (
<figureCitation id="F6D0C7E3B915FFD087451B5BFDADF8D9" box="[535,631,1800,1827]" captionStart="FIGURE 4" captionStartId="9.[152,255,1856,1880]" captionTargetBox="[205,1382,181,1832]" captionTargetId="figure-24@9.[205,1382,181,1832]" captionTargetPageId="9" captionText="FIGURE 4. Vitruvius dobby sp. nov., structures of head and thorax: A–C—head (A—dorsal view, magnification 37×; B— lateral view, 45×, C—ventral view, 40×); D—thoracic sterna (23×); E—external scent efferent system of the metathoracic scent gland (30×); F—peritreme and ostiole (120×); G—detail of peritreme and evaporatorium (300×). Abbreviations: b = buccula; la1 = labiomere 1; lb = labrum; mes = mesosternum; mp = mandibular plate; mts = metasternum; mys = mycoid surface of evaporatorium; o = ostiole; pes = peritremal surface. Scale bars: A–E—1 mm; F—0.4 mm; G—0.1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210090" httpUri="https://zenodo.org/record/13210090/files/figure.png" pageId="2" pageNumber="421">Fig. 4D</figureCitation>
). Spiracles black, spiracle on ventrite II partly to almost completely exposed. Trichobothria paired, inconspicuous, located laterally between an imaginary line connecting spiracles and connexival margins, transverse (
<figureCitation id="F6D0C7E3B915FFD087531B03FD5BF891" box="[513,641,1872,1899]" captionStart="FIGURE 5" captionStartId="10.[152,255,1553,1577]" captionTargetBox="[151,1435,181,1529]" captionTargetId="figure-37@10.[151,1435,181,1529]" captionTargetPageId="10" captionText="FIGURE 5. Vitruvius dobby sp. nov., structures of abdomen: A—spiracle and trichobothria on ventrite VI (magnification 85×); B—detail of trichobothria on ventrite VI (500×); C–F—male genital capsule (C—caudal view, 55×; D—lateral view, 60×; E—dorsal view, 37×; F—ventral view, 42×). Abbreviations: pa = paramere; pla = posterolateral angle of genital capsule; sp = spiracle; tr = trichobothrium; x = proctiger (segment X). Scale bars: A, D—0.5 mm, B—0.1 mm; C, E–F—1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210094" httpUri="https://zenodo.org/record/13210094/files/figure.png" pageId="2" pageNumber="421">Figs 5A–B</figureCitation>
).
</paragraph>
<paragraph id="6E54DB66B915FFD185951B27FF10FF03" blockId="2.[151,1437,576,2042]" lastBlockId="3.[151,1437,151,1005]" lastPageId="3" lastPageNumber="422" pageId="2" pageNumber="421">
<emphasis id="5C9F0774B915FFD085951B27FEADF875" bold="true" box="[199,375,1908,1935]" italics="true" pageId="2" pageNumber="421">Male genitalia.</emphasis>
Genital capsule trapezoidal, broader apically near posterolateral angles (
<figureCitation id="F6D0C7E3B915FFD081EB1B27FAEFF875" box="[1209,1333,1908,1935]" captionStart="FIGURE 5" captionStartId="10.[152,255,1553,1577]" captionTargetBox="[151,1435,181,1529]" captionTargetId="figure-37@10.[151,1435,181,1529]" captionTargetPageId="10" captionText="FIGURE 5. Vitruvius dobby sp. nov., structures of abdomen: A—spiracle and trichobothria on ventrite VI (magnification 85×); B—detail of trichobothria on ventrite VI (500×); C–F—male genital capsule (C—caudal view, 55×; D—lateral view, 60×; E—dorsal view, 37×; F—ventral view, 42×). Abbreviations: pa = paramere; pla = posterolateral angle of genital capsule; sp = spiracle; tr = trichobothrium; x = proctiger (segment X). Scale bars: A, D—0.5 mm, B—0.1 mm; C, E–F—1 mm. Orig. Petr Kment." figureDoi="http://doi.org/10.5281/zenodo.13210094" httpUri="https://zenodo.org/record/13210094/files/figure.png" pageId="2" pageNumber="421">Figs 5E–F</figureCitation>
). Genital cup distinctly exposed dorsally; ventral rim broad; dorsal rim concave with clearly visible, spine-like sclerotized processes on each side of segment X visible in dorsal and caudal views (
<figureCitation id="F6D0C7E3B915FFD086AF1BEFFB86F82D" box="[1021,1116,1980,2007]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="2" pageNumber="421">Figs 1B</figureCitation>
,
<figureCitation id="F6D0C7E3B915FFD0813F1BEEFB48F82D" box="[1133,1170,1981,2007]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="2" pageNumber="421">3B</figureCitation>
: black arrows; 5C–D). Phallotheca dorsoventrally bilobate, basal portion slightly sclerotized; spermathecal reservoir broad, clearly visible (
<figureCitation id="F6D0C7E3B914FFD185CD1CC4FF26FF4B" box="[159,252,151,177]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="3" pageNumber="422">Figs 1C</figureCitation>
,
<figureCitation id="F6D0C7E3B914FFD184591CC4FEF5FF4B" box="[267,303,151,177]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="3" pageNumber="422">3C</figureCitation>
). Phallus with two pairs of conjunctival processes, internal process bearing a sclerotized hook-like apical portion. Vesica small, inconspicuous. Parameres robust, complex, with a finger-like, small process (
<figureCitation id="F6D0C7E3B914FFD180681CE8FA4DFF2F" box="[1338,1431,187,213]" captionStart="FIGURE 1" captionStartId="4.[152,255,1822,1846]" captionTargetBox="[203,1344,199,1769]" captionTargetId="figure-37@4.[183,1404,179,1796]" captionTargetPageId="4" captionText="FIGURE 1. Vitruvius insignis Distant, 1901. A—habitus (1—dorsal view; 2—ventral view); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = fingerlike apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210084" httpUri="https://zenodo.org/record/13210084/files/figure.png" pageId="3" pageNumber="422">Figs 1D</figureCitation>
,
<figureCitation id="F6D0C7E3B914FFD185C51C8CFF67FF03" box="[151,189,223,249]" captionStart="FIGURE 3" captionStartId="8.[152,255,1822,1846]" captionTargetBox="[213,1350,191,1794]" captionTargetId="figure-37@8.[207,1381,181,1798]" captionTargetPageId="8" captionText="FIGURE 3. Vitruvius dobby sp. nov.A—habitus (1—dorsal; 2—ventral); B—genital capsule (ventral, lateral, dorsal and caudal views); C—phallus (dorsoventral and lateral views); D—right paramere. Abbreviations: ap = finger-like apical process of the paramere; ecp = external conjunctival processes; icp = interior conjunctival process; sap = sclerotized hook-like apical portion of the interior conjunctival process; sr = spermathecal reservoir (outlined in a dashed line); black arrows = spine-like sclerotized processes on each side of segment X. Scale bar measurements indicated besides each corresponding scale bar. Orig. Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210088" httpUri="https://zenodo.org/record/13210088/files/figure.png" pageId="3" pageNumber="422">3D</figureCitation>
).
</paragraph>
<paragraph id="6E54DB66B914FFD185951D51FF1DFD0B" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185951D51FE4EFEE7" bold="true" box="[199,404,258,285]" italics="true" pageId="3" pageNumber="422">Female genitalia.</emphasis>
Genital plates broad (
<figureCitation id="F6D0C7E3B914FFD187C71D50FD36FEE7" box="[661,748,259,285]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">Fig 2D</figureCitation>
); valvifers VIII rounded apically, reduced, partially covered by abdominal segment VII, mesial margins contacting medially, pilose; valvifers IX fused, elongate, trapezoid, internally with a projection on each mesial angle; laterotergites IX rectangular, elongate, pilose, coarsely punctured; sternite X rectangular; laterotergites VIII coarsely punctured, somewhat triangular, with rounded apical angles. Internal genitalia (
<figureCitation id="F6D0C7E3B914FFD184361DC0FE66FE57" box="[356,444,403,429]" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">Fig. 2E</figureCitation>
):
<emphasis id="5C9F0774B914FFD1849D1DC0FD46FE57" box="[463,668,403,429]" italics="true" pageId="3" pageNumber="422">ductus receptaculi</emphasis>
elongate, portion before globular vesicular area strongly sclerotized, except basal third. Ring sclerites (rs) elongate oval. Vesicular area membranous, openings not following vesicular axis, slightly displaced laterally. Internal rod of vesicular area strongly sclerotized, parallel-sided, acuminate at base, slightly dilated at apex. When vesicular area is turgid, internal rod of vesicular area bent.
<emphasis id="5C9F0774B914FFD181201DACFB24FDE3" box="[1138,1278,511,537]" italics="true" pageId="3" pageNumber="422">Pars distalis</emphasis>
less than onehalf length of interior rod of vesicular area. Proximal annular flange developed, projecting transversally, strongly sclerotized.
<emphasis id="5C9F0774B914FFD184721E14FE2BFD9B" box="[288,497,583,609]" italics="true" pageId="3" pageNumber="422">Pars intermedialis</emphasis>
robust, basal part in contact with proximal annular flange elongate. Distal annular flange of similar appearance as proximal annular flange, fused to apical receptacle.Apical receptacle elongate, basal portion with a medial expansion, globular-like, constrained at centre, apical part slightly globose, with three, distally directed finger-like projections, one of which clearly surpasses proximal annular flange, the other two shorter (
<figureCitation id="F6D0C7E3B914FFD1803D1EE0FF60FD0B" captionStart="FIGURE 2" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[151,1436,185,1799]" captionTargetId="figure-24@5.[151,1436,181,1799]" captionTargetPageId="5" captionText="FIGURE 2. Vitruvius insignis Distant, 1901. A—lectotype (1—dorsal view; 2—labels); B—external scent efferent system of the metathotacic glands; C—female genital plates; D—Vitruvius dobby sp. nov., female genital plates; E—V. insignis, female genital plates (internal view) and internal genitalia. Abbreviations: aaf = anterior annular flange; ar = apical receptacle. dr = ductus receptaculi; ir = internal rod of vesicular area; paf = proximal annular flange; pd = pars distalis; pi = pars intermedialis; rs = ring sclerites; va = vesicular area. Scale bar measurements indicated besides each corresponding scale bar. b, c and d of same scale. Orig. Lectotype Maria Tavano (Museo Civico di Storia Naturale di Genova); rest Marcos Roca-Cusachs." figureDoi="http://doi.org/10.5281/zenodo.13210086" httpUri="https://zenodo.org/record/13210086/files/figure.png" pageId="3" pageNumber="422">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB914FFD185951EA8FE2DFC5F" pageId="3" pageNumber="422" type="discussion">
<paragraph id="6E54DB66B914FFD185951EA8FE2DFC5F" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185951EA8FEDCFCEF" bold="true" box="[199,262,763,789]" pageId="3" pageNumber="422">Note.</emphasis>
Both species are very similar in external appearance; when specimens of both species are observed simultaneously, however there are some characters that exhibit slight, but constant differences between members of the two species. These differences, however, are difficult to discern without having specimens of both species to compare. Ultimately, the only reliable way to distinguish between the two species is by examining the male genitalia, which are distinctive.
</paragraph>
</subSubSection>
<subSubSection id="26F188EDB914FFD185951FFCFBA1FC17" pageId="3" pageNumber="422" type="etymology">
<paragraph id="6E54DB66B914FFD185951FFCFBA1FC17" blockId="3.[151,1437,151,1005]" pageId="3" pageNumber="422">
<emphasis id="5C9F0774B914FFD185951FFCFE94FC33" bold="true" box="[199,334,943,969]" pageId="3" pageNumber="422">Etymology.</emphasis>
The genus was presumably named after
<taxonomicName id="A9EBA0E5B914FFD186721FFCFC51FC33" authorityName="Distant" authorityYear="1901" box="[800,907,943,969]" class="Insecta" genus="Vitruvius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">Vitruvius</taxonomicName>
, a Roman architect and engineer during the 1
<superScript id="999E762EB914FFD180C31FFCFA46FC47" attach="none" box="[1425,1436,943,957]" fontSize="6" pageId="3" pageNumber="422">st</superScript>
century BC, known for his ten-volume work titled
<emphasis id="5C9F0774B914FFD187821F80FC5AFC17" box="[720,896,979,1005]" italics="true" pageId="3" pageNumber="422">De architectura</emphasis>
. Gender is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>