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<document id="7A4884168EEAD548697396EE32AF5672" ID-DOI="10.1111/j.1096-3642.2008.00415.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630461491835" checkinUser="felipe" docAuthor="Wu, Shu-Ping, Hwang, Chung-Chi &amp; Lin, Yao-Sung" docDate="2008" docId="E45287EFFFE5430BFF75F92283C854B4" docLanguage="en" docName="j.1096-3642.2008.00415.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Satsuma lini Wu &amp; Hwang &amp; Lin 2008, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="456" masterDocId="186BFF97FFF54318FF92FFF482155506" masterDocTitle="Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan" masterLastPageNumber="493" masterPageNumber="437" pageNumber="453" updateTime="1699279363994" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4803C40712D39F41EC4BFAE6C64811CF">Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan</mods:title>
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<mods:namePart id="7CAE545BF1A27B3D34BA862E116857C3">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="18962FC35D55A4E0309A2A447BA2B39D">Hwang, Chung-Chi</mods:namePart>
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<subSubSection id="24E16572FFE54308FF75F922808B53EB" box="[231,670,1750,1773]" pageId="16" pageNumber="453" type="nomenclature">
<paragraph id="6C4436F9FFE54308FF75F922808B53EB" blockId="16.[231,670,1750,1773]" box="[231,670,1750,1773]" pageId="16" pageNumber="453">
<heading id="370C8195FFE54308FF75F922808B53EB" box="[231,670,1750,1773]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="453" reason="2">
<taxonomicName id="ABFB4D7AFFE54308FF75F922838453EC" ID-CoL="79QBB" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[231,401,1750,1771]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="453" phylum="Mollusca" rank="species" species="lini" status="sp. nov.">SATSUMA LINI</taxonomicName>
<emphasis id="5E8FEAEBFFE54308FE09F92E801553EA" bold="true" box="[411,512,1751,1772]" pageId="16" pageNumber="453">
<taxonomicNameLabel id="45BC5790FFE54308FE09F92E801553EA" box="[411,512,1751,1772]" pageId="16" pageNumber="453" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4C02A7CFFE54308FD80F923807E53EA" box="[530,619,1751,1772]" captionStart="Figure 11" captionStartId="18.[144,223,717,736]" captionTargetBox="[369,1201,198,688]" captionTargetId="figure-263@18.[367,1201,196,688]" captionTargetPageId="18" captionText="Figure 11. Satsuma lini sp. nov. AD, holotype (TMMT 0602, shell height = 17.6 mm). E, F, paratype (TMMT 0621, shell height = 17 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446725" httpUri="https://zenodo.org/record/5446725/files/figure.png" pageId="16" pageNumber="453">FIGS 11</figureCitation>
,
<figureCitation id="F4C02A7CFFE54308FDE8F923808353EB" box="[634,662,1751,1773]" captionStart="Figure 12" captionStartId="18.[144,223,1493,1512]" captionTargetBox="[150,755,813,1463]" captionTargetId="figure-297@18.[148,756,812,1464]" captionTargetPageId="18" captionText="Figure 12. Reproductive system of Satsuma lini sp. nov. (holotype, TMMT 0602), immature genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446727" httpUri="https://zenodo.org/record/5446727/files/figure.png" pageId="16" pageNumber="453">12</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24E16572FFE5430BFF02F90983C854B4" lastPageId="19" lastPageNumber="456" pageId="16" pageNumber="453" type="materials_examined">
<paragraph id="6C4436F9FFE54308FF02F909837D5214" blockId="16.[144,758,1789,1903]" box="[144,360,1789,1810]" pageId="16" pageNumber="453">
<emphasis id="5E8FEAEBFFE54308FF02F909837D5214" box="[144,360,1789,1810]" italics="true" pageId="16" pageNumber="453">Material examined</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE54308FF02F8E886F55332" blockId="16.[144,758,1789,1903]" lastBlockId="16.[806,1421,1536,1588]" pageId="16" pageNumber="453">
<emphasis id="5E8FEAEBFFE54308FF02F8E883575237" box="[144,322,1820,1841]" italics="true" pageId="16" pageNumber="453">Type specimen:</emphasis>
<materialsCitation id="DC933CA4FFE54308FEC1F8E886F55332" accessionNumber="A21015" collectionCode="ANSP" pageId="16" pageNumber="453" specimenCode="TMMT 0602, TMMT 0621, BMNH 20060753" specimenCount="2" typeStatus="holotype">
<typeStatus id="B340885BFFE54308FEC1F8E883A85234" box="[339,445,1820,1842]" pageId="16" pageNumber="453" type="holotype">Holotype</typeStatus>
:
<specimenCode id="3C5D9E82FFE54308FE42F8E880725234" box="[464,615,1820,1842]" collectionCode="TMMT" pageId="16" pageNumber="453">TMMT 0602</specimenCode>
(from type locality, dry shell, tissue in alcohol, dissected). Three
<typeStatus id="B340885BFFE54308FF02F8AD83115268" box="[144,260,1881,1902]" pageId="16" pageNumber="453" type="paratype">paratypes</typeStatus>
: all from type locality,
<specimenCode id="3C5D9E82FFE54308FD9CF8AD80895269" box="[526,668,1881,1903]" collectionCode="TMMT" pageId="16" pageNumber="453">TMMT 0621</specimenCode>
(immature, in alcohol);
<collectionCode id="0AEAAE3CFFE54308FC63F9F486225313" box="[1009,1079,1536,1557]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="16" pageNumber="453">ANSP</collectionCode>
A21015 (shell with tissue in alcohol);
<specimenCode id="3C5D9E82FFE54308FC1DF9EB86425332" box="[911,1111,1566,1588]" collectionCode="BMNH" pageId="16" pageNumber="453">BMNH 20060753</specimenCode>
(dry shell).
</materialsCitation>
</paragraph>
<paragraph id="6C4436F9FFE54308FCB4F98481A25382" blockId="16.[806,1421,1647,1730]" box="[806,951,1647,1669]" pageId="16" pageNumber="453">
<emphasis id="5E8FEAEBFFE54308FCB4F98481A25382" box="[806,951,1647,1669]" italics="true" pageId="16" pageNumber="453">
<typeStatus id="B340885BFFE54308FCB4F984814F5383" box="[806,858,1648,1669]" pageId="16" pageNumber="453">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE54308FCB4F97A819453C4" blockId="16.[806,1421,1647,1730]" pageId="16" pageNumber="453">
<materialsCitation id="DC933CA4FFE54308FCB4F97A819453C4" country="Taiwan" location="Nantou County" pageId="16" pageNumber="453" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
Shanlinxi,
<collectingRegion id="AE3FF81BFFE54308FC3EF97A867053A5" box="[940,1125,1678,1699]" country="Taiwan" name="Nantou" pageId="16" pageNumber="453">Nantou County</collectingRegion>
, central
<collectingCountry id="14EC7669FFE54308FB4BF97A873853A2" box="[1241,1325,1678,1700]" name="Taiwan" pageId="16" pageNumber="453">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFE54308FAD7F97A879353A5" box="[1349,1414,1678,1700]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="16" pageNumber="453">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFE54308FCB4F958816253C4" box="[806,887,1708,1730]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="16" pageNumber="453">Table 1</tableCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="6C4436F9FFE54308FCB4F90A81835215" blockId="16.[806,1421,1790,1903]" box="[806,918,1790,1811]" pageId="16" pageNumber="453">
<emphasis id="5E8FEAEBFFE54308FCB4F90A81835215" box="[806,918,1790,1811]" italics="true" pageId="16" pageNumber="453">Diagnosis</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE5430AFCB4F8E881B25642" blockId="16.[806,1421,1790,1903]" lastBlockId="18.[806,1422,815,1173]" lastPageId="18" lastPageNumber="455" pageId="16" pageNumber="453">
Shell and soft body white with grey pedal stripe; periphery angulated but not keeled; junction between columellar and inferior lips angulated. Because of a
<emphasis id="5E8FEAEBFFE7430AFCB4FCDB81B25642" box="[806,935,815,836]" italics="true" pageId="18" pageNumber="455">Description</emphasis>
</paragraph>
<caption id="38846671FFE44309FF36F8FF80E7523D" ID-DOI="http://doi.org/10.5281/zenodo.5446723" ID-Zenodo-Dep="5446723" httpUri="https://zenodo.org/record/5446723/files/figure.png" pageId="17" pageNumber="454" startId="17.[164,243,1803,1822]" targetBox="[168,1441,197,1771]" targetPageId="17">
<paragraph id="6C4436F9FFE44309FF36F8FF80E7523D" blockId="17.[164,1441,1803,1851]" pageId="17" pageNumber="454">
<emphasis id="5E8FEAEBFFE44309FF36F8FF830C5218" bold="true" box="[164,281,1803,1822]" pageId="17" pageNumber="454">Figure 10.</emphasis>
Reproductive system of
<taxonomicName id="ABFB4D7AFFE44309FDA0F8FF80F8521B" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1906" box="[562,749,1803,1822]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="454" phylum="Mollusca" rank="species" species="insignis">
<emphasis id="5E8FEAEBFFE44309FDA0F8FF80F8521B" box="[562,749,1803,1822]" italics="true" pageId="17" pageNumber="454">Satsuma insignis</emphasis>
</taxonomicName>
(TMMT P-0280-1, Tengzhi). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations.
</paragraph>
</caption>
<caption id="38846671FFE7430AFF02FD39865B57FB" ID-DOI="http://doi.org/10.5281/zenodo.5446725" ID-Zenodo-Dep="5446725" httpUri="https://zenodo.org/record/5446725/files/figure.png" pageId="18" pageNumber="455" startId="18.[144,223,717,736]" targetBox="[369,1201,198,688]" targetPageId="18">
<paragraph id="6C4436F9FFE7430AFF02FD39865B57FB" blockId="18.[144,1421,716,766]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFF02FD39801957E6" bold="true" box="[144,524,717,736]" pageId="18" pageNumber="455">
Figure 11.
<taxonomicName id="ABFB4D7AFFE7430AFE82FD3983BF57E6" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[272,426,717,736]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="455" phylum="Mollusca" rank="species" species="lini" status="sp. nov.">
<emphasis id="5E8FEAEBFFE7430AFE82FD3983BF57E6" bold="true" box="[272,426,717,736]" italics="true" pageId="18" pageNumber="455">Satsuma lini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFE7430AFE21FD39801957E6" box="[435,524,717,736]" pageId="18" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AD, holotype (TMMT 0602, shell height = 17.6 mm). E, F, paratype (TMMT 0621, shell height = 17 mm). G, living specimen. Arrow indicates the pre-apertural constriction.
</paragraph>
</caption>
<paragraph id="6C4436F9FFE7430AFCB4FCB987645193" blockId="18.[806,1422,815,1173]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFCB4FCB981745664" box="[806,865,845,866]" italics="true" pageId="18" pageNumber="455">Shell</emphasis>
(
<figureCitation id="F4C02A7CFFE7430AFCE6FCB981D45665" box="[884,961,845,867]" captionStart="Figure 11" captionStartId="18.[144,223,717,736]" captionTargetBox="[369,1201,198,688]" captionTargetId="figure-263@18.[367,1201,196,688]" captionTargetPageId="18" captionText="Figure 11. Satsuma lini sp. nov. AD, holotype (TMMT 0602, shell height = 17.6 mm). E, F, paratype (TMMT 0621, shell height = 17 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446725" httpUri="https://zenodo.org/record/5446725/files/figure.png" pageId="18" pageNumber="455">Fig. 11</figureCitation>
): Dextral, conical, medium sized. Apex obtuse. Whorls inflated. Periphery bluntly angulated. Base expanded. Pre-apertural constriction behind outer and inferior lips present. Surface smooth, glossy with spiral striae. Shell colour whitish. Periostracum thin. Aperture diagonal, ovate-lunate, angulated between outer lip and inferior lip. Peristome thin, expanded, reflected at curved inferior lip. Superior columellar lip reflected. Umbilicus covered by columellar lip mostly, crevice-like. Columellar lip vertical to subvertical, angulated at junction with inferior lip.
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4FB3786635077" blockId="18.[806,1421,1219,1394]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFCB4FB3781DC51DE" box="[806,969,1219,1240]" italics="true" pageId="18" pageNumber="455">Band or stain:</emphasis>
Polymorphism exists in this taxon. The majority of individuals do not possess band or colour stain, and display a whitish coloration. A rare form exists with pink stain and smear from the third to the fourth band position. Band formula = 0 0 0 0 0 0 0 0; 0 0 (0 0) 0 0 0 0 (very rare).
</paragraph>
<caption id="38846671FFE7430AFF02FA2180275325" ID-DOI="http://doi.org/10.5281/zenodo.5446727" ID-Zenodo-Dep="5446727" httpUri="https://zenodo.org/record/5446727/files/figure.png" pageId="18" pageNumber="455" startId="18.[144,223,1493,1512]" targetBox="[150,755,813,1463]" targetPageId="18">
<paragraph id="6C4436F9FFE7430AFF02FA2180275325" blockId="18.[144,758,1493,1571]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFF02FA21831050EE" bold="true" box="[144,261,1493,1512]" pageId="18" pageNumber="455">Figure 12.</emphasis>
Reproductive system of
<emphasis id="5E8FEAEBFFE7430AFDB4FA2182AA5300" bold="true" pageId="18" pageNumber="455">
<taxonomicName id="ABFB4D7AFFE7430AFDB4FA2180D350EE" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[550,710,1493,1512]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="455" phylum="Mollusca" rank="species" species="lini" status="sp. nov.">
<emphasis id="5E8FEAEBFFE7430AFDB4FA2180D350EE" bold="true" box="[550,710,1493,1512]" italics="true" pageId="18" pageNumber="455">Satsuma lini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFE7430AFD44FA2282AA5300" pageId="18" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype, TMMT 0602), immature genitalia. Scale bar = 5 mm. See text for abbreviations.
</paragraph>
</caption>
<paragraph id="6C4436F9FFE7430AFF02F99F839F53B8" blockId="18.[144,759,1643,1726]" pageId="18" pageNumber="455">lack of knowledge of its genitalia, distinguishing this species from similar species based on shell characteristics only is difficult.</paragraph>
<paragraph id="6C4436F9FFE7430AFF02F90983125214" blockId="18.[144,758,1789,1903]" box="[144,263,1789,1810]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFF02F90983125214" box="[144,263,1789,1810]" italics="true" pageId="18" pageNumber="455">Etymology</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE7430AFF02F8E883815269" blockId="18.[144,758,1789,1903]" pageId="18" pageNumber="455">
L.
<taxonomicName id="ABFB4D7AFFE7430AFF21F8E882CE5237" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[179,219,1820,1841]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="455" phylum="Mollusca" rank="species" species="lini">
<emphasis id="5E8FEAEBFFE7430AFF21F8E882CE5237" box="[179,219,1820,1841]" italics="true" pageId="18" pageNumber="455">lini</emphasis>
</taxonomicName>
: the name is dedicated to the late Professor Lin, Fei-Jan (19342004), a pioneering evolutionary biologist from
<collectingCountry id="14EC7669FFE7430AFEAAF8AD83855269" box="[312,400,1881,1903]" name="Taiwan" pageId="18" pageNumber="455">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4FA6B879B50F5" blockId="18.[806,1422,1439,1523]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFCB4FA6B860850B3" box="[806,1053,1439,1461]" italics="true" pageId="18" pageNumber="455">Reproductive system</emphasis>
(
<figureCitation id="F4C02A7CFFE7430AFBAEFA6B869F50B0" box="[1084,1162,1439,1462]" captionStart="Figure 12" captionStartId="18.[144,223,1493,1512]" captionTargetBox="[150,755,813,1463]" captionTargetId="figure-297@18.[148,756,812,1464]" captionTargetPageId="18" captionText="Figure 12. Reproductive system of Satsuma lini sp. nov. (holotype, TMMT 0602), immature genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446727" httpUri="https://zenodo.org/record/5446727/files/figure.png" pageId="18" pageNumber="455">Fig. 12</figureCitation>
): No reproductively mature specimens were collected. An immature genitalia is figured to demonstrate its basic configuration.
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4F9D481A45333" blockId="18.[806,1421,1568,1682]" box="[806,945,1568,1589]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFCB4F9D481A45333" box="[806,945,1568,1589]" italics="true" pageId="18" pageNumber="455">Distribution</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4F9CB874F5394" blockId="18.[806,1421,1568,1682]" pageId="18" pageNumber="455">
<materialsCitation id="DC933CA4FFE7430AFCB4F9CB87405394" country="Taiwan" elevation="1700" location="Shanlinxi" pageId="18" pageNumber="455" specimenCount="1" stateProvince="Nantou">
The species was found only in a narrow area of mid-altitude forest (
<quantity id="AB039B1CFFE7430AFB8DF9A986AC5375" box="[1055,1209,1629,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="18" pageNumber="455" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="E7D6D1CAFFE7430AFB8DF9A986AC5375" box="[1055,1209,1629,1651]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="18" pageNumber="455" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">16001800 m</elevation>
</quantity>
) near
<location id="69246022FFE7430AFA85F9A9879D5375" LSID="urn:lsid:plazi:treatment:E45287EFFFE5430BFF75F92283C854B4:69246022FFE7430AFA85F9A9879D5375" box="[1303,1416,1629,1651]" country="Taiwan" name="Shanlinxi" pageId="18" pageNumber="455" stateProvince="Nantou">Shanlinxi</location>
,
<collectingRegion id="AE3FF81BFFE7430AFCB4F98981CC5397" box="[806,985,1660,1682]" country="Taiwan" name="Nantou" pageId="18" pageNumber="455">Nantou County</collectingRegion>
, central
<collectingCountry id="14EC7669FFE7430AFBD3F98886805394" box="[1089,1173,1660,1682]" name="Taiwan" pageId="18" pageNumber="455">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFE7430AFB35F98886FD5397" box="[1191,1256,1660,1682]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="18" pageNumber="455">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFE7430AFB6AF988875F5394" box="[1272,1354,1660,1682]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="18" pageNumber="455">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4F934819E53D3" blockId="18.[806,1421,1728,1903]" box="[806,907,1728,1749]" pageId="18" pageNumber="455">
<emphasis id="5E8FEAEBFFE7430AFCB4F934819E53D3" box="[806,907,1728,1749]" italics="true" pageId="18" pageNumber="455">Remarks</emphasis>
</paragraph>
<paragraph id="6C4436F9FFE7430AFCB4F92A864B5269" blockId="18.[806,1421,1728,1903]" pageId="18" pageNumber="455">
Inhabits typically above
<specimenCount id="7AFDFD70FFE7430AFBF0F92B868653F2" box="[1122,1171,1759,1780]" pageId="18" pageNumber="455" type="generic">
<quantity id="AB039B1CFFE7430AFBF0F92B869A53F2" box="[1122,1167,1759,1780]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="455" unit="m" value="3.0">3 m</quantity>
.
</specimenCount>
Adults were found during winter (
<figureCitation id="F4C02A7CFFE7430AFC71F90986535215" box="[995,1094,1789,1811]" captionStart="Figure 11" captionStartId="18.[144,223,717,736]" captionTargetBox="[369,1201,198,688]" captionTargetId="figure-263@18.[367,1201,196,688]" captionTargetPageId="18" captionText="Figure 11. Satsuma lini sp. nov. AD, holotype (TMMT 0602, shell height = 17.6 mm). E, F, paratype (TMMT 0621, shell height = 17 mm). G, living specimen. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446725" httpUri="https://zenodo.org/record/5446725/files/figure.png" pageId="18" pageNumber="455">Fig. 11G</figureCitation>
). Only
<specimenCount id="7AFDFD70FFE7430AFB35F909874C5215" box="[1191,1369,1789,1811]" pageId="18" pageNumber="455" type="generic">four specimens</specimenCount>
are available; two of these have adult shell morphology but genital development is incomplete. The other individuals are sub-adults.
</paragraph>
<paragraph id="6C4436F9FFE6430BFF2EFF3283C854B4" blockId="19.[164,779,198,434]" pageId="19" pageNumber="456">
This species is morphologically similar to
<taxonomicName id="ABFB4D7AFFE6430BFD0EFF32811655DD" authorityName=", Ohara &amp; Otani" authorityYear="2002" box="[668,771,198,219]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="albida">
<emphasis id="5E8FEAEBFFE6430BFD0EFF32811655DD" box="[668,771,198,219]" italics="true" pageId="19" pageNumber="456">S. albida</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFB4D7AFFE6430BFF36FF11833E55FF" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[164,299,228,250]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="swinhoei" status="sp. nov.">
<emphasis id="5E8FEAEBFFE6430BFF36FF11833E55FF" box="[164,299,228,250]" italics="true" pageId="19" pageNumber="456">S. swinhoei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFE6430BFEA4FF11839A55FC" box="[310,399,229,250]" pageId="19" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="ABFB4D7AFFE6430BFE42FF11805855FC" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1906" box="[464,589,229,250]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="insignis">
<emphasis id="5E8FEAEBFFE6430BFE42FF11805855FC" box="[464,589,229,250]" italics="true" pageId="19" pageNumber="456">S. insignis</emphasis>
</taxonomicName>
, but its genetic relationship is closer to
<taxonomicName id="ABFB4D7AFFE6430BFE54FEF78057541F" baseAuthorityName="PILSBRY &amp; HIRASE" baseAuthorityYear="1906" box="[454,578,259,281]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="insignis">
<emphasis id="5E8FEAEBFFE6430BFE54FEF78057541F" box="[454,578,259,281]" italics="true" pageId="19" pageNumber="456">S. insignis</emphasis>
</taxonomicName>
(see Phylogeny). Compared with
<taxonomicName id="ABFB4D7AFFE6430BFEC9FED683D65431" authorityName=", Ohara &amp; Otani" authorityYear="2002" box="[347,451,290,311]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="albida">
<emphasis id="5E8FEAEBFFE6430BFEC9FED683D65431" box="[347,451,290,311]" italics="true" pageId="19" pageNumber="456">S. albida</emphasis>
</taxonomicName>
,
<taxonomicName id="ABFB4D7AFFE6430BFE43FED6800D5431" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[465,536,290,311]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="lini">
<emphasis id="5E8FEAEBFFE6430BFE43FED6800D5431" box="[465,536,290,311]" italics="true" pageId="19" pageNumber="456">S. lini</emphasis>
</taxonomicName>
has a larger, thinner and more rigid shell, higher spire and more oblique aperture. The species differs from
<taxonomicName id="ABFB4D7AFFE6430BFDCAFEAB80F15472" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[600,740,351,372]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="456" phylum="Mollusca" rank="species" species="swinhoei">
<emphasis id="5E8FEAEBFFE6430BFDCAFEAB80F15472" box="[600,740,351,372]" italics="true" pageId="19" pageNumber="456">S. swinhoei</emphasis>
</taxonomicName>
in having a periphery that is not keeled, smaller shell diameter and higher spire.
</paragraph>
</subSubSection>
</treatment>
</document>