Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan Author Wu, Shu-Ping Author Hwang, Chung-Chi Author Lin, Yao-Sung text Zoological Journal of the Linnean Society 2008 2008-10-31 154 3 437 493 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x journal article 10.1111/j.1096-3642.2008.00415.x 0024-4082 5446700 AE7FA6D7-DB18-4868-9698-822F799FFCBD SATSUMA LINI SP. NOV. ( FIGS 11 , 12 ) Material examined Type specimen: Holotype : TMMT 0602 (from type locality, dry shell, tissue in alcohol, dissected). Three paratypes : all from type locality, TMMT 0621 (immature, in alcohol); ANSP A21015 (shell with tissue in alcohol); BMNH 20060753 (dry shell). Type locality Shanlinxi, Nantou County , central Taiwan ( Fig. 1 , Table 1 ) Diagnosis Shell and soft body white with grey pedal stripe; periphery angulated but not keeled; junction between columellar and inferior lips angulated. Because of a Description Figure 10. Reproductive system of Satsuma insignis (TMMT P-0280-1, Tengzhi). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations. Figure 11. Satsuma lini sp. nov. A–D, holotype (TMMT 0602, shell height = 17.6 mm). E, F, paratype (TMMT 0621, shell height = 17 mm). G, living specimen. Arrow indicates the pre-apertural constriction. Shell ( Fig. 11 ): Dextral, conical, medium sized. Apex obtuse. Whorls inflated. Periphery bluntly angulated. Base expanded. Pre-apertural constriction behind outer and inferior lips present. Surface smooth, glossy with spiral striae. Shell colour whitish. Periostracum thin. Aperture diagonal, ovate-lunate, angulated between outer lip and inferior lip. Peristome thin, expanded, reflected at curved inferior lip. Superior columellar lip reflected. Umbilicus covered by columellar lip mostly, crevice-like. Columellar lip vertical to subvertical, angulated at junction with inferior lip. Band or stain: Polymorphism exists in this taxon. The majority of individuals do not possess band or colour stain, and display a whitish coloration. A rare form exists with pink stain and smear from the third to the fourth band position. Band formula = 0 0 0 0 0 0 0 0; 0 0 (0 0) 0 0 0 0 (very rare). Figure 12. Reproductive system of Satsuma lini sp. nov. (holotype, TMMT 0602), immature genitalia. Scale bar = 5 mm. See text for abbreviations. lack of knowledge of its genitalia, distinguishing this species from similar species based on shell characteristics only is difficult. Etymology L. lini : the name is dedicated to the late Professor Lin, Fei-Jan (1934–2004), a pioneering evolutionary biologist from Taiwan . Reproductive system ( Fig. 12 ): No reproductively mature specimens were collected. An immature genitalia is figured to demonstrate its basic configuration. Distribution The species was found only in a narrow area of mid-altitude forest ( 1600–1800 m ) near Shanlinxi , Nantou County , central Taiwan ( Fig. 1 , Table 1 ) . Remarks Inhabits typically above 3 m . Adults were found during winter ( Fig. 11G ). Only four specimens are available; two of these have adult shell morphology but genital development is incomplete. The other individuals are sub-adults. This species is morphologically similar to S. albida , S. swinhoei sp. nov. and S. insignis , but its genetic relationship is closer to S. insignis (see Phylogeny). Compared with S. albida , S. lini has a larger, thinner and more rigid shell, higher spire and more oblique aperture. The species differs from S. swinhoei in having a periphery that is not keeled, smaller shell diameter and higher spire.