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<document id="61A6BEDE3E3AE39520D8977F64CFA8B1" ID-DOI="10.5281/zenodo.277211" ID-GBIF-Dataset="02822034-b581-4195-a509-5ee435369d0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="277211" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460276828081" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2011" docId="D34C87B84D2F2630FF44F9996292FEF0" docLanguage="en" docName="zt02819p050.pdf" docOrigin="Zootaxa 2819" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pycnoderma Grube 1877" docType="key" docVersion="9" lastPageNumber="31" masterDocId="2F75FFC04D32262EFFD3FFA9670EFFD2" masterDocTitle="Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="30" updateTime="1698248591664" updateUser="plazi">
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<mods:title id="AE4BF77DF7E4AC0FCA3B323A096215FB">Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)</mods:title>
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<treatment id="D34C87B84D2F2630FF44F9996292FEF0" ID-DOI="http://doi.org/10.5281/zenodo.6183607" ID-GBIF-Taxon="119396150" ID-Zenodo-Dep="6183607" LSID="urn:lsid:plazi:treatment:D34C87B84D2F2630FF44F9996292FEF0" httpUri="http://treatment.plazi.org/id/D34C87B84D2F2630FF44F9996292FEF0" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="13FF65254D2F2633FF44F99965A9F998" box="[151,679,1584,1611]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="5B5A36AE4D2F2633FF44F99965A9F998" blockId="29.[151,679,1584,1611]" box="[151,679,1584,1611]" pageId="29" pageNumber="30">
<heading id="001281C24D2F2633FF44F99965A9F998" bold="true" box="[151,679,1584,1611]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="6991EABC4D2F2633FF44F99965A9F998" bold="true" box="[151,679,1584,1611]" pageId="29" pageNumber="30">
Key to species of
<taxonomicName id="9CE54D2D4D2F2633FEBEF99865A9F998" ID-CoL="84T3C" authority="Grube, 1877" authorityName="Grube" authorityYear="1877" box="[365,679,1584,1611]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="genus">
<emphasis id="6991EABC4D2F2633FEBEF998650AF999" bold="true" box="[365,516,1585,1611]" italics="true" pageId="29" pageNumber="30">Pycnoderma</emphasis>
Grube, 1877
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13FF65254D2F2630FF44F9DE6292FEF0" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="key">
<keyStep id="E0112E0B4D2F2633FF44F9DE6292F978" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F9DE6292F95E" blockId="29.[151,1436,1655,2029]" box="[151,1436,1655,1676]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F9DE6292F95E" box="[151,1436,1655,1676]" pageId="29" pageNumber="30">1 Tunic with few or no sediment grains; posterior neurospines entire............................................. 2</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2F2633FF44F93C6292F978" blockId="29.[151,1436,1655,2029]" box="[151,1436,1685,1706]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F93C6292F978" box="[151,1436,1685,1706]" pageId="29" pageNumber="30">- Tunic covered by small sediment grains; posterior neurospines bifid, or bidentate.................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2F2633FF44F91B6292F8D0" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F91B6292F915" blockId="29.[151,1436,1655,2029]" box="[151,1436,1714,1735]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F91B6292F915" box="[151,1436,1714,1735]" pageId="29" pageNumber="30">2(1) Tunic hyaline; chaetiger 1 with neurochaetae close to notochaetae; papillae thin, sparse, body rugose................... 3</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2F2633FF44F9666292F8D0" blockId="29.[151,1436,1655,2029]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F9666292F8D0" pageId="29" pageNumber="30">- Tunic opaque; chaetiger 1 with neurochaetae clearly separated from notochaetae, as a lateral fan; papillae abundant, body pilose or pubescent.................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2F2633FF44F8A36292F8EE" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F8A36292F8CD" blockId="29.[151,1436,1655,2029]" box="[151,1436,1802,1823]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F8A36292F8CD" box="[151,1436,1802,1823]" pageId="29" pageNumber="30">
3(2) Chaetigers 17 long, smooth; with three rows of elongate papillae...................................
<taxonomicName id="9CE54D2D4D2F2633FAD4F8A36251F8CD" ID-CoL="4QMBS" box="[1287,1375,1802,1823]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="dannyi" status="sp. nov.">
<emphasis id="6991EABC4D2F2633FAD4F8A36251F8CD" box="[1287,1375,1802,1823]" italics="true" pageId="29" pageNumber="30">P. dannyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2F2633FAB5F8A36292F8CD" box="[1382,1436,1802,1823]" pageId="29" pageNumber="30" rank="species">
<emphasis id="6991EABC4D2F2633FAB5F8A36292F8CD" bold="true" box="[1382,1436,1802,1823]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2F2633FF44F88E6292F8EE" blockId="29.[151,1436,1655,2029]" box="[151,1436,1831,1853]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F88E6292F8EE" box="[151,1436,1831,1853]" pageId="29" pageNumber="30">
- Chaetigers 17 not markedly longer than following ones; with 68 irregular rows of tiny papillae..
<taxonomicName id="9CE54D2D4D2F2633FB72F8816292F8EE" ID-CoL="4QMBR" authority="Grube, 1877" authorityName="Grube" authorityYear="1877" box="[1185,1436,1831,1853]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="congoense">
<emphasis id="6991EABC4D2F2633FB72F8816213F8EF" box="[1185,1309,1832,1853]" italics="true" pageId="29" pageNumber="30">P. congoense</emphasis>
Grube, 1877
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2F2633FF44F8EC6292F846" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F8EC6298F8A5" blockId="29.[151,1436,1655,2029]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F8EC6298F8A5" pageId="29" pageNumber="30">4(2) Cephalic cage chaetae as long as first 1114 chaetigers; median notochaetae with many long articles, each about 7 times longer than wide........................................................................................... 5</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2F2633FF44F8D66292F846" blockId="29.[151,1436,1655,2029]" box="[151,1436,1919,1940]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F8D66292F846" box="[151,1436,1919,1940]" pageId="29" pageNumber="30">- Cephalic cage chaetae as long as first 78 chaetigers......................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2F2633FF44F8346292F81D" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F8346292F860" blockId="29.[151,1436,1655,2029]" box="[151,1436,1949,1970]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F8346292F860" box="[151,1436,1949,1970]" pageId="29" pageNumber="30">
5(4) Neurochaetae with two articles...........................................
<taxonomicName id="9CE54D2D4D2F2633FBCFF8346231F860" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[1052,1343,1949,1970]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="ferruginea" status="comb. nov.">
<emphasis id="6991EABC4D2F2633FBCFF8346397F860" box="[1052,1177,1949,1970]" italics="true" pageId="29" pageNumber="30">P. ferruginea</emphasis>
(
<bibRefCitation id="3F744B5F4D2F2633FB76F8346239F860" author="Gallardo" box="[1189,1335,1949,1970]" pageId="29" pageNumber="30" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2F2633FA97F8346292F860" box="[1348,1436,1949,1970]" pageId="29" pageNumber="30">
<emphasis id="6991EABC4D2F2633FA97F8346292F860" bold="true" box="[1348,1436,1949,1970]" pageId="29" pageNumber="30">n. comb.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2F2633FF44F8136292F81D" blockId="29.[151,1436,1655,2029]" box="[151,1436,1978,2000]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F8136292F81D" box="[151,1436,1978,2000]" pageId="29" pageNumber="30">
- Neurochaetae with three articles, median one smaller...........................................
<taxonomicName id="9CE54D2D4D2F2633FB35F8126251F802" ID-CoL="4QMBT" box="[1254,1375,1979,2000]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="escobarae" status="sp. nov.">
<emphasis id="6991EABC4D2F2633FB35F8126251F802" box="[1254,1375,1979,2000]" italics="true" pageId="29" pageNumber="30">P. escobarae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2F2633FAB5F8136292F81D" box="[1382,1436,1978,1999]" pageId="29" pageNumber="30" rank="species">
<emphasis id="6991EABC4D2F2633FAB5F8136292F81D" bold="true" box="[1382,1436,1978,1999]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2F2630FF44F87E6292FF7E" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<paragraph id="5B5A36AE4D2F2633FF44F87E6292F83E" blockId="29.[151,1436,1655,2029]" box="[151,1436,2007,2029]" pageId="29" pageNumber="30">
<keyLead id="E014959B4D2F2633FF44F87E6292F83E" box="[151,1436,2007,2029]" pageId="29" pageNumber="30">
6(4) Median notochaetae as long as body width.....................................
<taxonomicName id="9CE54D2D4D2F2633FBE6F8716230F83E" ID-CoL="5b697208-1041-4395-858d-e9408826ac31" authority="Treadwell, 1901" authorityName="Treadwell" authorityYear="1901" box="[1077,1342,2007,2029]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="29" pageNumber="30" phylum="Annelida" rank="species" species="glabra" status="comb. nov.">
<emphasis id="6991EABC4D2F2633FBE6F8716382F83F" box="[1077,1164,2008,2029]" italics="true" pageId="29" pageNumber="30">P. glabra</emphasis>
(
<bibRefCitation id="3F744B5F4D2F2633FB4AF87E6239F83E" author="Treadwell" box="[1177,1335,2007,2028]" pageId="29" pageNumber="30" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fish Commission for 1900, 2, 181 - 210." type="journal article" year="1901">Treadwell, 1901</bibRefCitation>
)
</taxonomicName>
<emphasis id="6991EABC4D2F2633FA97F87E6292F83E" bold="true" box="[1348,1436,2007,2028]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="72A257C74D2F2633FA97F87E6292F83E" box="[1348,1436,2007,2028]" pageId="29" pageNumber="30">n. comb.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2C2630FF44FF3E6292FF7E" blockId="30.[151,1437,151,290]" box="[151,1436,151,173]" pageId="30" pageNumber="31">
<keyLead id="E014959B4D2C2630FF44FF3E6292FF7E" box="[151,1436,151,173]" pageId="30" pageNumber="31">
- Median notochaetae as long as 1/3 body width...............................
<taxonomicName id="9CE54D2D4D2C2630FBC0FF316231FF7E" authority="Fauvel, 1932" authorityName="Fauvel" authorityYear="1932" box="[1043,1343,151,173]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="talehsapensis" status="comb. nov.">
<emphasis id="6991EABC4D2C2630FBC0FF3163A5FF7F" box="[1043,1195,152,173]" italics="true" pageId="30" pageNumber="31">P. talehsapensis</emphasis>
(
<bibRefCitation id="3F744B5F4D2C2630FB6BFF3E6236FF7E" author="Fauvel" box="[1208,1336,151,172]" pageId="30" pageNumber="31" refString="Fauvel, P. (1932) Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum, Calcutta, 12, 1 - 262." type="journal article" year="1932">Fauvel, 1932</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2C2630FA97FF3E6292FF7E" box="[1348,1436,151,172]" pageId="30" pageNumber="31">
<emphasis id="6991EABC4D2C2630FA97FF3E6292FF7E" bold="true" box="[1348,1436,151,172]" pageId="30" pageNumber="31">n. comb.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E0112E0B4D2C2630FF44FF1C6292FEF0" pageId="30" pageNumber="31">
<paragraph id="5B5A36AE4D2C2630FF44FF1C6292FF35" blockId="30.[151,1437,151,290]" pageId="30" pageNumber="31">
<keyLead id="E014959B4D2C2630FF44FF1C6292FF35" pageId="30" pageNumber="31">
7(1) First few chaetigers without dorsal lobes; sediment grains sparse, leaving wide tunic areas without sediment grains; first neurohooks from chaetiger 7; posterior neurospines bifid...............................................
<taxonomicName id="9CE54D2D4D2C2630FAD0FF7B626EFF35" ID-CoL="4QMBV" box="[1283,1376,210,231]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="glasbyi" status="sp. nov.">
<emphasis id="6991EABC4D2C2630FAD0FF7B626EFF35" box="[1283,1376,210,231]" italics="true" pageId="30" pageNumber="31">P. glasbyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2C2630FAB5FF7B6292FF35" box="[1382,1436,210,231]" pageId="30" pageNumber="31" rank="species">
<emphasis id="6991EABC4D2C2630FAB5FF7B6292FF35" bold="true" box="[1382,1436,210,231]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="5B5A36AE4D2C2630FF44FF466292FEF0" blockId="30.[151,1437,151,290]" pageId="30" pageNumber="31">
<keyLead id="E014959B4D2C2630FF44FF466292FEF0" pageId="30" pageNumber="31">
- First few chaetigers with large dorsal lobes; sediment grains densely packed, completely covering the tunic; posterior neurospines bidentate..........................................................
<taxonomicName id="9CE54D2D4D2C2630FBEBFEA46230FEF0" authority="Hartman, 1961" authorityName="Hartman" authorityYear="1961" box="[1080,1342,269,290]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="31" phylum="Annelida" rank="species" species="gracilis" status="comb. nov.">
<emphasis id="6991EABC4D2C2630FBEBFEA46396FEF0" box="[1080,1176,269,290]" italics="true" pageId="30" pageNumber="31">P. gracilis</emphasis>
(
<bibRefCitation id="3F744B5F4D2C2630FB76FEA46239FEF0" author="Hartman" box="[1189,1335,269,290]" pageId="30" pageNumber="31" refString="Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 25, 1 - 226." type="journal article" year="1961">Hartman, 1961</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72A257C74D2C2630FA97FEA46292FEF0" box="[1348,1436,269,290]" pageId="30" pageNumber="31">
<emphasis id="6991EABC4D2C2630FA97FEA46292FEF0" bold="true" box="[1348,1436,269,290]" pageId="30" pageNumber="31">n. comb.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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