311 lines
50 KiB
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311 lines
50 KiB
XML
<document id="A9AA24C2E48B8324F6AF0761CECD1605" ID-DOI="10.5281/zenodo.277211" ID-GBIF-Dataset="02822034-b581-4195-a509-5ee435369d0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="277211" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460276828081" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2011" docId="D34C87B84D11260BFF44F99D63A5FB85" docLanguage="en" docName="zt02819p050.pdf" docOrigin="Zootaxa 2819" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pycnoderma ferruginea Gallardo 1968, n. comb." docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="2F75FFC04D32262EFFD3FFA9670EFFD2" masterDocTitle="Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="36" updateTime="1698248591664" updateUser="plazi">
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<mods:title id="9542991E3728696512E2BA4EC529376E">Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)</mods:title>
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<mods:namePart id="7E340EE90A307E930483D94FB42C561E">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="358C6B08F92B285FA095B4A81C3AF384">Zootaxa</mods:title>
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<mods:date id="0A350F56083DA0E8C582636BA0381D1F">2011</mods:date>
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<mods:identifier id="C39A3401363F797B3ED8C45AE22CCA68" type="DOI">10.5281/zenodo.277211</mods:identifier>
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<treatment id="D34C87B84D11260BFF44F99D63A5FB85" ID-DOI="http://doi.org/10.5281/zenodo.6183615" ID-GBIF-Taxon="119396166" ID-Zenodo-Dep="6183615" LSID="urn:lsid:plazi:treatment:D34C87B84D11260BFF44F99D63A5FB85" httpUri="http://treatment.plazi.org/id/D34C87B84D11260BFF44F99D63A5FB85" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
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<subSubSection id="13FF65254D11260DFF44F99D660DF9BD" pageId="35" pageNumber="36" type="nomenclature">
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<paragraph id="5B5A36AE4D11260DFF44F99D65F2F99C" blockId="35.[151,764,1588,1647]" box="[151,764,1588,1614]" pageId="35" pageNumber="36">
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<heading id="001281C24D11260DFF44F99D65F2F99C" bold="true" box="[151,764,1588,1614]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
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<emphasis id="6991EABC4D11260DFF44F99D65F2F99C" bold="true" box="[151,764,1588,1614]" pageId="35" pageNumber="36">
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<taxonomicName id="9CE54D2D4D11260DFF44F99D6584F99C" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[151,650,1588,1614]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="ferruginea" status="comb. nov.">
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<emphasis id="6991EABC4D11260DFF44F99D66B9F99C" bold="true" box="[151,439,1588,1614]" italics="true" pageId="35" pageNumber="36">Pycnoderma ferruginea</emphasis>
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(
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<bibRefCitation id="3F744B5F4D11260DFE16F99D658CF99C" author="Gallardo" box="[453,642,1588,1614]" pageId="35" pageNumber="36" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="72A257C74D11260DFD41F99D65F2F99C" box="[658,764,1588,1614]" pageId="35" pageNumber="36">n. comb.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5B5A36AE4D11260DFF44F9FE660DF9BD" blockId="35.[151,764,1588,1647]" box="[151,259,1623,1647]" pageId="35" pageNumber="36">
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<figureCitation id="C3DE2A2B4D11260DFF44F9FE660DF9BD" box="[151,259,1623,1647]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="35" pageNumber="36">Figure 16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="13FF65254D11260DFF44F93664DBF906" pageId="35" pageNumber="36" type="reference_group">
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<paragraph id="5B5A36AE4D11260DFF44F93665F0F967" blockId="35.[151,981,1695,1748]" box="[151,766,1695,1717]" pageId="35" pageNumber="36">
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<treatmentCitationGroup id="7BF511804D11260DFF44F93665F0F967" box="[151,766,1695,1717]" pageId="35" pageNumber="36">
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<taxonomicName id="9CE54D2D4D11260DFF44F936651CF967" ID-CoL="N4JL" authority="Gallardo, 1968:107" authorityName="Gallardo" authorityPageNumber="107" authorityYear="1968" box="[151,530,1695,1717]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="ferruginea">
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<emphasis id="6991EABC4D11260DFF44F9366649F967" box="[151,327,1695,1717]" italics="true" pageId="35" pageNumber="36">Brada ferruginea</emphasis>
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<treatmentCitation id="DA4410BF4D11260DFE9EF936651CF967" author="Gallardo" box="[333,530,1695,1717]" page="107" pageId="35" pageNumber="36" year="1968">
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<bibRefCitation id="3F744B5F4D11260DFE9EF93666E6F967" author="Gallardo" box="[333,488,1695,1717]" pageId="35" pageNumber="36" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
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:107
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</treatmentCitation>
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</taxonomicName>
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–108, Pl. 49,
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<figureCitation id="C3DE2A2B4D11260DFD4AF93665F4F967" box="[665,762,1695,1717]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="16.[151,250,1582,1605]" captionStartId-1="18.[151,250,1245,1268]" captionTargetBox-0="[173,1411,248,1569]" captionTargetBox-1="[169,1411,301,1219]" captionTargetId-0="figure@16.[169,1417,237,1577]" captionTargetId-1="figure@18.[169,1417,301,1224]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionText-0="FIGURE 5. Piromis eruca (Claparède, 1868). A. Neotype (MNHN-A 507), complete in lateral view. B. Same, anterior end, lateral view. C. Same, anterior end, dorsal view. D. Paraneotype, head, palps and branchiae detached (BS: branchial scars, DL: dorsal lip, LL: lateral lip). E. Non-type specimen (MNHN-A 185), exposed anterior end, palps and branchiae detached (BS: branchial scars, Ca: caruncle, Pr: prostomium, PS: palp scar). F. Paraneotype, notochaetal regions, basal, median, distal and an enlargement of the tip. G. Same, neurochaetae from a median chaetiger, insert showing one tip. H. Same, neurochaetae from a posterior chaetiger, insert showing one tip. Scale bars: A: 3 mm, B: 1 mm, C: 1 mm, D – E: 0.5 mm, F: 50 µm, G: 200 µm, H: 25 µm." captionText-1="FIGURE 6. Piromis fauchaldi n. sp. A. Holotype (LACM- 2511), complete. B. Same, anterior end, dorsal view. C. Paratype (LACM-AHF 1733), anterior end, ventral view showing some epizoic hydroids. D. Holotype, head, palps and branchiae removed (BS: branchial scars, Ca: caruncle, Pr: prostomium, PS: palp scar). E. Paratype (LACM-AHF- 2512), notochaetae from an anterior chaetiger showing basal, median and distal regions. F. Same, neurochaetae from a median chaetiger, insert showing chaetal tips. G. Same, neurochaetae from a posterior chaetiger, insert showing chaetal tips. Scale bars: A: 2 mm, B – C: 1 mm, D: 250 µm, E: 50 µm, F: 75 µm, G: 100 µm." httpUri-0="https://zenodo.org/record/277216/files/figure.png" httpUri-1="https://zenodo.org/record/277217/files/figure.png" pageId="35" pageNumber="36">Figs. 5–6</figureCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="5B5A36AE4D11260DFF44F91664DBF906" blockId="35.[151,981,1695,1748]" box="[151,981,1725,1748]" pageId="35" pageNumber="36">
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<treatmentCitationGroup id="7BF511804D11260DFF44F91664DBF906" box="[151,981,1725,1748]" pageId="35" pageNumber="36">
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<taxonomicName id="9CE54D2D4D11260DFF44F9166529F901" authority="Gallardo, 1968:109" authorityName="Gallardo" authorityPageNumber="109" authorityYear="1968" box="[151,551,1725,1748]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="congoensis">
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<emphasis id="6991EABC4D11260DFF44F9166652F906" box="[151,348,1727,1748]" italics="true" pageId="35" pageNumber="36">Piromis congoensis</emphasis>
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<treatmentCitation id="DA4410BF4D11260DFEB0F9146529F901" author="Gallardo" box="[355,551,1725,1747]" page="109" pageId="35" pageNumber="36" year="1968">
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<bibRefCitation id="3F744B5F4D11260DFEB0F91466F2F901" author="Gallardo" box="[355,508,1725,1747]" pageId="35" pageNumber="36" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
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:109
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</treatmentCitation>
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||
</taxonomicName>
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–110, Pl. 51,
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<figureCitation id="C3DE2A2B4D11260DFD7DF9146405F901" box="[686,779,1725,1747]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="8.[151,250,1839,1862]" captionStartId-1="10.[151,250,1857,1880]" captionStartId-2="12.[151,250,1737,1760]" captionStartId-3="14.[151,250,1034,1057]" captionStartId-4="16.[151,250,1582,1605]" captionStartId-5="18.[151,250,1245,1268]" captionStartId-6="20.[151,250,1329,1352]" captionTargetBox-0="[178,1414,841,1814]" captionTargetBox-1="[169,1414,329,1828]" captionTargetBox-2="[173,1416,913,1714]" captionTargetBox-3="[169,1409,182,1008]" captionTargetBox-4="[173,1411,248,1569]" captionTargetBox-5="[169,1411,301,1219]" captionTargetBox-6="[172,1408,481,1304]" captionTargetId-0="figure@8.[169,1417,841,1818]" captionTargetId-1="figure@10.[169,1417,329,1836]" captionTargetId-2="figure@12.[169,1417,913,1716]" captionTargetId-3="figure@14.[169,1417,182,1013]" captionTargetId-4="figure@16.[169,1417,237,1577]" captionTargetId-5="figure@18.[169,1417,301,1224]" captionTargetId-6="figure@20.[169,1417,481,1309]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="12" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="18" captionTargetPageId-6="20" captionText-0="FIGURE 1. Piromis arenosus Kinberg, 1867. A. Syntypes in lateral view. B. Thicker syntype, anterior end, dorsal view. C. Same, posterior end, lateral view. D. Non-type specimen (LAMNH- 5210), anterior end dissected, slightly collapsed, palps and most branchiae removed (BS: branchial scars, Ca: caruncle, DL: dorsal lip, LL: lateral lip, Pr: prostomium, PS: palp scar). E. Thicker syntype, notochaetae portions, basal, medial and distal with an insert of the tip. F. Non-type specimen (LAMNH-AHF- 2506), neurochaetae from a median chaetiger with an insert of one tip. G. Same, neurochaetae from a posterior chaetiger with an insert of a tip. Scale bars: A: 6 mm, B: 1.5 mm, C – D: 1 mm, E: 25 µm, F: 90 µm, G: 40 µm." captionText-1="FIGURE 2. Piromis amoureuxi n. sp. A. Holotype (MNHN- 900), twisted. B. Same, anterior end in dorsal view. C. Same, median segments in lateral view, showing sediment particles dorsally and laterally. D. Paratype (MNHN- 900 a), anterior end, palps and most branchiae detached (BD: branchial scars, LL: lateral lip, PS: palp scar). E. Holotype, anterior notopodium. F. Same, close-up of notochaetal regions, left, median and distal. G. Same, neurochaetae from a median chaetiger. H. Same, neurochaetae from a posterior chaetiger. Scale bars: A: 1 mm, B – C: 0.5 mm, D: 0.3 mm, E: 100 µm, F: 10 µm, G: 50 µm, H: 25 µm." captionText-2="FIGURE 3. Piromis brisegnoi n. sp. A. Holotype (LAMNH-AHF- 2509), lateral view. B. Same, anterior end, lateral view showing abundant chaetae and long papillae. C. Non-type specimen, anterior end, frontal view, palps and branchiae removed (BS: branchial scars, LL: lateral lip, PS: palp scar, VV: ventral lip). D. Holotype, notochaetae from an anterior chaetiger. E. Same, neurochaetae from a median chaetiger, insert showing the tips. F. Same, neurochaetae from a posterior chaetiger, insert showing the tip. Scale bars: A: 5 mm, B: 1.5 mm, C: 0.5 mm, D: 150 µm, E: 60 µm, F: 50 µm." captionText-3="FIGURE 4. Piromis capulata (Moore, 1909) n. comb. A. Holotype (CAS- 19721), complete in lateral view. B. Same, anterior end, lateral view. C. Same, anterior end, dorsal view, notice the eroded areas. D. Non-type specimen (USNM- 40484), anterior end, dorsal view. E. Same, anterior end, palps and most branchiae detached (BS: branchial scars, LL: lateral lip). F. Same, right neuropodium showing the gonopodial lobe with an asterisk in the base. G. Holotype, notochaetal regions, basal, median, distal. H. Same, median neuropodia with an insert showing the tips. Scale bars: A: 5 mm, B: 2.3 mm, C: 1.5 mm, D – E: 2 mm, F: 325 µm, G: 100 µm, H: 50 µm." captionText-4="FIGURE 5. Piromis eruca (Claparède, 1868). A. Neotype (MNHN-A 507), complete in lateral view. B. Same, anterior end, lateral view. C. Same, anterior end, dorsal view. D. Paraneotype, head, palps and branchiae detached (BS: branchial scars, DL: dorsal lip, LL: lateral lip). E. Non-type specimen (MNHN-A 185), exposed anterior end, palps and branchiae detached (BS: branchial scars, Ca: caruncle, Pr: prostomium, PS: palp scar). F. Paraneotype, notochaetal regions, basal, median, distal and an enlargement of the tip. G. Same, neurochaetae from a median chaetiger, insert showing one tip. H. Same, neurochaetae from a posterior chaetiger, insert showing one tip. Scale bars: A: 3 mm, B: 1 mm, C: 1 mm, D – E: 0.5 mm, F: 50 µm, G: 200 µm, H: 25 µm." captionText-5="FIGURE 6. Piromis fauchaldi n. sp. A. Holotype (LACM- 2511), complete. B. Same, anterior end, dorsal view. C. Paratype (LACM-AHF 1733), anterior end, ventral view showing some epizoic hydroids. D. Holotype, head, palps and branchiae removed (BS: branchial scars, Ca: caruncle, Pr: prostomium, PS: palp scar). E. Paratype (LACM-AHF- 2512), notochaetae from an anterior chaetiger showing basal, median and distal regions. F. Same, neurochaetae from a median chaetiger, insert showing chaetal tips. G. Same, neurochaetae from a posterior chaetiger, insert showing chaetal tips. Scale bars: A: 2 mm, B – C: 1 mm, D: 250 µm, E: 50 µm, F: 75 µm, G: 100 µm." captionText-6="FIGURE 7. Piromis kisemboanus (Augener, 1918) n. comb., non-type specimens. A. Complete (MNHN- 507), lateral view. B. Same, anterior end, lateral view. C. Same, anterior end, dorsal view. D. Same, medium parapodium. E. Another specimen (USNM-unnumb.), notochaetae from an anterior notopodium showing articulation in basal, medial and distal regions. F. Another specimen (MNHN- 507), anterior neurochaetae, insert showing chaetal tips. G. Another specimen (USNM-unnumb.), posterior neurochaetae, insert showing chaetal tips. Scale bars: A: 2 mm, B – C: 0.5 mm, D: 0.25 mm, E: 66 µm, F: 50 µm, G: 100 µm." httpUri-0="https://zenodo.org/record/277212/files/figure.png" httpUri-1="https://zenodo.org/record/277213/files/figure.png" httpUri-2="https://zenodo.org/record/277214/files/figure.png" httpUri-3="https://zenodo.org/record/277215/files/figure.png" httpUri-4="https://zenodo.org/record/277216/files/figure.png" httpUri-5="https://zenodo.org/record/277217/files/figure.png" httpUri-6="https://zenodo.org/record/277218/files/figure.png" pageId="35" pageNumber="36">Figs. 1–7</figureCitation>
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||
(
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||
<emphasis id="6991EABC4D11260DFCC9F916644EF906" box="[794,832,1727,1748]" italics="true" pageId="35" pageNumber="36">non</emphasis>
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||
Grube, 1877).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="13FF65254D11260AFF44F8A9643AFF06" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="materials_examined">
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||
<paragraph id="5B5A36AE4D11260DFF44F8A96406F8B2" blockId="35.[151,1436,1791,2033]" pageId="35" pageNumber="36">
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||
<emphasis id="6991EABC4D11260DFF44F8A96572F8CA" bold="true" box="[151,636,1791,1816]" pageId="35" pageNumber="36">
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||
<typeStatus id="845E880C4D11260DFF44F8A967DCF8CA" box="[151,210,1792,1816]" pageId="35" pageNumber="36">Type</typeStatus>
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material. Western Tropical Pacific.
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</emphasis>
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<typeStatus id="845E880C4D11260DFD50F8A965E5F8CA" box="[643,747,1792,1816]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
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of
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<taxonomicName id="9CE54D2D4D11260DFCC1F8A96388F8CA" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[786,1158,1792,1816]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="ferruginea">
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<emphasis id="6991EABC4D11260DFCC1F8A964DBF8CA" box="[786,981,1792,1816]" italics="true" pageId="35" pageNumber="36">Brada ferruginea</emphasis>
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<bibRefCitation id="3F744B5F4D11260DFC0EF8A96388F8CA" author="Gallardo" box="[989,1158,1792,1816]" pageId="35" pageNumber="36" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
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</taxonomicName>
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(LACM-AHF -312), off West side of Hon Lon Island, Bay of Nha Trang,
|
||
<collectingCountry id="23F2763E4D11260DFD6AF88D6428F8EE" box="[697,806,1828,1852]" name="Vietnam" pageId="35" pageNumber="36">Viet Nam</collectingCountry>
|
||
, Naga Expediton, R/V
|
||
<emphasis id="6991EABC4D11260DFBFAF88C639DF8EE" box="[1065,1171,1829,1852]" italics="true" pageId="35" pageNumber="36">Mao Tien</emphasis>
|
||
, Sta. 117 (12°11ʹ21ʺ N, 109°16ʹ15ʺ E),
|
||
<quantity id="9C1D9B4B4D11260DFE96F8E1668EF8B2" box="[325,384,1864,1888]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="m" value="12.0">12 m</quantity>
|
||
,
|
||
<date id="2F5B106E4D11260DFE5FF8E1652CF8B2" box="[396,546,1864,1888]" pageId="35" pageNumber="36" value="1960-02-10">10 Feb. 1960</date>
|
||
, V.A. Gallardo, coll.
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D11260AFF16F8C5643AFF06" blockId="35.[151,1436,1791,2033]" lastBlockId="36.[151,1437,151,392]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
<emphasis id="6991EABC4D11260DFF16F8C5640CF857" bold="true" box="[197,770,1900,1925]" pageId="35" pageNumber="36">Additional material. Western Tropical Pacific.</emphasis>
|
||
One mature female (LACM-AHF-2515), broken in two pieces,
|
||
<quantity id="9C1D9B4B4D11260DFF38F839662AF87A" box="[235,292,1936,1960]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="35" pageNumber="36" unit="km" value="4.0">4 km</quantity>
|
||
N Hon Ho island,
|
||
<collectingCountry id="23F2763E4D11260DFE2BF8396569F87A" box="[504,615,1936,1960]" name="Vietnam" pageId="35" pageNumber="36">Viet Nam</collectingCountry>
|
||
, Naga Expediton, R/V
|
||
<emphasis id="6991EABC4D11260DFCBFF83864D8F87A" box="[876,982,1937,1960]" italics="true" pageId="35" pageNumber="36">Mao Tien</emphasis>
|
||
, Sta. 153I (12°19ʹ20ʺ N, 109°21ʹ40ʺ E),
|
||
<quantity id="9C1D9B4B4D11260DFF44F81D67DCF81E" box="[151,210,1972,1996]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="35" pageNumber="36" unit="m" value="43.0">43 m</quantity>
|
||
, mud,
|
||
<date id="2F5B106E4D11260DFECDF81D66BCF81E" box="[286,434,1972,1996]" pageId="35" pageNumber="36" value="1960-02-24">24 Feb. 1960</date>
|
||
(cephalic cage chaetae length equivalent to the anterior end up to chaetiger 18). Anterior fragment (LACM-AHF-2516),
|
||
<quantity id="9C1D9B4B4D11260DFE25F8716520F822" box="[502,558,2008,2032]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="2.0">2 km</quantity>
|
||
NW Hon Ho
|
||
<collectingCountry id="23F2763E4D11260DFD1AF871641EF822" box="[713,784,2008,2032]" name="Iceland" pageId="35" pageNumber="36">Island</collectingCountry>
|
||
, Bay of Nha Trang,
|
||
<collectingCountry id="23F2763E4D11260DFC22F8716351F822" box="[1009,1119,2008,2032]" name="Vietnam" pageId="35" pageNumber="36">Viet Nam</collectingCountry>
|
||
, Naga Expedition, R/V
|
||
<emphasis id="6991EABC4D11260AFABAF87067C8FF62" italics="true" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">Mao Tien</emphasis>
|
||
, Sta. 15411 (12°18ʹ0 6ʺ N, 109°21ʹ27ʺ E),
|
||
<quantity id="9C1D9B4B4D16260AFD7BFF3E65EDFF7D" box="[680,739,151,175]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="36" pageNumber="37" unit="m" value="43.0">43 m</quantity>
|
||
, mud,
|
||
<date id="2F5B106E4D16260AFCE3FF3E64C8FF7D" box="[816,966,151,175]" pageId="36" pageNumber="37" value="1960-02-24">24 Feb. 1960</date>
|
||
, V.A. Gallardo, coll. (cephalic cage chaetae length equivalent to the anterior end up to chaetiger 14).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FF65254D16260BFF16FF7666DEFECE" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="5B5A36AE4D16260AFF16FF76620AFE55" blockId="36.[151,1437,151,392]" pageId="36" pageNumber="37">
|
||
<emphasis id="6991EABC4D16260AFF16FF766659FF2A" bold="true" box="[197,343,223,248]" pageId="36" pageNumber="37">Description.</emphasis>
|
||
<typeStatus id="845E880C4D16260AFE8EFF7666CBFF25" box="[349,453,223,247]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
(LACM-AHF-312) broken into three parts, anterior fragment reddish, especially anteriorly, with most chaetae directed forward, median and posterior fragments pale (
|
||
<figureCitation id="C3DE2A2B4D16260AFBE0FEAD6388FECE" box="[1075,1158,260,284]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
A), all cylindrical; tunic thin, densely papillated, fine red sediment particles anteriorly; papillae mostly small, thin, cirriform, abundant all over the body, and few longer, capitate, isolated, along body in 2–3 regular rows, and others close to parapodia.
|
||
<typeStatus id="845E880C4D16260AFF44FEC667F1FE55" box="[151,255,367,391]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
46(15+4+27) mm long,
|
||
<quantity id="9C1D9B4B4D16260AFDC7FEC66564FE55" box="[532,618,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="37" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide, cephalic cage
|
||
<quantity id="9C1D9B4B4D16260AFC87FEC664A7FE55" box="[852,937,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="36" pageNumber="37" unit="mm" value="5.5">5.5 mm</quantity>
|
||
long, 65(28+7+30) chaetigers.
|
||
</paragraph>
|
||
<caption id="0F9A66264D16260AFF44FB246304FACD" httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37" targetBox="[175,1413,448,1138]" targetPageId="36">
|
||
<paragraph id="5B5A36AE4D16260AFF44FB246304FACD" blockId="36.[151,1436,1165,1311]" pageId="36" pageNumber="37">
|
||
<emphasis id="6991EABC4D16260AFF44FB246617FB71" bold="true" box="[151,281,1165,1188]" pageId="36" pageNumber="37">FIGURE 16</emphasis>
|
||
.
|
||
<taxonomicName id="9CE54D2D4D16260AFEF4FB2765C6FB76" authority="(Gallardo, 1968)" authorityName="Gallardo" authorityYear="1968" baseAuthorityName="Gallardo" baseAuthorityYear="1968" box="[295,712,1166,1188]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="ferruginea" status="comb. nov.">
|
||
<emphasis id="6991EABC4D16260AFEF4FB276517FB76" box="[295,537,1166,1188]" italics="true" pageId="36" pageNumber="37">Pycnoderma ferruginea</emphasis>
|
||
(Gallardo, 1968)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72A257C74D16260AFD03FB276429FB71" box="[720,807,1166,1187]" pageId="36" pageNumber="37">
|
||
<emphasis id="6991EABC4D16260AFD03FB276429FB71" bold="true" box="[720,807,1166,1187]" pageId="36" pageNumber="37">n. comb.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A. Holotype (LACM-AHF-312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF-2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B5A36AE4D16260AFF16FAEF6367F9E4" blockId="36.[151,1436,1350,2023]" pageId="36" pageNumber="37">
|
||
Cephalic hood partially exposed in
|
||
<typeStatus id="845E880C4D16260AFD87FAEF65B7FA8C" box="[596,697,1350,1374]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
, short, margin papillated. Anterior end observed in another specimen (LACM-AHF-11153); cephalic hood very short, margin papillated. Prostomium depressed, eyes not visible; caruncle short, not reaching posterior margin of branchial plate (
|
||
<figureCitation id="C3DE2A2B4D16260AFCBEFA2764B1FA74" box="[877,959,1422,1446]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
E), lateral ridges short, prominent, extending towards lateral bases of branchiae. Palps lost; palp keels rounded. Lateral lips wide, well-developed; dorsal and ventral lips reduced. Branchiae mostly lost, cirriform, pale (LACM-AHF-11154), sessile on branchial plate, widely separated medially into two groups, over a tongue-like protuberance, branchial groups posteriorly coalescent, with about 100 filaments per side. Size relationships between palps and branchiae unknown.
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D16260AFF16F9EB6477F884" blockId="36.[151,1436,1350,2023]" pageId="36" pageNumber="37">
|
||
Cephalic cage chaetae more than twice as long as body width (
|
||
<figureCitation id="C3DE2A2B4D16260AFC53F9EB64DCF988" box="[896,978,1602,1626]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
B). Only chaetiger 1 involved in cephalic cage, notochaetae of chaetiger 2 less than twice as long as following ones. Cephalic cage notochaetae arranged in a short dorsal row, neurochaetae as a long lateral row (
|
||
<figureCitation id="C3DE2A2B4D16260AFD31F923643DF970" box="[738,819,1674,1698]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
D); 8–10 notochaetae and 12–15 neurochaetae per bundle, longest about 1/4 as long as notochaetae. Anterior dorsal margin of chaetiger 1 short, papillated (
|
||
<figureCitation id="C3DE2A2B4D16260AFAF4F9076275F914" box="[1319,1403,1710,1734]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
C). Anterior chaetigers without long papillae. Chaetigers 1–3 decreasing in size anterior to posterior, second parapodia displaced dorsally. Post-cephalic cage chaetigers not elongated; a white rounded dorsal spot present on posterior margin of chaetiger 1. Chaetal transition from cephalic cage to body chaetae gradual; compound neurohooks from chaetiger 16. Gonopodial lobes in chaetiger 5, low, pale, rounded.
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D16260AFF16F8CB6204F84C" blockId="36.[151,1436,1350,2023]" pageId="36" pageNumber="37">Parapodia poorly developed, lateral. Median neuropodia lateral. Notopodia low, thin lobes, with 3–4 long capitate papillae. Neuropodia slightly longer lobes, with some superior and inferior long capitate papillae.</paragraph>
|
||
<paragraph id="5B5A36AE4D16260BFF16F80365D3FF25" blockId="36.[151,1436,1350,2023]" lastBlockId="37.[151,1436,151,1113]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||
Median notochaetae in short transverse row, 6–8 per bundle, 3/5 as long as body width, posterior notochaetae twice as long as body width; notochaetae all multiarticulated capillaries with very long articles (
|
||
<figureCitation id="C3DE2A2B4D16260AFB1BF8676217F834" box="[1224,1305,1998,2022]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="36" pageNumber="37">Fig. 16</figureCitation>
|
||
F), continuing to tip. Neurochaetae similar to notochaetae in anterior chaetigers, arranged in transverse row. Neurohooks from chaetiger 16, 6–8 per bundle, each with feebly marked, anchylosed short articles basally, followed by 2–5 long articles, with long, lanceolate, aristate blade (
|
||
<figureCitation id="C3DE2A2B4D17260BFDBFFF7665B0FF25" box="[620,702,223,247]" captionStart="FIGURE 16" captionStartId="36.[151,250,1165,1188]" captionTargetBox="[175,1413,448,1138]" captionTargetId="figure@36.[169,1417,445,1145]" captionTargetPageId="36" captionText="FIGURE 16. Pycnoderma ferruginea (Gallardo, 1968) n. comb. A. Holotype (LACM-AHF- 312), dorsal view. B. Same, anterior end, dorsal view. C. Same, close-up to show dorsally displaced chaetiger 1. D. Same, anterior end, lateral view. E. Nontype specimen (LACM-AHF- 2516), anterior end exposed, eroded, branchiae and palps lost (Ca: caruncle, BS: branchial scars, DL: dorsal lip, LL: lateral lip, PS: palp scar, VL: ventral lip). F. Holotype, parapodium from chaetiger 22 (insert: neurochaetal tips). Scale bars: A: 2.25 mm, B: 1.1 mm, C: 1 mm, D: 0.5 mm, E: 0.2 mm, F: 250 µm." httpUri="https://zenodo.org/record/277227/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D17260BFF16FEAD66DEFECE" blockId="37.[151,1436,151,1113]" box="[197,464,260,284]" pageId="37" pageNumber="38">Posterior end unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FF65254D17260BFF16FE8167E6FBE6" pageId="37" pageNumber="38" type="discussion">
|
||
<paragraph id="5B5A36AE4D17260BFF16FE8165CCFE7E" blockId="37.[151,1436,151,1113]" pageId="37" pageNumber="38">
|
||
<emphasis id="6991EABC4D17260BFF16FE816637FE92" bold="true" box="[197,313,296,320]" pageId="37" pageNumber="38">Remarks.</emphasis>
|
||
<taxonomicName id="9CE54D2D4D17260BFE91FE81641AFEED" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[322,788,295,320]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea" status="comb. nov.">
|
||
<emphasis id="6991EABC4D17260BFE91FE816540FE92" box="[322,590,296,320]" italics="true" pageId="37" pageNumber="38">Pycnoderma ferruginea</emphasis>
|
||
(
|
||
<bibRefCitation id="3F744B5F4D17260BFD8CFE8E6402FEED" author="Gallardo" box="[607,780,295,319]" pageId="37" pageNumber="38" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72A257C74D17260BFCCEFE81648BFE92" box="[797,901,296,320]" pageId="37" pageNumber="38">
|
||
<emphasis id="6991EABC4D17260BFCCEFE81648BFE92" bold="true" box="[797,901,296,320]" pageId="37" pageNumber="38">n. comb.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was placed in the genus
|
||
<taxonomicName id="9CE54D2D4D17260BFB7AFE8163E1FE92" box="[1193,1263,296,320]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">
|
||
<emphasis id="6991EABC4D17260BFB7AFE8163E1FE92" box="[1193,1263,296,320]" italics="true" pageId="37" pageNumber="38">Brada</emphasis>
|
||
</taxonomicName>
|
||
because of the presence of ventral lobes in chaetiger 5. However, the cephalic cage, the body shape, and the
|
||
<typeStatus id="845E880C4D17260BFB61FEE463ECFEB7" box="[1202,1250,333,357]" pageId="37" pageNumber="38">type</typeStatus>
|
||
of neurochaetae are very different from those of
|
||
<emphasis id="6991EABC4D17260BFDD9FED96558FE5A" box="[522,598,368,392]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="9CE54D2D4D17260BFDD9FED9655DFE5A" box="[522,595,368,392]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">Brada</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and this species is therefore here referred to
|
||
<emphasis id="6991EABC4D17260BFBB3FED963FBFE5A" box="[1120,1269,368,392]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="9CE54D2D4D17260BFBB3FED963FEFE5A" box="[1120,1264,368,392]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">Pycnoderma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and hence the new combination:
|
||
<taxonomicName id="9CE54D2D4D17260BFEB4FE3C65B3FE7E" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[359,701,404,428]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">
|
||
<emphasis id="6991EABC4D17260BFEB4FE3C66F7FE7E" box="[359,505,404,428]" italics="true" pageId="37" pageNumber="38">P. ferruginea</emphasis>
|
||
(
|
||
<bibRefCitation id="3F744B5F4D17260BFDD4FE3D65BBFE7E" author="Gallardo" box="[519,693,404,428]" pageId="37" pageNumber="38" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D17260BFF16FE1E620CFD1E" blockId="37.[151,1436,151,1113]" pageId="37" pageNumber="38">
|
||
The description of
|
||
<taxonomicName id="9CE54D2D4D17260BFE48FE10655CFE02" box="[411,594,440,464]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="talehsapensis">
|
||
<emphasis id="6991EABC4D17260BFE48FE10655CFE02" box="[411,594,440,464]" italics="true" pageId="37" pageNumber="38">B. talehsapensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="3F744B5F4D17260BFDAFFE1E6415FE1D" author="Fauvel" box="[636,795,439,463]" pageId="37" pageNumber="38" refString="Fauvel, P. (1932) Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum, Calcutta, 12, 1 - 262." type="journal article" year="1932">Fauvel (1932)</bibRefCitation>
|
||
and its redescription (
|
||
<bibRefCitation id="3F744B5F4D17260BFBC5FE1E63ABFE1D" author="Fauvel" box="[1046,1189,439,463]" pageId="37" pageNumber="38" refString="Fauvel, P. (1953) The Fauna of India including Pakistan, Ceylon, Burma and Malaya. Annelida Polychaeta. Allahabad: Indian Press, 12 + 507 pp." type="book" year="1953">Fauvel 1953</bibRefCitation>
|
||
) were not overlooked by
|
||
<bibRefCitation id="3F744B5F4D17260BFF6FFE75667BFE26" author="Gallardo" box="[188,373,476,500]" pageId="37" pageNumber="38" refString="Gallardo, V. A. (1968 [1967]) Polychaeta from the Bay of Nha Trang, South Viet Nam. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 1959 - 1961. Scripps Institution of Oceanography NAGA Report, 4, 35 - 279." type="journal article" year="1968">Gallardo (1968)</bibRefCitation>
|
||
, but he confused the species when comparing his specimens to Fauvel’s species, which is a close relative. The two species differ in the relative size of the cephalic cage and in the pigmentation pattern of the branchial filaments. In
|
||
<emphasis id="6991EABC4D17260BFE4EFD8C6534FDEE" box="[413,570,548,572]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="9CE54D2D4D17260BFE4EFD8C6538FDEE" box="[413,566,548,572]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">B. ferruginea</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
chaetal length of the
|
||
<typeStatus id="845E880C4D17260BFCE3FD8D649CFDEE" box="[816,914,548,572]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
is about the same over the first 13 chaetigers; this relative size of cephalic cage chaetae is confirmed in two other specimens being
|
||
<quantity id="9C1D9B4B4D17260BFB9CFDEE639EFD8D" box="[1103,1168,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="38" unit="mm" value="1.0">1 mm</quantity>
|
||
wide, which have these chaetae being as long as reaching chaetigers 12 or 17. In contrast, in
|
||
<taxonomicName id="9CE54D2D4D17260BFC49FDC4635EFD56" box="[922,1104,620,644]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="talehsapensis">
|
||
<emphasis id="6991EABC4D17260BFC49FDC4635EFD56" box="[922,1104,620,644]" italics="true" pageId="37" pageNumber="38">B. talehsapensis</emphasis>
|
||
</taxonomicName>
|
||
the cephalic cage chaetae are shorter, with their length being about the same as the length of the anterior end up to chaetiger 7. Further, the branchial filaments are homogeneously pale in
|
||
<taxonomicName id="9CE54D2D4D17260BFDA4FD1C6407FD1E" box="[631,777,692,716]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">
|
||
<emphasis id="6991EABC4D17260BFDA4FD1C6407FD1E" box="[631,777,692,716]" italics="true" pageId="37" pageNumber="38">P. ferruginea</emphasis>
|
||
</taxonomicName>
|
||
whereas they are banded in
|
||
<emphasis id="6991EABC4D17260BFB9BFD1C620CFD1E" box="[1096,1282,692,716]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="9CE54D2D4D17260BFB9BFD1C63F0FD1E" box="[1096,1278,692,716]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="talehsapensis">P. talehsapensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D17260BFF16FD7E64CFFC8E" blockId="37.[151,1436,151,1113]" pageId="37" pageNumber="38">
|
||
These two species were included in
|
||
<taxonomicName id="9CE54D2D4D17260BFD8BFD716590FD22" box="[600,670,728,752]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">
|
||
<emphasis id="6991EABC4D17260BFD8BFD716590FD22" box="[600,670,728,752]" italics="true" pageId="37" pageNumber="38">Brada</emphasis>
|
||
</taxonomicName>
|
||
by Fauvel, because they have on the fifth chaetiger “… the nephridial papillae characteristic of the genus
|
||
<taxonomicName id="9CE54D2D4D17260BFD81FD556596FCC6" authority="Fauvel 1932:185" authorityName="Fauvel" authorityPageNumber="185" authorityYear="1932" box="[594,664,764,788]" class="Polychaeta" family="Flabelligeridae" genus="Brada" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="genus">
|
||
<emphasis id="6991EABC4D17260BFD81FD556596FCC6" box="[594,664,764,788]" italics="true" pageId="37" pageNumber="38">Brada</emphasis>
|
||
</taxonomicName>
|
||
ʺ (
|
||
<bibRefCitation id="3F744B5F4D17260BFD67FD556476FCC6" author="Fauvel" box="[692,888,764,788]" pageId="37" pageNumber="38" refString="Fauvel, P. (1932) Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum, Calcutta, 12, 1 - 262." type="journal article" year="1932">Fauvel 1932:185</bibRefCitation>
|
||
). Four paragraphs above, however, he indicated that its “… general appearance is rather unusual for the genus, the body being generally short with few segments, whilst in this species it is long, vermiform and has numerous segments.ʺ
|
||
</paragraph>
|
||
<paragraph id="5B5A36AE4D17260BFF16FCCE67E6FBE6" blockId="37.[151,1436,151,1113]" pageId="37" pageNumber="38">
|
||
On the other hand,
|
||
<taxonomicName id="9CE54D2D4D17260BFE71FCC0653BFC52" box="[418,565,872,896]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">
|
||
<emphasis id="6991EABC4D17260BFE71FCC0653BFC52" box="[418,565,872,896]" italics="true" pageId="37" pageNumber="38">P. ferruginea</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="9CE54D2D4D17260BFD64FCC06449FC52" box="[695,839,872,896]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="escobarae" status="sp. nov.">
|
||
<emphasis id="6991EABC4D17260BFD64FCC06449FC52" box="[695,839,872,896]" italics="true" pageId="37" pageNumber="38">P. escobarae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="72A257C74D17260BFC9CFCC1649EFC52" box="[847,912,872,896]" pageId="37" pageNumber="38" rank="species">
|
||
<emphasis id="6991EABC4D17260BFC9CFCC1649EFC52" bold="true" box="[847,912,872,896]" pageId="37" pageNumber="38">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(see above). Both species have opaque tunics with papillae and notochaetae with very long articles. They differ because
|
||
<taxonomicName id="9CE54D2D4D17260BFBCAFC2463BFFC76" box="[1049,1201,908,932]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">
|
||
<emphasis id="6991EABC4D17260BFBCAFC2463BFFC76" box="[1049,1201,908,932]" italics="true" pageId="37" pageNumber="38">P. ferruginea</emphasis>
|
||
</taxonomicName>
|
||
has twice as many notochaetae, the body is more papillose, and the papillae are longer. They also differ in the relative number of articles per neurospine because in
|
||
<taxonomicName id="9CE54D2D4D17260BFE26FC7C6589FC3E" box="[501,647,980,1004]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="ferruginea">
|
||
<emphasis id="6991EABC4D17260BFE26FC7C6589FC3E" box="[501,647,980,1004]" italics="true" pageId="37" pageNumber="38">P. ferruginea</emphasis>
|
||
</taxonomicName>
|
||
there are only two articles whereas in
|
||
<emphasis id="6991EABC4D17260BFBEFFC7C63DCFC3E" box="[1084,1234,980,1004]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="9CE54D2D4D17260BFBEFFC7C63C0FC3E" box="[1084,1230,980,1004]" class="Polychaeta" family="Flabelligeridae" genus="Pycnoderma" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="escobarae">P. escobarae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
neurospines have mostly three articles with a smaller median one. In addition, these two species occur in different oceans and water depths.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13FF65254D17260BFF16FB9663A5FB85" box="[197,1195,1087,1113]" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="5B5A36AE4D17260BFF16FB9663A5FB85" blockId="37.[151,1436,151,1113]" box="[197,1195,1087,1113]" pageId="37" pageNumber="38">
|
||
<emphasis id="6991EABC4D17260BFF16FB966650FB8A" bold="true" box="[197,350,1087,1112]" pageId="37" pageNumber="38">Distribution.</emphasis>
|
||
Restricted to the Gulf of Nha Trang,
|
||
<collectingCountry id="23F2763E4D17260BFCD2FB96646DFB85" box="[769,867,1087,1111]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
|
||
, in shallow water (
|
||
<quantity id="9C1D9B4B4D17260BFBE5FB966391FB85" box="[1078,1183,1087,1111]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="37" pageNumber="38" unit="m" value="21.0" valueMax="30.0" valueMin="12.0">12–30 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |