158 lines
21 KiB
XML
158 lines
21 KiB
XML
<document id="8D30BF6B1FC8C01ACF1DBB0F8CE6DBDC" ID-CLB-Dataset="20365" ID-DOI="10.11646/zootaxa.5104.3.1" ID-GBIF-Dataset="b05ecf1d-1574-4c8a-8df7-12fdd1f7ab22" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332042" ID-ZooBank="F081D236-D592-4829-A3E9-0CA58EF39E43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594429951" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="D31B87CF1E65FFF9CBA9FF6AEF2EF7C2" docLanguage="en" docName="zootaxa.5104.3.1.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium tichomirovae Shavrin 2022, sp. n." docType="treatment" docVersion="8" lastPageNumber="335" masterDocId="2F22FFB71E47FFDBCB3EFFFDED05F16A" masterDocTitle="A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records" masterLastPageNumber="346" masterPageNumber="301" pageNumber="335" updateTime="1699316848316" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C2E2236BE880A9102A4FF176DE1AE7DC">A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records</mods:title>
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<mods:namePart id="D4826F905BC0C8B31408BCEBDF8399E4">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="DB7AFFB50D9FFAAA1C557D3EFF51F0FE">Zootaxa</mods:title>
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<mods:date id="E68751F551E56E0FCE6F26E4FC26E994">2022</mods:date>
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<mods:number id="579576D07DBC60ACA5AE4BF7C2C8E4A5">2022-02-28</mods:number>
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<mods:number id="B98CC6638619B5D4487BEA1B60914D9B">5104</mods:number>
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<mods:identifier id="E51E95BED50AC723B63DCE9C28093CAC" type="DOI">10.11646/zootaxa.5104.3.1</mods:identifier>
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<treatment id="D31B87CF1E65FFF9CBA9FF6AEF2EF7C2" ID-DOI="http://doi.org/10.5281/zenodo.6333136" ID-GBIF-Taxon="193878638" ID-Zenodo-Dep="6333136" LSID="urn:lsid:plazi:treatment:D31B87CF1E65FFF9CBA9FF6AEF2EF7C2" httpUri="http://treatment.plazi.org/id/D31B87CF1E65FFF9CBA9FF6AEF2EF7C2" lastPageNumber="335" pageId="34" pageNumber="335">
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<subSubSection id="13A865521E65FFF9CBA9FF6AEF0AF1DB" box="[151,527,151,178]" pageId="34" pageNumber="335" type="nomenclature">
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<paragraph id="5B0D36D91E65FFF9CBA9FF6AEF0AF1DB" blockId="34.[151,527,151,214]" box="[151,527,151,178]" pageId="34" pageNumber="335">
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<heading id="004581B51E65FFF9CBA9FF6AEF0AF1DB" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="34" pageNumber="335" reason="1">
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<emphasis id="69C6EACB1E65FFF9CBA9FF6AEF0AF1DB" bold="true" box="[151,527,151,178]" pageId="34" pageNumber="335">
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<taxonomicName id="9CB24D5A1E65FFF9CBA9FF6AECC0F1D8" ID-CoL="9GCTX" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,453,151,178]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="tichomirovae" status="sp. nov.">
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<emphasis id="69C6EACB1E65FFF9CBA9FF6AECC0F1D8" bold="true" box="[151,453,151,178]" italics="true" pageId="34" pageNumber="335">Anthobium tichomirovae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72F557B01E65FFF9CAF3FF6AEF0AF1DB" box="[461,527,151,177]" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E65FFF9CBA9FF46EC95F1BF" box="[151,400,187,214]" pageId="34" pageNumber="335" type="description">
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<paragraph id="5B0D36D91E65FFF9CBA9FF46EC95F1BF" blockId="34.[151,527,151,214]" box="[151,400,187,214]" pageId="34" pageNumber="335">
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(
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<figureCitation id="C3892A5C1E65FFF9CBA1FF46EDF8F1BF" box="[159,253,187,213]" captionStart="FIGURE 81" captionStartId="25.[151,250,720,744]" captionTargetBox="[156,1436,190,696]" captionTargetId="figure-16@25.[151,1436,181,696]" captionTargetPageId="25" captionText="FIGURE 81. Distribution of Anthobium tectum (circles) and A. tichomirovae (square). Black symbols: material studied by author, white symbols: previously published material not seen by author." figureDoi="http://doi.org/10.5281/zenodo.6332068" httpUri="https://zenodo.org/record/6332068/files/figure.png" pageId="34" pageNumber="335">Figs. 81</figureCitation>
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,
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<figureCitation id="C3892A5C1E65FFF9CA37FF46EC22F1BC" box="[265,295,187,214]" captionStart="FIGURES 94–98" captionStartId="36.[151,264,1931,1955]" captionTargetBox="[171,1408,280,1896]" captionTargetId="figure-71@36.[159,1429,273,1908]" captionTargetPageId="36" captionText="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332079" httpUri="https://zenodo.org/record/6332079/files/figure.png" pageId="34" pageNumber="335">95</figureCitation>
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,
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<figureCitation id="C3892A5C1E65FFF9CA0DFF46EC8DF1BF" box="[307,392,187,213]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="34" pageNumber="335">99–100</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E65FFF9CBA9FEFEE840F0E3" pageId="34" pageNumber="335" type="materials_examined">
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<paragraph id="5B0D36D91E65FFF9CBA9FEFEE840F0E3" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
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<emphasis id="69C6EACB1E65FFF9CBA9FEFEECC3F077" bold="true" box="[151,454,259,285]" pageId="34" pageNumber="335">Type material examined:</emphasis>
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<materialsCitation id="EBDA3C841E65FFF9CAF1FEFEE840F0E3" ID-GBIF-Occurrence="3697351410" collectingDate="1990-07-16" collectionCode="ZMM" country="Tajikistan" location="Tadzikiсtaн" pageId="34" pageNumber="335" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="8409887B1E65FFF9CAF1FEFEEF32F077" box="[463,567,259,285]" pageId="34" pageNumber="335" type="holotype">Holotype</typeStatus>
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<specimenCount id="4DB4FD501E65FFF9C97FFEF9EF53F074" box="[577,598,260,286]" pageId="34" pageNumber="335" type="male">♂</specimenCount>
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[dissected; right protarsus missing; apical part of right paramere slightly deformed]: ‘
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<collectingCountry id="23A576491E65FFF9CA1EFEDAECC2F02B" box="[288,455,295,321]" name="Tajikistan" pageId="34" pageNumber="335">Таджикистан</collectingCountry>
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. С[еверный]. | склон Гиссар- | ского хр. УЩ. Кан | чоч. 2500 м. н.У.м. |
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<date id="2F0C10191E65FFF9CE32FEDAE89DF02B" box="[1292,1432,295,321]" pageId="34" pageNumber="335" value="1990-07-16">
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<collectingDate id="3F48E9F11E65FFF9CE32FEDAE89DF02B" box="[1292,1432,295,321]" pageId="34" pageNumber="335" value="1990-07-16">16.VII.1990</collectingDate>
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</date>
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. КУЗЬминых [Tajikistan, North slope of Gissar Mt. Kanchoch Gorge.
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<quantity id="9C4A9B3C1E65FFF9C895FEB6E904F00C" box="[939,1025,331,358]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="335" unit="m" value="2500.0">2500 m</quantity>
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a.s.l. […] Kuzminykh leg.]’ <hand - written>, ‘HOLOTYPE |
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<taxonomicName id="9CB24D5A1E65FFF9CA8BFE92EF35F0E3" authorityName="Leach" authorityYear="1819" box="[437,560,367,393]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="genus">
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<emphasis id="69C6EACB1E65FFF9CA8BFE92EF35F0E3" box="[437,560,367,393]" italics="true" pageId="34" pageNumber="335">Anthobium</emphasis>
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</taxonomicName>
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<taxonomicName id="9CB24D5A1E65FFF9C97DFE92EFD3F0E3" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[579,726,367,393]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="tichomirovae" status="sp. nov.">
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<emphasis id="69C6EACB1E65FFF9C97DFE92EFD3F0E3" box="[579,726,367,393]" italics="true" pageId="34" pageNumber="335">tichomirovae</emphasis>
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</taxonomicName>
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<emphasis id="69C6EACB1E65FFF9C9E3FE92EE18F0E3" bold="true" box="[733,797,367,393]" pageId="34" pageNumber="335">
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<taxonomicNameLabel id="72F557B01E65FFF9C9E3FE92EE18F0E3" box="[733,797,367,393]" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2022’ <red, printed> (
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<collectionCode id="3DA3AE1C1E65FFF9CFD1FE92E83DF0E3" box="[1263,1336,367,393]" collectionName="ZMM" pageId="34" pageNumber="335">ZMM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="13A865521E65FFF9CBF9FE6EECB9F491" pageId="34" pageNumber="335" type="description">
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<paragraph id="5B0D36D91E65FFF9CBF9FE6EE9E7F0BB" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
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<emphasis id="69C6EACB1E65FFF9CBF9FE6EEC5CF0C7" bold="true" box="[199,345,403,429]" pageId="34" pageNumber="335">Description.</emphasis>
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Measurements: HL: 0.45; HW: 0.70; AL: 1.42; OL: 0.17; PL: 0.65; PW: 1.22; ESL: 1.50; EW: 1.65; AW: 1.57; MTbL: 0.70; MTrL: 0.42 (MTrL 1–4: 0.25; MTrL 5: 0.17); AedL: 0.62; TL: 4.20.
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</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FE26EF76F237" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
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Habitus as in
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<figureCitation id="C3892A5C1E65FFF9CA58FE26ECBEF09C" box="[358,443,475,502]" captionStart="FIGURES 94–98" captionStartId="36.[151,264,1931,1955]" captionTargetBox="[171,1408,280,1896]" captionTargetId="figure-71@36.[159,1429,273,1908]" captionTargetPageId="36" captionText="FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332079" httpUri="https://zenodo.org/record/6332079/files/figure.png" pageId="34" pageNumber="335">Fig. 95</figureCitation>
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. Head, maxillary palpomeres, antennomeres 4–11 and abdomen reddish-brown; pronotum brown; lateral and basal portions of pronotum, elytra and apical part of abdomen yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Head with distinct, transverse and very dense microsculpture, finer in middle, diagonal between anteocellar foveae and infraorbital ridges; neck with dense and transverse meshes; pronotum with fine transverse microreticulation, finer in mediobasal third; apical part of scutellum without sculpture; abdomen with dense and transverse microsculpture. Head with moderately fine, dense punctation, finer and sparser in middle between anteocellar foveae, coarser on infraorbital ridges; neck with sparse and fine punctation; pronotum with very dense punctation, slightly coarser and deeper than that on head, finer and sparser in mediobasal third; punctation of elytra as that on pronotum, finer and sparser in parascutellar portion and middle, coarser and deeper in apical portion, on each elytron forming very tangled and vague six longitudinal rows of punctures; abdominal tergites with indistinct, very fine and sparse punctation.
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</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FC9AEC8EF578" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">Head 1.5 times as broad as long, with moderately wide anteocellar foveae, reaching level of apical margin of eye; postocular ridges somewhat smooth. Antennomere 2 slightly narrower than basal antennomere, 3 slightly longer than 2, 4 shorter and broader than 3, 5–7 slightly broader than 4, 8–9 slightly shorter and broader than 7, 10 shorter than 9, apical antennomere 1.4 times as long as preapical antennomere, from middle gradually narrowed toward rounded apex.</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FBE1E90FF517" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">Pronotum convex, 1.8 times as broad as long, 1.7 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin widely rounded and distinctly protruded anteriad; lateral portions deeply impressed, with moderately wide and deep oval spot in front of middle.</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FB7BEF49F5A8" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">Elytra slightly broader than long, more than twice as long as pronotum, reaching basal margin of abdominal tergite IV, with straight apical margins.</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FB31EF99F58F" blockId="34.[151,1437,259,1704]" box="[199,668,1227,1254]" pageId="34" pageNumber="335">Metatarsus 1.6 times as long as metatibia.</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FB13EEDFF427" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
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Male. Aedeagus with wide basal bulb, gradually narrowed toward widely rounded apex; parameres exceeding apex of median lobe, widened apically, with two short apical and one preapical setae; internal sac very long, spirally folded in basal portion (
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<figureCitation id="C3892A5C1E65FFF9CA9DFACEECFDF427" box="[419,504,1331,1357]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="34" pageNumber="335">Fig. 99</figureCitation>
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). Lateral aspect of aedeagus as in
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<figureCitation id="C3892A5C1E65FFF9C84BFACEEED3F427" box="[885,982,1331,1357]" captionStart="FIGURES 99–104" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[183,1403,190,1910]" captionTargetId="figure-16@37.[174,1413,181,1919]" captionTargetPageId="37" captionText="FIGURES 99–104. Aedeagus of Anthobium tichomirovae (Figs. 99–100; holotype), A. umbrinum (Figs. 101–102; holotype) and A. vitreum (Figs. 103–104; holotype): 99, 101, 103—parameral view, 100, 102, 104—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332081" httpUri="https://zenodo.org/record/6332081/files/figure.png" pageId="34" pageNumber="335">Fig. 100</figureCitation>
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.
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</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FAABEC94F41A" blockId="34.[151,1437,259,1704]" box="[199,401,1366,1392]" pageId="34" pageNumber="335">Female unknown.</paragraph>
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<paragraph id="5B0D36D91E65FFF9CBF9FA85ECB9F491" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
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<emphasis id="69C6EACB1E65FFF9CBF9FA85ECB5F4F9" bold="true" box="[199,432,1400,1427]" pageId="34" pageNumber="335">Comparative notes.</emphasis>
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Regarding the shape of the relatively narrow aedeagus with widely truncate apex and long parameres, this species is similar to the Chinese
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<taxonomicName id="9CB24D5A1E65FFF9C9FAFA66EE74F4DF" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[708,881,1435,1462]" class="Insecta" family="Staphylinidae" genus="Anthobium" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="cornatipenis" status="sp. nov.">
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<emphasis id="69C6EACB1E65FFF9C9FAFA66EFD0F4DF" box="[708,725,1435,1461]" italics="true" pageId="34" pageNumber="335">A</emphasis>
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.
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<emphasis id="69C6EACB1E65FFF9C9DBFA61EE74F4DF" box="[741,881,1436,1461]" italics="true" pageId="34" pageNumber="335">cornatipenis</emphasis>
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</taxonomicName>
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<emphasis id="69C6EACB1E65FFF9C844FA66EEB9F4DF" bold="true" box="[890,956,1435,1461]" pageId="34" pageNumber="335">
|
||
<taxonomicNameLabel id="72F557B01E65FFF9C844FA66EEB9F4DF" box="[890,956,1435,1461]" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it is distinguished by a paler and larger body, longer antennomeres 6–10, distinctly protruding anterior angles of the pronotum, and wider apical portions of the parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A865521E65FFF9CBF9F9FEEDEEF708" pageId="34" pageNumber="335" type="distribution">
|
||
<paragraph id="5B0D36D91E65FFF9CBF9F9FEE893F777" blockId="34.[151,1437,259,1704]" box="[199,1430,1539,1566]" pageId="34" pageNumber="335">
|
||
<emphasis id="69C6EACB1E65FFF9CBF9F9FEEC64F777" bold="true" box="[199,353,1539,1565]" pageId="34" pageNumber="335">Distribution.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="8409887B1E65FFF9C9C0F9F9EE2BF774" box="[766,814,1540,1566]" pageId="34" pageNumber="335">type</typeStatus>
|
||
locality in Gissar mountain range,
|
||
<collectingCountry id="23A576491E65FFF9CF86F9FEE823F777" box="[1208,1318,1539,1565]" name="Tajikistan" pageId="34" pageNumber="335">Tajikistan</collectingCountry>
|
||
(
|
||
<figureCitation id="C3892A5C1E65FFF9CE0BF9FEE88DF777" box="[1333,1416,1539,1565]" captionStart="FIGURE 81" captionStartId="25.[151,250,720,744]" captionTargetBox="[156,1436,190,696]" captionTargetId="figure-16@25.[151,1436,181,696]" captionTargetPageId="25" captionText="FIGURE 81. Distribution of Anthobium tectum (circles) and A. tichomirovae (square). Black symbols: material studied by author, white symbols: previously published material not seen by author." figureDoi="http://doi.org/10.5281/zenodo.6332068" httpUri="https://zenodo.org/record/6332068/files/figure.png" pageId="34" pageNumber="335">Fig. 81</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5B0D36D91E65FFF9CBF9F9DBEDEEF708" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
|
||
<emphasis id="69C6EACB1E65FFF9CBF9F9DBEC83F755" bold="true" box="[199,390,1573,1600]" pageId="34" pageNumber="335">Natural history.</emphasis>
|
||
The specimen was collected at an elevation of
|
||
<quantity id="9C4A9B3C1E65FFF9C8A3F9DBEEF7F72A" box="[925,1010,1574,1601]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="34" pageNumber="335" unit="m" value="2500.0">2500 m</quantity>
|
||
a.s.l. Detailed ecological data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13A865521E65FFF9CBF9F996EF2EF7C2" pageId="34" pageNumber="335" type="etymology">
|
||
<paragraph id="5B0D36D91E65FFF9CBF9F996EF2EF7C2" blockId="34.[151,1437,259,1704]" pageId="34" pageNumber="335">
|
||
<emphasis id="69C6EACB1E65FFF9CBF9F996EC4BF7EF" bold="true" box="[199,334,1643,1669]" pageId="34" pageNumber="335">Etymology.</emphasis>
|
||
Patronymic, the species is named in memory of the famous Russian staphylinidologist Anna L`vovna Tichomirova (1940–1987).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |