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<document id="C24327CCAC3F39A26E888916DD4BB5BF" ID-CLB-Dataset="299971" ID-DOI="10.11646/zootaxa.5481.1.8" ID-GBIF-Dataset="1f5f39c9-63c9-4838-ab70-cef9388a710f" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742772" ID-ZooBank="C1DC50E5-B45B-4161-8D0C-BAB786F66316" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1721033636255" checkinUser="plazi" docAuthor="Boderau, Mathieu, Ngo-Muller, Valerie &amp; Nel, André" docDate="2024" docId="CD1187F3B004FFA3F4E1683B6C9D84B7" docLanguage="en" docName="zootaxa.5481.1.8.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Perlomyia cantalensis Boderau, Ngo-Muller &amp; Nel, 2024, sp. nov." docType="treatment" docUuid="D68FB615-936B-4598-ADC3-7CACD6A1EFC3" docUuidSource="ZooBank" docVersion="5" lastPageNumber="136" masterDocId="3128FF8BB005FFA6F4766E0C68208632" masterDocTitle="First representatives of the stonefly genus Perlomyia and the beaded lacewing genus Isoscelipteron from the Upper Miocene of France (Plecoptera: Leuctridae and Neuroptera: Berothidae), with biogeographical considerations" masterLastPageNumber="140" masterPageNumber="131" pageNumber="132" updateTime="1721244612684" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFEC45D7729DF1DEDB663952AE10793F">First representatives of the stonefly genus Perlomyia and the beaded lacewing genus Isoscelipteron from the Upper Miocene of France (Plecoptera: Leuctridae and Neuroptera: Berothidae), with biogeographical considerations</mods:title>
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<mods:namePart id="E08B447F396AB24E7E5A84E4014B8A41">Boderau, Mathieu</mods:namePart>
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<mods:affiliation id="7F73B5C5C4D56D15632808C43AF158EC">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 45 rue Buffon 75005 Paris, France</mods:affiliation>
<mods:nameIdentifier id="E4848CC0943E89EC6C615AB99D7A1EF2" type="email">mathieuboderau@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A76C448A0DFD357B5402D82A50855C98">Ngo-Muller, Valerie</mods:namePart>
<mods:nameIdentifier id="FD7218032098D6EF4021989E667AED22" type="ORCID">0000-0002-9456-6667</mods:nameIdentifier>
<mods:affiliation id="CC48BDF3B40E91282731E31ACC3BAD34">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 45 rue Buffon 75005 Paris, France</mods:affiliation>
<mods:nameIdentifier id="FAB7107D38FDF4CBD8642765E5B9CCE7" type="email">vngomuller@gmail.com</mods:nameIdentifier>
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<mods:namePart id="868EC1C8C9B97444679DF424571DFCCF">Nel, André</mods:namePart>
<mods:nameIdentifier id="B729A6EC01245893042CBCEF1E9B9BB2" type="ORCID">0000-0002-4241-7651</mods:nameIdentifier>
<mods:affiliation id="08EF0774D0E4D06CE2370346F8EE4EF3">Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 45 rue Buffon 75005 Paris, France</mods:affiliation>
<mods:nameIdentifier id="1994A361CA8DB0F6754359C166EB486B" type="email">anel@mnhn.fr</mods:nameIdentifier>
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<mods:date id="0FA6F7D393656EFB613D238E61D67E2C">2024</mods:date>
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<mods:number id="5B1D0B8AF69D749962026E7904F4F052">2024-07-15</mods:number>
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<treatment id="CD1187F3B004FFA3F4E1683B6C9D84B7" ID-DOI="http://doi.org/10.5281/zenodo.12762118" ID-GBIF-Taxon="235594773" ID-Zenodo-Dep="12762118" LSID="urn:lsid:zoobank.org:act:D68FB615-936B-4598-ADC3-7CACD6A1EFC3" httpUri="http://treatment.plazi.org/id/CD1187F3B004FFA3F4E1683B6C9D84B7" lastPageId="5" lastPageNumber="136" pageId="1" pageNumber="132">
<subSubSection id="0DA2656EB004FFA7F4E1683B6A278063" box="[151,519,1591,1618]" pageId="1" pageNumber="132" type="nomenclature">
<paragraph id="450736E5B004FFA7F4E1683B6A278063" blockId="1.[151,989,1591,1689]" box="[151,519,1591,1618]" pageId="1" pageNumber="132">
<heading id="1E4F8189B004FFA7F4E1683B6A278063" bold="true" box="[151,519,1591,1618]" fontSize="11" level="1" pageId="1" pageNumber="132" reason="1">
<emphasis id="77CCEAF7B004FFA7F4E1683B6A278063" bold="true" box="[151,519,1591,1618]" pageId="1" pageNumber="132">
<taxonomicName id="82B84D66B004FFA7F4E1683B69818060" authorityName="Boderau &amp; Ngo-Muller &amp; Nel" authorityYear="2024" box="[151,417,1591,1618]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="132" phylum="Arthropoda" rank="species" species="cantalensis" status="sp. nov.">
<emphasis id="77CCEAF7B004FFA7F4E1683B69818060" bold="true" box="[151,417,1591,1618]" italics="true" pageId="1" pageNumber="132">Perlomyia cantalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CFF578CB004FFA7F5DF683B6A278063" box="[425,519,1591,1617]" pageId="1" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="0DA2656EB004FFA7F4E1685768D980AB" pageId="1" pageNumber="132" type="description">
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urn:lsid:zoobank.org:act:
<uuid id="311E0C30B004FFA7F5DC68576BFD8047" box="[426,989,1627,1653]" pageId="1" pageNumber="132">A122D4A3-1FED-400A-863F-D107AB4D4A6C</uuid>
</uri>
</paragraph>
<paragraph id="450736E5B004FFA7F4E1687368D980AB" blockId="1.[151,989,1591,1689]" box="[151,249,1663,1689]" pageId="1" pageNumber="132">
<figureCitation id="DD832A60B004FFA7F4E1687368D980AB" box="[151,249,1663,1689]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[152,255,1228,1252]" captionStartId-1="3.[152,255,1424,1448]" captionStartId-2="4.[152,255,1097,1121]" captionStartId-3="4.[152,255,2002,2026]" captionTargetBox-0="[151,1435,181,1204]" captionTargetBox-1="[151,1435,183,1400]" captionTargetBox-2="[151,1435,181,1073]" captionTargetBox-3="[159,1429,1152,1971]" captionTargetId-0="figure-30@2.[151,1435,181,1204]" captionTargetId-1="figure-17@3.[151,1435,181,1400]" captionTargetId-2="figure-30@4.[151,1435,181,1073]" captionTargetId-3="figure-52@4.[151,1436,1145,1978]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURE 1. Perlomyia cantalensis sp. nov., holotype IF-STR-0104. Photograph of general habitus. Scale bar = 2 mm." captionText-1="FIGURE 2. Perlomyia cantalensis sp. nov., holotype IF-STR-0104. Photographs. A, forewing. B, leg. Scale bars = 2 mm (A), 1 mm (B)." captionText-2="FIGURE 3. Perlomyia cantalensis sp. nov., holotype IF-STR-0104. Photograph of hind wing. Scale bar = 2 mm." captionText-3="FIGURE 4. Perlomyia cantalensis sp. nov., holotype IF-STR-0104. Reconstruction of terminalia.. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12742776" figureDoi-1="http://doi.org/10.5281/zenodo.12742778" figureDoi-2="http://doi.org/10.5281/zenodo.12742782" figureDoi-3="http://doi.org/10.5281/zenodo.12742784" httpUri-0="https://zenodo.org/record/12742776/files/figure.png" httpUri-1="https://zenodo.org/record/12742778/files/figure.png" httpUri-2="https://zenodo.org/record/12742782/files/figure.png" httpUri-3="https://zenodo.org/record/12742784/files/figure.png" pageId="1" pageNumber="132">Figs 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0DA2656EB004FFA7F4E168CB69878118" pageId="1" pageNumber="132" type="materials_examined">
<paragraph id="450736E5B004FFA7F4E168CB69878118" blockId="1.[151,1437,1735,2014]" pageId="1" pageNumber="132">
<emphasis id="77CCEAF7B004FFA7F4E168CB692680D3" bold="true" box="[151,262,1735,1761]" pageId="1" pageNumber="132">Material.</emphasis>
<materialsCitation id="F5D03CB8B004FFA7F57D68CB69878118" ID-GBIF-Occurrence="4906760302" country="France" location="Museum National d'Histoire Naturelle" municipality="Paris" pageId="1" pageNumber="132" specimenCount="1" typeStatus="holotype">
<typeStatus id="9A038847B004FFA7F57D68CB695380D3" box="[267,371,1735,1761]" pageId="1" pageNumber="132" type="holotype">Holotype</typeStatus>
IF-STR-0104 (a complete body, with leg fragments, a forewing and a broken hind wing, which is not surprising because these insects are especially fragile), provisionally housed in the
<location id="4067603EB004FFA7F02A68E76922811B" LSID="urn:lsid:plazi:treatment:CD1187F3B004FFA3F4E1683B6C9D84B7:4067603EB004FFA7F02A68E76922811B" country="France" municipality="Paris" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="132">Muséum National d’Histoire Naturelle</location>
,
<collectingMunicipality id="A563AC9FB004FFA7F57B69036968811B" box="[269,328,1807,1833]" pageId="1" pageNumber="132">Paris</collectingMunicipality>
,
<collectingCountry id="3DAF7675B004FFA7F525691C69838118" box="[339,419,1808,1834]" name="France" pageId="1" pageNumber="132">France</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0DA2656EB004FFA7F4B1693F6B79817C" box="[199,857,1843,1870]" pageId="1" pageNumber="132" type="etymology">
<paragraph id="450736E5B004FFA7F4B1693F6B79817C" blockId="1.[151,1437,1735,2014]" box="[199,857,1843,1870]" pageId="1" pageNumber="132">
<emphasis id="77CCEAF7B004FFA7F4B1693F6968817F" bold="true" box="[199,328,1843,1869]" pageId="1" pageNumber="132">Etymology</emphasis>
. Named after the Cantal department of
<collectingCountry id="3DAF7675B004FFA7F77369386B75817C" box="[773,853,1844,1870]" name="France" pageId="1" pageNumber="132">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0DA2656EB004FFA7F4B1695B6A0F81A4" pageId="1" pageNumber="132" type="distribution">
<paragraph id="450736E5B004FFA7F4B1695B6A0F81A4" blockId="1.[151,1437,1735,2014]" pageId="1" pageNumber="132">
<emphasis id="77CCEAF7B004FFA7F4B1695B69E68143" bold="true" box="[199,454,1879,1905]" pageId="1" pageNumber="132">Locality and horizon</emphasis>
. Latest Miocene, diatom paleomaar, Sainte-Reine (quarry of Foufouilloux, Virargues village), near Murat, Cantal,
<collectingCountry id="3DAF7675B004FFA7F5AD69706A0B81A4" box="[475,555,1916,1942]" name="France" pageId="1" pageNumber="132">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0DA2656EB004FFA7F4B169936CB781EF" pageId="1" pageNumber="132" type="diagnosis">
<paragraph id="450736E5B004FFA7F4B169936CB781EF" blockId="1.[151,1437,1735,2014]" pageId="1" pageNumber="132">
<emphasis id="77CCEAF7B004FFA7F4B169936962818B" bold="true" box="[199,322,1951,1977]" pageId="1" pageNumber="132">Diagnosis.</emphasis>
Wings infuscate; hind wing ra-rp crossvein located slightly basad to fork of RP; cerci without visible expansion, sternite IX elongate but relatively short, not extending below the male genitalia.
</paragraph>
</subSubSection>
<caption id="11C7666DB007FFA4F4EE6AC06D7082D6" ID-DOI="http://doi.org/10.5281/zenodo.12742776" ID-Zenodo-Dep="12742776" box="[152,1360,1228,1253]" httpUri="https://zenodo.org/record/12742776/files/figure.png" pageId="2" pageNumber="133" startId="2.[152,255,1228,1252]" targetBox="[151,1435,181,1204]" targetPageId="2" targetType="figure">
<paragraph id="450736E5B007FFA4F4EE6AC06D7082D6" blockId="2.[152,1360,1228,1253]" box="[152,1360,1228,1253]" pageId="2" pageNumber="133">
<emphasis id="77CCEAF7B007FFA4F4EE6AC0693982D6" bold="true" box="[152,281,1228,1252]" pageId="2" pageNumber="133">FIGURE 1.</emphasis>
<taxonomicName id="82B84D66B007FFA4F5696AC06A2182D6" authorityName="Boderau &amp; Ngo-Muller &amp; Nel" authorityYear="2024" box="[287,513,1228,1252]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="species" species="cantalensis" status="sp. nov.">
<emphasis id="77CCEAF7B007FFA4F5696AC06A2182D6" box="[287,513,1228,1252]" italics="true" pageId="2" pageNumber="133">Perlomyia cantalensis</emphasis>
</taxonomicName>
<emphasis id="77CCEAF7B007FFA4F6716AC16A7782D7" bold="true" box="[519,599,1229,1253]" pageId="2" pageNumber="133">
<taxonomicNameLabel id="6CFF578CB007FFA4F6716AC16A7782D7" box="[519,599,1229,1253]" pageId="2" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype IF-STR-0104. Photograph of general habitus. Scale bar = 2 mm.
</paragraph>
</caption>
<subSubSection id="0DA2656EB007FFA4F4B16B3A6CB78133" pageId="2" pageNumber="133" type="description">
<paragraph id="450736E5B007FFA4F4B16B3A6CB78133" blockId="2.[151,1437,1334,1973]" pageId="2" pageNumber="133">
<emphasis id="77CCEAF7B007FFA4F4B16B3A69798362" bold="true" box="[199,345,1334,1360]" pageId="2" pageNumber="133">Description.</emphasis>
Body dark brown,
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long; wings infuscate. Head
<quantity id="82409B00B007FFA4F79C6B3B6C638363" box="[1002,1091,1335,1361]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="133" unit="mm" value="0.8">0.8 mm</quantity>
long,
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wide; antennae and mouthparts not discernable. Thorax
<quantity id="82409B00B007FFA4F6106B566A9E8347" box="[614,702,1370,1397]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="133" unit="mm" value="1.5">1.5 mm</quantity>
long,
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wide; fragments of legs unpreserved except for a complete detached leg, showing second tarsomere shorter than first and third tarsomeres. Forewing 8.0 mm long,
<quantity id="82409B00B007FFA4F4E16BAF68CF838F" box="[151,239,1443,1469]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="133" unit="mm" value="2.3">2.3 mm</quantity>
wide; area between ScP and costa
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wide; ScP
<quantity id="82409B00B007FFA4F72E6BAF6B8F838F" box="[856,943,1443,1469]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="133" unit="mm" value="4.1">4.1 mm</quantity>
long, simple; R/RA straight; RP separating from RA
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from wing base, at same point where M separates from R; RP with two long straight branches, separating
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distad of base of RP, without crossvein in-between; crossvein between RA and RP aligned with distal vein between ScP and RA; M forked 2.0 mm from its base, without crossvein between its branches; branches of RP and of M anteriorly curved; eight oblique crossveins between M and CuA; 10 oblique crossveins between CuA and CuP; CuA simple, nearly straight with a weak posterior curve at its apex; branches of PCu sigmoidally curved, with a very small basal cell in-between, anal vein very short. Hind wing
<quantity id="82409B00B007FFA4F04368766CAC80A7" box="[1077,1164,1658,1685]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="133" unit="mm" value="6.3">6.3 mm</quantity>
long, ca.
<quantity id="82409B00B007FFA4F08868776D7680A7" box="[1278,1366,1659,1685]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="133" unit="mm" value="1.9">1.9 mm</quantity>
wide; ScP
<quantity id="82409B00B007FFA4F4BF68926900808B" box="[201,288,1694,1721]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="133" unit="mm" value="2.5">2.5 mm</quantity>
long, simple; RP with two long simple branches; ra-rp crossvein located slightly basad to fork of RP; crossvein m-cu distad of fork of Cu; three anal veins. Abdomen
<quantity id="82409B00B007FFA4F70768CF6BE880EF" box="[881,968,1731,1757]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="133" unit="mm" value="4.2">4.2 mm</quantity>
long,
<quantity id="82409B00B007FFA4F06768CF6C4880EF" box="[1041,1128,1731,1757]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="133" unit="mm" value="0.8">0.8 mm</quantity>
wide; cerci without visible expansion, sternite IX elongate but relatively short, not extending below the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="0DA2656EB007FFA3F4B169076C9D84B7" lastPageId="5" lastPageNumber="136" pageId="2" pageNumber="133" type="discussion">
<paragraph id="450736E5B007FFA4F4B169076C1D8187" blockId="2.[151,1437,1334,1973]" pageId="2" pageNumber="133">
<emphasis id="77CCEAF7B007FFA4F4B1690769098117" bold="true" box="[199,297,1803,1829]" pageId="2" pageNumber="133">Remark</emphasis>
. The pattern of the forewing venation of this fossil is nearly identical to that of the extant genus
<taxonomicName id="82B84D66B007FFA4F1336907691E817A" authority="Stephen, 1836" authorityName="Stephen" authorityYear="1836" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B007FFA4F13369076DBC8116" box="[1349,1436,1803,1828]" italics="true" pageId="2" pageNumber="133">Leuctra</emphasis>
Stephen, 1836
</taxonomicName>
, supporting an attribution to the family
<taxonomicName id="82B84D66B007FFA4F76369236BB0817B" authorityName="Klapalek" authorityYear="1905" box="[789,912,1839,1865]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="family">Leuctridae</taxonomicName>
. More precisely, it belongs to the
<taxonomicName id="82B84D66B007FFA4F15269236DBC817B" authorityName="Klapalek" authorityYear="1905" box="[1316,1436,1839,1865]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="133" phylum="Arthropoda" rank="family">Leuctridae</taxonomicName>
because of the following characters: needle-like habitus, absence of gills, absence of X-patterned crossveins in forewings, second tarsal segment shorter than first and third segments, in forewing CuA simple, and antero-distal cell lacking a crossvein (Béthoux
<emphasis id="77CCEAF7B007FFA4F66569976A6C8187" box="[531,588,1947,1973]" italics="true" pageId="2" pageNumber="133">et al.</emphasis>
2015,
<bibRefCitation id="21294B14B007FFA4F6EE69976B7B8187" author="Chen, Z. - T. &amp; Du, Y. - Z." box="[664,859,1947,1973]" pageId="2" pageNumber="133" pagination="59 - 74" refId="ref4759" refString="Chen, Z. - T. &amp; Du, Y. - Z. (2018) A checklist and adult key to the Chinese stonefly (Plecoptera) genera. Zootaxa, 4375 (1), 59 - 74. https: // doi. org / 10.11646 / zootaxa. 4375.1.2" type="journal article" year="2018">Chen &amp; Du 2018</bibRefCitation>
;
<bibRefCitation id="21294B14B007FFA4F71169976C118187" author="Chen, Z. - T. &amp; Liu, H. - L." box="[871,1073,1947,1973]" pageId="2" pageNumber="133" pagination="649 - 654" refId="ref4814" refString="Chen, Z. - T. &amp; Liu, H. - L. (2022) A new needle stonefly with parasitic mites from the Eocene Baltic amber. Acta Palaeontologica Polonica, 67, 649 - 654. https: // doi. org / 10.4202 / app. 00984.2022" type="journal article" year="2022">Chen &amp; Liu 2022</bibRefCitation>
).
</paragraph>
<caption id="11C7666DB006FFA5F4EE6B9C692083FE" ID-DOI="http://doi.org/10.5281/zenodo.12742778" ID-Zenodo-Dep="12742778" httpUri="https://zenodo.org/record/12742778/files/figure.png" pageId="3" pageNumber="134" startId="3.[152,255,1424,1448]" targetBox="[151,1435,183,1400]" targetPageId="3" targetType="figure">
<paragraph id="450736E5B006FFA5F4EE6B9C692083FE" blockId="3.[151,1436,1424,1484]" pageId="3" pageNumber="134">
<emphasis id="77CCEAF7B006FFA5F4EE6B9C6938839A" bold="true" box="[152,280,1424,1448]" pageId="3" pageNumber="134">FIGURE 2.</emphasis>
<taxonomicName id="82B84D66B006FFA5F5686B9C6A20839A" authorityName="Boderau &amp; Ngo-Muller &amp; Nel" authorityYear="2024" box="[286,512,1424,1448]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="cantalensis" status="sp. nov.">
<emphasis id="77CCEAF7B006FFA5F5686B9C6A20839A" box="[286,512,1424,1448]" italics="true" pageId="3" pageNumber="134">Perlomyia cantalensis</emphasis>
</taxonomicName>
<emphasis id="77CCEAF7B006FFA5F6706B9C6A76839A" bold="true" box="[518,598,1424,1448]" pageId="3" pageNumber="134">
<taxonomicNameLabel id="6CFF578CB006FFA5F6706B9C6A76839A" box="[518,598,1424,1448]" pageId="3" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype IF-STR-0104. Photographs. A, forewing. B, leg. Scale bars = 2 mm (A), 1 mm (B).
</paragraph>
</caption>
<paragraph id="450736E5B006FFA5F4B168126A8D80B3" blockId="3.[151,1437,1566,1989]" pageId="3" pageNumber="134">
The new fossil is unlikely to fit in the monogeneric subfamily
<taxonomicName id="82B84D66B006FFA5F7F568126C11800A" authorityName="Zwick" authorityYear="1973" box="[899,1073,1566,1592]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="subFamily" subFamily="Megaleuctrinae">Megaleuctrinae</taxonomicName>
with
<taxonomicName id="82B84D66B006FFA5F00768126DB8800B" authority="Neave, 1934" authorityName="Neave" authorityYear="1934" box="[1137,1432,1566,1593]" class="Insecta" family="Leuctridae" genus="Megaleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F00768126D20800A" box="[1137,1280,1566,1592]" italics="true" pageId="3" pageNumber="134">Megaleuctra</emphasis>
Neave, 1934
</taxonomicName>
. This genus is characterized by large body size (about
<quantity id="82409B00B006FFA5F68E684E6B69806F" box="[760,841,1602,1629]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="134" unit="mm" value="15.0">15 mm</quantity>
) and the presence of six anal veins in hind wing, vs. only three in the new fossil (
<bibRefCitation id="21294B14B006FFA5F5A0686B6A8180B3" author="Ham, S. A. &amp; Bae, Y. J." box="[470,673,1639,1665]" pageId="3" pageNumber="134" pagination="336 - 341" refId="ref5260" refString="Ham, S. A. &amp; Bae, Y. J. (2002) The stonefly genus Megaleuctra (Plecoptera: Leuctridae) new to East Palearctic region, with description of Megaleuctra saebat new species. Entomological News, 113, 336 - 341." type="journal article" year="2002">Ham &amp; Bae 2002</bibRefCitation>
).
</paragraph>
<paragraph id="450736E5B006FFA5F4B168866A4281F6" blockId="3.[151,1437,1566,1989]" pageId="3" pageNumber="134">
The male sternite IX of the new fossil is strongly produced as in
<taxonomicName id="82B84D66B006FFA5F7D168866CE98097" authority="(Ricker, 1943)" baseAuthorityName="Ricker" baseAuthorityYear="1943" box="[935,1225,1674,1701]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F7D168866C3C8096" box="[935,1052,1674,1700]" italics="true" pageId="3" pageNumber="134">Perlomyia</emphasis>
(
<bibRefCitation id="21294B14B006FFA5F05D68866CE18097" author="Ricker, W. E." box="[1067,1217,1674,1701]" pageId="3" pageNumber="134" pagination="1 - 145" refId="ref6358" refString="Ricker, W. E. (1943) Stoneflies of Southwestern British Columbia. Indiana University Publication in Science Series, 12, 1 - 145." type="journal article" year="1943">Ricker, 1943</bibRefCitation>
)
</taxonomicName>
. The veins RP and M of forewing with a common origin is a character of the new fossil only present in the extant genus
<taxonomicName id="82B84D66B006FFA5F15E68A26DBD80FA" baseAuthorityName="Ricker" baseAuthorityYear="1943" box="[1320,1437,1710,1736]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F15E68A26DBD80FA" box="[1320,1437,1710,1736]" italics="true" pageId="3" pageNumber="134">Perlomyia</emphasis>
</taxonomicName>
(Needham &amp; Claassen, 1925: pl. 32, fig. 5;
<bibRefCitation id="21294B14B006FFA5F60068DE6BA380DF" author="Nelson, C. H. &amp; Hanson, J. F." box="[630,899,1746,1773]" pageId="3" pageNumber="134" pagination="187 - 199" refId="ref6179" refString="Nelson, C. H. &amp; Hanson, J. F. (1973) The genus Perlomyia (Plecoptera: Leuctridae). Journal of the Kansas Entomological Society, 46, 187 - 199. [https: // www. jstor. org / stable / 25082566]" type="journal article" year="1973">Nelson &amp; Hanson, 1973</bibRefCitation>
), unlike in
<taxonomicName id="82B84D66B006FFA5F78868DF6C7580DE" authorityName="Stephen" authorityYear="1836" box="[1022,1109,1747,1772]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F78868DF6C7580DE" box="[1022,1109,1747,1772]" italics="true" pageId="3" pageNumber="134">Leuctra</emphasis>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F01768DE6DB880DF" authority="Despax, 1929" authorityName="Despax" authorityYear="1929" box="[1121,1432,1746,1773]" class="Insecta" family="Leuctridae" genus="Pachyleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F01768DE6CD680DE" box="[1121,1270,1746,1772]" italics="true" pageId="3" pageNumber="134">Pachyleuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F08B68DF6DB880DF" author="Despax, R." box="[1277,1432,1747,1773]" pageId="3" pageNumber="134" pagination="298 - 301" refId="ref4965" refString="Despax, R. (1929) Description sommaire d'un sous-genre nouveau et deux especes nouvelles du genre Leuctra Stephan (Plecoptera). Bulletin de la Societe Entomologique de France, 34, 298 - 301. https: // doi. org / 10.3406 / bsef. 1920.26647" type="journal article" year="1929">Despax, 1929</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F4E168FA6A728123" authority="Shepard &amp; Baumann, 1995" authorityName="Shepard &amp; Baumann" authorityYear="1995" box="[151,594,1782,1809]" class="Insecta" family="Leuctridae" genus="Calileuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F4E168FA69378122" box="[151,279,1782,1808]" italics="true" pageId="3" pageNumber="134">Calileuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F56868FA6A728123" author="Shepard, W. D. &amp; Baumann, R. W." box="[286,594,1782,1809]" pageId="3" pageNumber="134" pagination="124 - 134" refId="ref6527" refString="Shepard, W. D. &amp; Baumann, R. W. (1995) Calileuctra, a new genus, and two new species of stoneflies from California (Plecoptera: Leuctridae). Great Basin Naturalist, 55, 124 - 134. [https: // www. jstor. org / stable / 41712876] https: // doi. org / 10.5962 / bhl. part. 22805" type="journal article" year="1995">Shepard &amp; Baumann, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F62B68FA6BA98123" authority="Hanson, 1941" authorityName="Hanson" authorityYear="1941" box="[605,905,1782,1809]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F62B68FA6AC48122" box="[605,740,1782,1808]" italics="true" pageId="3" pageNumber="134">Paraleuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F69D68FB6BA98123" author="Hanson, J. F." box="[747,905,1783,1809]" pageId="3" pageNumber="134" pagination="57 - 66" refId="ref5308" refString="Hanson, J. F. (1941) Studies on the Plecoptera of North America. 2. Bulletin of the Brooklyn Entomological Society, 36, 57 - 66." type="journal article" year="1941">Hanson, 1941</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F7E368FA6D388123" authority="Stark &amp; Kyzar, 2000" authorityName="Stark &amp; Kyzar" authorityYear="2000" box="[917,1304,1782,1809]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F7E368FA6C068122" box="[917,1062,1782,1808]" italics="true" pageId="3" pageNumber="134">Pomoleuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F05B68FA6D388123" author="Stark, B. P. &amp; Kyzar, J. W." box="[1069,1304,1782,1809]" pageId="3" pageNumber="134" pagination="119 - 135" refId="ref6758" refString="Stark, B. P. &amp; Kyzar, J. W. (2000) Systematics of Nearctic Paraleuctra with description of a new genus (Plecoptera: Leuctridae). Tijdschrift voor Entomologie, 144, 119 - 135. https: // doi. org / 10.1163 / 22119434 - 99900060" type="journal article" year="2000">Stark &amp; Kyzar, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F15568FA69118107" authority="Ricker, 1952" authorityName="Ricker" authorityYear="1952" class="Insecta" family="Leuctridae" genus="Zealeuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F15568FA6DBC8122" box="[1315,1436,1782,1808]" italics="true" pageId="3" pageNumber="134">Zealeuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F4E1691669118107" author="Ricker, W. E." box="[151,305,1818,1845]" pageId="3" pageNumber="134" pagination="1 - 200" refId="ref6386" refString="Ricker, W. E. (1952) Systematic studies in Plecoptera. Indiana University Publication in Science Series, 18, 1 - 200." type="journal article" year="1952">Ricker, 1952</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F53469166A648107" authority="Ricker, 1943" authorityName="Ricker" authorityYear="1943" box="[322,580,1818,1845]" class="Insecta" family="Leuctridae" genus="Moselia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F534691669BC8106" box="[322,412,1818,1844]" italics="true" pageId="3" pageNumber="134">Moselia</emphasis>
<bibRefCitation id="21294B14B006FFA5F5DD69166A648107" author="Ricker, W. E." box="[427,580,1818,1845]" pageId="3" pageNumber="134" pagination="1 - 145" refId="ref6358" refString="Ricker, W. E. (1943) Stoneflies of Southwestern British Columbia. Indiana University Publication in Science Series, 12, 1 - 145." type="journal article" year="1943">Ricker, 1943</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F62069176B478107" authority="Ricker, 1943" authorityName="Ricker" authorityYear="1943" box="[598,871,1818,1845]" class="Insecta" family="Leuctridae" genus="Despaxia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F62069176AE08106" box="[598,704,1819,1844]" italics="true" pageId="3" pageNumber="134">Despaxia</emphasis>
<bibRefCitation id="21294B14B006FFA5F6BB69166B478107" author="Ricker, W. E." box="[717,871,1818,1845]" pageId="3" pageNumber="134" pagination="1 - 145" refId="ref6358" refString="Ricker, W. E. (1943) Stoneflies of Southwestern British Columbia. Indiana University Publication in Science Series, 12, 1 - 145." type="journal article" year="1943">Ricker, 1943</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F70F69166CF58107" authority="Klapalek, 1912" authorityName="Klapalek" authorityYear="1912" box="[889,1237,1818,1845]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F70F69166C338106" box="[889,1043,1818,1844]" italics="true" pageId="3" pageNumber="134">Rhopalopsole</emphasis>
<bibRefCitation id="21294B14B006FFA5F05669166CF58107" author="Klapalek, F." box="[1056,1237,1818,1845]" pageId="3" pageNumber="134" pagination="342 - 351" refId="ref5473" refString="Klapalek, F. (1912) H. Sauter's Formosa Ausbeute. Entomologische Mitteilungen, 1, 342 - 351. https: // doi. org / 10.5962 / bhl. part. 25903" type="journal article" year="1912">Klapálek, 1912</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="82B84D66B006FFA5F0916916696B816B" authority="Consiglio, 1957" authorityName="Consiglio" authorityYear="1957" class="Insecta" family="Leuctridae" genus="Tyrrhenoleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F09169166DBC8106" box="[1255,1436,1818,1844]" italics="true" pageId="3" pageNumber="134">Tyrrhenoleuctra</emphasis>
<bibRefCitation id="21294B14B006FFA5F4E16932696B816B" author="Consiglio, C." box="[151,331,1854,1881]" pageId="3" pageNumber="134" refId="ref4895" refString="Consiglio, C. (1957) Tyrrhenoleuctra nomen novum (Plecoptera). Bollettino della Societa Entomologica Italiana, 87, 17." type="journal volume" year="1957">Consiglio, 1957</bibRefCitation>
</taxonomicName>
(replacement name for
<emphasis id="77CCEAF7B006FFA5F62F69326AE6816A" box="[601,710,1854,1880]" italics="true" pageId="3" pageNumber="134">Strobiella</emphasis>
Klapálek, 1901), †
<taxonomicName id="82B84D66B006FFA5F7D669326C92816B" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[928,1202,1854,1881]" class="Insecta" family="Leuctridae" genus="Baltileuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F7D669326C06816A" box="[928,1062,1854,1880]" italics="true" pageId="3" pageNumber="134">Baltileuctra</emphasis>
Chen, 2018
</taxonomicName>
, †
<taxonomicName id="82B84D66B006FFA5F0BA693268EF814F" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" class="Insecta" family="Leuctridae" genus="Euroleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F0BA69326D72816A" box="[1228,1362,1854,1880]" italics="true" pageId="3" pageNumber="134">Euroleuctra</emphasis>
Chen, 2018
</taxonomicName>
and †
<taxonomicName id="82B84D66B006FFA5F56E696E6AEA814F" authority="Caruso &amp; Wichard, 2011" authorityName="Caruso &amp; Wichard" authorityYear="2011" box="[280,714,1890,1917]" class="Insecta" family="Leuctridae" genus="Palaeopsole" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B006FFA5F56E696E6982814E" box="[280,418,1890,1916]" italics="true" pageId="3" pageNumber="134">Palaeopsole</emphasis>
<bibRefCitation id="21294B14B006FFA5F5DA696F6AEA814F" author="Caruso, C. &amp; Wichard, W." box="[428,714,1890,1917]" pageId="3" pageNumber="134" pagination="69 - 77" refId="ref4595" refString="Caruso, C. &amp; Wichard, W. (2011) Paleogeographic distribution of Leuctridae and Nemouridae genera preserved in Baltic amber, with the description of Palaeopsole weiterschani n. gen., n. sp. (Plecoptera). Entomologie heute, 23, 69 - 77." type="journal article" year="2011">Caruso &amp; Wichard, 2011</bibRefCitation>
</taxonomicName>
(all from Baltic amber, Eocene) (
<bibRefCitation id="21294B14B006FFA5F026696E6CDB814F" author="Klapalek, F." box="[1104,1275,1890,1917]" pageId="3" pageNumber="134" pagination="342 - 351" refId="ref5473" refString="Klapalek, F. (1912) H. Sauter's Formosa Ausbeute. Entomologische Mitteilungen, 1, 342 - 351. https: // doi. org / 10.5962 / bhl. part. 25903" type="journal article" year="1912">Klapálek 1912</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F17F696E6DB8814F" author="Ricker, W. E." box="[1289,1432,1890,1917]" pageId="3" pageNumber="134" pagination="1 - 145" refId="ref6358" refString="Ricker, W. E. (1943) Stoneflies of Southwestern British Columbia. Indiana University Publication in Science Series, 12, 1 - 145." type="journal article" year="1943">Ricker 1943</bibRefCitation>
,
<bibRefCitation id="21294B14B006FFA5F4E1698A68F28193" author="Ricker, W. E." box="[151,210,1926,1953]" pageId="3" pageNumber="134" pagination="1 - 200" refId="ref6386" refString="Ricker, W. E. (1952) Systematic studies in Plecoptera. Indiana University Publication in Science Series, 18, 1 - 200." type="journal article" year="1952">1952</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F496698A69B18193" author="Consiglio, C." box="[224,401,1926,1953]" pageId="3" pageNumber="134" pagination="81 - 91" refId="ref4867" refString="Consiglio, C. (1956) Note sul genere Strobliella Klapalek (Plecoptera). Memorie della Societa Entomologica Italiana, 35, 81 - 91." type="journal article" year="1956">Consiglio 1956</bibRefCitation>
,
<bibRefCitation id="21294B14B006FFA5F5E8698A69F98193" author="Consiglio, C." box="[414,473,1926,1953]" pageId="3" pageNumber="134" refId="ref4895" refString="Consiglio, C. (1957) Tyrrhenoleuctra nomen novum (Plecoptera). Bollettino della Societa Entomologica Italiana, 87, 17." type="journal volume" year="1957">1957</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F591698A6A8E8192" author="Berthelemy, C." box="[487,686,1926,1952]" pageId="3" pageNumber="134" pagination="175 - 198" refId="ref4530" refString="Berthelemy, C. (1968) Contribution a la connaissance des Leuctridae (Plecoptera). Annales de Limnologie, 4, 175 - 198. https: // doi. org / 10.1051 / limn / 1968016" type="journal article" year="1968">Berthélemy 1968</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F6CA698A6BCF8193" author="Shepard, W. D. &amp; Baumann, R. W." box="[700,1007,1926,1953]" pageId="3" pageNumber="134" pagination="124 - 134" refId="ref6527" refString="Shepard, W. D. &amp; Baumann, R. W. (1995) Calileuctra, a new genus, and two new species of stoneflies from California (Plecoptera: Leuctridae). Great Basin Naturalist, 55, 124 - 134. [https: // www. jstor. org / stable / 41712876] https: // doi. org / 10.5962 / bhl. part. 22805" type="journal article" year="1995">Shepard &amp; Baumann 1995</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F78B698A6CBF8193" author="Shimizu, T." box="[1021,1183,1926,1953]" pageId="3" pageNumber="134" pagination="285 - 303" refId="ref6601" refString="Shimizu, T. (2000) Paraleuctra (Insecta: Plecoptera: Leuctridae) from Japan, with taxonomic notes on the Japanese Leuctridae. Species Diversity, 5, 285 - 303. https: // doi. org / 10.12782 / specdiv. 5.285" type="journal article" year="2000">Shimizu 2000</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F0DA698A6DB88193" author="Stark, B. P. &amp; Kyzar, J. W." box="[1196,1432,1926,1953]" pageId="3" pageNumber="134" pagination="119 - 135" refId="ref6758" refString="Stark, B. P. &amp; Kyzar, J. W. (2000) Systematics of Nearctic Paraleuctra with description of a new genus (Plecoptera: Leuctridae). Tijdschrift voor Entomologie, 144, 119 - 135. https: // doi. org / 10.1163 / 22119434 - 99900060" type="journal article" year="2000">Stark &amp; Kyzar 2000</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F4E169A7698A81F7" author="Caruso, C. &amp; Wichard, W." box="[151,426,1962,1989]" pageId="3" pageNumber="134" pagination="69 - 77" refId="ref4595" refString="Caruso, C. &amp; Wichard, W. (2011) Paleogeographic distribution of Leuctridae and Nemouridae genera preserved in Baltic amber, with the description of Palaeopsole weiterschani n. gen., n. sp. (Plecoptera). Entomologie heute, 23, 69 - 77." type="journal article" year="2011">Caruso &amp; Wichard 2011</bibRefCitation>
;
<bibRefCitation id="21294B14B006FFA5F5C069A66A6281F6" author="Chen, Z. - T." box="[438,578,1962,1988]" pageId="3" pageNumber="134" pagination="281 - 287" refId="ref4647" refString="Chen, Z. - T. (2018 a) Baltileuctra gen. nov., a new genus of Leuctridae (Insecta: Plecoptera) in Baltic amber. Zootaxa, 4407 (2), 281 - 287. https: // doi. org / 10.11646 / zootaxa. 4407.2.9" type="journal article" year="2018">Chen 2018a</bibRefCitation>
,b).
</paragraph>
<caption id="11C7666DB001FFA2F4EE6A456D3C8253" ID-DOI="http://doi.org/10.5281/zenodo.12742782" ID-Zenodo-Dep="12742782" box="[152,1308,1097,1121]" httpUri="https://zenodo.org/record/12742782/files/figure.png" pageId="4" pageNumber="135" startId="4.[152,255,1097,1121]" targetBox="[151,1435,181,1073]" targetPageId="4" targetType="figure">
<paragraph id="450736E5B001FFA2F4EE6A456D3C8253" blockId="4.[152,1308,1097,1121]" box="[152,1308,1097,1121]" pageId="4" pageNumber="135">
<emphasis id="77CCEAF7B001FFA2F4EE6A4569388253" bold="true" box="[152,280,1097,1121]" pageId="4" pageNumber="135">FIGURE 3.</emphasis>
<taxonomicName id="82B84D66B001FFA2F5696A456A218253" authorityName="Boderau &amp; Ngo-Muller &amp; Nel" authorityYear="2024" box="[287,513,1097,1121]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="cantalensis" status="sp. nov.">
<emphasis id="77CCEAF7B001FFA2F5696A456A218253" box="[287,513,1097,1121]" italics="true" pageId="4" pageNumber="135">Perlomyia cantalensis</emphasis>
</taxonomicName>
<emphasis id="77CCEAF7B001FFA2F6716A456A778253" bold="true" box="[519,599,1097,1121]" pageId="4" pageNumber="135">
<taxonomicNameLabel id="6CFF578CB001FFA2F6716A456A778253" box="[519,599,1097,1121]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype IF-STR-0104. Photograph of hind wing. Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="11C7666DB001FFA2F4EE69DE6D6E81DB" ID-DOI="http://doi.org/10.5281/zenodo.12742784" ID-Zenodo-Dep="12742784" box="[152,1358,2002,2026]" httpUri="https://zenodo.org/record/12742784/files/figure.png" pageId="4" pageNumber="135" startId="4.[152,255,2002,2026]" targetBox="[159,1429,1152,1971]" targetPageId="4" targetType="figure">
<paragraph id="450736E5B001FFA2F4EE69DE6D6E81DB" blockId="4.[152,1358,2002,2026]" box="[152,1358,2002,2026]" pageId="4" pageNumber="135">
<emphasis id="77CCEAF7B001FFA2F4EE69DE693881D8" bold="true" box="[152,280,2002,2026]" pageId="4" pageNumber="135">FIGURE 4.</emphasis>
<taxonomicName id="82B84D66B001FFA2F56969DE6A2181D8" authorityName="Boderau &amp; Ngo-Muller &amp; Nel" authorityYear="2024" box="[287,513,2002,2026]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="cantalensis" status="sp. nov.">
<emphasis id="77CCEAF7B001FFA2F56969DE6A2181D8" box="[287,513,2002,2026]" italics="true" pageId="4" pageNumber="135">Perlomyia cantalensis</emphasis>
</taxonomicName>
<emphasis id="77CCEAF7B001FFA2F67169DE6A7781D8" bold="true" box="[519,599,2002,2026]" pageId="4" pageNumber="135">
<taxonomicNameLabel id="6CFF578CB001FFA2F67169DE6A7781D8" box="[519,599,2002,2026]" pageId="4" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype IF-STR-0104. Reconstruction of terminalia.. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="450736E5B000FFA3F4B16E9B69C687BB" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="136">
Furthermore, the new fossil strongly differs from
<taxonomicName id="82B84D66B000FFA3F6816E9B6BA88683" authorityName="Stark &amp; Kyzar" authorityYear="2000" box="[759,904,151,177]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F6816E9B6BA88683" box="[759,904,151,177]" italics="true" pageId="5" pageNumber="136">Pomoleuctra</emphasis>
</taxonomicName>
in the structure of the branches of PCu that are sigmoidally curved and with a very small basal cell in-between in the new fossil vs. simply posteriorly curved with a broad cell in-between in the latter (
<bibRefCitation id="21294B14B000FFA3F65B6ED36B3486CB" author="Stark, B. P. &amp; Kyzar, J. W." box="[557,788,223,249]" pageId="5" pageNumber="136" pagination="119 - 135" refId="ref6758" refString="Stark, B. P. &amp; Kyzar, J. W. (2000) Systematics of Nearctic Paraleuctra with description of a new genus (Plecoptera: Leuctridae). Tijdschrift voor Entomologie, 144, 119 - 135. https: // doi. org / 10.1163 / 22119434 - 99900060" type="journal article" year="2000">Stark &amp; Kyzar 2000</bibRefCitation>
).
<taxonomicName id="82B84D66B000FFA3F7536ED36B8586CB" authorityName="Shepard &amp; Baumann" authorityYear="1995" box="[805,933,223,249]" class="Insecta" family="Leuctridae" genus="Calileuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F7536ED36B8586CB" box="[805,933,223,249]" italics="true" pageId="5" pageNumber="136">Calileuctra</emphasis>
</taxonomicName>
differs from the new fossil in the branches of PCu distally fused and not sigmoidal and in the presence of only three crossveins between M and CuA in forewing (
<bibRefCitation id="21294B14B000FFA3F4E96F2B69F68773" author="Shepard, W. D. &amp; Baumann, R. W." box="[159,470,294,321]" pageId="5" pageNumber="136" pagination="124 - 134" refId="ref6527" refString="Shepard, W. D. &amp; Baumann, R. W. (1995) Calileuctra, a new genus, and two new species of stoneflies from California (Plecoptera: Leuctridae). Great Basin Naturalist, 55, 124 - 134. [https: // www. jstor. org / stable / 41712876] https: // doi. org / 10.5962 / bhl. part. 22805" type="journal article" year="1995">Shepard &amp; Baumann 1995</bibRefCitation>
).
<taxonomicName id="82B84D66B000FFA3F59A6F2B6A818773" authorityName="Consiglio" authorityYear="1957" box="[492,673,295,321]" class="Insecta" family="Leuctridae" genus="Tyrrhenoleuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F59A6F2B6A818773" box="[492,673,295,321]" italics="true" pageId="5" pageNumber="136">Tyrrhenoleuctra</emphasis>
</taxonomicName>
and †
<taxonomicName id="82B84D66B000FFA3F69D6F2B6B518773" authorityName="Chen" authorityYear="2018" box="[747,881,295,321]" class="Insecta" family="Leuctridae" genus="Baltileuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F69D6F2B6B518773" box="[747,881,295,321]" italics="true" pageId="5" pageNumber="136">Baltileuctra</emphasis>
</taxonomicName>
also have only three-four such veins (
<bibRefCitation id="21294B14B000FFA3F1586F2B68F38757" author="Consiglio, C." pageId="5" pageNumber="136" pagination="81 - 91" refId="ref4867" refString="Consiglio, C. (1956) Note sul genere Strobliella Klapalek (Plecoptera). Memorie della Societa Entomologica Italiana, 35, 81 - 91." type="journal article" year="1956">Consiglio 1956</bibRefCitation>
).
<taxonomicName id="82B84D66B000FFA3F49E6F4769628757" authorityName="Ricker" authorityYear="1943" box="[232,322,331,357]" class="Insecta" family="Leuctridae" genus="Moselia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F49E6F4769628757" box="[232,322,331,357]" italics="true" pageId="5" pageNumber="136">Moselia</emphasis>
</taxonomicName>
and
<taxonomicName id="82B84D66B000FFA3F50C6F4769C48756" authorityName="Ricker" authorityYear="1943" box="[378,484,331,356]" class="Insecta" family="Leuctridae" genus="Despaxia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F50C6F4769C48756" box="[378,484,331,356]" italics="true" pageId="5" pageNumber="136">Despaxia</emphasis>
</taxonomicName>
are excluded because the crossvein m-cu is distad of the fork of Cu in hind wing in the new fossil (vs. proximad).
</paragraph>
<paragraph id="450736E5B000FFA3F4B16F9F6D18842B" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="136">
In †
<taxonomicName id="82B84D66B000FFA3F4856F9F6959879F" authorityName="Chen" authorityYear="2018" box="[243,377,403,429]" class="Insecta" family="Leuctridae" genus="Baltileuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F4856F9F6959879F" box="[243,377,403,429]" italics="true" pageId="5" pageNumber="136">Baltileuctra</emphasis>
</taxonomicName>
the ra-rp crossvein is located distally to the fork of RP in forewings but basal to it in hind wing, a character uncommon among
<taxonomicName id="82B84D66B000FFA3F5866FBB6BA287E3" authority="(Chen &amp; Liu 2022: 652)" baseAuthorityName="Chen &amp; Liu" baseAuthorityPageNumber="652" baseAuthorityYear="2022" box="[496,898,438,465]" class="Insecta" family="Leuctridae" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="family">
Leuctridae (
<bibRefCitation id="21294B14B000FFA3F6006FBB6B5987E3" author="Chen, Z. - T. &amp; Liu, H. - L." box="[630,889,438,465]" pageId="5" pageNumber="136" pagination="649 - 654" refId="ref4814" refString="Chen, Z. - T. &amp; Liu, H. - L. (2022) A new needle stonefly with parasitic mites from the Eocene Baltic amber. Acta Palaeontologica Polonica, 67, 649 - 654. https: // doi. org / 10.4202 / app. 00984.2022" type="journal article" year="2022">Chen &amp; Liu 2022: 652</bibRefCitation>
)
</taxonomicName>
. In the new fossil, the hind wing ra-rp crossvein is also located slightly basad to the fork of RP, while in the extant
<taxonomicName id="82B84D66B000FFA3F71F6FD76BFE87C7" baseAuthorityName="Ricker" baseAuthorityYear="1943" box="[873,990,475,501]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F71F6FD76BFE87C7" box="[873,990,475,501]" italics="true" pageId="5" pageNumber="136">Perlomyia</emphasis>
</taxonomicName>
, the hind wing ra-rp crossvein is located opposite the fork of RP (
<bibRefCitation id="21294B14B000FFA3F5DC6FF36A97842B" author="Nelson, C. H. &amp; Hanson, J. F." box="[426,695,511,537]" pageId="5" pageNumber="136" pagination="187 - 199" refId="ref6179" refString="Nelson, C. H. &amp; Hanson, J. F. (1973) The genus Perlomyia (Plecoptera: Leuctridae). Journal of the Kansas Entomological Society, 46, 187 - 199. [https: // www. jstor. org / stable / 25082566]" type="journal article" year="1973">Nelson &amp; Hanson 1973</bibRefCitation>
). †
<taxonomicName id="82B84D66B000FFA3F6AF6FF36B7F842B" authorityName="Chen" authorityYear="2018" box="[729,863,511,537]" class="Insecta" family="Leuctridae" genus="Baltileuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="77CCEAF7B000FFA3F6AF6FF36B7F842B" box="[729,863,511,537]" italics="true" pageId="5" pageNumber="136">Baltileuctra</emphasis>
</taxonomicName>
has very long cerci, unlike the new fossil.
</paragraph>
<paragraph id="450736E5B000FFA3F4B16C2F6C9D84B7" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="136">
The new fossil has simple cerci, its sternite IX is relatively short, while the extant species have a longer sternite IX and/or cerci with expansions (
<bibRefCitation id="21294B14B000FFA3F6706C4B6AB18453" author="Frison, T. H." box="[518,657,583,609]" pageId="5" pageNumber="136" pagination="256 - 265" refId="ref5211" refString="Frison, T. H. (1936) Some new species of stoneflies from Oregon (Plecoptera). Annals of the Entomological Society of America, 29 (2), 256 - 265. https: // doi. org / 10.1093 / aesa / 29.2.256" type="journal article" year="1936">Frison 1936</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F6EA6C4B6B068453" author="Kawai, T." box="[668,806,583,609]" pageId="5" pageNumber="136" pagination="1 - 211" refId="ref5425" refString="Kawai, T. (1967) Plecoptera (Insecta). Fauna Japonica. Biogeographical Society of Japan, Tokyo, 1 - 211." type="book chapter" year="1967">Kawai 1967</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F7466C4B6C188453" author="Nelson, C. H. &amp; Hanson, J. F." box="[816,1080,583,609]" pageId="5" pageNumber="136" pagination="187 - 199" refId="ref6179" refString="Nelson, C. H. &amp; Hanson, J. F. (1973) The genus Perlomyia (Plecoptera: Leuctridae). Journal of the Kansas Entomological Society, 46, 187 - 199. [https: // www. jstor. org / stable / 25082566]" type="journal article" year="1973">Nelson &amp; Hanson 1973</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F0356C4B6CCF8453" author="Zhiltzova, L. A." box="[1091,1263,583,609]" pageId="5" pageNumber="136" pagination="359 - 364" refId="ref6868" refString="Zhiltzova, L. A. (1974) Rare genera of the family Leuctridae (lnsecta, Plecoptera) in the fauna of the USSR. Zoologicheskii Zhurnal, 53, 359 - 364." type="journal article" year="1974">Zhiltzova 1974</bibRefCitation>
,
<bibRefCitation id="21294B14B000FFA3F08F6C4A6D148453" author="Zhiltzova, L. A." box="[1273,1332,582,609]" pageId="5" pageNumber="136" pagination="221 - 230" refId="ref6904" refString="Zhiltzova, L. A. (1975) Rhopalopsole, a new genus of stoneflies for the USSR (Plecoptera, Leuctridae). Zoologicheskii Zhurnal, 54, 221 - 230." type="journal article" year="1975">1975</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F1496C4B68F384B7" author="Shimizu, T." pageId="5" pageNumber="136" pagination="285 - 303" refId="ref6601" refString="Shimizu, T. (2000) Paraleuctra (Insecta: Plecoptera: Leuctridae) from Japan, with taxonomic notes on the Japanese Leuctridae. Species Diversity, 5, 285 - 303. https: // doi. org / 10.12782 / specdiv. 5.285" type="journal article" year="2000">Shimizu 2000</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F4A86C6769EC84B7" author="Sivec, I. &amp; Stark, B. P." box="[222,460,619,645]" pageId="5" pageNumber="136" pagination="94 - 103" refId="ref6648" refString="Sivec, I. &amp; Stark, B. P. (2012 a) Seven new species of Perlomyia (Plecoptera: Leuctridae) from Japan. Illiesia, 8, 94 - 103." type="journal article" year="2012">Sivec &amp; Stark 2012a</bibRefCitation>
,b;
<bibRefCitation id="21294B14B000FFA3F59C6C676AED84B7" author="Muranyi, D. &amp; Jeon, M. J. &amp; Hwang, J. M. &amp; Seo, H. Y." box="[490,717,619,645]" pageId="5" pageNumber="136" pagination="145 - 154" refId="ref5741" refString="Muranyi, D., Jeon, M. J., Hwang, J. M. &amp; Seo, H. Y. (2014) Korean species of the genus Perlomyia Banks, 1906 (Plecoptera: Leuctridae). Zootaxa, 3881 (2), 145 - 154. https: // doi. org / 10.11646 / zootaxa. 3881.2.3" type="journal article" year="2014">
Murányi
<emphasis id="77CCEAF7B000FFA3F6246C676AAA84B7" box="[594,650,619,645]" italics="true" pageId="5" pageNumber="136">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F6A16C676BCF84B7" author="Muranyi, D. &amp; Hwang, J. M." box="[727,1007,619,645]" pageId="5" pageNumber="136" pagination="43 - 61" refId="ref5807" refString="Muranyi, D. &amp; Hwang, J. M. (2017) Four new species and further contributions to the Leuctridae (Plecoptera) of the Korean Peninsula. Zootaxa, 4282 (1), 43 - 61. https: // doi. org / 10.11646 / zootaxa. 4282.1.2" type="journal article" year="2017">Murányi &amp; Hwang 2017</bibRefCitation>
;
<bibRefCitation id="21294B14B000FFA3F78C6C676C9184B7" author="Nakamine, H." box="[1018,1201,619,645]" pageId="5" pageNumber="136" pagination="91 - 94" refId="ref5861" refString="Nakamine, H. (2022) A new species of Perlomyia (Insecta: Plecoptera: Leuctridae) from Japan. Species Diversity, 27, 91 - 94. https: // doi. org / 10.12782 / specdiv. 27.91" type="journal article" year="2022">Nakamine 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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