266 lines
37 KiB
XML
266 lines
37 KiB
XML
<document id="2BA79CBBFD7B13ED305FAF962900154F" ID-DOI="10.11646/zootaxa.3429.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="10096509" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629801692040" checkinUser="felipe" docAuthor="Jäger, Peter" docDate="2012" docId="C84887861139FFB8FF71FF0154CFFDF8" docLanguage="en" docName="zt03429p063.pdf" docOrigin="Zootaxa 3429 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3429.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ctenus holthoffi Jäger, 2012, spec. nov." docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="3471FFFE111BFF9DFFE6F83B5749FFC4" masterDocTitle="Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat & Gray 2001 (Arachnida: Araneae: Ctenidae)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="35" updateTime="1699579940834" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="21A86D8A61C82C4D15E05998C9356433" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5C1F2A2700BE54ED9ACB06E11C4AD494">
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<mods:title id="013634E03A98E25EFBD6E1D5732297F5">Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat & Gray 2001 (Arachnida: Araneae: Ctenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A5AE013CA05AE82AF72F813D99A8B21E" type="personal">
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<mods:role id="F980283142B4211AB6BA2DA8C17AD315">
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<mods:roleTerm id="4ADCAB228A78E5390FF0B6E25E86D61F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F885D78CF08563F1806B4EED969AD8C2">Jäger, Peter</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4ACDDE5C131CFBEB5093554351047421">text</mods:typeOfResource>
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<mods:titleInfo id="CCFD7AC7FB5CD8C3B18FE84F7820BF84">
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<mods:title id="34F3422105BF2FF7A7014C59140BA9FD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="41A7C771B7DDD1637F05AEC0A36F332C">
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<mods:date id="1995052CCBBFEE2B31CB485996F07D75">2012</mods:date>
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<mods:detail id="A7D50F3F8F94BDE57B596682AB576161" type="pubDate">
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<mods:number id="CBB6E4759862D67FDC9F3D8C607C2855">2012-08-20</mods:number>
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</mods:detail>
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<mods:detail id="AA06F96D2BC924311EB43E2E72CEC0E7" type="volume">
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<mods:number id="CF8E0373711920783DE54011DB02D064">3429</mods:number>
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</mods:detail>
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<mods:detail id="82853ADE72621A3BF467589513942C48" type="issue">
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<mods:number id="D6F4E5C8B63DBC7A76E32261E80315BA">1</mods:number>
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</mods:detail>
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<mods:extent id="D85C7D3EC36D8446572F8F65DE040852" unit="page">
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<mods:start id="610A28362FA281DAEBD71C575D1D0CB5">1</mods:start>
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<mods:end id="05F398FDF2F494080D6261DB155446CC">63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="D40792DC99E84865746E8CD3C9EFE6A7">
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<mods:url id="7A487F1E980DADD286003C929168BA55">http://dx.doi.org/10.11646/zootaxa.3429.1.1</mods:url>
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</mods:location>
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<mods:classification id="E22681D16D8EB260E5B1A2ECD0115E34">journal article</mods:classification>
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<mods:identifier id="25D66FD48A3718A2B7A0875CAF1F7089" type="DOI">10.11646/zootaxa.3429.1.1</mods:identifier>
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<mods:identifier id="4ABCE10FD8D3ABF1163DFBB895FE3C6A" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="24165AF7A5F11E2DE2502635382E0745" type="Zenodo-Dep">10096509</mods:identifier>
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</mods:mods>
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<treatment id="C84887861139FFB8FF71FF0154CFFDF8" LSID="urn:lsid:plazi:treatment:C84887861139FFB8FF71FF0154CFFDF8" httpUri="http://treatment.plazi.org/id/C84887861139FFB8FF71FF0154CFFDF8" lastPageId="37" lastPageNumber="38" pageId="34" pageNumber="35">
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<subSubSection id="08FB651B1139FFBFFF71FF015692F890" box="[151,475,1850,1876]" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="405E36901139FFBFFF71FF015692F890" blockId="34.[151,475,1850,1909]" box="[151,475,1850,1876]" pageId="34" pageNumber="35">
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<heading id="1B1681FC1139FFBFFF71FF015692F890" bold="true" box="[151,475,1850,1876]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
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<emphasis id="7295EA821139FFBFFF71FF015692F890" bold="true" box="[151,475,1850,1876]" pageId="34" pageNumber="35">
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<taxonomicName id="87E14D131139FFBFFF71FF015615F890" ID-CoL="6BNZR" authorityName="Jager" authorityYear="2012" box="[151,348,1850,1876]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="holthoffi" status="sp. nov.">
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<emphasis id="7295EA821139FFBFFF71FF015615F890" bold="true" box="[151,348,1850,1876]" italics="true" pageId="34" pageNumber="35">Ctenus holthoffi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="69A657F91139FFBFFE85FF015692F890" box="[355,475,1850,1876]" pageId="34" pageNumber="35" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="08FB651B1139FFBEFF71FF66542AFEDF" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
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<paragraph id="405E36901139FFBFFF71FF665613F8B1" blockId="34.[151,475,1850,1909]" box="[151,346,1885,1909]" pageId="34" pageNumber="35">
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<figureCitation id="D8DA2A151139FFBFFF71FF6657A5F8B1" box="[151,236,1885,1909]" captionStart="FIGURE 39" captionStartId="14.[151,250,1909,1932]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure-80@14.[151,1436,193,1888]" captionTargetPageId="14" captionText="FIGURE 39. Records of Ctenus spp. in South and Southeast Asia. 1 Between Chitre and Chandrung, Nepal. 2–5 Burma (2" figureDoi="http://doi.org/10.5281/zenodo.10096523" httpUri="https://zenodo.org/record/10096523/files/figure.png" pageId="34" pageNumber="35">Figs 39</figureCitation>
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,
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<figureCitation id="D8DA2A151139FFBFFF1EFF665613F8B1" box="[248,346,1885,1909]" captionStart-0="FIGURES 146–153" captionStart-1="FIGURES 154–162" captionStartId-0="36.[151,264,1933,1956]" captionStartId-1="37.[151,264,1894,1917]" captionTargetBox-0="[204,1386,193,1911]" captionTargetBox-1="[163,1436,639,1863]" captionTargetId-0="figure-59@36.[204,1386,193,1918]" captionTargetId-1="figure-219@37.[151,1436,628,1873]" captionTargetPageId-0="36" captionTargetPageId-1="37" captionText-0="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." captionText-1="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi-0="http://doi.org/10.5281/zenodo.10096557" figureDoi-1="http://doi.org/10.5281/zenodo.10096559" httpUri-0="https://zenodo.org/record/10096557/files/figure.png" httpUri-1="https://zenodo.org/record/10096559/files/figure.png" pageId="34" pageNumber="35">146–162</figureCitation>
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</paragraph>
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<paragraph id="405E36901138FFBEFF71F8AC542AFEDF" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
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hand, by night, P. Jäger & J. Martens leg.
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<date id="345F10501138FFBEFD8AF8AC55A4FF6B" box="[620,749,151,175]" pageId="35" pageNumber="36" value="2010-03-16">16.III.2010</date>
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.
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<materialsCitation id="F0893CCD1138FFBEFD1EF8A352D1FF6B" box="[760,1432,151,176]" collectionCode="SMF" pageId="35" pageNumber="36" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<emphasis id="7295EA821138FFBEFD1EF8A354ADFF74" bold="true" box="[760,996,151,176]" pageId="35" pageNumber="36">
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<typeStatus id="9F5A88321138FFBEFD1EF8A3543DFF74" box="[760,884,152,176]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="56E7FD191138FFBEFC66F8AC54ADFF74" box="[896,996,151,176]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
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||
</emphasis>
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(
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<collectionCode id="26F0AE551138FFBEFC15F8AC5367FF6B" box="[1011,1070,151,175]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="35" pageNumber="36">SMF</collectionCode>
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) with same data as for holotype
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</materialsCitation>
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.
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<materialsCitation id="F0893CCD1138FFBEFF71F8805651FF33" collectingDate="2010-03-16" collectionCode="SMF" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<emphasis id="7295EA821138FFBEFF71F88057A3FF10" bold="true" box="[151,234,187,212]" pageId="35" pageNumber="36">
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||
<specimenCount id="56E7FD191138FFBEFF71F88057A3FF10" box="[151,234,187,212]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
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||
</emphasis>
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||
(
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<collectionCode id="26F0AE551138FFBEFEE6F8805674FF17" box="[256,317,187,211]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="35" pageNumber="36">SMF</collectionCode>
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), with same data as for holotype, but: primary forest, dry leaf litter and soil, at day, sieving,
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<date id="345F10501138FFBEFF71F8E45651FF33" box="[151,280,223,247]" pageId="35" pageNumber="36" value="2010-03-16">
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<collectingDate id="241BE9B81138FFBEFF71F8E45651FF33" box="[151,280,223,247]" pageId="35" pageNumber="36" value="2010-03-16">16.III.2010</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="F0893CCD1138FFBEFEC4F8E452CAFF33" box="[290,1411,223,248]" collectingDate="2010-03-12" collectionCode="SMF" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<emphasis id="7295EA821138FFBEFEC4F8E45639FF3C" bold="true" box="[290,368,223,248]" pageId="35" pageNumber="36">
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||
<specimenCount id="56E7FD191138FFBEFEC4F8E45639FF3C" box="[290,368,223,248]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
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||
</emphasis>
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||
(
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||
<collectionCode id="26F0AE551138FFBEFE99F8E456F5FF33" box="[383,444,223,247]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="35" pageNumber="36">SMF</collectionCode>
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), with same data as for holotype, but: coffee plantation and primary forest,
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<date id="345F10501138FFBEFAE4F8E452CAFF33" box="[1282,1411,223,247]" pageId="35" pageNumber="36" value="2010-03-12">
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||
<collectingDate id="241BE9B81138FFBEFAE4F8E452CAFF33" box="[1282,1411,223,247]" pageId="35" pageNumber="36" value="2010-03-12">12.III.2010</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="F0893CCD1138FFBEFA68F8E45417FEDF" collectingDate="2010-03-19" collectionCode="SMF" pageId="35" pageNumber="36" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<emphasis id="7295EA821138FFBEFA68F8E457AFFEDF" bold="true" pageId="35" pageNumber="36">
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||
<specimenCount id="56E7FD191138FFBEFA68F8E457AFFEDF" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
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||
</emphasis>
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||
(
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||
<collectionCode id="26F0AE551138FFBEFF13F938567BFEDF" box="[245,306,259,283]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="35" pageNumber="36">SMF</collectionCode>
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), with same data as for holotype, but:
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<date id="345F10501138FFBEFD3BF9385417FEDF" box="[733,862,259,283]" pageId="35" pageNumber="36" value="2010-03-19">
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||
<collectingDate id="241BE9B81138FFBEFD3BF9385417FEDF" box="[733,862,259,283]" pageId="35" pageNumber="36" value="2010-03-19">19.III.2010</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="08FB651B1138FFBEFF21F91D532EFE42" pageId="35" pageNumber="36" type="materials_examined">
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||
<paragraph id="405E36901138FFBEFF21F91D532EFE42" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
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||
Additional material examined.
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<materialsCitation id="F0893CCD1138FFBEFDD8F91D532AFE42" collectingDate="2012-04-30" collectionCode="SMF" collectorName="P. Jager" country="Laos" elevation="200" latitude="17.631617" location="Tham Kamouk" longLatPrecision="1" longitude="105.12503" municipality="Gnommalat District" pageId="35" pageNumber="36" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Khammouan">
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||
<emphasis id="7295EA821138FFBEFDD8F91D544BFEFB" bold="true" box="[574,770,294,319]" pageId="35" pageNumber="36">
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||
<specimenCount id="56E7FD191138FFBEFDD8F91D55C6FEFB" box="[574,655,294,319]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
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||
,
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||
<specimenCount id="56E7FD191138FFBEFD7AF91D544BFEFB" box="[668,770,294,319]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
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||
</emphasis>
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||
(
|
||
<collectionCode id="26F0AE551138FFBEFCF4F91C5406FEFB" box="[786,847,295,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="35" pageNumber="36">SMF</collectionCode>
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||
),
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<collectingCountry id="38F676001138FFBEFC82F91C54D4FEFA" box="[868,925,295,318]" name="Laos" pageId="35" pageNumber="36">Laos</collectingCountry>
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||
,
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<collectingRegion id="8225F8721138FFBEFC4FF91C53E1FEFB" box="[937,1192,295,319]" country="Laos" name="Khammouan" pageId="35" pageNumber="36">Khammouan Province</collectingRegion>
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||
,
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||
<collectingMunicipality id="A03AACEA1138FFBEFB52F91C52D1FEFB" box="[1204,1432,295,319]" pageId="35" pageNumber="36">Gnommalat District</collectingMunicipality>
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||
,
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||
<locationDeviation id="51EC2FB21138FFBEFF71F9715680FEA6" box="[151,457,330,354]" location="Ban Thatot" pageId="35" pageNumber="36">
|
||
<quantity id="87199B751138FFBEFF71F97157A1FEA6" box="[151,232,330,354]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="35" pageNumber="36" unit="km" value="2.5">2.5 km</quantity>
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WNW Ban Thatot
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||
</locationDeviation>
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||
,
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||
<location id="453E604B1138FFBEFE31F97155CFFEA6" LSID="urn:lsid:plazi:treatment:C84887861139FFB8FF71FF0154CFFDF8:453E604B1138FFBEFE31F97155CFFEA6" box="[471,646,330,354]" country="Laos" latitude="17.631617" longLatPrecision="1" longitude="105.12503" municipality="Gnommalat District" name="Tham Kamouk" pageId="35" pageNumber="36" stateProvince="Khammouan">Tham Kamouk</location>
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||
,
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||
<geoCoordinate id="25D550571138FFBEFD73F970547AFEA6" box="[661,819,330,354]" degrees="17" direction="north" minutes="37.897" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="17.631617">N 17°37.897´</geoCoordinate>
|
||
,
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||
<geoCoordinate id="25D550571138FFBEFCA4F97054A5FEA6" box="[834,1004,330,354]" degrees="105" direction="east" minutes="07.502" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="105.12503">E 105°07.502´</geoCoordinate>
|
||
, ca.
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||
<quantity id="87199B751138FFBEFBC3F9715326FEA6" box="[1061,1135,330,354]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="200.0">
|
||
<elevation id="CBCCD1A31138FFBEFBC3F9715326FEA6" box="[1061,1135,330,354]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="200.0">200 m</elevation>
|
||
</quantity>
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||
altitude, in front of cave, secondary forest, leaf litter between rocks, by hand, by sunset,
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||
<collectorName id="ED1453461138FFBEFCB1F95454E4FE43" box="[855,941,367,391]" pageId="35" pageNumber="36">P. Jäger</collectorName>
|
||
leg.
|
||
<date id="345F10501138FFBEFC02F955532AFE42" box="[996,1123,366,390]" pageId="35" pageNumber="36" value="2012-04-30">
|
||
<collectingDate id="241BE9B81138FFBEFC02F955532AFE42" box="[996,1123,366,390]" pageId="35" pageNumber="36" value="2012-04-30">30.IV.2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08FB651B1138FFBEFF21F9A9545DFE35" pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph id="405E36901138FFBEFF21F9A9545DFE35" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
|
||
<emphasis id="7295EA821138FFBEFF21F9A95607FE6E" bold="true" box="[199,334,402,426]" inLineHeading="true" pageId="35" pageNumber="36" reason="1">Etymology.</emphasis>
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This species is named in honour of the former conductor of the Paderborn cathedral choir, Mr Theodor Holthoff, for his patience with a crowd of young boys and a good time especially when we played football in a special room of the cathedral; name in genitive case.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="08FB651B1138FFBEFF21F9C65364FD0C" pageId="35" pageNumber="36" type="diagnosis">
|
||
<paragraph id="405E36901138FFBEFF21F9C65364FD0C" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
|
||
<emphasis id="7295EA821138FFBEFF21F9C65608FDD2" bold="true" box="[199,321,509,534]" inLineHeading="true" pageId="35" pageNumber="36" reason="1">Diagnosis.</emphasis>
|
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Medium-sized
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<taxonomicName id="87E14D131138FFBEFE1BF9C65529FDD1" authorityName="Keyserling" authorityYear="1877" box="[509,608,509,533]" class="Arachnida" family="Ctenidae" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Ctenidae</taxonomicName>
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||
(total length male 11.8, female 13.0–15.0). Distinguished from other
|
||
<taxonomicName id="87E14D131138FFBEFF71FA1A5654FDFD" box="[151,285,545,569]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="7295EA821138FFBEFF71FA1A57ADFDFD" box="[151,228,545,569]" italics="true" pageId="35" pageNumber="36">Ctenus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
by the modified leg coxae III and IV in males each with ventral hump and covered by small stout spines (
|
||
<figureCitation id="D8DA2A151138FFBEFF16FA7F56DAFD98" box="[240,403,580,604]" captionStart="FIGURES 154–162" captionStartId="37.[151,264,1894,1917]" captionTargetBox="[163,1436,639,1863]" captionTargetId="figure-219@37.[151,1436,628,1873]" captionTargetPageId="37" captionText="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi="http://doi.org/10.5281/zenodo.10096559" httpUri="https://zenodo.org/record/10096559/files/figure.png" pageId="35" pageNumber="36">Figs 157–158</figureCitation>
|
||
), those on coxae III indistinct; moreover, palpal tibia provided with strong RTA curved inwards (
|
||
<figureCitation id="D8DA2A151138FFBEFEE6FA53562DFD44" box="[256,356,616,640]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="35" pageNumber="36">Fig. 147</figureCitation>
|
||
); tegulum with large longitudinally elongated central tegular apophysis (
|
||
<figureCitation id="D8DA2A151138FFBEFB7AFA5353B6FD44" box="[1180,1279,616,640]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="35" pageNumber="36">Fig. 147</figureCitation>
|
||
). Females are recognised by their huge lateral teeth almost extending over the entire length of epigyne (
|
||
<figureCitation id="D8DA2A151138FFBEFB6BFAB75262FD60" box="[1165,1323,652,676]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="35" pageNumber="36">Figs 149–151</figureCitation>
|
||
,
|
||
<figureCitation id="D8DA2A151138FFBEFADEFAB7522AFD60" box="[1336,1379,652,676]" captionStart="FIGURES 154–162" captionStartId="37.[151,264,1894,1917]" captionTargetBox="[163,1436,639,1863]" captionTargetId="figure-219@37.[151,1436,628,1873]" captionTargetPageId="37" captionText="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi="http://doi.org/10.5281/zenodo.10096559" httpUri="https://zenodo.org/record/10096559/files/figure.png" pageId="35" pageNumber="36">162</figureCitation>
|
||
) and their widely separated spermathecae (> two diameter of spermathecae) (
|
||
<figureCitation id="D8DA2A151138FFBEFC58FA8B5369FD0C" box="[958,1056,688,712]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="35" pageNumber="36">Fig. 152</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08FB651B1138FFB8FF21FAE854E1FDD3" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="405E36901138FFBEFF21FAE85484FB95" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
|
||
<emphasis id="7295EA821138FFBEFF21FAE856D4FD28" bold="true" box="[199,413,723,748]" pageId="35" pageNumber="36">
|
||
<emphasis id="7295EA821138FFBEFF21FAE85611FD28" box="[199,344,723,748]" inLineHeading="true" pageId="35" pageNumber="36" reason="1">Description.</emphasis>
|
||
Male
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="9F5A88321138FFBEFE48FAE8555EFD2F" box="[430,535,723,747]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
). PL 6.3, PW 4.8, AW 2.4, OL 5.8, OW 3.5. Eye diameters and interdistances: AME 0.30, ALE 0.19, PME 0.38, PLE 0.35, AME–AME 0.13, AME–ALE 0.31, PME–PME 0.22, PME–PLE 0.34, AME–PME 0.07, ALE–PLE 0.15, clypeus AME 0.13, clypeus ALE 0.50. Palp and leg measurements: palp 7.6 (2.8, 1.1, 1.5, -, 2.2), I 19.5 (5.2, 2.6, 5.2, 4.7, 1.8), II 17.6 (4.8, 2.6, 4.4, 4.3, 1.5), III 15.3 (4.2, 2.2, 3.5, 4.1, 1.3), IV 22.3 (5.5, 2.4, 5.3, 7.2, 1.9). Leg formula 4123. Spination of palp and legs: palp 141, 100, 1100; femora I p021, d111, r112, II p112, d111, r 112, III p1(0)12, d111, r112, IV p112, d111, r002; patellae 101; tibiae I p010, r110, v22222, II p010, r110, v22222, III p1(2)1, d111, r11, v222; IV p11, d111, r11, v222; metatarsi I–II p111, r111, v223, III p112, d010, r112, v222, IV p112, d010, r112,
|
||
<date id="345F10501138FFBEFCC2FBF55424FC22" box="[804,877,974,998]" pageId="35" pageNumber="36" value="2222-05">v2222</date>
|
||
. Chelicerae with 3 promarginal, 5 retromarginal teeth, and with elongated patch of several tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 6 bristles. Sparse scopula on all tarsi and metatarsi I–III. Leg claw I and IV with 3 secondary teeth. Position of tarsal organ: I 0.88, II 0.85, III 0.78, IV 0.77.
|
||
</paragraph>
|
||
<paragraph id="405E36901138FFBEFF21FC6754D4FAC7" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
|
||
Palp as in diagnosis (
|
||
<figureCitation id="D8DA2A151138FFBEFE51FC675511FBB0" box="[439,600,1116,1140]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="35" pageNumber="36">Figs 146–138</figureCitation>
|
||
). RTA with relatively broad base, tip with scratches distally in retrolateral view. Cymbium tip slightly conical, retro-proximally widened and with small pocket. Embolus arising at 8 to 8.30- o’clock-position from tegulum, semicircular in ventral view, short with broad base, its tip slightly bifurcated and situated in distal half of tegulum. Conductor arising at 12-o’clock-position subdistally. Tegular apophysis arising at 6-o’clock-position from tegulum, distinctly excavated on dorsal side.
|
||
</paragraph>
|
||
<paragraph id="405E36901138FFBEFF21FD3455AFF9AD" blockId="35.[151,1437,151,1927]" pageId="35" pageNumber="36">
|
||
Colour (
|
||
<figureCitation id="D8DA2A151138FFBEFEC1FD345683FAE3" box="[295,458,1295,1319]" captionStart="FIGURES 154–162" captionStartId="37.[151,264,1894,1917]" captionTargetBox="[163,1436,639,1863]" captionTargetId="figure-219@37.[151,1436,628,1873]" captionTargetPageId="37" captionText="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi="http://doi.org/10.5281/zenodo.10096559" httpUri="https://zenodo.org/record/10096559/files/figure.png" pageId="35" pageNumber="36">Figs 154–158</figureCitation>
|
||
). Reddish-brown with dark pattern. Dorsal prosoma with distinct light median band consisting of white dense hairs, laterally with radial marble-like pattern; distinctly marked fovea partly hidden behind hairs. Sternum and ventral coxae reddish-brown without pattern; labium and gnathocoxae reddish-brown with grey and light distal parts. Chelicerae as dorsal prosoma with longitudinal lines in proximal half and with dark distal half. Legs reddish-brown with femora and hind legs darker; femora and partly other segments with white hairs dorsally; legs III + IV with distal tibia and metatarsi dark. Dorsal opisthosoma greyish-brown, with large light area and lateral dark patches in anterior half; posterior half with light median band and pairs of dark spots. Lateral opisthosoma without distinct pattern. Ventral opisthosoma dark with four posteriorly converging rows of white patches consisting of white hairs; epiandrium and muscle sigilla light. Anterior lateral spinnerets dark, posterior lateral and median spinnerets and anal tubercle light.
|
||
</paragraph>
|
||
<paragraph id="405E36901138FFBEFF21FE4E5394F836" blockId="35.[151,1437,151,1927]" lastBlockId="35.[151,1436,1938,2035]" pageId="35" pageNumber="36">
|
||
<emphasis id="7295EA821138FFBEFF21FE4E55FAF948" box="[199,691,1652,1677]" inLineHeading="true" pageId="35" pageNumber="36" reason="1">
|
||
<emphasis id="7295EA821138FFBEFF21FE4E5657F949" bold="true" box="[199,286,1653,1677]" pageId="35" pageNumber="36">Female</emphasis>
|
||
(
|
||
<typeStatus id="9F5A88321138FFBEFED6FE4F56DBF948" box="[304,402,1652,1676]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
|
||
together with
|
||
<typeStatus id="9F5A88321138FFBEFDA6FE4F55EEF948" box="[576,679,1652,1676]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
PL 6.1, PW 4.9, AW 3.2, OL 6.9, OW 4.3. Eye diameters and interdistances: AME 0.31, ALE 0.28, PME 0.40, PLE 0.33, AME–AME 0.17, AME–ALE 0.39, PME–PME 0.22, PME–PLE 0.46, AME–PME 0.09, ALE–PLE 0.12, clypeus AME 0.15, clypeus ALE 0.56. Palp and leg measurements: palp 6.2 (2.2, 1.2, 1.3, -, 1.5), I 15.1 (4.3, 2.4, 3.8, 3.3, 1.3), II 14.4 (4.2, 2.3, 3.6, 3.1, 1.2), III 12.9 (3.9, 1.8, 2.9, 3.1, 1.2), IV 19.2 (5.1, 2.1, 4.6, 5.7, 1.7). Leg formula 4123. Spination of palp and legs: palp 131, 100, 1211, 1013; femora I p021, d111, r111, II p112, d111, r 111, III p112(1), d111, r 112, IV p1(0)12, d111, r002(1); patellae I–II 000, III–IV 101; tibiae I–II v22222, III–IV p11, d111, r11, v222; metatarsi I–II v222, III p112, d010, r112, v222, IV p112, d010, r112,
|
||
<date id="345F10501138FFBEFDB8FF5455EFF843" box="[606,678,1903,1927]" pageId="35" pageNumber="36" value="2122-05">v2122</date>
|
||
. Chelicerae with 3 promarginal, 5(4) retromarginal teeth, and with elongated patch of several tiny denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 6–7 thin bristles. Ventral tarsi and metatarsi I–II with sparse scopula. Leg claw I with 3 secondary teeth, leg claw IV with 5 (3 large and 2 small). Position of tarsal organ: I 0.84, II 0.79, III 0.66, IV 0.72.
|
||
</paragraph>
|
||
<caption id="149E6618113FFFB9FF71FFB654A0F823" ID-DOI="http://doi.org/10.5281/zenodo.10096557" ID-Zenodo-Dep="10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,264,1933,1956]" targetBox="[204,1386,193,1911]" targetPageId="36">
|
||
<paragraph id="405E3690113FFFB9FF71FFB654A0F823" blockId="36.[151,1436,1933,2023]" pageId="36" pageNumber="37">
|
||
<emphasis id="7295EA82113FFFB9FF71FFB655EFF867" bold="true" box="[151,678,1933,1956]" pageId="36" pageNumber="37">
|
||
FIGURES 146–153.
|
||
<taxonomicName id="87E14D13113FFFB9FE98FFB65564F867" authorityName="Jager" authorityYear="2012" box="[382,557,1933,1955]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="holthoffi" status="sp. nov.">
|
||
<emphasis id="7295EA82113FFFB9FE98FFB65564F867" bold="true" box="[382,557,1933,1955]" italics="true" pageId="36" pageNumber="37">Ctenus holthoffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69A657F9113FFFB9FDDFFFB555EFF867" box="[569,678,1934,1955]" pageId="36" pageNumber="37" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="405E3690113EFFB8FF21F8AC57B6FEA0" blockId="37.[151,1436,151,572]" pageId="37" pageNumber="38">
|
||
Copulatory organ as in diagnosis (
|
||
<figureCitation id="D8DA2A15113EFFB8FD87F8AC544DFF6B" box="[609,772,151,175]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="37" pageNumber="38">Figs 149–153</figureCitation>
|
||
,
|
||
<figureCitation id="D8DA2A15113EFFB8FCF3F8AC540AFF6B" box="[789,835,151,175]" captionStart="FIGURES 154–162" captionStartId="37.[151,264,1894,1917]" captionTargetBox="[163,1436,639,1863]" captionTargetId="figure-219@37.[151,1436,628,1873]" captionTargetPageId="37" captionText="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi="http://doi.org/10.5281/zenodo.10096559" httpUri="https://zenodo.org/record/10096559/files/figure.png" pageId="37" pageNumber="38">162</figureCitation>
|
||
). Epigynal field anterio-laterally with two separate patches, close to the base of lateral teeth with two pairs of slit sense organs. Median plate roughly transverse oval, lateral teeth arising at anterior part and with small basal field of scratches. Epigyne rounded posteriorly. Internal duct system with rather short lateral folds diverging posteriorly. Transverse ovoid spermathecae directed towards each other, internally with a small chamber between spermathecae and fertilisation ducts, the latter pointing medially.
|
||
</paragraph>
|
||
<paragraph id="405E3690113EFFB8FF21F95454CFFE6F" blockId="37.[151,1436,151,572]" pageId="37" pageNumber="38">
|
||
Colour (
|
||
<figureCitation id="D8DA2A15113EFFB8FEC2F954568AFE43" box="[292,451,367,391]" captionStart="FIGURES 154–162" captionStartId="37.[151,264,1894,1917]" captionTargetBox="[163,1436,639,1863]" captionTargetId="figure-219@37.[151,1436,628,1873]" captionTargetPageId="37" captionText="FIGURES 154–162. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (154–158 holotype male, 159–162 paratype female). 154–156, 159–161 Habitus (154, 159 dorsal; 155, 160 ventral; 156 lateral; 161 frontal). 157–158 Hump on" figureDoi="http://doi.org/10.5281/zenodo.10096559" httpUri="https://zenodo.org/record/10096559/files/figure.png" pageId="37" pageNumber="38">Figs 159–161</figureCitation>
|
||
). As in male, but median dorsal opisthosomal band and bands on frontal chelicerae less distinct. Lateral and dorsal opisthosoma with additional dark spots.
|
||
</paragraph>
|
||
<paragraph id="405E3690113EFFB8FF21F98C54E1FDD3" blockId="37.[151,1436,151,572]" pageId="37" pageNumber="38">
|
||
<emphasis id="7295EA82113EFFB8FF21F98C5608FE14" bold="true" box="[199,321,439,464]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Variation.</emphasis>
|
||
Second female
|
||
<typeStatus id="9F5A8832113EFFB8FE1AF9835528FE14" box="[508,609,440,464]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
|
||
: PL 7.0, OL 8.3. Epigyne larger and epigynal field fused with anterior patches (
|
||
<figureCitation id="D8DA2A15113EFFB8FF18F9E7562CFE30" box="[254,357,476,500]" captionStart="FIGURES 146–153" captionStartId="36.[151,264,1933,1956]" captionTargetBox="[204,1386,193,1911]" captionTargetId="figure-59@36.[204,1386,193,1918]" captionTargetPageId="36" captionText="FIGURES 146–153. Ctenus holthoffi spec. nov., from Tad Fane, Champasak, Laos (146–148 holotype male, 149–153 paratype females). 146–148 Left male palp (146 prolateral; 147 ventral; 148 retrolateral). 149–150 Epigyne, ventral; 151 Epigyne, posterior; 152 Vulva, dorsal; 153 Schematic course of duct system, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10096557" httpUri="https://zenodo.org/record/10096557/files/figure.png" pageId="37" pageNumber="38">Fig. 150</figureCitation>
|
||
). Only one pair of slit sense organs was recognised in the undissected epigyne. Colour was generally darker with a stronger contrast between light and dark parts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08FB651B113EFFB8FF21FA1854CFFDF8" box="[199,902,547,572]" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="405E3690113EFFB8FF21FA1854CFFDF8" blockId="37.[151,1436,151,572]" box="[199,902,547,572]" pageId="37" pageNumber="38">
|
||
<emphasis id="7295EA82113EFFB8FF21FA185629FDF8" bold="true" box="[199,352,547,572]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="9F5A8832113EFFB8FD86FA1F55D9FDF8" box="[608,656,548,572]" pageId="37" pageNumber="38">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="D8DA2A15113EFFB8FD1CFA1F5406FDF8" box="[762,847,548,572]" captionStart="FIGURE 39" captionStartId="14.[151,250,1909,1932]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure-80@14.[151,1436,193,1888]" captionTargetPageId="14" captionText="FIGURE 39. Records of Ctenus spp. in South and Southeast Asia. 1 Between Chitre and Chandrung, Nepal. 2–5 Burma (2" figureDoi="http://doi.org/10.5281/zenodo.10096523" httpUri="https://zenodo.org/record/10096523/files/figure.png" pageId="37" pageNumber="38">Fig. 39</figureCitation>
|
||
: 12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |