260 lines
36 KiB
XML
260 lines
36 KiB
XML
<document id="07C7E731003A5076A4FBA44EE2356F1A" ID-CLB-Dataset="4617" ID-DOI="10.5281/zenodo.5331298" ID-GBIF-Dataset="3f9f1bfd-4da1-46f0-a1e8-e9e57320bcb3" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169405387" checkinUser="diego" docAuthor="Krikken, Jan & Huijbregts, Johannes" docDate="2012" docId="C83387FAFFCF056909FBFAEB9DC2FCEC" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.425-431.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Onthophagus rutriceps Krikken & Huijbregts 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="431" masterDocId="340AFF82FFCE056F0857FF809E3CFF9F" masterDocTitle="An unusual new Onthophagus from New Guinea (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="431" masterPageNumber="425" pageNumber="426" updateTime="1699185740713" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="91BAE85C552FD2DAB72334B664D28396">
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<mods:title id="E9B2BA513D0963EDF42A3AAC479E0591">An unusual new Onthophagus from New Guinea (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="90FD26484D0348B8EF2B666ADABBD378" type="personal">
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<mods:namePart id="C2033F8C8B789F8AFD22254EC3CA4E9C">Krikken, Jan</mods:namePart>
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<mods:roleTerm id="50A7542CBD393C6ECCE40695EE3861D2">Author</mods:roleTerm>
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<mods:namePart id="C00FFF4A9C661BEE3CEE3B89C7E28866">Huijbregts, Johannes</mods:namePart>
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<mods:typeOfResource id="C9E01D4ABC4E492F9CF8C081ACFF80D5">text</mods:typeOfResource>
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<mods:title id="14C9F2693890D0149928BE5D21F543B5">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="FA465E45B6DFDB4FB5CED216AE74BEC5">
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<mods:date id="D4AF22C9A61458A01BF3BADC9611A721">2012</mods:date>
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<mods:number id="CF8C895E1E6B2DDC5FC4E199964148CB">2012-12-17</mods:number>
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<mods:number id="E3ADB1FD62F66B5DCD2B58C3B8CDFAEC">52</mods:number>
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<mods:classification id="9625DEC37C57A26AB0E651854BBB497B">journal article</mods:classification>
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<mods:identifier id="77BACA14B240CADC971FBDFC31A19B45" type="DOI">10.5281/zenodo.5331298</mods:identifier>
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<treatment id="C83387FAFFCF056909FBFAEB9DC2FCEC" ID-DOI="http://doi.org/10.5281/zenodo.5399031" ID-GBIF-Taxon="185224786" ID-Zenodo-Dep="5399031" LSID="urn:lsid:plazi:treatment:C83387FAFFCF056909FBFAEB9DC2FCEC" httpUri="http://treatment.plazi.org/id/C83387FAFFCF056909FBFAEB9DC2FCEC" lastPageId="6" lastPageNumber="431" pageId="1" pageNumber="426">
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<subSubSection id="08806567FFCF056E09FBFAEB9D3AFA1B" box="[428,774,1387,1412]" pageId="1" pageNumber="426" type="nomenclature">
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<paragraph id="402536ECFFCF056E09FBFAEB9D3AFA1B" blockId="1.[428,774,1387,1412]" box="[428,774,1387,1412]" pageId="1" pageNumber="426">
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<heading id="1B6D8180FFCF056E09FBFAEB9D3AFA1B" bold="true" box="[428,774,1387,1412]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="426" reason="3">
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<emphasis id="72EEEAFEFFCF056E09FBFAEB9D3AFA1B" bold="true" box="[428,774,1387,1412]" pageId="1" pageNumber="426">
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<taxonomicName id="879A4D6FFFCF056E09FBFAEB9C96FA1B" ID-CoL="7XP4K" authority="Krikken & Huijbregts, 2012" authorityName="Krikken & Huijbregts" authorityYear="2012" box="[428,682,1387,1412]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="rutriceps" status="sp. nov.">
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<emphasis id="72EEEAFEFFCF056E09FBFAEB9C96FA1B" bold="true" box="[428,682,1387,1412]" italics="true" pageId="1" pageNumber="426">Onthophagus rutriceps</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="69DD5785FFCF056E0AE6FAEC9D3AFA1B" box="[689,774,1388,1412]" pageId="1" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="08806567FFCF056E0A72FA0A9CB1FA02" box="[549,653,1418,1437]" pageId="1" pageNumber="426" type="description">
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<paragraph id="402536ECFFCF056E0A72FA0A9CB1FA02" blockId="1.[549,653,1418,1437]" box="[549,653,1418,1437]" pageId="1" pageNumber="426">
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(
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<figureCitation id="D8A12A69FFCF056E0A7CFA0A9CBAFA02" box="[555,646,1418,1437]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="4.[151,190,1537,1556]" captionStartId-1="5.[113,152,983,1002]" captionStartId-2="5.[113,152,1494,1513]" captionTargetBox-0="[175,1115,232,1430]" captionTargetBox-1="[124,1080,219,936]" captionTargetBox-2="[113,1088,1124,1472]" captionTargetId-0="figure-11@4.[151,1126,215,1435]" captionTargetId-1="figure-13@5.[110,1090,219,941]" captionTargetId-2="figure-70@5.[113,1088,1124,1474]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="Figs. 1–9. Habitus of Onthophagus rutriceps sp. nov., male holotype (1–5), female paratype (6–9), body length ca. 13 and 10 mm, respectively. Oblique, dorsal, lateral, ventral, frontodorsal views." captionText-1="Figs. 10–15. Body parts and labels of Onthophagus rutriceps sp. nov., male holotype (10–11), female paratype (12–13), male paratype (14, 15, ex NMPC). 10 – head, full-face view; 11–12 – protibia upper side; 13 – metatibia underside; 14 – parameres, lateral view; 15 – text of four labels (Bennigsen male). Scale lines equal 1 mm." captionText-2="Figs. 16–17. Field photos. Onthophagus rutriceps sp. nov., male left (16), female right (17). (Photo T. H. Larsen)." figureDoi-0="http://doi.org/10.5281/zenodo.5331300" figureDoi-1="http://doi.org/10.5281/zenodo.5331304" figureDoi-2="http://doi.org/10.5281/zenodo.5331306" httpUri-0="https://zenodo.org/record/5331300/files/figure.png" httpUri-1="https://zenodo.org/record/5331304/files/figure.png" httpUri-2="https://zenodo.org/record/5331306/files/figure.png" pageId="1" pageNumber="426">Figs. 1–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="08806567FFCF056E0826FA349DAAF988" pageId="1" pageNumber="426" type="materials_examined">
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<paragraph id="402536ECFFCF056E0826FA349DAAF988" blockId="1.[113,1090,1459,1559]" pageId="1" pageNumber="426">
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<emphasis id="72EEEAFEFFCF056E0826FA349ECAFA58" bold="true" box="[113,246,1459,1479]" pageId="1" pageNumber="426">Type material.</emphasis>
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<materialsCitation id="F0F23CB1FFCF056E08ADFA349D67FA7D" ID-GBIF-Occurrence="3340021303" collectingDate="2011-07" collectionCode="RMNH" collectorName="T. Larsen" country="Papua New Guinea" elevation="775" latitude="-5.9577775" location="Transect" longLatPrecision="21" longitude="146.87361" municipality="Huon Peninsula" pageId="1" pageNumber="426" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
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<typeStatus id="9F21884EFFCF056E08ADFA349F72FA59" box="[250,334,1460,1479]" pageId="1" pageNumber="426" type="holotype">HOLOTYPE</typeStatus>
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: male (
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<collectionCode id="268BAE29FFCF056E09DAFA349FEDFA58" box="[397,465,1460,1479]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="426" type="Museum">RMNH</collectionCode>
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), from “
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<collectingCountry id="388D767CFFCF056E0A40FA349CD6FA58" box="[535,746,1460,1479]" name="Papua New Guinea" pageId="1" pageNumber="426">PAPUA NEW GUINEA</collectingCountry>
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:
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<collectingRegion id="825EF80EFFCF056E0AA5FA349D57FA58" box="[754,875,1460,1479]" country="Papua New Guinea" name="Morobe" pageId="1" pageNumber="426">Morobe Prov.</collectingRegion>
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: Saruwaged Mtns.,
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<collectingMunicipality id="A041AC96FFCF056E0C46FA349EFAFA7D" pageId="1" pageNumber="426">Huon Peninsula</collectingMunicipality>
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, YUS
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<location id="45456037FFCF056E0954FA4F9F72FA7D" LSID="urn:lsid:plazi:treatment:C83387FAFFCF056909FBFAEB9DC2FCEC:45456037FFCF056E0954FA4F9F72FA7D" box="[259,334,1487,1506]" country="Papua New Guinea" latitude="-5.9577775" longLatPrecision="21" longitude="146.87361" municipality="Huon Peninsula" name="Transect" pageId="1" pageNumber="426" stateProvince="Morobe">Transect</location>
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,
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<geoCoordinate id="25AE502BFFCF056E090EFA4F9F8DFA7D" box="[345,433,1485,1506]" degrees="5" direction="south" minutes="57" orientation="latitude" pageId="1" pageNumber="426" precision="15" seconds="28" value="-5.9577775">
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5
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<superScript id="B7EF9BA4FFCF056E0933FA4D9F56FA46" attach="none" box="[356,362,1485,1497]" fontSize="5" pageId="1" pageNumber="426">o</superScript>
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57’28”S
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</geoCoordinate>
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<geoCoordinate id="25AE502BFFCF056E09EEFA4F9C16FA7D" box="[441,554,1485,1506]" degrees="146" direction="east" minutes="52" orientation="longitude" pageId="1" pageNumber="426" precision="15" seconds="25" value="146.87361">
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146
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<superScript id="B7EF9BA4FFCF056E098EFA4D9FE3FA46" attach="none" box="[473,479,1485,1497]" fontSize="5" pageId="1" pageNumber="426">o</superScript>
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52’25”E
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</geoCoordinate>
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,
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<quantity id="87629B09FFCF056E0A62FA4F9CB3FA7D" box="[565,655,1487,1506]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.5" metricValueMin="6.0" pageId="1" pageNumber="426" unit="m" value="775.0" valueMax="950.0" valueMin="600.0">
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<elevation id="CBB7D1DFFFCF056E0A62FA4F9CB3FA7D" box="[565,655,1487,1506]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.5" metricValueMin="6.0" pageId="1" pageNumber="426" unit="m" value="775.0" valueMax="950.0" valueMin="600.0">600-950m</elevation>
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||
</quantity>
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||
,
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||
<date id="3424102CFFCF056E0ACDFA4F9CCDFA7D" box="[666,753,1487,1506]" pageId="1" pageNumber="426" value="2011-07">
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||
<collectingDate id="2460E9C4FFCF056E0ACDFA4F9CCDFA7D" box="[666,753,1487,1506]" pageId="1" pageNumber="426" value="2011-07">July 2011</collectingDate>
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</date>
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||
,
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<collectorName id="ED6F533AFFCF056E0AABFA4F9D6EFA7D" box="[764,850,1487,1506]" pageId="1" pageNumber="426">T. Larsen</collectorName>
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||
”
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</materialsCitation>
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||
.
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||
<materialsCitation id="F0F23CB1FFCF056E0B31FA4F9C04FA63" ID-GBIF-Occurrence="3340021302" collectingDate="2011-07" collectionCode="NAIC, CMNC" collectorName="T. Larsen" country="Papua New Guinea" elevation="775" latitude="-5.9577775" location="Transect" longLatPrecision="21" longitude="146.87361" municipality="Huon Peninsula" pageId="1" pageNumber="426" specimenCount="13" specimenCount-male="13" stateProvince="Morobe" typeStatus="paratype">
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||
<typeStatus id="9F21884EFFCF056E0B31FA4F9D81FA7F" box="[870,957,1487,1506]" pageId="1" pageNumber="426" type="paratype">PARATYPES</typeStatus>
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:
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<specimenCount id="569CFD65FFCF056E0B9CFA4F9A26FA7D" box="[971,1050,1487,1506]" pageId="1" pageNumber="426" type="male">13 males</specimenCount>
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and females, same label data as holotype (
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<collectionCode id="268BAE29FFCF056E09E7FA699FDBFA63" box="[432,487,1513,1532]" country="Papua New Guinea" httpUri="http://grbio.org/cool/7mi7-dyvm" name="National Agricultural Insect Collection" pageId="1" pageNumber="426">NAIC</collectionCode>
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,
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<collectionCode id="268BAE29FFCF056E09B8FA699C0EFA63" box="[495,562,1513,1532]" pageId="1" pageNumber="426">CMNC</collectionCode>
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)
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</materialsCitation>
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||
;
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<materialsCitation id="F0F23CB1FFCF056E0A68FA699DAEF988" ID-GBIF-Occurrence="3340021301" collectionCode="NMPC" collectorName="v. Bennigsen" country="Papua New Guinea" location="Kani Mountains" municipality="German New Guinea" pageId="1" pageNumber="426" specimenCount="1" specimenCount-male="1" stateProvince="Madang" typeStatus="paratype">
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<specimenCount id="569CFD65FFCF056E0A68FA699C47FA63" box="[575,635,1513,1532]" pageId="1" pageNumber="426" type="male">1 male</specimenCount>
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||
, New Guinea [probably former
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<collectingMunicipality id="A041AC96FFCF056E0BD0FA699A02FA63" box="[903,1086,1513,1533]" pageId="1" pageNumber="426">German New Guinea</collectingMunicipality>
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||
: now
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<collectingRegion id="825EF80EFFCF056E08CCF9849F24F988" box="[155,280,1540,1559]" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="426">Madang Prov.</collectingRegion>
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||
:
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<location id="45456037FFCF056E0949F9849F94F988" LSID="urn:lsid:plazi:treatment:C83387FAFFCF056909FBFAEB9DC2FCEC:45456037FFCF056E0949F9849F94F988" box="[286,424,1540,1559]" country="Papua New Guinea" municipality="German New Guinea" name="Kani Mountains" pageId="1" pageNumber="426" stateProvince="Madang">Kani Mountains</location>
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(cf.
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<figureCitation id="D8A12A69FFCF056E0987F9849C32F988" box="[464,526,1540,1559]" captionStart="Figs" captionStartId="5.[113,152,983,1002]" captionTargetBox="[124,1080,219,936]" captionTargetId="figure-13@5.[110,1090,219,941]" captionTargetPageId="5" captionText="Figs. 10–15. Body parts and labels of Onthophagus rutriceps sp. nov., male holotype (10–11), female paratype (12–13), male paratype (14, 15, ex NMPC). 10 – head, full-face view; 11–12 – protibia upper side; 13 – metatibia underside; 14 – parameres, lateral view; 15 – text of four labels (Bennigsen male). Scale lines equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331304" httpUri="https://zenodo.org/record/5331304/files/figure.png" pageId="1" pageNumber="426">Fig. 15</figureCitation>
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||
), year ca. 1900],
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||
<collectorName id="ED6F533AFFCF056E0AF5F9849D2CF988" box="[674,784,1540,1559]" pageId="1" pageNumber="426">v. Bennigsen</collectorName>
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||
[lgt.] (
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||
<collectionCode id="268BAE29FFCF056E0B1CF9849DB0F988" box="[843,908,1540,1559]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="426">NMPC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="08806567FFCC056D08C0FF5B9CDEFCAC" pageId="2" pageNumber="427" type="diagnosis">
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||
<paragraph id="402536ECFFCC056D08C0FF5B9CDEFCAC" blockId="2.[151,1127,219,1555]" pageId="2" pageNumber="427">
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||
<emphasis id="72EEEAFEFFCC056D08C0FF5B9F30FF6C" bold="true" box="[151,268,219,243]" pageId="2" pageNumber="427">Diagnosis.</emphasis>
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Head of male with tip slightly bent upward, extended as “shovel”, apical crest shortly rounded; entire head surface lacking protrusions. Head of female shorter, with bisinuate, shortly reflexed clypeal apex and a pair of simple, pillar-like erect interocular horns; clypeofrontal ridge distinct, arcuate. Intercoxal lobe of metasternum just behind mesosternum with cariniform prow-shaped protrusion in both sexes (rounded in profile, pointed in ventral view). Head brown-black, remainder of body predominantly orange-yellow or more rufous, with symmetric brown-black markings. Pronotum of major males with forwardcurved postdiscal projection (“handle”), anterior pronotal declivity broadly deplanate (but surface not delimited by extra ridges). Anterolateral angles of pronotum acute. Pronotal base medially obtusely angular. Elytral striae virtually impunctate, odd discal interstriae convex, intervening even discal interstriae less convex and broader; interstrial surface (like most of dorsum) with indistinct micropunctation only. Legs robust. External protibial denticles of female not separated by serration, separating emarginations widely, evenly rounded; proximal serration limited; note other sexual differences in protibial shape (see below). Apical crests of meso- and metatibiae multi-sinuate-dentate, fringed with setae, lacking the more common shorter spines. Inner side of male protibial apex projecting angularly. Eyes darkish, foramina narrow, widely separated (interocular distance 10–11 single eye-widths). Dorsum virtually glabrous. Known body length
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<quantity id="87629B09FFCC056D0982FD7C9C00FC8C" box="[469,572,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="2" pageNumber="427" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
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. Parameres (
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||
<figureCitation id="D8A12A69FFCC056D0A96FD7B9D30FC8B" box="[705,780,763,788]" captionStart="Figs" captionStartId="5.[113,152,983,1002]" captionTargetBox="[124,1080,219,936]" captionTargetId="figure-13@5.[110,1090,219,941]" captionTargetPageId="5" captionText="Figs. 10–15. Body parts and labels of Onthophagus rutriceps sp. nov., male holotype (10–11), female paratype (12–13), male paratype (14, 15, ex NMPC). 10 – head, full-face view; 11–12 – protibia upper side; 13 – metatibia underside; 14 – parameres, lateral view; 15 – text of four labels (Bennigsen male). Scale lines equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331304" httpUri="https://zenodo.org/record/5331304/files/figure.png" pageId="2" pageNumber="427">Fig. 14</figureCitation>
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) tapering, bent downward, distal (downward) upper side (“roof”) narrow, parallel-sided.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08806567FFCC056908C0FCBB9FE5FD0C" lastPageId="6" lastPageNumber="431" pageId="2" pageNumber="427" type="description">
|
||
<paragraph id="402536ECFFCC056D08C0FCBB9D3AFC4C" blockId="2.[151,1127,219,1555]" pageId="2" pageNumber="427">
|
||
<emphasis id="72EEEAFEFFCC056D08C0FCBB9F27FCCC" bold="true" box="[151,283,827,851]" pageId="2" pageNumber="427">Description</emphasis>
|
||
(
|
||
<typeStatus id="9F21884EFFCC056D097FFCBB9FB6FCCC" box="[296,394,827,851]" pageId="2" pageNumber="427" type="holotype">holotype</typeStatus>
|
||
, male). Body length ca.
|
||
<quantity id="87629B09FFCC056D0ADEFCBC9CEAFCCC" box="[649,726,828,852]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="427" unit="mm" value="13.0">13 mm</quantity>
|
||
. Habitus robust, convex; colour generally yellow to orange, with symmetric brown-black markings; largely shiny, several parts more or less matt (sericeous, microreticulate). Dorsal side and pygidium virtually glabrous, seemingly almost impunctate (micropunctation and microsetae present, but inconspicuous); ventral side and legs with numerous long, yellowish setae.
|
||
</paragraph>
|
||
<paragraph id="402536ECFFCC056D08E2FC5B9C24FB4B" blockId="2.[151,1127,219,1555]" pageId="2" pageNumber="427">Head black, shiny, with numerous inconspicuous micropunctures. Clypeus shovel-like: elongate forward, surface concave behind apex, sides marginate (well-defined margin finely punctate); lateral border virtually straight, converging to slightly reflexed tip, apical crest shortly rounded; clypeofrontal transition lacking ridge. Clypeogenal transition at border virtually continuous, fine suture curving from border onto frons; genal border very widely rounded, outer margin (anterior corner) crowded with fine punctation. Eye foramina narrowly elliptic, with ca 11 facet rows across widest point. Ratio interocular distance / maximum (transverse, single) eye width ca 10.</paragraph>
|
||
<paragraph id="402536ECFFCC056D08E2FB5C9CE9F98C" blockId="2.[151,1127,219,1555]" pageId="2" pageNumber="427">Pronotum moderately convex, broadly deplanate in front of short, forward-curved postdiscal projection; this projection slightly tapering, apex rounded, upper side of projection with slight longitudinal impression. Anterolateral surface of pronotum gradually, moderately declivous to very shallowly concave corner, anterolateral angle acute; anterior and lateral border of pronotum narrowly marginate; anterior section of lateral border virtually straight (full-face view), posterior section slightly sinuate; posterolateral angle rounded off to generally widely rounded, immarginate pronotal base; median section of base obtusely angular, finely marginate. Most of pronotal surface shiny orange, postdiscal projection and its immediate periphery black, basomedian surface also narrowly black; margins of pronotum narrowly infuscated; much of surface with inconspicuous micropunctation.</paragraph>
|
||
<paragraph id="402536ECFFCD056C08C7FF5B9D49FE0C" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="428">Elytra broad, largely yellow, more or less shiny; base and apex with broad transverse brownblack marking; scutellum indistinct, shape of elytral base and apex unmodified; epipleuron black, orange midway, glabrous. Elytra with 8 fine, distinct, straight to slightly curved striae, stria 7 hardly sinuate in front, extending to shiny humeral umbone; strial punctures obsolescent, hardly distinct. Elytral interstriae 1, 3, 5, 7, 8 convex, more shiny than less convex and broader interstriae 2, 4 and 6; interstrial micropunctation hardly distinct.</paragraph>
|
||
<paragraph id="402536ECFFCD056C08C7FE1C9D51FC8C" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="428">Antenna with 9 antennomeres; club lamellae unmodified, yellow-brown, covered with light pubescence; scapus in front with fine, longitudinal ridge. Mentum in front deeply excised; labial palpi with very small segment 4 distinct. Prosternum, mesosternum, mesepimeron black. Propectoral sides with fine coxal-marginal ridge widely curving forward, but not reaching border; surface abundantly hemipunctate-setose, laterally yellow. Intercoxal lobe of metasternum distinctly prow-shaped in front, flanked by numerous fine seta-bearing punctures; metasternal disc shiny, yellowish, virtually glabrous, smooth (micropunctation and their microsetae aside); flanks and adjacent metepisterna sericeous, and abundantly, finely hemipunctate-setose. Abdominal ventrites largely yellow, subsericeous, sides with row of fine, seta-bearing punctures; anal ventrite glabrous, medially black. Pygidium very slightly convex, yellow, shiny, apex black; margins narrowly raised, well defined, black; pygidial base with fine transverse ridge; surface with inconspicuous micropunctation.</paragraph>
|
||
<paragraph id="402536ECFFCD056C08C7FC9C9C06FB4C" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="428">
|
||
Legs robust, tibiae and tarsi generally shiny brown, femora largely yellow (base and apex infuscated); coxae largely black. Protibia with 3+1 broad external denticles, separated by some serration; proximal serration consisting of ca. 5 separate small denticles, remainder of proximal edge continuous; apico-external denticle oblique to tibial axis; terminal spur slightly curved, elongate-tapering; protibial underside with longitudinal crest, ending apico-internally at accrete, projecting angle; protarsus unmodified. Profemoral underside with numerous punctures bearing long seta, mesofemoral underside with fewer seta-bearing punctures, metafemoral underside glabrous. Meso- and metatibiae dilated distad to oblong, sinuatedentate apical crest, which is fringed with long setae; external tibial side with 3 distinct sets of seta-bearing fossorial protrusions, remaining surface with numerous long setae. Terminal spurs on meso- and metatibiae elongate-tapering, their tips blunted-curved (metatibial spurs of
|
||
<typeStatus id="9F21884EFFCD056C08DBFBFB9ED4FB0C" box="[140,232,1147,1171]" pageId="3" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
largely worn away). Tarsi generally slender, with unmodified sickle-shaped claws; meso- and metatarsomeres 1 long, straight, unmodified; length proportions of metatarsomeres
|
||
<quantity id="87629B09FFCD056C0826FB3B9E89FB4C" box="[113,181,1211,1235]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="3" pageNumber="428" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
|
||
approximate integers: 30/10/7/5/12.
|
||
</paragraph>
|
||
<paragraph id="402536ECFFCD056C08C7FB5B9FFCFAAC" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="428">
|
||
Body part measurements of male
|
||
<typeStatus id="9F21884EFFCD056C0A5DFB5B9C54FB6C" box="[522,616,1243,1267]" pageId="3" pageNumber="428" type="holotype">holotype</typeStatus>
|
||
in mm (dorsal view): Maximum width of head 3.3. Median length of pronotum 4.6, maximum width 6.2. Sutural length of elytra 4.4, maximum width combined 6.6.
|
||
</paragraph>
|
||
<paragraph id="402536ECFFCD056C08C7FABB9C67FA2C" blockId="3.[113,1089,219,1587]" pageId="3" pageNumber="428">
|
||
Body part measurements of female
|
||
<typeStatus id="9F21884EFFCD056C0A47FABC9C50FACB" box="[528,620,1340,1364]" pageId="3" pageNumber="428" type="paratype">paratype</typeStatus>
|
||
pictured (dorsal view): Maximum width of head 3.2. Median length of pronotum 4.2, maximum width 5.8. Sutural length of elytra 4.3, maximum width combined 6.3. Length proportions of terminal spur
|
||
<emphasis id="72EEEAFEFFCD056C0B19FAFD9DAEFA0B" box="[846,914,1405,1428]" italics="true" pageId="3" pageNumber="428">versus</emphasis>
|
||
metatarsomeres
|
||
<quantity id="87629B09FFCD056C0826FA1B9E89FA2C" box="[113,181,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="3" pageNumber="428" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
|
||
approximate integers: 25/32/10/6/5/11.
|
||
</paragraph>
|
||
<paragraph id="402536ECFFCD05690826FA3B9C4BFE2C" blockId="3.[113,1089,219,1587]" lastBlockId="6.[151,1128,219,883]" lastPageId="6" lastPageNumber="431" pageId="3" pageNumber="428">
|
||
<emphasis id="72EEEAFEFFCD056C0826FA3B9FD0FA4C" bold="true" box="[113,492,1467,1492]" pageId="3" pageNumber="428">Variation and sexual dimorphism.</emphasis>
|
||
Females with pair of long, erect pillar-like horns between eyes, tip of horns rounded off, horns circular on cross-section, their base set slightly away from eye foraminal edge (distance from eyes virtually equal to distance between them). Clypeus short, edge generally rounded, but tip distinctly bisinuate, margin narrowly reflexed. Clypeofrontal ridge distinct, widely evenly arcuate, angularly reaching distinct clypeogenal suture. Clypeal surface transversely braided rugulate, frons in front of horns transversely rugulate-punctate. Pronotal discal protrusion shifted forward, somewhat transverse, not projecting forward as “handle”; anterior discal surface more convex than in male; pronotal punctation minute, generally more distinct than in male. Female protibia lacking apico-internal projection; upper side with well defined depressed parts externally delimited by extra ridge along non-serrate inter-dental emarginations.
|
||
</paragraph>
|
||
<caption id="14E56664FFCA056B08C0F9819D7BF9B0" ID-DOI="http://doi.org/10.5281/zenodo.5331300" ID-Zenodo-Dep="5331300" httpUri="https://zenodo.org/record/5331300/files/figure.png" pageId="4" pageNumber="429" startId="4.[151,190,1537,1556]" targetBox="[175,1115,232,1430]" targetPageId="4">
|
||
<paragraph id="402536ECFFCA056B08C0F9819D7BF9B0" blockId="4.[151,1127,1537,1583]" pageId="4" pageNumber="429">
|
||
Figs. 1–9. Habitus of
|
||
<taxonomicName id="879A4D6FFFCA056B0918F9829C29F98B" authority="Krikken & Huijbregts, 2012" authorityName="Krikken & Huijbregts" authorityYear="2012" box="[335,533,1538,1556]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="429" phylum="Arthropoda" rank="species" species="rutriceps" status="sp. nov.">
|
||
<emphasis id="72EEEAFEFFCA056B0918F9829C29F98B" box="[335,533,1538,1556]" italics="true" pageId="4" pageNumber="429">Onthophagus rutriceps</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69DD5785FFCA056B0A4DF9829C62F98A" box="[538,606,1538,1557]" pageId="4" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype (1–5), female paratype (6–9), body length ca. 13 and 10 mm, respectively. Oblique, dorsal, lateral, ventral, frontodorsal views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="14E56664FFCB056A0826FC579A3DFBBF" ID-DOI="http://doi.org/10.5281/zenodo.5331304" ID-Zenodo-Dep="5331304" httpUri="https://zenodo.org/record/5331304/files/figure.png" pageId="5" pageNumber="430" startId="5.[113,152,983,1002]" targetBox="[124,1080,219,936]" targetPageId="5">
|
||
<paragraph id="402536ECFFCB056A0826FC579A3DFBBF" blockId="5.[113,1088,983,1056]" pageId="5" pageNumber="430">
|
||
Figs. 10–15. Body parts and labels of
|
||
<taxonomicName id="879A4D6FFFCB056A0993FC589CB7FC75" authority="Krikken & Huijbregts, 2012" authorityName="Krikken & Huijbregts" authorityYear="2012" box="[452,651,984,1002]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="430" phylum="Arthropoda" rank="species" species="rutriceps" status="sp. nov.">
|
||
<emphasis id="72EEEAFEFFCB056A0993FC589CB7FC75" box="[452,651,984,1002]" italics="true" pageId="5" pageNumber="430">Onthophagus rutriceps</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69DD5785FFCB056A0AC4FC579CE6FC75" box="[659,730,983,1002]" pageId="5" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype (10–11), female paratype (12–13), male paratype (14, 15, ex NMPC). 10 – head, full-face view; 11–12 – protibia upper side; 13 – metatibia underside; 14 – parameres, lateral view; 15 – text of four labels (Bennigsen male). Scale lines equal 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="14E56664FFCB056A0826FA569A00FA75" ID-DOI="http://doi.org/10.5281/zenodo.5331306" ID-Zenodo-Dep="5331306" box="[113,1084,1494,1514]" httpUri="https://zenodo.org/record/5331306/files/figure.png" pageId="5" pageNumber="430" startId="5.[113,152,1494,1513]" targetBox="[113,1088,1124,1472]" targetPageId="5">
|
||
<paragraph id="402536ECFFCB056A0826FA569A00FA75" blockId="5.[113,1084,1494,1514]" box="[113,1084,1494,1514]" pageId="5" pageNumber="430">
|
||
Figs. 16–17. Field photos.
|
||
<taxonomicName id="879A4D6FFFCB056A0901FA579C27FA76" authority="Krikken & Huijbregts, 2012" authorityName="Krikken & Huijbregts" authorityYear="2012" box="[342,539,1495,1513]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="430" phylum="Arthropoda" rank="species" species="rutriceps" status="sp. nov.">
|
||
<emphasis id="72EEEAFEFFCB056A0901FA579C27FA76" box="[342,539,1495,1513]" italics="true" pageId="5" pageNumber="430">Onthophagus rutriceps</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69DD5785FFCB056A0A77FA579C59FA75" box="[544,613,1495,1514]" pageId="5" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male left (16), female right (17). (Photo T. H. Larsen).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="402536ECFFC8056908E2FE3B9F1FFDCC" blockId="6.[151,1128,219,883]" pageId="6" pageNumber="431">Male clypeal extension and length of pronotal projection less in smaller males, real minors with reduced, simply prominent discal protrusion. Extension of elytral brown-black markings varies, basal and apical markings may be broadly connected over lateral declivity. Pronotum may be entirely yellow-orange. Frontal horns of minor females shorter, but still very distinct.</paragraph>
|
||
<paragraph id="402536ECFFC8056908E2FDDB9FE5FD0C" blockId="6.[151,1128,219,883]" pageId="6" pageNumber="431">
|
||
Colours vary slightly: yellow may be more rufous, black may be brownish-black. Approximate body length
|
||
<quantity id="87629B09FFC80569093BFDFC9FE9FD0C" box="[364,469,636,660]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="6" pageNumber="431" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08806567FFC8056908C0FD1B9A35FCAB" pageId="6" pageNumber="431" type="biology_ecology">
|
||
<paragraph id="402536ECFFC8056908C0FD1B9A35FCAB" blockId="6.[151,1128,219,883]" pageId="6" pageNumber="431">
|
||
<emphasis id="72EEEAFEFFC8056908C0FD1B9ECDFD2C" bold="true" box="[151,241,667,691]" pageId="6" pageNumber="431">Biology.</emphasis>
|
||
<taxonomicName id="879A4D6FFFC8056908A2FD1D9FDBFD2B" authority="Krikken & Huijbregts, 2012" authorityName="Krikken & Huijbregts" authorityYear="2012" box="[245,487,669,692]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="431" phylum="Arthropoda" rank="species" species="rutriceps" status="sp. nov.">
|
||
<emphasis id="72EEEAFEFFC8056908A2FD1D9FDBFD2B" box="[245,487,669,692]" italics="true" pageId="6" pageNumber="431">Onthophagus rutriceps</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="69DD5785FFC8056909BCFD1C9C07FD2C" box="[491,571,668,691]" pageId="6" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
|
||
occurred from
|
||
<quantity id="87629B09FFC805690A80FD1B9D70FD2C" box="[727,844,667,691]" metricMagnitude="2" metricUnit="m" metricValue="8.25" metricValueMax="9.5" metricValueMin="7.0" pageId="6" pageNumber="431" unit="m" value="825.0" valueMax="950.0" valueMin="700.0">700–950 m</quantity>
|
||
a. s.l. in primary rainforest. Although the species was most abundant at about
|
||
<quantity id="87629B09FFC805690AF0FD3C9CD0FD4C" box="[679,748,700,724]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="431" unit="m" value="700.0">700 m</quantity>
|
||
, it was not collected at lower elevation (
|
||
<quantity id="87629B09FFC805690898FD5B9F24FD6B" box="[207,280,731,756]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="431" unit="m" value="600.0">600 m</quantity>
|
||
), which may have been due to habitat disturbance. It did not occur at all above
|
||
<specimenCount id="569CFD65FFC8056908C0FD7B9EDDFC8C" box="[151,225,763,787]" pageId="6" pageNumber="431" type="generic">
|
||
<quantity id="87629B09FFC8056908C0FD7B9EE1FC8C" box="[151,221,763,787]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="431" unit="m" value="950.0">950 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Despite trapping with rotting fruit and carrion at the same sites, the species was only attracted to pitfall traps baited with human dung (T. H. Larsen, pers. comm. 2012).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08806567FFC8056908C0FCBC9D41FCCB" box="[151,893,827,852]" pageId="6" pageNumber="431" type="etymology">
|
||
<paragraph id="402536ECFFC8056908C0FCBC9D41FCCB" blockId="6.[151,1128,219,883]" box="[151,893,827,852]" pageId="6" pageNumber="431">
|
||
<emphasis id="72EEEAFEFFC8056908C0FCBC9F24FCCB" bold="true" box="[151,280,828,852]" pageId="6" pageNumber="431">Etymology.</emphasis>
|
||
Latin contraction, meaning shovel-headed
|
||
<emphasis id="72EEEAFEFFC805690AB2FCBD9D41FCCB" box="[741,893,829,852]" italics="true" pageId="6" pageNumber="431">
|
||
<taxonomicName id="879A4D6FFFC805690AB2FCBD9D44FCCB" authorityName="Latreille" authorityYear="1802" box="[741,888,829,852]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="431" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08806567FFC8056908C0FCDB9DC2FCEC" box="[151,1022,859,883]" pageId="6" pageNumber="431" type="distribution">
|
||
<paragraph id="402536ECFFC8056908C0FCDB9DC2FCEC" blockId="6.[151,1128,219,883]" box="[151,1022,859,883]" pageId="6" pageNumber="431">
|
||
<emphasis id="72EEEAFEFFC8056908C0FCDB9F16FCEC" bold="true" box="[151,298,859,883]" pageId="6" pageNumber="431">Distribution.</emphasis>
|
||
<collectingCountry id="388D767CFFC805690967FCDC9C3EFCEC" box="[304,514,859,883]" name="Papua New Guinea" pageId="6" pageNumber="431">Papua New Guinea</collectingCountry>
|
||
: Huon Peninsula and possibly adjacent regions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |