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<document id="B3C1911E18AF7CA0534F387EF8122A81" ID-CLB-Dataset="113069" ID-DOI="10.11646/zootaxa.5111.1.1" ID-GBIF-Dataset="f3992bb2-9962-4f3c-8f5b-131cfe552c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6340729" ID-ZooBank="836B0F69-037C-4D0F-80DB-94FE454F48E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646828425946" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2022" docId="C822F213FF6124DBCB88A25571D9226A" docLanguage="en" docName="zootaxa.5111.1.1.pdf" docOrigin="Zootaxa 5111 (1)" docSource="http://zoobank.org/836b0f69-037c-4d0f-80db-94fe454f48e3" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tribasodites vertexalis Yin 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="173" masterDocId="341B8A6BFFCB2477CB1FA14C7567265F" masterDocTitle="The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="171" updateTime="1699317591893" updateUser="ExternalLinkService">
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<mods:title id="1CA17EAFB5EF9523DB17EE3D2EBD235B">The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="548BAB01E2A116AB57FAE09690BB905C">Yin, Zi-Wei</mods:namePart>
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<mods:date id="EAE027186D526D10987702B05D1AB1ED">2022</mods:date>
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<mods:number id="757997D1E5A374158C5B7E5DB07C1058">2022-03-09</mods:number>
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<mods:number id="69A2E10F15BD8FF9A60200874DB7B7AF">5111</mods:number>
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<mods:classification id="55CC35DA10671C30C25C259F3650C898">journal article</mods:classification>
<mods:identifier id="5659781CBFE99A13857595250FD830F7" type="CLB-Dataset">113069</mods:identifier>
<mods:identifier id="9DA336DB19D7BFCCE63FE83BDA38DE8E" type="DOI">10.11646/zootaxa.5111.1.1</mods:identifier>
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<treatment id="C822F213FF6124DBCB88A25571D9226A" ID-DOI="http://doi.org/10.5281/zenodo.6964434" ID-GBIF-Taxon="198579348" ID-Zenodo-Dep="6964434" LSID="urn:lsid:plazi:treatment:C822F213FF6124DBCB88A25571D9226A" httpUri="http://treatment.plazi.org/id/C822F213FF6124DBCB88A25571D9226A" lastPageId="172" lastPageNumber="173" pageId="170" pageNumber="171">
<subSubSection id="0891108EFF6124DDCB88A255776D256C" box="[151,522,793,820]" pageId="170" pageNumber="171" type="nomenclature">
<paragraph id="40344305FF6124DDCB88A255776D256C" blockId="170.[151,597,793,892]" box="[151,522,793,820]" pageId="170" pageNumber="171">
<heading id="1B7CF469FF6124DDCB88A255776D256C" bold="true" box="[151,522,793,820]" fontSize="11" level="2" pageId="170" pageNumber="171" reason="4">
<emphasis id="72FF9F17FF6124DDCB88A255776D256C" bold="true" box="[151,522,793,820]" pageId="170" pageNumber="171">
<taxonomicName id="878B3886FF6124DDCB88A25574C1256B" ID-CoL="9GDH8" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[151,422,793,820]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="vertexalis" status="sp. nov.">
<emphasis id="72FF9F17FF6124DDCB88A25574C1256B" bold="true" box="[151,422,793,820]" italics="true" pageId="170" pageNumber="171">Tribasodites vertexalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69CC226CFF6124DDCAB2A255776D256C" box="[429,522,793,819]" pageId="170" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6124DDCB88A27177322526" pageId="170" pageNumber="171" type="description">
<paragraph id="40344305FF6124DDCB88A27174E32508" blockId="170.[151,597,793,892]" box="[151,388,829,855]" pageId="170" pageNumber="171">
(
<figureCitation id="D8B05F80FF6124DDCB80A27175912508" box="[159,246,829,855]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="170" pageNumber="171">Figs 73</figureCitation>
,
<figureCitation id="D8B05F80FF6124DDCA1DA27174542508" box="[258,307,829,855]" captionStart="FIGURE 86" captionStartId="188.[151,250,1881,1905]" captionTargetBox="[147,1439,181,1857]" captionTargetId="figure-19@188.[147,1439,181,1857]" captionTargetPageId="188" captionText="FIGURE 86. Distribution of Tibetan Batrisini. A. Tribasodites dilophus sp. nov. (circle), T. elongatus sp. nov. (triangle) and T. grandiceps sp. nov. (square). B. T. gyirong sp. nov. (circle), T. kiypu sp. nov. (triangle) and T. mirabilis sp. nov. (square). C. T. prolixicornis sp. nov. (circle), T. Tribasodites pugiunculus sp. nov. (triangle) T. vertexalis sp. nov. (square) and T. yatung sp. nov. (diamond)." figureDoi="http://doi.org/10.5281/zenodo.6340942" httpUri="https://zenodo.org/record/6340942/files/figure.png" pageId="170" pageNumber="171">86C</figureCitation>
,
<figureCitation id="D8B05F80FF6124DDCA21A271741B2508" box="[318,380,829,855]" captionStart="FIGURE 100" captionStartId="202.[151,250,1591,1615]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure-19@202.[151,1436,181,1567]" captionTargetPageId="202" captionText="FIGURE 100. Environment of and collecting sites at Nyingchi area (A–E) and Chamdo (ăƀ) (F). A. 80K. B. 96K. C–E. Xiachayu (T察ø). F. pass of Jueba Mountain (ẅ巴山口), Markam (Ẽ康). Photo Credit: A by Zhong Peng; B by Xiao-Bin Song; F by Wen-Xuan Bi." figureDoi="http://doi.org/10.5281/zenodo.6340975" httpUri="https://zenodo.org/record/6340975/files/figure.png" pageId="170" pageNumber="171">100B</figureCitation>
)
</paragraph>
<paragraph id="40344305FF6124DDCB88A22D77322526" blockId="170.[151,597,793,892]" box="[151,597,865,892]" pageId="170" pageNumber="171">
Chinese common name:
<emphasis id="72FF9F17FF6124DDCAB2A22E77322526" bold="true" box="[429,597,866,889]" pageId="170" pageNumber="171">ḔĦûḃà甲</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6124DDCB88A2E876D525BD" pageId="170" pageNumber="171" type="materials_examined">
<paragraph id="40344305FF6124DDCB88A2E876D525BD" blockId="170.[151,1437,932,2039]" pageId="170" pageNumber="171">
<emphasis id="72FF9F17FF6124DDCB88A2E8745825E1" bold="true" box="[151,319,932,958]" pageId="170" pageNumber="171">Type material</emphasis>
(
<materialsCitation id="F0E34958FF6124DDCA4DA2E876D525BD" ID-GBIF-Occurrence="3879233490" collectingDate="2017-03-17" collectionCode="SNUC" collectorName="X. - B. Song" country="China" county="Medog County" elevation="1460" latitude="29.583559" location="China" longLatPrecision="1" longitude="95.46618" pageId="170" pageNumber="171" specimenCount="2" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<specimenCount id="568D888CFF6124DDCA4DA2E874EF25E0" box="[338,392,932,959]" pageId="170" pageNumber="171" type="generic">1 ex.</specimenCount>
).
<emphasis id="72FF9F17FF6124DDCA80A2E877D125E1" bold="true" box="[415,694,932,958]" pageId="170" pageNumber="171">
<typeStatus id="9F30FDA7FF6124DDCA80A2E8772525E1" box="[415,578,932,958]" pageId="170" pageNumber="171" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="389C0395FF6124DDC950A2E877D625E1" box="[591,689,932,958]" name="China" pageId="170" pageNumber="171">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="568D888CFF6124DDC9A1A2E977B225E0" box="[702,725,933,959]" pageId="170" pageNumber="171" type="male">♂</specimenCount>
, ‘
<location id="455415DEFF6124DDC9F9A2E8765525E1" LSID="urn:lsid:plazi:treatment:C822F213FF6124DBCB88A25571D9226A:455415DEFF6124DDC9F9A2E8765525E1" box="[742,818,932,958]" country="China" county="Medog County" latitude="29.583559" longLatPrecision="1" longitude="95.46618" name="China" pageId="170" pageNumber="171" stateProvince="Xizang">China</location>
:
<collectingRegion id="824F8DE7FF6124DDC822A2E876F625E1" box="[829,913,932,958]" country="China" name="Xizang" pageId="170" pageNumber="171">Xizang</collectingRegion>
,
<collectingCounty id="A9553B89FF6124DDC882A2E8712F25E1" box="[925,1096,932,958]" pageId="170" pageNumber="171">Mêdog County</collectingCounty>
, 96K,
<quantity id="8773EEE0FF6124DDCF8CA2E8718D25E0" box="[1171,1258,932,959]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="170" pageNumber="171" unit="m" value="1460.0">
<elevation id="CBA6A436FF6124DDCF8CA2E8718D25E0" box="[1171,1258,932,959]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="170" pageNumber="171" unit="m" value="1460.0">1460 m</elevation>
</quantity>
,
<geoCoordinate id="25BF25C2FF6124DDCFE8A2E870FF25E0" box="[1271,1432,932,959]" degrees="29" direction="north" minutes="35" orientation="latitude" pageId="170" pageNumber="171" precision="1" seconds="0.81" value="29.583559">29°35’0.81”N</geoCoordinate>
,
<geoCoordinate id="25BF25C2FF6124DDCB88A284742425BC" box="[151,323,968,995]" degrees="95" direction="east" minutes="27" orientation="longitude" pageId="170" pageNumber="171" precision="1" seconds="58.26" value="95.46618">95°27’58.26”E</geoCoordinate>
,
<date id="343565C5FF6124DDCA51A28474AD25BD" box="[334,458,968,994]" pageId="170" pageNumber="171" value="2017-03-17">
<collectingDate id="24719C2DFF6124DDCA51A28474AD25BD" box="[334,458,968,994]" pageId="170" pageNumber="171" value="2017-03-17">17.iii.2017</collectingDate>
</date>
,
<collectorName id="ED7E26D3FF6124DDCAC9A285773225BD" box="[470,597,968,995]" pageId="170" pageNumber="171">X.-B. Song</collectorName>
leg. [
<emphasis id="72FF9F17FF6124DDC98AA285766225BF" bold="true" box="[661,773,969,992]" pageId="170" pageNumber="171">ƱẪẸṘ</emphasis>
96K]’ (
<collectionCode id="269ADBC0FF6124DDC84AA28476C125BD" box="[853,934,968,994]" pageId="170" pageNumber="171">SNUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6124DDCBD8A2A0715E235D" pageId="170" pageNumber="171" type="diagnosis">
<paragraph id="40344305FF6124DDCBD8A2A0715E235D" blockId="170.[151,1437,932,2039]" pageId="170" pageNumber="171">
<emphasis id="72FF9F17FF6124DDCBD8A2A074252259" bold="true" box="[199,322,1004,1030]" pageId="170" pageNumber="171">Diagnosis.</emphasis>
Male. Body length
<quantity id="8773EEE0FF6124DDC938A2A077F72258" box="[551,656,1004,1031]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="170" pageNumber="171" unit="mm" value="2.4">2.40 mm</quantity>
. Head rounded triangular, as broad as pronotum, tempus moderately long, vertex and frons strongly modified, each with large projection at middle; antenna lacking modifications. Pronotum with laterally carinate median and lateral longitudinal sulci, discal carinae composed row of two small discal and one large antebasal spine; lateral margin denticulate at anterior half, with distinct marginal spine. Discal stria of elytron shallow and long, extending posteriorly to apical 1/4 of elytral length; disc finely punctate. Fore legs simple; mesotrochanter with apically curved ventral spine, mesotibia with distinct apical spine; metatrochanter with broad, curved ventral projection. Aedeagus strongly asymmetric, median lobe constricted at base, ventral stalk strongly broad at middle, dorsal lobe weakly sclerotized, parameres flattened, membranous.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6124DBCBD8A44071D825B2" lastPageId="172" lastPageNumber="173" pageId="170" pageNumber="171" type="description">
<paragraph id="40344305FF6124DDCBD8A44076422315" blockId="170.[151,1437,932,2039]" pageId="170" pageNumber="171">
<emphasis id="72FF9F17FF6124DDCBD8A440743E2379" bold="true" box="[199,345,1292,1318]" pageId="170" pageNumber="171">Description.</emphasis>
<emphasis id="72FF9F17FF6124DDCA40A44074FE2379" box="[351,409,1292,1318]" italics="true" pageId="170" pageNumber="171">Male</emphasis>
. Body (
<figureCitation id="D8B05F80FF6124DDCAEEA44077312379" box="[497,598,1292,1318]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="170" pageNumber="171">Fig. 73A</figureCitation>
) length
<quantity id="8773EEE0FF6124DDC9AEA440767F2378" box="[689,792,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="170" pageNumber="171" unit="mm" value="2.4">2.40 mm</quantity>
; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body covered with relatively dense pubescence.
</paragraph>
<paragraph id="40344305FF6124DDCBD8A41876C12140" blockId="170.[151,1437,932,2039]" pageId="170" pageNumber="171">
Head (
<figureCitation id="D8B05F80FF6124DDCA0EA41874132331" box="[273,372,1364,1390]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="170" pageNumber="171">Fig. 73B</figureCitation>
) rounded triangular, much wider than long, length
<quantity id="8773EEE0FF6124DDC8B4A41871772330" box="[939,1040,1364,1391]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="170" pageNumber="171" unit="mm" value="0.48">0.48 mm</quantity>
, width across eyes
<quantity id="8773EEE0FF6124DDCFFAA418702B2330" box="[1253,1356,1364,1391]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="170" pageNumber="171" unit="mm" value="0.59">0.59 mm</quantity>
; vertex with large projection at middle, area anterior and lateral projection impressed and with pair of nude foveae, with tufts of setae at anterolateral margins of impression, postocular area with oval patch covered by dense, short setae; frons anteriorly demarcated from clypeus by complete frontal-clypeal ridge, which merged at middle and extending anteriorly to apex of clypeus, with strongly raised tubercle covered by long posteriorly pointed setae at middle; clypeus with smooth surface, its entire anterior margin carinate and raised; ocular-mandibular carina complete, carina branched below eye, extended ventrally and then anteriorly to posteroventral articulation of mandible. Venter with rough surface; small gular foveae (posterior tentorial pits) originating from shared oval opening, with thin median carina extending from opening anteriorly to mouthparts. Compound eyes prominent, composed of approximately 55 ommatidia. Antenna lacking modification, length
<quantity id="8773EEE0FF6124DDC9DDA7D4764020EC" box="[706,807,1688,1715]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="170" pageNumber="171" unit="mm" value="1.26">1.26 mm</quantity>
, club (
<figureCitation id="D8B05F80FF6124DDC86EA7D476B220ED" box="[881,981,1688,1714]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="170" pageNumber="171">Fig. 73C</figureCitation>
) loosely formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2–7 each slightly elongate, sub-moniliform, 8 smallest, 9 much larger than 8, elongate, 10 slightly broader and as long as 9, 11 largest, approximately as long as 9 and 10 combined, sub-fusiform, with indistinct line separating apical 2/5 from basal 3/5.
</paragraph>
<paragraph id="40344305FF6124DCCBD8A66571D3268A" blockId="170.[151,1437,932,2039]" lastBlockId="171.[151,1437,151,537]" lastPageId="171" lastPageNumber="172" pageId="170" pageNumber="171">
Pronotum (
<figureCitation id="D8B05F80FF6124DDCA59A66474CD211D" box="[326,426,1832,1858]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="170" pageNumber="171">Fig. 73B</figureCitation>
) slightly wider than long, length
<quantity id="8773EEE0FF6124DDC83CA66476ED211C" box="[803,906,1832,1859]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="170" pageNumber="171" unit="mm" value="0.53">0.53 mm</quantity>
, width
<quantity id="8773EEE0FF6124DDC8C1A6647123211C" box="[990,1092,1832,1859]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="170" pageNumber="171" unit="mm" value="0.58">0.58 mm</quantity>
, widest at middle; lateral margins rounded at middle, convergent apically and parallel at basal 1/4; disc moderately convex, finely punctate, median longitudinal sulcus with carinate sides, followed posteriorly by oval impression and short mediobasal carina, with pair of mesally carinate lateral longitudinal sulci, discal carinae composed of row of two distinct discal and one large antebasal spine; lateral margin with large marginal and five to six smaller spines; lateral antebasal foveae small and asetose; with small outer and inner pair of basolateral foveae. Prosternum with anterior part longer than coxal part, with small lateral procoxal foveae; thin hypomeral ridge complete, extending from base to middle of anterior part, with punctiform lateral antebasal hypomeral pit; margin of coxal cavity broadly carinate.
</paragraph>
<paragraph id="40344305FF6024DCCBD8A193714A273A" blockId="171.[151,1437,151,537]" pageId="171" pageNumber="172">
Elytra wider than long, length
<quantity id="8773EEE0FF6024DCC935A19377F426A6" box="[554,659,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="171" pageNumber="172" unit="mm" value="0.73">0.73 mm</quantity>
, width
<quantity id="8773EEE0FF6024DCC9F5A193763426A6" box="[746,851,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="171" pageNumber="172" unit="mm" value="0.97">0.97 mm</quantity>
, widest at middle, narrowing form middle towards apex; each elytron with three large, asetose basal foveae; discal stria shallow and long, extending posteriorly from outer basal fovea to apical 1/4 of elytral length; humerus broadly prominent, with small denticle, subhumeral fovea small, carinate marginal stria extending from fovea to posterior margin of elytron.
</paragraph>
<paragraph id="40344305FF6024DCCBD8A023745D2446" blockId="171.[151,1437,151,537]" pageId="171" pageNumber="172">Mesoventrite short, demarcated from metaventrite by transverse carinae; median mesoventral foveae relatively broadly separated, large lateral mesoventral foveae not forked internally, with short mesoventral process. Metaventrite slightly prominent admesally, impressed at middle, with well-developed lateral mesocoxal and two lateral metaventral foveae, posterior margin with small and narrow split at middle. Marginal carina of meso- and metaventrite complete.</paragraph>
<caption id="14F4138DFF6024DCCB88A6DC762121B0" ID-DOI="http://doi.org/10.5281/zenodo.6340910" ID-Zenodo-Dep="6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="171" pageNumber="172" startId="171.[151,250,1936,1960]" targetBox="[152,1436,561,1911]" targetPageId="171">
<paragraph id="40344305FF6024DCCB88A6DC762121B0" blockId="171.[151,1437,1936,2032]" pageId="171" pageNumber="172">
<emphasis id="72FF9F17FF6024DCCB88A6DC744521F7" bold="true" box="[151,290,1936,1960]" pageId="171" pageNumber="172">FIGURE 73.</emphasis>
Morphology of
<taxonomicName id="878B3886FF6024DCCAD1A6DC77D221F7" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[462,693,1936,1960]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="vertexalis" status="sp. nov.">
<emphasis id="72FF9F17FF6024DCCAD1A6DC77D221F7" box="[462,693,1936,1960]" italics="true" pageId="171" pageNumber="172">Tribasodites vertexalis</emphasis>
</taxonomicName>
<emphasis id="72FF9F17FF6024DCC9A1A6DC767721F7" bold="true" box="[702,784,1936,1960]" pageId="171" pageNumber="172">
<taxonomicNameLabel id="69CC226CFF6024DCC9A1A6DC767721F7" box="[702,784,1936,1960]" pageId="171" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="72FF9F17FF6024DCC840A6DC761021F7" bold="true" box="[863,887,1936,1960]" pageId="171" pageNumber="172">A.</emphasis>
Dorsal habitus.
<emphasis id="72FF9F17FF6024DCCF3CA6DC715D21F7" bold="true" box="[1059,1082,1936,1960]" pageId="171" pageNumber="172">B.</emphasis>
Head and pronotum.
<emphasis id="72FF9F17FF6024DCCE02A6DC705121F7" bold="true" box="[1309,1334,1936,1960]" pageId="171" pageNumber="172">C.</emphasis>
Antennomeres 9–11.
<emphasis id="72FF9F17FF6024DCCA07A6F874562193" bold="true" box="[280,305,1972,1996]" pageId="171" pageNumber="172">D.</emphasis>
Mesotrochanter.
<emphasis id="72FF9F17FF6024DCCAFEA6F8749F2193" bold="true" box="[481,504,1972,1996]" pageId="171" pageNumber="172">E.</emphasis>
Mesotibia.
<emphasis id="72FF9F17FF6024DCC96FA6F877E42193" bold="true" box="[624,643,1972,1996]" pageId="171" pageNumber="172">F.</emphasis>
Metatrochanter.
<emphasis id="72FF9F17FF6024DCC830A6F876072193" bold="true" box="[815,864,1972,1996]" pageId="171" pageNumber="172">G–I.</emphasis>
Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I.
</paragraph>
</caption>
<paragraph id="40344305FF6724DBCBD8A1DB779326A6" blockId="172.[151,1437,151,1077]" pageId="172" pageNumber="173">
Legs moderately elongate; protrochanter with exceptionally long seta on ventral margin; mesotrochanter with apically curved ventral spine (
<figureCitation id="D8B05F80FF6724DBCAEBA1F77739268A" box="[500,606,187,213]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="172" pageNumber="173">Fig. 73D</figureCitation>
), mesotibia (
<figureCitation id="D8B05F80FF6724DBC9ECA1F7763E268A" box="[755,857,187,213]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="172" pageNumber="173">Fig. 73E</figureCitation>
) with triangular apical spine; metatrochanter (
<figureCitation id="D8B05F80FF6724DBCE70A1F775A226A6" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="172" pageNumber="173">Fig. 73F</figureCitation>
) with broad, curved projection on ventral margin.
</paragraph>
<paragraph id="40344305FF6724DBCBD8A04F76A72462" blockId="172.[151,1437,151,1077]" pageId="172" pageNumber="173">
Abdomen widest at lateral margins of tergite 1 (IV), length
<quantity id="8773EEE0FF6724DBC876A04F76B72742" box="[873,976,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="172" pageNumber="173" unit="mm" value="0.63">0.63 mm</quantity>
, width
<quantity id="8773EEE0FF6724DBCF3BA04F71EB2742" box="[1060,1164,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="172" pageNumber="173" unit="mm" value="0.88">0.88 mm</quantity>
. Tergite 1 (IV) less than twice as long as 2 (V), basal sulcus separated by one pair of mediobasal and two pairs of basolateral foveae, with pair of long discal carinae, inner marginal carina hardly oblique, outer one thicker, both complete; tergite 2 (V) slightly longer than 3 (VI), 4 (VII) shorter than tergites 2 and 3 combined; tergites 2–4 (V–VII) each with one pair of basolateral foveae, 4 with small marginal tubercles, tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle. Sternite 2 (IV) with one pair of small mediobasal and three pairs of basolateral foveae; midlength of sternites 2–5 (IV–VII) gradually shorter, 3–4 each with two pairs of small basolateral foveae and small tubercle at middle, 5 with one pair of basolateral foveae, sternite 6 (VIII) transverse, posterior margin broadly emarginate, sternite 7 (IX) composed of several membranous structures.
</paragraph>
<paragraph id="40344305FF6724DBCBD8A30B705F24F6" blockId="172.[151,1437,151,1077]" pageId="172" pageNumber="173">
Aedeagus (
<figureCitation id="D8B05F80FF6724DBCA5AA30B74A5243E" box="[325,450,583,609]" captionStart="FIGURE 73" captionStartId="171.[151,250,1936,1960]" captionTargetBox="[152,1436,561,1911]" captionTargetId="figure-169@171.[151,1436,561,1912]" captionTargetPageId="171" captionText="FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I." figureDoi="http://doi.org/10.5281/zenodo.6340910" httpUri="https://zenodo.org/record/6340910/files/figure.png" pageId="172" pageNumber="173">Fig. 73G–I</figureCitation>
)
<quantity id="8773EEE0FF6724DBCACEA30A7752243E" box="[465,565,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="172" pageNumber="173" unit="mm" value="0.35">0.35 mm</quantity>
long, strongly asymmetric; median lobe constricted at base, with rounded triangular foramen, ventral stalk strongly broad at middle, flattened apically and narrowing towards apex, dorsal lobe weakly sclerotized, extended to right side (in ventral view), parameres membranous, broad and plate-like.
</paragraph>
<paragraph id="40344305FF6724DBCBD8A3FF74F82492" blockId="172.[151,1437,151,1077]" box="[199,415,691,717]" pageId="172" pageNumber="173">
<emphasis id="72FF9F17FF6724DBCBD8A3FF747C2492" box="[199,283,691,717]" italics="true" pageId="172" pageNumber="173">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="40344305FF6724DBCBD8A39A71D825B2" blockId="172.[151,1437,151,1077]" pageId="172" pageNumber="173">
<emphasis id="72FF9F17FF6724DBCBD8A39A74D324AE" bold="true" box="[199,436,726,753]" pageId="172" pageNumber="173">Comparative notes.</emphasis>
Several different forms of aedeagi have been found within the possibly heterogenous
<taxonomicName id="878B3886FF6724DBCB88A3B77446254A" authorityName="Jeannel" authorityYear="1960" box="[151,289,763,789]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FF6724DBCB88A3B77446254A" box="[151,289,763,789]" italics="true" pageId="172" pageNumber="173">Tribasodites</emphasis>
</taxonomicName>
. This species possesses an aedeagus of a form typical for the genera
<emphasis id="72FF9F17FF6724DBCF3CA3B771DC254A" box="[1059,1211,763,789]" italics="true" pageId="172" pageNumber="173">Hingstoniella</emphasis>
,
<taxonomicName id="878B3886FF6724DBCFD6A3B77009254A" authority="Yin, 2022" authorityName="Yin" authorityYear="2022" box="[1225,1390,763,789]" class="Insecta" family="Staphylinidae" genus="Besuchetaceus" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="209" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="72FF9F17FF6724DBCFD6A3B77009254A" box="[1225,1390,763,789]" italics="true" pageId="172" pageNumber="173">Besuchetaceus</emphasis>
</taxonomicName>
Yin &amp; Li, and
<emphasis id="72FF9F17FF6724DBCA13A253741B2566" box="[268,380,799,825]" italics="true" pageId="172" pageNumber="173">Sinotrisus</emphasis>
Yin &amp; Li, creating concerns about the relationships between these groups. At this moment no discrete character can be found to support a placement of this species elsewhere other than in
<taxonomicName id="878B3886FF6724DBCFAEA20F705C2502" authorityName="Jeannel" authorityYear="1960" box="[1201,1339,835,861]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="72FF9F17FF6724DBCFAEA20F705C2502" box="[1201,1339,835,861]" italics="true" pageId="172" pageNumber="173">Tribasodites</emphasis>
</taxonomicName>
. Among the Tibetan species, the habitus of
<taxonomicName id="878B3886FF6724DBC90AA22B77F825DE" authorityName="Yin" authorityYear="2022" box="[533,671,871,897]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="vertexalis">
<emphasis id="72FF9F17FF6724DBC90AA22B774225DF" box="[533,549,871,896]" italics="true" pageId="172" pageNumber="173">T</emphasis>
.
<emphasis id="72FF9F17FF6724DBC92DA22B77F825DE" box="[562,671,871,897]" italics="true" pageId="172" pageNumber="173">vertexalis</emphasis>
</taxonomicName>
is relatively much stouter than the others, the male vertex is strongly modified with a large central excavation, and has lateral rounded areas covered by dense short setae. These characters in combination with the simple male antennae, the discal carina of the pronotum composed of three large spines, and the triangular apical spines of the mesotibiae readily separates this species from the others.
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6724DBCBD8A2BA76CE224E" box="[199,937,1014,1041]" pageId="172" pageNumber="173" type="distribution">
<paragraph id="40344305FF6724DBCBD8A2BA76CE224E" blockId="172.[151,1437,151,1077]" box="[199,937,1014,1041]" pageId="172" pageNumber="173">
<emphasis id="72FF9F17FF6724DBCBD8A2BA7406224F" bold="true" box="[199,353,1014,1040]" pageId="172" pageNumber="173">Distribution.</emphasis>
Mêdog County,
<collectingRegion id="824F8DE7FF6724DBC903A2BB773E224E" box="[540,601,1015,1041]" country="China" name="Xizang" pageId="172" pageNumber="173">Tibet</collectingRegion>
, SW
<collectingCountry id="389C0395FF6724DBC98BA2BB77B0224E" box="[660,727,1015,1041]" name="China" pageId="172" pageNumber="173">China</collectingCountry>
(
<figureCitation id="D8B05F80FF6724DBC9F9A2BB7637224E" box="[742,848,1015,1041]" captionStart="FIGURE 86" captionStartId="188.[151,250,1881,1905]" captionTargetBox="[147,1439,181,1857]" captionTargetId="figure-19@188.[147,1439,181,1857]" captionTargetPageId="188" captionText="FIGURE 86. Distribution of Tibetan Batrisini. A. Tribasodites dilophus sp. nov. (circle), T. elongatus sp. nov. (triangle) and T. grandiceps sp. nov. (square). B. T. gyirong sp. nov. (circle), T. kiypu sp. nov. (triangle) and T. mirabilis sp. nov. (square). C. T. prolixicornis sp. nov. (circle), T. Tribasodites pugiunculus sp. nov. (triangle) T. vertexalis sp. nov. (square) and T. yatung sp. nov. (diamond)." figureDoi="http://doi.org/10.5281/zenodo.6340942" httpUri="https://zenodo.org/record/6340942/files/figure.png" pageId="172" pageNumber="173">Figs 86C</figureCitation>
,
<figureCitation id="D8B05F80FF6724DBC843A2BB76FB224E" box="[860,924,1015,1041]" captionStart="FIGURE 100" captionStartId="202.[151,250,1591,1615]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure-19@202.[151,1436,181,1567]" captionTargetPageId="202" captionText="FIGURE 100. Environment of and collecting sites at Nyingchi area (A–E) and Chamdo (ăƀ) (F). A. 80K. B. 96K. C–E. Xiachayu (T察ø). F. pass of Jueba Mountain (ẅ巴山口), Markam (Ẽ康). Photo Credit: A by Zhong Peng; B by Xiao-Bin Song; F by Wen-Xuan Bi." figureDoi="http://doi.org/10.5281/zenodo.6340975" httpUri="https://zenodo.org/record/6340975/files/figure.png" pageId="172" pageNumber="173">100B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0891108EFF6724DBCBD8A55771D9226A" box="[199,1214,1051,1077]" pageId="172" pageNumber="173" type="etymology">
<paragraph id="40344305FF6724DBCBD8A55771D9226A" blockId="172.[151,1437,151,1077]" box="[199,1214,1051,1077]" pageId="172" pageNumber="173">
<emphasis id="72FF9F17FF6724DBCBD8A5577429226A" bold="true" box="[199,334,1051,1077]" pageId="172" pageNumber="173">Etymology.</emphasis>
The species epithet refers to the strongly modified male vertex of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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