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<mods:title id="BC627744AF30862A4AD221368276ACF8">Archaeococcoid scale insects (Hemiptera: Coccoidea) from the tropical high mountains of the Andean Cordillera, South America</mods:title>
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<subSubSection id="08A7EED0FFBAFFAFFF41F8AEFC9E2838" box="[151,807,1891,1917]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="4002BD5BFFBAFFAFFF41F8AEFC9E2838" blockId="3.[151,807,1891,1917]" box="[151,807,1891,1917]" pageId="3" pageNumber="4">
<heading id="1B4A0A37FFBAFFAFFF41F8AEFC9E2838" bold="true" box="[151,807,1891,1917]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="72C96149FFBAFFAFFF41F8AEFC9E2838" bold="true" box="[151,807,1891,1917]" pageId="3" pageNumber="4">
The family
<taxonomicName id="87BDC6D8FFBAFFAFFEF0F8AEFE502838" box="[294,489,1891,1917]" class="Insecta" family="Monophlebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Monophlebidae</taxonomicName>
in the Neotropical region
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="08A7EED0FFBAFFA8FF41F860FF5A2D8A" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="4002BD5BFFBAFFA8FF41F860FCA72F9D" blockId="3.[151,1437,1965,2030]" lastBlockId="4.[151,1437,152,1995]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
The
<taxonomicName id="87BDC6D8FFBAFFAFFF1AF860FE3D2882" box="[204,388,1965,1991]" class="Insecta" family="Monophlebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Monophlebidae</taxonomicName>
is a large family with a worldwide distribution. The classification of the Monophlebinae (now
<taxonomicName id="87BDC6D8FFBAFFAFFF08F819FE1928AB" box="[222,416,2004,2030]" class="Insecta" family="Monophlebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Monophlebidae</taxonomicName>
) was first established by
<bibRefCitation id="242CC0AAFFBAFFAFFD0DF819FC1028AB" author="Morrison" box="[731,937,2004,2030]" pageId="3" pageNumber="38" refString="Morrison, H. (1927) Descriptions of new genera and species belonging to the coccid family Margarodidae. Proceedings of the Biological Society of Washington, 40, 99 - 109." type="journal article" year="1927">Morrison (1927)</bibRefCitation>
, and discussed further in his remarkable monograph (
<bibRefCitation id="242CC0AAFFBDFFA8FEFAFF55FE512FF7" author="Morrison" box="[300,488,152,178]" pageId="4" pageNumber="38" refString="Morrison, H. (1928) A classification of the higher groups and genera of the coccid family Margarodidae. United States Department of Agriculture, Washington D. C. Technical Bulletin, 52, 1 - 239." type="journal article" year="1928">Morrison, 1928</bibRefCitation>
). Since then there have been additional significant studies, the latest by
<bibRefCitation id="242CC0AAFFBDFFA8FAE5FF55FEF82F9D" author="Unruh" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 a) Molecular data reveal convergent reproductive strategies in iceryne scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8 - 50, DOI: 10.1111 / j. 1365 - 3113.2007.00404. x." type="journal article" year="2008" yearSuffix="a">Unruh &amp; Gullan (2008a</bibRefCitation>
,
<bibRefCitation id="242CC0AAFFBDFFA8FE98FF73FE222F9D" author="Gullan" box="[334,411,190,216]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 b) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa, 1803, 1 - 106." type="journal article" year="2008" yearSuffix="b">2008b</bibRefCitation>
), and
<bibRefCitation id="242CC0AAFFBDFFA8FE37FF73FCA32F9D" author="Williams" box="[481,794,190,216]" pageId="4" pageNumber="38" refString="Williams, D. J. &amp; Gullan, P. J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini). Journal of Natural History, 42, 77 - 128." type="journal article" year="2008">Williams &amp; Gullan (2008)</bibRefCitation>
.
</paragraph>
<paragraph id="4002BD5BFFBDFFA8FF13FF28FF5A2D8A" blockId="4.[151,1437,152,1995]" pageId="4" pageNumber="5">
Adult female monophlebids are usually rather large, and important characteristic at all growth stages is their mobility, due to the presence of well-developed brown or black legs. The adult females are characterised by the following morphological features: antennae 711 segmented; ventral cicatrices present on the abdomen, usually with 1 or 3 posterior to the vulva, but in some cases many more; multilocular pores often numerous, but rather variable in the structure and configuration of the loculi; multilocular pores also sometimes in groups, forming either an ovisac pore band or marsupial pore band; hairs, hair-like setae, flagellate setae, spiniform setae and spines present or absent; thoracic spiracles usually without pores in atrium; abdominal spiracles varying in number from 2 to 7 pairs, each with or without pores in atrium; anal opening dorsal, with a simple anal tube either with only a sclerotised ring or also with pores at inner end. Identification of monophlebid species is often very difficult. Therefore, for taxonomic purposes, the simultaneous description of the adult female and male, and the first-instar nymph of a given species helps to establish the best characters for its identification and those of related species (Foldi, 2006;
<bibRefCitation id="242CC0AAFFBDFFA8FB00FD43FF6F2D8A" author="Foldi" pageId="4" pageNumber="38" refString="Foldi, I. &amp; Watson, G. W. (2001) A new pest scale insect on avocado trees in Colombia, Laurencella colombiana, sp. n. (Hemiptera: Coccoidae: Margarodidae). Annales de la Societe Entomologique de France (N. S.), 37, 367 - 374." type="journal article" year="2001">Foldi &amp; Watson, 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="08A7EED0FFBDFFA9FF13FD11FBB02DB3" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="4002BD5BFFBDFFA8FF13FD11FC172BDA" blockId="4.[151,1437,152,1995]" pageId="4" pageNumber="5">
The taxonomy of the
<taxonomicName id="87BDC6D8FFBDFFA8FE1EFD11FD3A2DB3" box="[456,643,732,758]" class="Insecta" family="Monophlebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Monophlebidae</taxonomicName>
in the Neotropical region has been greatly modified by the recent major revision of the tribe Iceryini (
<bibRefCitation id="242CC0AAFFBDFFA8FD8DFCCFFC322C59" author="Unruh" box="[603,907,770,796]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 a) Molecular data reveal convergent reproductive strategies in iceryne scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8 - 50, DOI: 10.1111 / j. 1365 - 3113.2007.00404. x." type="journal article" year="2008" yearSuffix="a">Unruh &amp; Gullan, 2008a</bibRefCitation>
), which is the largest and the most widely distributed tribe in the family (Foldi, 2001;
<bibRefCitation id="242CC0AAFFBDFFA8FD61FCE4FCC02C06" author="Ben-Dov" box="[695,889,809,835]" pageId="4" pageNumber="37" refString="Ben-Dov, Y. (2005) A Systematic Catalogue of the Scale Insect Family Margarodidae (Hemiptera: Coccoidea) of the World. Intercept Limited. 400 p." type="book" year="2005">Ben-Dov, 2005</bibRefCitation>
).
<bibRefCitation id="242CC0AAFFBDFFA8FC58FCE4FB792C06" author="Unruh" box="[910,1216,809,835]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 a) Molecular data reveal convergent reproductive strategies in iceryne scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8 - 50, DOI: 10.1111 / j. 1365 - 3113.2007.00404. x." type="journal article" year="2008" yearSuffix="a">Unruh &amp; Gullan (2008a)</bibRefCitation>
reconstructed the phylogeny of the Iceryini on the basis of molecular data and established numerous new combinations, new synonymies, considered several species to be
<emphasis id="72C96149FFBDFFA8FD6CFCBAFCE62CD5" box="[698,863,887,912]" italics="true" pageId="4" pageNumber="5">incertae sedis</emphasis>
, and designated
<typeStatus id="9F0603F9FFBDFFA8FBFFFCBBFB1C2CD5" box="[1065,1189,886,912]" pageId="4" pageNumber="5" type="lectotype">lectotypes</typeStatus>
. They recognised 74 species in the tribe Iceryini belonging to only five genera:
<taxonomicName id="87BDC6D8FFBDFFA8FCBEFC50FBC12CF2" authority="Cockerell" authorityName="Cockerell" box="[872,1144,925,951]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FCBEFC50FC402CF3" box="[872,1017,925,950]" italics="true" pageId="4" pageNumber="5">Crypticerya</emphasis>
Cockerell
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FB5EFC50FA212CF2" authority="Morrison" authorityName="Morrison" box="[1160,1432,925,951]" class="Insecta" family="Margarodidae" genus="Echinicerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FB5EFC50FAA22CF3" box="[1160,1307,925,950]" italics="true" pageId="4" pageNumber="5">Echinicerya</emphasis>
Morrison
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FF41FC09FDF32C9B" authority="Pesson &amp; Bielenin" authorityName="Pesson &amp; Bielenin" box="[151,586,964,990]" class="Insecta" family="Margarodidae" genus="Gigantococcus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FF41FC09FEEB2C98" box="[151,338,964,989]" italics="true" pageId="4" pageNumber="5">Gigantococcus</emphasis>
Pesson &amp; Bielenin
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FD81FC09FCE82C9B" authority="Fernald" authorityName="Fernald" box="[599,849,964,990]" class="Insecta" family="Margarodidae" genus="Gueriniella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FD81FC09FD512C98" box="[599,744,964,989]" italics="true" pageId="4" pageNumber="5">Gueriniella</emphasis>
Fernald
</taxonomicName>
and
<taxonomicName id="87BDC6D8FFBDFFA8FC46FC09FA242C9B" authority="Signoret. Unruh &amp; Gullan (2008a)" authorityName="Signoret. Unruh &amp; Gullan" authorityYear="2008" box="[912,1437,964,990]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FC46FC09FC592C98" box="[912,992,964,989]" italics="true" pageId="4" pageNumber="5">Icerya</emphasis>
Signoret.
<bibRefCitation id="242CC0AAFFBDFFA8FBB2FC09FA8B2C9B" author="Unruh" box="[1124,1330,964,990]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 a) Molecular data reveal convergent reproductive strategies in iceryne scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8 - 50, DOI: 10.1111 / j. 1365 - 3113.2007.00404. x." type="journal article" year="2008" yearSuffix="a">Unruh &amp; Gullan</bibRefCitation>
(2008a)
</taxonomicName>
redescribed and illustrated the
<typeStatus id="9F0603F9FFBDFFA8FDDCFC27FD842B41" box="[522,573,1002,1028]" pageId="4" pageNumber="5">type</typeStatus>
species of each genus, and provided a key to the genera based on adult females. In addition,
<bibRefCitation id="242CC0AAFFBDFFA8FE4DFBDCFD722B6E" author="Unruh" box="[411,715,1041,1067]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 b) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa, 1803, 1 - 106." type="journal article" year="2008" yearSuffix="b">Unruh &amp; Gullan (2008b)</bibRefCitation>
published an identification guide to all species in the tribe Iceryini, and the genus
<taxonomicName id="87BDC6D8FFBDFFA8FE65FBF5FDFB2B14" box="[435,578,1080,1105]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FE65FBF5FDFB2B14" box="[435,578,1080,1105]" italics="true" pageId="4" pageNumber="5">Crypticerya</emphasis>
</taxonomicName>
has been revised recently by Unruh (2008). Recently,
<bibRefCitation id="242CC0AAFFBDFFA8FB00FBF5FF5F2B3D" author="Kondo" pageId="4" pageNumber="38" refString="Kondo, T. &amp; Unruh, C. M. (2009) A new species of Crypticerya Cockerell (Hemiptera: Monophlebidae) from Colombia, with a key to species of the tribe Iceryini found in South America. Neotropical Entomology, 38, 92 - 100." type="journal article" year="2009">Kondo &amp; Unruh (2009)</bibRefCitation>
described a polyphagus new species,
<taxonomicName id="87BDC6D8FFBDFFA8FD62FB92FB762B3D" authority="Kondo &amp; Unruh" authorityName="Kondo &amp; Unruh" box="[692,1231,1118,1144]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="multicicatrices">
<emphasis id="72C96149FFBDFFA8FD62FB92FC472B3D" box="[692,1022,1119,1144]" italics="true" pageId="4" pageNumber="5">Crypticerya multicicatrices</emphasis>
Kondo &amp; Unruh
</taxonomicName>
, from
<collectingCountry id="38AAFDCBFFBDFFA8FAC8FB93FA2E2B3D" box="[1310,1431,1118,1144]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
, with a key to the adult females of South American iceryine species.
</paragraph>
<paragraph id="4002BD5BFFBDFFA8FF13FB61FCD6288E" blockId="4.[151,1437,152,1995]" pageId="4" pageNumber="5">
The fauna of the Neotropical region now includes 33 known monophlebid species in 7 genera and 2 tribes: 1. Tribe Iceryini:
<taxonomicName id="87BDC6D8FFBDFFA8FEBCFB1EFDCE2BA8" authority="Cockerell" authorityName="Cockerell" box="[362,631,1235,1261]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FEBCFB1EFE412BA9" box="[362,504,1235,1260]" italics="true" pageId="4" pageNumber="5">Crypticerya</emphasis>
Cockerell
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FD52FB1EFC332BA8" authority="Morrison" authorityName="Morrison" box="[644,906,1235,1261]" class="Insecta" family="Margarodidae" genus="Echinicerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FD52FB1EFCAA2BA9" box="[644,787,1235,1260]" italics="true" pageId="4" pageNumber="5">Echinicerya</emphasis>
Morrison
</taxonomicName>
and
<taxonomicName id="87BDC6D8FFBDFFA8FC13FB1EFB3A2BA8" authority="Signoret" authorityName="Signoret" box="[965,1155,1235,1261]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FC13FB1EFBA82BA9" box="[965,1041,1235,1260]" italics="true" pageId="4" pageNumber="5">Icerya</emphasis>
Signoret
</taxonomicName>
; and 2. Tribe Llaveiini:
<taxonomicName id="87BDC6D8FFBDFFA8FF41FB34FED12A56" authority="Foldi" authorityName="Foldi" box="[151,360,1273,1299]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FF41FB34FE9A2A57" box="[151,291,1273,1298]" italics="true" pageId="4" pageNumber="5">Laurencella</emphasis>
Foldi
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FEA0FB34FD922A56" authority="Llave" authorityName="Llave" box="[374,555,1273,1299]" class="Insecta" family="Margarodidae" genus="Llaveia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FEA0FB34FE772A57" box="[374,462,1273,1298]" italics="true" pageId="4" pageNumber="5">Llaveia</emphasis>
(Llave)
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FDEFFB34FCFF2A56" authority="Foldi" authorityName="Foldi" box="[569,838,1273,1299]" class="Insecta" family="Margarodidae" genus="Neohodgsonius" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FDEFFB34FD572A57" box="[569,750,1273,1298]" italics="true" pageId="4" pageNumber="5">Neohodgsonius</emphasis>
(Foldi)
</taxonomicName>
and
<taxonomicName id="87BDC6D8FFBDFFA8FCA8FB34FB3F2A56" authority="Townsend" authorityName="Townsend" box="[894,1158,1273,1299]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FCA8FB34FBBE2A57" box="[894,1031,1273,1298]" italics="true" pageId="4" pageNumber="5">Protortonia</emphasis>
Townsend
</taxonomicName>
(Foldi, 2001,
<bibRefCitation id="242CC0AAFFBDFFA8FAFDFB34FF6E2A7F" author="Ben-Dov" pageId="4" pageNumber="37" refString="Ben-Dov, Y. (2005) A Systematic Catalogue of the Scale Insect Family Margarodidae (Hemiptera: Coccoidea) of the World. Intercept Limited. 400 p." type="book" year="2005">Ben-Dov, 2005</bibRefCitation>
,
<bibRefCitation id="242CC0AAFFBDFFA8FF32FAEDFDA52A7F" author="Williams" box="[228,540,1312,1338]" pageId="4" pageNumber="38" refString="Williams, D. J. &amp; Gullan, P. J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini). Journal of Natural History, 42, 77 - 128." type="journal article" year="2008">Williams &amp; Gullan, 2008</bibRefCitation>
,
<bibRefCitation id="242CC0AAFFBDFFA8FDFFFAEDFCF72A7F" author="Unruh" box="[553,846,1312,1338]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 a) Molecular data reveal convergent reproductive strategies in iceryne scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re-interpretation of morphology and a revised generic classification. Systematic Entomology, 33, 8 - 50, DOI: 10.1111 / j. 1365 - 3113.2007.00404. x." type="journal article" year="2008" yearSuffix="a">Unruh &amp; Gullan, 2008a</bibRefCitation>
,
<bibRefCitation id="242CC0AAFFBDFFA8FC8AFAEDFCD62A7F" author="Unruh" box="[860,879,1312,1338]" pageId="4" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 b) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa, 1803, 1 - 106." type="journal article" year="2008" yearSuffix="b">b</bibRefCitation>
). The record of the tribe Monophlebini in the Neotropics is based on
<taxonomicName id="87BDC6D8FFBDFFA8FE71FA8AFD452A25" authority="Hempel" authorityName="Hempel" box="[423,764,1350,1376]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="niveus">
<emphasis id="72C96149FFBDFFA8FE71FA8AFD202A25" box="[423,665,1351,1376]" italics="true" pageId="4" pageNumber="5">Monophlebus niveus</emphasis>
Hempel
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FC95FA8BFC332A25" box="[835,906,1350,1376]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
(
<bibRefCitation id="242CC0AAFFBDFFA8FC4CFA8BFBFA2A25" author="Hempel" box="[922,1091,1350,1376]" pageId="4" pageNumber="38" refString="Hempel, A. (1920) Descriptions of new and little known Coccidae. Revista do MuseuPaulista, 12, 355 - 365." type="journal article" year="1920">Hempel, 1920</bibRefCitation>
) and, although the identity of this species has not been established, it is unlikely to be congeneric with the
<typeStatus id="9F0603F9FFBDFFA8FBEBFAA0FBC82AC2" box="[1085,1137,1389,1415]" pageId="4" pageNumber="5">type</typeStatus>
species of
<taxonomicName id="87BDC6D8FFBDFFA8FB2FFAA0FED52AEB" authority="Guerin-Meneville" authorityName="Guerin-Meneville" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FB2FFAA0FA252AC3" box="[1273,1436,1389,1414]" italics="true" pageId="4" pageNumber="5">Monophlebus</emphasis>
Guérin-Méneville
</taxonomicName>
,
<taxonomicName id="87BDC6D8FFBDFFA8FEA1FA59FD242AEB" authority="Burmeister" authorityName="Burmeister" box="[375,669,1428,1454]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="72C96149FFBDFFA8FEA1FA59FDAD2AE8" box="[375,532,1428,1453]" italics="true" pageId="4" pageNumber="5">M. atripennis</emphasis>
Burmeister
</taxonomicName>
from Java. The largest of these genera is
<taxonomicName id="87BDC6D8FFBDFFA8FB57FA59FAB42AE8" box="[1153,1293,1428,1453]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FB57FA59FAB42AE8" box="[1153,1293,1428,1453]" italics="true" pageId="4" pageNumber="5">Crypticerya</emphasis>
</taxonomicName>
, which now includes 16 species in the Neotropical region excluding the Nearctic part of
<collectingCountry id="38AAFDCBFFBDFFA8FBBCFA76FB762A90" box="[1130,1231,1467,1493]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
: 1.
<taxonomicName id="87BDC6D8FFBDFFA8FB2BFA76FE9A2ABE" authority="Newstead" authorityName="Newstead" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="abrahami">
<emphasis id="72C96149FFBDFFA8FB2BFA76FA242A91" box="[1277,1437,1467,1492]" italics="true" pageId="4" pageNumber="5">C. abrahami</emphasis>
(Newstead)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FEA6FA2CFE542ABE" box="[368,493,1505,1531]" name="Venezuela" pageId="4" pageNumber="5">Venezuela</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FDFCFA2CFD342ABE" box="[554,653,1505,1531]" name="Guyana" pageId="4" pageNumber="5">Guyana</collectingCountry>
; 2.
<taxonomicName id="87BDC6D8FFBDFFA8FD6CFA2CFC532ABE" authority="Hempel" authorityName="Hempel" box="[698,1002,1505,1531]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="72C96149FFBDFFA8FD6CFA2CFCD42ABF" box="[698,877,1505,1530]" italics="true" pageId="4" pageNumber="5">C. brasiliensis</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FBE1FA2CFB0D2ABE" box="[1079,1204,1505,1531]" name="Argentina" pageId="4" pageNumber="5">Argentina</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FB17FA2CFAB62ABE" box="[1217,1295,1505,1531]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FACAFA2CFA212ABE" box="[1308,1432,1505,1531]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FF41F9C5FF4F2967" box="[151,246,1544,1570]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FEE7F9C5FED52967" box="[305,364,1544,1570]" name="Peru" pageId="4" pageNumber="5">Peru</collectingCountry>
; 3.
<taxonomicName id="87BDC6D8FFBDFFA8FE4EF9C5FDCB2967" authority="Hempel" authorityName="Hempel" box="[408,626,1544,1570]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="72C96149FFBDFFA8FE4EF9C5FE412964" box="[408,504,1544,1569]" italics="true" pageId="4" pageNumber="5">C. flava</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FD6BF9C5FCB32967" box="[701,778,1544,1570]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
; 4.
<taxonomicName id="87BDC6D8FFBDFFA8FCE0F9C5FBE82967" authority="Hempel" authorityName="Hempel" box="[822,1105,1544,1570]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="flocculosa">
<emphasis id="72C96149FFBDFFA8FCE0F9C5FC6C2964" box="[822,981,1544,1569]" italics="true" pageId="4" pageNumber="5">C. flocculosa</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FB4DF9C5FB512967" box="[1179,1256,1544,1570]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
; 5.
<taxonomicName id="87BDC6D8FFBDFFA8FAC2F9C5FEBE290D" authority="Hempel" authorityName="Hempel" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="genistae">
<emphasis id="72C96149FFBDFFA8FAC2F9C5FA252964" box="[1300,1436,1544,1569]" italics="true" pageId="4" pageNumber="5">C. genistae</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE86F9E3FE2E290D" box="[336,407,1582,1608]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
and recently spread to the Caribben and southeastern
<collectingCountry id="38AAFDCBFFBDFFA8FBC1F9E3FBEB290D" box="[1047,1106,1582,1608]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
(
<bibRefCitation id="242CC0AAFFBDFFA8FBB7F9E3FAF7290D" author="Hodges" box="[1121,1358,1582,1608]" pageId="4" pageNumber="38" refString="Hodges, G. S., Hodges, A. C. &amp; Unruh, C. M. (2008) A new exotic pest for Florida's natural areas: Crypticerya genistae (Hemiptera: Monophlebidae). Florida Entomologist, 91, 335 - 337." type="journal article" year="2008">
Hodges
<emphasis id="72C96149FFBDFFA8FB12F9E2FB41290D" box="[1220,1272,1583,1608]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., 2008
</bibRefCitation>
); 6.
<taxonomicName id="87BDC6D8FFBDFFA8FA57F9E2FE1A292A" authority="Hempel" authorityName="Hempel" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="72C96149FFBDFFA8FA57F9E2FE95292B" italics="true" pageId="4" pageNumber="5">C. luederwaldti</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE3AF998FD8E292A" box="[492,567,1621,1647]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
; 7.
<taxonomicName id="87BDC6D8FFBDFFA8FDB7F998FCDD292A" authority="Morrison" authorityName="Morrison" box="[609,868,1621,1647]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="72C96149FFBDFFA8FDB7F998FD65292B" box="[609,732,1621,1646]" italics="true" pageId="4" pageNumber="5">C. minima</emphasis>
(Morrison)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FC7AF998FB9C292A" box="[940,1061,1621,1647]" name="Argentina" pageId="4" pageNumber="5">Argentina</collectingCountry>
; 8.
<taxonomicName id="87BDC6D8FFBDFFA8FB98F998FEBD29D3" authority="Riley &amp; Howard" authorityName="Riley &amp; Howard" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="montserratensis">
<emphasis id="72C96149FFBDFFA8FB98F998FA95292B" box="[1102,1324,1621,1646]" italics="true" pageId="4" pageNumber="5">C. montserratensis</emphasis>
(Riley &amp; Howard)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE87F9B1FE7729D3" box="[337,462,1660,1686]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FE0DF9B1FDFE29D3" box="[475,583,1660,1686]" name="Grenada" pageId="4" pageNumber="5">Grenada</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FD82F9B1FD0C29D3" box="[596,693,1660,1686]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FD15F9B1FCF729D3" box="[707,846,1660,1686]" name="Montserrat" pageId="4" pageNumber="5">Montserrat</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FC8DF9B1FC0529D3" box="[859,956,1660,1686]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FC2CF9B1FB2829D3" box="[1018,1169,1660,1686]" name="Puerto Rico" pageId="4" pageNumber="5">Puerto Rico</collectingCountry>
; 9.
<taxonomicName id="87BDC6D8FFBDFFA8FB69F9B1FEEE29F8" authority="Kondo &amp; Unruh" authorityName="Kondo &amp; Unruh" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="multicicatrices">
<emphasis id="72C96149FFBDFFA8FB69F9B1FA2429D0" box="[1215,1437,1660,1685]" italics="true" pageId="4" pageNumber="5">C. multicicatrices</emphasis>
Kondo &amp; Unruh
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE49F96EFDAF29F8" box="[415,534,1699,1725]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
; 10.
<taxonomicName id="87BDC6D8FFBDFFA8FD98F96EFC1329F8" authority="Riley &amp; Howard" authorityName="Riley &amp; Howard" box="[590,938,1699,1725]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="palmeri">
<emphasis id="72C96149FFBDFFA8FD98F96EFD7229F9" box="[590,715,1699,1724]" italics="true" pageId="4" pageNumber="5">C. palmeri</emphasis>
(Riley &amp; Howard)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FC27F96EFBD029F8" box="[1009,1129,1699,1725]" name="Argentina" pageId="4" pageNumber="5">Argentina</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FBA3F96EFB0D29F8" box="[1141,1204,1699,1725]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FB3BF96EFAF329F8" box="[1261,1354,1699,1725]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
; 11.
<taxonomicName id="87BDC6D8FFBDFFA8FA57F96EFE2D29A6" authority="Newstead" authorityName="Newstead" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pimentae">
<emphasis id="72C96149FFBDFFA8FA57F96EFEBD29A7" italics="true" pageId="4" pageNumber="5">C. pimentae</emphasis>
(Newstead)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE09F904FD8729A6" box="[479,574,1737,1763]" name="Jamaica" pageId="4" pageNumber="5">Jamaica</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FDAFF904FD0129A6" box="[633,696,1737,1763]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
. 12. C.
<emphasis id="72C96149FFBDFFA8FCC3F904FCED29A7" box="[789,852,1737,1762]" italics="true" pageId="4" pageNumber="5">rileyi</emphasis>
(Cockerell) from
<collectingCountry id="38AAFDCBFFBDFFA8FBF8F904FB3529A6" box="[1070,1164,1737,1763]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
; 13.
<taxonomicName id="87BDC6D8FFBDFFA8FB10F904FEBF284F" authority="Riley &amp; Howard" authorityName="Riley &amp; Howard" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="rosae">
<emphasis id="72C96149FFBDFFA8FB10F904FA9229A7" box="[1222,1323,1737,1762]" italics="true" pageId="4" pageNumber="5">C. rosae</emphasis>
(Riley &amp; Howard)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FE83F93DFE68284F" box="[341,465,1776,1802]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FE34F93DFDFC284F" box="[482,581,1776,1802]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FD85F93DFD53284F" box="[595,746,1776,1802]" name="Puerto Rico" pageId="4" pageNumber="5">Puerto Rico</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FCFCF93DFCD1284F" box="[810,872,1776,1802]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
(Florida); 14.
<taxonomicName id="87BDC6D8FFBDFFA8FBF2F93DFA88284F" authority="Morrison" authorityName="Morrison" box="[1060,1329,1776,1802]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="72C96149FFBDFFA8FBF2F93DFB27284C" box="[1060,1182,1776,1801]" italics="true" pageId="4" pageNumber="5">C. similis</emphasis>
(Morrison)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FA57F93DFEBA2875" name="El Salvador" pageId="4" pageNumber="5">El Salvador</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBDFFA8FED9F8DBFED42875" box="[271,365,1814,1840]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FE7FF8DBFDAB2875" box="[425,530,1814,1840]" name="Trinidad and Tobago" pageId="4" pageNumber="5">Trinidad</collectingCountry>
; 15.
<taxonomicName id="87BDC6D8FFBDFFA8FD9AF8DAFCCA2875" authority="Leonardi" authorityName="Leonardi" box="[588,883,1814,1840]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subandina">
<emphasis id="72C96149FFBDFFA8FD9AF8DAFD552875" box="[588,748,1815,1840]" italics="true" pageId="4" pageNumber="5">C. subandina</emphasis>
(Leonardi)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FC68F8DBFB812875" box="[958,1080,1814,1840]" name="Argentina" pageId="4" pageNumber="5">Argentina</collectingCountry>
, and 16.
<taxonomicName id="87BDC6D8FFBDFFA8FB72F8DAFA252875" authority="Cockerell" authorityName="Cockerell" box="[1188,1436,1814,1840]" class="Insecta" family="Margarodidae" genus="Crypticerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="zeteki">
<emphasis id="72C96149FFBDFFA8FB72F8DAFAB52875" box="[1188,1292,1815,1840]" italics="true" pageId="4" pageNumber="5">C. zeteki</emphasis>
(Cockerell)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FF0FF8F0FEF72812" box="[217,334,1853,1879]" name="Colombia" pageId="4" pageNumber="5">Colombia</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBDFFA8FE5FF8F0FE5E2812" box="[393,487,1853,1879]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
Canal Zone.
<taxonomicName id="87BDC6D8FFBDFFA8FD5FF8F0FCA12813" box="[649,792,1853,1878]" class="Insecta" family="Margarodidae" genus="Echinicerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FD5FF8F0FCA12813" box="[649,792,1853,1878]" italics="true" pageId="4" pageNumber="5">Echinicerya</emphasis>
</taxonomicName>
includes only
<taxonomicName id="87BDC6D8FFBDFFA8FC1FF8F0FB732812" authority="Morrison" authorityName="Morrison" box="[969,1226,1853,1879]" class="Insecta" family="Margarodidae" genus="Echinicerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="72C96149FFBDFFA8FC1FF8F0FBEA2813" box="[969,1107,1853,1878]" italics="true" pageId="4" pageNumber="5">E. anomala</emphasis>
Morrison
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBDFFA8FAC2F8F0FA202812" box="[1300,1433,1853,1879]" name="Guatemala" pageId="4" pageNumber="5">Guatemala</collectingCountry>
. The most widespread monophlebid species in the Neotropics is
<taxonomicName id="87BDC6D8FFBDFFA8FC54F8A9FB1B283B" authority="Maskell" authorityName="Maskell" box="[898,1186,1892,1918]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="purchasi">
<emphasis id="72C96149FFBDFFA8FC54F8A9FB802838" box="[898,1081,1892,1917]" italics="true" pageId="4" pageNumber="5">Icerya purchasi</emphasis>
Maskell
</taxonomicName>
, found in most South American countries, but native to
<collectingCountry id="38AAFDCBFFBDFFA8FDFAF847FD2528E1" box="[556,668,1930,1956]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
. Two other
<taxonomicName id="87BDC6D8FFBDFFA8FCFEF846FCCA28E1" box="[808,883,1931,1956]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBDFFA8FCFEF846FCCA28E1" box="[808,883,1931,1956]" italics="true" pageId="4" pageNumber="5">Icerya</emphasis>
</taxonomicName>
species have been recorded in the Neotropics, namely
<taxonomicName id="87BDC6D8FFBDFFA8FF22F87CFD86288E" authority="Westwood" authorityName="Westwood" box="[244,575,1969,1995]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="seychellarum">
<emphasis id="72C96149FFBDFFA8FF22F87CFE11288F" box="[244,424,1969,1994]" italics="true" pageId="4" pageNumber="5">I. seychellarum</emphasis>
(Westwood)
</taxonomicName>
and
<taxonomicName id="87BDC6D8FFBDFFA8FDAEF87CFCD6288E" authority="Hempel." authorityName="Hempel." box="[632,879,1969,1995]" class="Insecta" family="Margarodidae" genus="Icerya" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="schrottkyi">
<emphasis id="72C96149FFBDFFA8FDAEF87CFCBA288F" box="[632,771,1969,1994]" italics="true" pageId="4" pageNumber="5">I. schrottkyi</emphasis>
Hempel.
</taxonomicName>
</paragraph>
<paragraph id="4002BD5BFFBCFFA9FF13FF55FBB02DB3" blockId="5.[151,1437,152,835]" pageId="5" pageNumber="6">
The tribe Llaveiini includes four genera:
<taxonomicName id="87BDC6D8FFBCFFA9FD1FFF55FCE42FF4" box="[713,861,152,177]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FD1FFF55FCE42FF4" box="[713,861,152,177]" italics="true" pageId="5" pageNumber="6">Laurencella</emphasis>
</taxonomicName>
includes
<taxonomicName id="87BDC6D8FFBCFFA9FC00FF55FAEE2FF7" authority="Foldi &amp; Watson" authorityName="Foldi &amp; Watson" box="[982,1367,152,178]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="colombiana">
<emphasis id="72C96149FFBCFFA9FC00FF55FB302FF4" box="[982,1161,152,177]" italics="true" pageId="5" pageNumber="6">L. colombiana</emphasis>
Foldi &amp; Watson
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FF41FF73FEA82F9D" box="[151,273,190,216]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
,
<taxonomicName id="87BDC6D8FFBCFFA9FECBFF72FE432F9D" authority="Foldi" authorityName="Foldi" box="[285,506,190,216]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="marikana">
<emphasis id="72C96149FFBCFFA9FECBFF72FE082F9D" box="[285,433,191,216]" italics="true" pageId="5" pageNumber="6">L. marikana</emphasis>
Foldi
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FD93FF73FD1D2F9D" box="[581,676,190,216]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
,
<taxonomicName id="87BDC6D8FFBCFFA9FD66FF72FC1E2F9D" authority="Hempel" authorityName="Hempel" box="[688,935,190,216]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="taunayi">
<emphasis id="72C96149FFBCFFA9FD66FF72FC922F9D" box="[688,811,191,216]" italics="true" pageId="5" pageNumber="6">L. taunayi</emphasis>
(Hempel)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FC24FF73FB852F9D" box="[1010,1084,190,216]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
and
<taxonomicName id="87BDC6D8FFBCFFA9FBAEFF72FADA2F9D" authority="Signoret" authorityName="Signoret" box="[1144,1379,190,216]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="uhleri">
<emphasis id="72C96149FFBCFFA9FBAEFF72FB592F9D" box="[1144,1248,191,216]" italics="true" pageId="5" pageNumber="6">L. uhleri</emphasis>
(Signoret)
</taxonomicName>
(see below).
<taxonomicName id="87BDC6D8FFBCFFA9FF29FF28FEE72FBB" box="[255,350,229,254]" class="Insecta" family="Margarodidae" genus="Llaveia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FF29FF28FEE72FBB" box="[255,350,229,254]" italics="true" pageId="5" pageNumber="6">Llaveia</emphasis>
</taxonomicName>
includes
<taxonomicName id="87BDC6D8FFBCFFA9FE0FFF28FD212FBA" authority="Llave" authorityName="Llave" box="[473,664,229,255]" class="Insecta" family="Margarodidae" genus="Llaveia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="axin">
<emphasis id="72C96149FFBCFFA9FE0FFF28FD8B2FBB" box="[473,562,229,254]" italics="true" pageId="5" pageNumber="6">L. axin</emphasis>
(Llave)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FD30FF28FCCB2FBA" box="[742,882,229,255]" name="Guatemala" pageId="5" pageNumber="6">Guatemala</collectingCountry>
,
<collectingCountry id="38AAFDCBFFBCFFA9FC56FF28FC662FBA" box="[896,991,229,255]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
(Nearctic and Neotropical), and
<taxonomicName id="87BDC6D8FFBCFFA9FA55FF28FE152E63" authority="Cockerell" authorityName="Cockerell" class="Insecta" family="Margarodidae" genus="Llaveia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="72C96149FFBCFFA9FA55FF28FEA12E60" italics="true" pageId="5" pageNumber="6">L. championi</emphasis>
(Cockerell)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FE2FFEC1FDE72E63" box="[505,606,268,294]" name="Panama" pageId="5" pageNumber="6">Panama</collectingCountry>
.
<taxonomicName id="87BDC6D8FFBCFFA9FDBCFEC1FC9F2E60" box="[618,806,268,293]" class="Insecta" family="Margarodidae" genus="Neohodgsonius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FDBCFEC1FC9F2E60" box="[618,806,268,293]" italics="true" pageId="5" pageNumber="6">Neohodgsonius</emphasis>
</taxonomicName>
includes
<taxonomicName id="87BDC6D8FFBCFFA9FC4BFEC1FB2A2E63" authority="Foldi" authorityName="Foldi" box="[925,1171,268,294]" class="Insecta" family="Margarodidae" genus="Neohodgsonius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cassicola">
<emphasis id="72C96149FFBCFFA9FC4BFEC1FB8D2E60" box="[925,1076,268,293]" italics="true" pageId="5" pageNumber="6">N. cassicola</emphasis>
(Foldi)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FB09FEC1FAD82E63" box="[1247,1377,268,294]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
. The species
<taxonomicName id="87BDC6D8FFBCFFA9FF23FEFFFD252E09" authority="Walker" authorityName="Walker" box="[245,668,306,332]" class="Insecta" family="Margarodidae" genus="Palaeococcus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="72C96149FFBCFFA9FF23FEFFFD922E0E" box="[245,555,306,331]" italics="true" pageId="5" pageNumber="6">Palaeococcus brasiliensis</emphasis>
(Walker)
</taxonomicName>
was described based on adult males from
<collectingCountry id="38AAFDCBFFBCFFA9FB4FFEFFFB5B2E09" box="[1177,1250,306,332]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
(
<bibRefCitation id="242CC0AAFFBCFFA9FB27FEFFFA2D2E09" author="Walker" box="[1265,1428,306,332]" pageId="5" pageNumber="38" refString="Walker, F. (1852) List of the specimens of homopterous insects in the collection of the British Museum, Part IV. London: British Museum (Natural History). 1188 pp." type="book" year="1852">Walker, 1852</bibRefCitation>
) and, although it is impossible to determine which genus this species belongs to, almost certainly it is not congeneric with the
<typeStatus id="9F0603F9FFBCFFA9FE58FE4DFE7B2EDF" box="[398,450,384,410]" pageId="5" pageNumber="6">type</typeStatus>
species of
<taxonomicName id="87BDC6D8FFBCFFA9FD9DFE4DFCC92EDF" authority="Cockerell" authorityName="Cockerell" box="[587,880,384,410]" class="Insecta" family="Margarodidae" genus="Palaeococcus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FD9DFE4DFD4B2EDC" box="[587,754,384,409]" italics="true" pageId="5" pageNumber="6">Palaeococcus</emphasis>
Cockerell
</taxonomicName>
from Europe. In addition,
<bibRefCitation id="242CC0AAFFBCFFA9FB63FE4DFF502E85" author="Williams" pageId="5" pageNumber="38" refString="Williams, D. J. &amp; Gullan, P. J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini). Journal of Natural History, 42, 77 - 128." type="journal article" year="2008">Williams &amp; Gullan (2008)</bibRefCitation>
have revised the Neotropical genus
<taxonomicName id="87BDC6D8FFBCFFA9FD78FE6AFC7E2E85" authority="Townsend" authorityName="Townsend" box="[686,967,422,448]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FD78FE6AFC872E85" box="[686,830,423,448]" italics="true" pageId="5" pageNumber="6">Protortonia</emphasis>
Townsend
</taxonomicName>
, describing and illustrating all known species, including all available instars, and this genus includes: 1.
<taxonomicName id="87BDC6D8FFBCFFA9FC7CFE00FBCA2EA2" authority="Ferris" authorityName="Ferris" box="[938,1139,461,487]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="72C96149FFBCFFA9FC7CFE00FBAA2EA3" box="[938,1043,461,486]" italics="true" pageId="5" pageNumber="6">P. azteca</emphasis>
(Ferris)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FB6AFE00FAA32EA2" box="[1212,1306,461,487]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
; 2.
<taxonomicName id="87BDC6D8FFBCFFA9FA93FE00FA252EA3" box="[1349,1436,461,486]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cacti">
<emphasis id="72C96149FFBCFFA9FA93FE00FA252EA3" box="[1349,1436,461,486]" italics="true" pageId="5" pageNumber="6">P. cacti</emphasis>
</taxonomicName>
(L.) from
<collectingCountry id="38AAFDCBFFBCFFA9FEDDFE39FE232D4B" box="[267,410,500,526]" name="Puerto Rico" pageId="5" pageNumber="6">Puerto Rico</collectingCountry>
; 3.
<emphasis id="72C96149FFBCFFA9FE15FE39FD852D48" box="[451,572,500,525]" italics="true" pageId="5" pageNumber="6">P. c ro t o n i s</emphasis>
Reyne from
<collectingCountry id="38AAFDCBFFBCFFA9FD0FFE39FC772D4B" box="[729,974,500,526]" name="Curaçao" pageId="5" pageNumber="6">Netherlands Antilles</collectingCountry>
; 4.
<taxonomicName id="87BDC6D8FFBCFFA9FC21FE39FB4A2D4B" authority="Foldi" authorityName="Foldi" box="[1015,1267,500,526]" class="Insecta" family="Monophlebidae" genus="Paramoandesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="72C96149FFBCFFA9FC21FE39FB172D48" box="[1015,1198,500,525]" italics="true" pageId="5" pageNumber="6">P. ecuadorensis</emphasis>
Foldi
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FAECFE39FA252D4B" box="[1338,1436,500,526]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
and
<collectingCountry id="38AAFDCBFFBCFFA9FF1FFDD7FEF82D71" box="[201,321,538,564]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
. 5.
<taxonomicName id="87BDC6D8FFBCFFA9FEBDFDD7FD852D71" authority="Fonseca" authorityName="Fonseca" box="[363,572,538,564]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="navesi">
<emphasis id="72C96149FFBCFFA9FEBDFDD7FE6D2D76" box="[363,468,538,563]" italics="true" pageId="5" pageNumber="6">P. navesi</emphasis>
Fonseca
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FD50FDD7FD6B2D71" box="[646,722,538,564]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
. 6.
<taxonomicName id="87BDC6D8FFBCFFA9FD2AFDD7FBA32D71" authority="Townsend" authorityName="Townsend" box="[764,1050,538,564]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="primitiva">
<emphasis id="72C96149FFBCFFA9FD2AFDD7FC3C2D76" box="[764,901,538,563]" italics="true" pageId="5" pageNumber="6">P. primitiva</emphasis>
(Townsend)
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FBB5FDD7FB792D71" box="[1123,1216,538,564]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
, and 7.
<taxonomicName id="87BDC6D8FFBCFFA9FAC8FDD7FE3C2D1E" authority="Williams &amp; Gullan" authorityName="Williams &amp; Gullan" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quernea">
<emphasis id="72C96149FFBCFFA9FAC8FDD7FA252D76" box="[1310,1436,538,563]" italics="true" pageId="5" pageNumber="6">P. quernea</emphasis>
Williams &amp; Gullan
</taxonomicName>
from
<collectingCountry id="38AAFDCBFFBCFFA9FE05FD8CFDE72D1E" box="[467,606,577,603]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
, collected at
<quantity id="874510BEFFBCFFA9FCD3FD8CFCDA2D1E" box="[773,867,577,603]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="m" value="3000.0">3000 m</quantity>
on
<taxonomicName id="87BDC6D8FFBCFFA9FC42FD8CFC452D1F" box="[916,1020,577,602]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="72C96149FFBCFFA9FC42FD8CFC452D1F" box="[916,1020,577,602]" italics="true" pageId="5" pageNumber="6">Quercus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="87BDC6D8FFBCFFA9FBECFD8CFB0C2D1E" box="[1082,1205,577,603]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
), on the slope of a volcano.
<bibRefCitation id="242CC0AAFFBCFFA9FED5FDA5FD822DC7" author="Williams" box="[259,571,616,642]" pageId="5" pageNumber="38" refString="Williams, D. J. &amp; Gullan, P. J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini). Journal of Natural History, 42, 77 - 128." type="journal article" year="2008">Williams &amp; Gullan (2008)</bibRefCitation>
provided keys to the genera in the tribe Llaveiini (based on adult female characters), and to the adult females of
<taxonomicName id="87BDC6D8FFBCFFA9FDACFD42FCB12DED" box="[634,776,655,680]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FDACFD42FCB12DED" box="[634,776,655,680]" italics="true" pageId="5" pageNumber="6">Protortonia</emphasis>
</taxonomicName>
species. They transferred
<taxonomicName id="87BDC6D8FFBCFFA9FB9CFD42FAC52DED" authority="Ferris" authorityName="Ferris" box="[1098,1404,654,680]" class="Insecta" family="Margarodidae" genus="Marchalina" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="72C96149FFBCFFA9FB9CFD42FA942DED" box="[1098,1325,655,680]" italics="true" pageId="5" pageNumber="6">Marchalina azteca</emphasis>
Ferris
</taxonomicName>
to
<taxonomicName id="87BDC6D8FFBCFFA9FF41FD78FE922D8B" box="[151,299,693,718]" class="Insecta" family="Margarodidae" genus="Protortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FF41FD78FE922D8B" box="[151,299,693,718]" italics="true" pageId="5" pageNumber="6">Protortonia</emphasis>
</taxonomicName>
(as
<taxonomicName id="87BDC6D8FFBCFFA9FEB4FD78FDF22D8A" authority="Ferris" authorityName="Ferris" box="[354,587,693,719]" class="Insecta" family="Margarodidae" genus="Marchalina" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="azteca">
<emphasis id="72C96149FFBCFFA9FEB4FD78FE6D2D8B" box="[354,468,693,718]" italics="true" pageId="5" pageNumber="6">P. azteca</emphasis>
(Ferris))
</taxonomicName>
, synonymised
<taxonomicName id="87BDC6D8FFBCFFA9FCDEFD78FB892D8A" authority="Ben-Dov" authorityName="Ben-Dov" box="[776,1072,693,719]" class="Insecta" family="Margarodidae" genus="Ecuadortonia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FCDEFD78FC0C2D8B" box="[776,949,693,718]" italics="true" pageId="5" pageNumber="6">Ecuadortonia</emphasis>
Ben-Dov
</taxonomicName>
with
<taxonomicName id="87BDC6D8FFBCFFA9FBADFD78FAD82D8A" authority="Foldi" authorityName="Foldi" box="[1147,1377,693,719]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FBADFD78FAAA2D8B" box="[1147,1299,693,718]" italics="true" pageId="5" pageNumber="6">Laurencella</emphasis>
Foldi
</taxonomicName>
, and transferred
<taxonomicName id="87BDC6D8FFBCFFA9FECBFD11FD882DB3" authority="Signoret" authorityName="Signoret" box="[285,561,732,758]" class="Insecta" family="Margarodidae" genus="Ortonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="uhleri">
<emphasis id="72C96149FFBCFFA9FECBFD11FE7E2DB0" box="[285,455,732,757]" italics="true" pageId="5" pageNumber="6">Ortonia uhleri</emphasis>
Signoret
</taxonomicName>
to
<taxonomicName id="87BDC6D8FFBCFFA9FD81FD11FD5D2DB0" box="[599,740,732,757]" class="Insecta" family="Margarodidae" genus="Laurencella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="72C96149FFBCFFA9FD81FD11FD5D2DB0" box="[599,740,732,757]" italics="true" pageId="5" pageNumber="6">Laurencella</emphasis>
</taxonomicName>
(as
<taxonomicName id="87BDC6D8FFBCFFA9FCC2FD11FBBC2DB3" authority="Signoret" authorityName="Signoret" box="[788,1029,732,758]" class="Insecta" family="Margarodidae" genus="Ortonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="uhleri">
<emphasis id="72C96149FFBCFFA9FCC2FD11FCC32DB0" box="[788,890,732,757]" italics="true" pageId="5" pageNumber="6">L. uhleri</emphasis>
(Signoret))
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="08A7EED0FFBCFFA9FF13FCCFFC2C2C06" pageId="5" pageNumber="6" type="discussion">
<paragraph id="4002BD5BFFBCFFA9FF13FCCFFC2C2C06" blockId="5.[151,1437,152,835]" pageId="5" pageNumber="6">
<emphasis id="72C96149FFBCFFA9FF13FCCFFE942C59" bold="true" box="[197,301,770,796]" pageId="5" pageNumber="6">Remark</emphasis>
. Readers can refer to the key in
<bibRefCitation id="242CC0AAFFBCFFA9FD63FCCFFC662C59" author="Unruh" box="[693,991,770,796]" pageId="5" pageNumber="38" refString="Unruh, C. M. &amp; Gullan, P. J. (2008 b) Identification guide to species in the scale insect tribe Iceryini (Coccoidea: Monophlebidae). Zootaxa, 1803, 1 - 106." type="journal article" year="2008" yearSuffix="b">Unruh &amp; Gullan (2008b)</bibRefCitation>
for identification of adult females of Iceryini and to
<bibRefCitation id="242CC0AAFFBCFFA9FE9FFCE4FDC42C06" author="Williams" box="[329,637,809,835]" pageId="5" pageNumber="38" refString="Williams, D. J. &amp; Gullan, P. J. (2008) A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini). Journal of Natural History, 42, 77 - 128." type="journal article" year="2008">Williams &amp; Gullan (2008)</bibRefCitation>
for genera of Llaveiini.
</paragraph>
</subSubSection>
</treatment>
</document>