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<document id="0FB7B51E81D1266C0E710CB488577DC7" ID-CLB-Dataset="7100" ID-DOI="10.5852/ejt.2021.748.1335" ID-GBIF-Dataset="04bf840b-8ba9-4059-8190-f2454630a714" ID-ISSN="2118-9773" ID-Zenodo-Dep="4736129" ID-ZooBank="AB54CD6A-B757-4E5A-8EB3-B9B466B74639" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1620111095485" checkinUser="plazi" docAuthor="Sousa, Rafael &amp; Seidel, Matthias" docDate="2021" docId="C10A5544FF80FFA1151CA5B3FC2AFC8A" docLanguage="en" docName="ejt-748_15-35.pdf" docOrigin="European Journal of Taxonomy 748 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.748.1335" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Inca irroratus Chevrolat 1833" docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="3D332D3CFF8FFFB0177CA434F970FFE8" masterDocTitle="Review of the Inca irroratus species group with description of two new species of Inca LePeletier &amp; Serville, 1828 (Coleoptera, Scarabaeidae, Cetoniinae)" masterLastPageNumber="35" masterPageNumber="15" pageNumber="30" updateTime="1698938226785" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="CA21C37D8EF0AED2ABE311C7AB557550">
<mods:title id="2B30CFAE31D2E14DBCDC50261FF9B7F4">Review of the Inca irroratus species group with description of two new species of Inca LePeletier &amp; Serville, 1828 (Coleoptera, Scarabaeidae, Cetoniinae)</mods:title>
</mods:titleInfo>
<mods:name id="1500B54EFFD8669CA4DBE9A074B19532" type="personal">
<mods:role id="87F8E500B8067D4DA637DB36015182C3">
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<mods:namePart id="6146F99BC61A4DA6C9AC458FBDB01A6F">Sousa, Rafael</mods:namePart>
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<mods:affiliation id="C4D9E6C28E1A6C4A2567E61FD132E961">Museu de Zoologia da Universidade de São Paulo, Avenida Nazaré 481, Ipiranga, 04263 - 000 São Paulo, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="468F8217DAA950C9DE4DE38DEF8D15AC" type="email">rafael.souza1988@gmail.com</mods:nameIdentifier>
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<mods:name id="C4D01AA4B58E90BF78597C82CAF2C32E" type="personal">
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<mods:namePart id="551AEC90859E6BEACD8014CD0E3A123E">Seidel, Matthias</mods:namePart>
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<mods:affiliation id="CF363B214A0AC34299C6494600237AB3">Centrum für Naturkunde, University of Hamburg, Martin-Luther-King-Platz 3, 20146, Hamburg, Germany.</mods:affiliation>
<mods:nameIdentifier id="9611E381668EA4734FE6814C71A01F51" type="email">matthias.seidel@uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="75FC96D3A79C79B32DA95AF7D1A11457">European Journal of Taxonomy</mods:title>
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<mods:part id="971911A8037F8E0CA1D3FBDF23828C21">
<mods:date id="E0FB69535E2479671F25CC220BCFE8C0">2021</mods:date>
<mods:detail id="B681BC190F6D2CDE80893A6BD2E9166F" type="pubDate">
<mods:number id="837164135D788B5BB7D13A00BC04745B">2021-04-30</mods:number>
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<mods:number id="CC189E2EF7D3AF20524309F359400312">748</mods:number>
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<mods:number id="D7F706ADBB9002AF7779727DE0ABE379">1</mods:number>
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<mods:classification id="1ADF1A5E223C22B1E21B516EDBAAE58A">journal article</mods:classification>
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<mods:identifier id="172771B7A267B38B61151DCAE0EFEEFA" type="DOI">10.5852/ejt.2021.748.1335</mods:identifier>
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<treatment id="C10A5544FF80FFA1151CA5B3FC2AFC8A" ID-DOI="http://doi.org/10.5281/zenodo.4737622" ID-GBIF-Taxon="181871975" ID-Zenodo-Dep="4737622" LSID="urn:lsid:plazi:treatment:C10A5544FF80FFA1151CA5B3FC2AFC8A" httpUri="http://treatment.plazi.org/id/C10A5544FF80FFA1151CA5B3FC2AFC8A" lastPageId="17" lastPageNumber="32" pageId="15" pageNumber="30">
<subSubSection id="01B9B7D9FF80FFBF151CA5B3FAA3FE49" box="[608,979,391,417]" pageId="15" pageNumber="30" type="nomenclature">
<paragraph id="491CE452FF80FFBF151CA5B3FAA3FE49" blockId="15.[605,983,391,454]" box="[608,979,391,417]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF151CA5B3FAA3FE49" box="[608,979,391,417]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="30" reason="2">
<taxonomicName id="8EA39FD1FF80FFBF151CA5B3FAA3FE49" authority="Chevrolat, 1833" authorityName="Chevrolat" authorityYear="1833" box="[608,979,391,417]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF151CA5B3FA7DFE49" bold="true" box="[608,781,391,417]" italics="true" pageId="15" pageNumber="30">Inca irroratus</emphasis>
<bibRefCitation id="2D3299A3FF80FFBF1468A5B3FAA3FE49" author="Chevrolat A." box="[788,979,391,417]" pageId="15" pageNumber="30" refId="ref9632" refString="Chevrolat A. 1833. Description d'especes nouvelles. Revue Entomologique - Coleopteres 10: unpaginated. Available from https: // www. biodiversitylibrary. org / item / 44561 # page / 349 / mode / 1 up [accessed 6 Apr. 2021]." type="journal volume" year="1833">Chevrolat, 1833</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="01B9B7D9FF80FFBF1521A59FFAA7FE2E" box="[605,983,427,454]" pageId="15" pageNumber="30" type="description">
<paragraph id="491CE452FF80FFBF1521A59FFAA7FE2E" blockId="15.[605,983,391,454]" box="[605,983,427,454]" pageId="15" pageNumber="30">
<figureCitation id="D198F8D7FF80FFBF1521A59FFA73FE2E" box="[605,771,427,454]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="15" pageNumber="30">Figs 1K–O, R</figureCitation>
,
<figureCitation id="D198F8D7FF80FFBF1473A598FA1FFE2E" box="[783,879,428,454]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. A–D, M. Inca axeli sp. nov. – E–H, N. Inca neglectus sp. nov. –I–L, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. M–O. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="15" pageNumber="30">2I–L, O</figureCitation>
,
<figureCitation id="D198F8D7FF80FFBF1407A598FACCFE2E" box="[891,956,428,454]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, G–N. I. axeli sp. nov. (A, D: ♀, G–N ♂). – B, E. I. neglectus sp. nov. – C, F. I. irroratus Chevrolat, 1833. A–C. Protibia, lateral view. D–F. Details of elytral epipleuron. G–N. Mouthparts. G–I. Right mandibles (dorsal, inner and ventral view). J–K. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="15" pageNumber="30">3C, F</figureCitation>
,
<figureCitation id="D198F8D7FF80FFBF14B4A59FFAA7FE2E" box="[968,983,427,454]" captionStart="Fig" captionStartId="16.[190,233,1998,2024]" captionTargetBox="[189,1399,1195,1959]" captionTargetId="figure-187@16.[189,1399,1195,1959]" captionTargetPageId="16" captionText="Fig. 5. Distribution map of species of the Inca irroratus group." figureDoi="http://doi.org/10.5281/zenodo.4736145" httpUri="https://zenodo.org/record/4736145/files/figure.png" pageId="15" pageNumber="30">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF80FFBF17C1A5DBFA60FDB9" pageId="15" pageNumber="30" type="reference_group">
<paragraph id="491CE452FF80FFBF17C1A5DBFBB9FDE0" blockId="15.[189,784,494,594]" box="[189,713,494,521]" pageId="15" pageNumber="30">
<treatmentCitationGroup id="69B3C37CFF80FFBF17C1A5DBFBB5FDE0" box="[189,709,494,521]" pageId="15" pageNumber="30">
<treatmentCitation id="C802C243FF80FFBF17C1A5DBFBB5FDE0" author="Chevrolat" box="[189,709,494,521]" pageId="15" pageNumber="30" year="1833">
<taxonomicName id="8EA39FD1FF80FFBF17C1A5DBFB53FDE0" ID-CoL="7FZBY" authority="Chevrolat, 1833" authorityName="Chevrolat" authorityYear="1833" box="[189,547,494,520]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irrorata">
<emphasis id="7BD73840FF80FFBF17C1A5DBF82EFDE0" box="[189,350,495,520]" italics="true" pageId="15" pageNumber="30">Ynca irrorata</emphasis>
<bibRefCitation id="2D3299A3FF80FFBF161AA5DAFB53FDE0" author="Chevrolat A." box="[358,547,494,520]" pageId="15" pageNumber="30" refId="ref9632" refString="Chevrolat A. 1833. Description d'especes nouvelles. Revue Entomologique - Coleopteres 10: unpaginated. Available from https: // www. biodiversitylibrary. org / item / 44561 # page / 349 / mode / 1 up [accessed 6 Apr. 2021]." type="journal volume" year="1833">Chevrolat, 1833</bibRefCitation>
</taxonomicName>
: unpaginated
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A620FB8DFDC5" blockId="15.[189,784,494,594]" box="[189,765,531,557]" pageId="15" pageNumber="30">
<treatmentCitationGroup id="69B3C37CFF80FFBF17C1A620FB8DFDC5" box="[189,765,531,557]" pageId="15" pageNumber="30">
<treatmentCitation id="C802C243FF80FFBF17C1A620FB8AFDC5" author="Burmeister H. &amp; Schaum H." box="[189,762,531,557]" page="380" pageId="15" pageNumber="30" year="1840">
<taxonomicName id="8EA39FD1FF80FFBF17C1A620FB8AFDC5" authority="Burmeister &amp; Schaum, 1840: 380" authorityName="Burmeister &amp; Schaum" authorityPageNumber="380" authorityYear="1840" box="[189,762,531,557]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF17C1A620F813FDC5" box="[189,355,532,557]" italics="true" pageId="15" pageNumber="30">Inca irroratus</emphasis>
<bibRefCitation id="2D3299A3FF80FFBF1617A627FB8AFDC5" author="Burmeister H. &amp; Schaum H." box="[363,762,531,557]" pageId="15" pageNumber="30" pagination="353 - 420" refId="ref9569" refString="Burmeister H. &amp; Schaum H. 1840. Kritische Revision der Lamellicornia melitophila. Erstes Stuck (Trichiadae). Zeitschrift fur die Entomologie 2: 353 - 420." type="journal article" year="1840">Burmeister &amp; Schaum, 1840: 380</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A60CFA60FDB9" blockId="15.[189,784,494,594]" box="[189,784,567,594]" pageId="15" pageNumber="30">
<treatmentCitationGroup id="69B3C37CFF80FFBF17C1A60CFA60FDB9" box="[189,784,567,594]" pageId="15" pageNumber="30">
<treatmentCitation id="C802C243FF80FFBF17C1A60CFBE3FDBA" author="Burmeister H." box="[189,659,567,594]" page="568" pageId="15" pageNumber="30" year="1847">
<taxonomicName id="8EA39FD1FF80FFBF17C1A60CFBE3FDBA" authority="Burmeister, 1847: 568" authorityName=", Burmeister" authorityPageNumber="568" authorityYear="1847" box="[189,659,567,594]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF17C1A60CF8F2FDB9" box="[189,386,567,593]" italics="true" pageId="15" pageNumber="30">Inca burmeisteri</emphasis>
<bibRefCitation id="2D3299A3FF80FFBF16F5A603FBE3FDBA" author="Burmeister H." box="[393,659,567,594]" pageId="15" pageNumber="30" refId="ref9541" refString="Burmeister H. 1847. Handbuch der Entomologie, volume 5. Coleoptera Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin." type="book" year="1847">Burmeister, 1847: 568</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="7BD73840FF80FFBF15E2A603FA60FDB9" bold="true" box="[670,784,567,593]" pageId="15" pageNumber="30">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF80FFBF17C1A64EFDC9FBE5" pageId="15" pageNumber="30" type="diagnosis">
<paragraph id="491CE452FF80FFBF17C1A64EF8B9FD7C" blockId="15.[189,457,634,660]" box="[189,457,634,660]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF17C1A64EF8B9FD7C" bold="true" box="[189,457,634,660]" fontSize="11" level="3" pageId="15" pageNumber="30" reason="3">
<emphasis id="7BD73840FF80FFBF17C1A64EF8B9FD7C" bold="true" box="[189,457,634,660]" pageId="15" pageNumber="30">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A69FFDC9FBE5" blockId="15.[188,1399,682,1037]" pageId="15" pageNumber="30">
<taxonomicName id="8EA39FD1FF80FFBF17C1A69FF815FD2C" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[189,357,683,708]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF17C1A69FF815FD2C" box="[189,357,683,708]" italics="true" pageId="15" pageNumber="30">Inca irroratus</emphasis>
</taxonomicName>
can be distinguished from the other two species of the group by: females with clypeus bearing a medial rounded fovea (fovea absent in
<taxonomicName id="8EA39FD1FF80FFBF158AA6FBFA39FD00" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[758,841,718,745]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF158AA6FBFA70FD00" box="[758,768,719,744]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1473A6FAFA39FD00" box="[783,841,718,744]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF142CA6FBFADAFD01" box="[848,938,719,745]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF149DA6FBFD1AFD00" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[993,1130,718,745]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF149DA6FBFA9BFD00" box="[993,1003,719,744]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1486A6FAFD1AFD00" box="[1018,1130,718,744]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF130DA6FBFDBFFD01" box="[1137,1231,719,745]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
); medial tooth well developed (almost indistinct in
<taxonomicName id="8EA39FD1FF80FFBF1500A6C0FA78FCE5" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[636,776,755,782]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1500A6C0FBF6FCE5" box="[636,646,756,781]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF15E4A6C7FA78FCE5" box="[664,776,755,781]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF1468A6C0FA06FCE6" box="[788,886,756,782]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
); clypeal horns in males with dorsolateral tooth short and rounded (long and acute in
<taxonomicName id="8EA39FD1FF80FFBF15B8A72CFA67FCD9" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[708,791,791,818]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF15B8A72CFBBCFCD9" box="[708,716,792,817]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF15A1A723FA67FCD9" box="[733,791,791,817]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF1463A72CFA0BFCDA" box="[799,891,792,818]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF14C5A72CFD4BFCD9" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[953,1083,792,818]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF14C5A72CFAB1FCD9" box="[953,961,792,817]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF14ADA72CFD4BFCD9" box="[977,1083,792,817]" italics="true" pageId="15" pageNumber="30">irroratus</emphasis>
</taxonomicName>
); pronotum with posterior angles rounded (acute in
<taxonomicName id="8EA39FD1FF80FFBF1696A709FB4DFCBE" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[490,573,828,854]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1696A709FB4DFCBE" box="[490,573,828,854]" italics="true" pageId="15" pageNumber="30">I. axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF1539A709FBD1FCBF" box="[581,673,829,855]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF15A1A709FA16FCBE" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[733,870,828,855]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF15A1A709FB96FCBE" box="[733,742,829,854]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF158AA708FA16FCBE" box="[758,870,828,854]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF1412A709FABCFCBF" box="[878,972,829,855]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
) and lateral margin slightly sinuous and deeply crenulated (strongly sinuous and shallowly crenulated in
<taxonomicName id="8EA39FD1FF80FFBF148BA755FD39FC93" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1015,1097,865,891]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF148BA755FA8FFC92" box="[1015,1023,865,890]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1373A755FD39FC93" box="[1039,1097,865,891]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF132FA755FDDFFC93" box="[1107,1199,865,891]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF1392A755FC06FC93" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1262,1398,865,891]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1392A755FD86FC92" box="[1262,1270,865,890]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF127AA755FC06FC93" box="[1286,1398,865,891]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF17C1A7B2F86AFC48" box="[189,282,902,928]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
); posterior tooth of protibia short and rounded (long and acute in
<taxonomicName id="8EA39FD1FF80FFBF1374A7B2FD2BFC77" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1032,1115,901,928]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1374A7B2FD62FC77" box="[1032,1042,902,927]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF135DA7B1FD2BFC77" box="[1057,1115,901,927]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF131CA7B2FDCAFC48" box="[1120,1210,902,928]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF1392A7B2FC06FC77" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1262,1398,901,928]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1392A7B2FD88FC77" box="[1262,1272,902,927]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF127AA7B1FC06FC77" box="[1286,1398,901,927]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF17C1A79EF86FFC2C" box="[189,287,938,964]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
); epipleuron with irregular and rounded marks on median area (indistinct marks in
<taxonomicName id="8EA39FD1FF80FFBF125CA79EFC07FC2C" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1312,1399,938,964]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF125CA79EFC5AFC2B" box="[1312,1322,938,963]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1241A79EFC07FC2C" box="[1341,1399,938,964]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF17C1A7FBF867FC01" box="[189,279,975,1001]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="8EA39FD1FF80FFBF1630A7FBF8A5FC00" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[332,469,974,1001]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1630A7FBF826FC00" box="[332,342,975,1000]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1619A7FAF8A5FC00" box="[357,469,974,1000]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF16A7A7FBFB49FC01" box="[475,569,975,1001]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
); outer distal process of parameres short and slightly rounded (long and acuminated in
<taxonomicName id="8EA39FD1FF80FFBF1616A7C7F8CCFBE5" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[362,444,1011,1037]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="axeli" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF1616A7C7F804FBE4" box="[362,372,1011,1036]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF16FEA7C7F8CCFBE5" box="[386,444,1011,1037]" italics="true" pageId="15" pageNumber="30">axeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF16BFA7C7FB51FBE5" box="[451,545,1011,1037]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
; long and rounded in
<taxonomicName id="8EA39FD1FF80FFBF145CA7C7FAD8FBE5" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[800,936,1011,1037]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF80FFBF145CA7C7FA5AFBE4" box="[800,810,1011,1036]" italics="true" pageId="15" pageNumber="30">I</emphasis>
.
<emphasis id="7BD73840FF80FFBF1444A7C7FAD8FBE5" box="[824,936,1011,1037]" italics="true" pageId="15" pageNumber="30">neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF80FFBF14CCA7C7FD7CFBE5" box="[944,1036,1011,1037]" pageId="15" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
) (see
<tableCitation id="0421D1E9FF80FFBF132DA7C7FDDAFBE5" box="[1105,1194,1011,1037]" captionStart="Table 2" captionStartId="12.[189,259,265,291]" captionTargetPageId="12" captionText="Table 2. Comparative table for the species of the irroratus group." httpUri="http://table.plazi.org/id/1DDCB4DAFF83FFBC17C1A53DFAB0FECC" pageId="15" pageNumber="30" tableUuid="1DDCB4DAFF83FFBC17C1A53DFAB0FECC">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF80FFBF17C1A001F8B5F92D" pageId="15" pageNumber="30" type="materials_examined">
<paragraph id="491CE452FF80FFBF17C1A001F8DDFBA7" blockId="15.[189,429,1077,1103]" box="[189,429,1077,1103]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF17C1A001F8DDFBA7" bold="true" box="[189,429,1077,1103]" fontSize="11" level="3" pageId="15" pageNumber="30" reason="3">
<emphasis id="7BD73840FF80FFBF17C1A001F8DDFBA7" bold="true" box="[189,429,1077,1103]" pageId="15" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF80FFBF1796A051FB89FB68" blockId="15.[189,1399,1125,1189]" box="[234,761,1125,1152]" pageId="15" pageNumber="30">
<emphasis id="7BD73840FF80FFBF1796A051FB56FB97" bold="true" box="[234,550,1125,1152]" pageId="15" pageNumber="30">
<typeStatus id="96185AF0FF80FFBF1796A051F822FB97" box="[234,338,1125,1151]" pageId="15" pageNumber="30" type="neotype">Neotype</typeStatus>
of
<taxonomicName id="8EA39FD1FF80FFBF1606A052FB56FB97" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[378,550,1125,1152]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF1606A052FB56FB97" bold="true" box="[378,550,1125,1152]" italics="true" pageId="15" pageNumber="30">Inca irroratus</emphasis>
</taxonomicName>
</emphasis>
(here designated)
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A0BEFC07FB4C" blockId="15.[189,1399,1125,1189]" box="[189,1399,1161,1189]" pageId="15" pageNumber="30">
<materialsCitation id="F9CBEE0FFF80FFBF17C1A0BEFC03FB4C" ID-GBIF-Occurrence="3097974323" box="[189,1395,1161,1189]" collectionCode="BMNH" country="Brazil" location="Rio de Janeiro" pageId="15" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<collectingCountry id="31B4A4C2FF80FFBF17C1A0BEF856FB4C" box="[189,294,1162,1188]" name="Brazil" pageId="15" pageNumber="30">BRAZIL</collectingCountry>
–
<emphasis id="7BD73840FF80FFBF163EA0BEF889FB4C" bold="true" box="[322,505,1162,1188]" pageId="15" pageNumber="30">
<collectingRegion id="8B672AB0FF80FFBF163EA0BEF889FB4C" box="[322,505,1162,1188]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
</emphasis>
•
<specimenCount id="5FA52FDBFF80FFBF156EA0BEFB30FB4C" box="[530,576,1162,1188]" pageId="15" pageNumber="30" type="male">1 ♂</specimenCount>
; “Rio. Jan. [Rio de Janeiro], Fry; 7517, Fry Coll. 1905-100.”;
<collectionCode id="2FB27C97FF80FFBF126BA0BEFC03FB4C" box="[1303,1395,1162,1188]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="15" pageNumber="30">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="491CE452FF80FFBF1796A0F9FA5DFB00" blockId="15.[186,1398,1228,1328]" box="[234,813,1228,1256]" pageId="15" pageNumber="30">
<emphasis id="7BD73840FF80FFBF1796A0F9FB2BFB0F" bold="true" box="[234,603,1228,1255]" pageId="15" pageNumber="30">
<typeStatus id="96185AF0FF80FFBF1796A0F9F817FB0F" box="[234,359,1229,1255]" pageId="15" pageNumber="30" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="8EA39FD1FF80FFBF16F3A0F9FB2BFB0F" authorityName=", Burmeister" authorityYear="1847" box="[399,603,1228,1255]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF16F3A0F9FB2BFB0F" bold="true" box="[399,603,1228,1255]" italics="true" pageId="15" pageNumber="30">Inca burmeisteri</emphasis>
</taxonomicName>
</emphasis>
(here designated)
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A0C5FD61FAD8" blockId="15.[186,1398,1228,1328]" pageId="15" pageNumber="30">
<materialsCitation id="F9CBEE0FFF80FFBF17C1A0C5FD7DFAD8" ID-GBIF-Occurrence="3097974308" collectionCode="MLUH" country="Brazil" location="Rio de Janeiro" pageId="15" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
<collectingCountry id="31B4A4C2FF80FFBF17C1A0C5F856FAE3" box="[189,294,1265,1291]" name="Brazil" pageId="15" pageNumber="30">BRAZIL</collectingCountry>
–
<emphasis id="7BD73840FF80FFBF1646A0C5F89CFAE3" bold="true" box="[314,492,1265,1291]" pageId="15" pageNumber="30">
<collectingRegion id="8B672AB0FF80FFBF1646A0C5F89CFAE3" box="[314,492,1265,1291]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
</emphasis>
•
<specimenCount id="5FA52FDBFF80FFBF157EA0C5FB5EFAE4" box="[514,558,1265,1292]" pageId="15" pageNumber="30" type="male">1 ♂</specimenCount>
; “Bras. [Brasil], Bske [Bescke] leg.,
<emphasis id="7BD73840FF80FFBF14B2A0C6FD2AFAE3" box="[974,1114,1266,1291]" italics="true" pageId="15" pageNumber="30">Burmeisteri</emphasis>
*, Besck,
<typeStatus id="96185AF0FF80FFBF13B4A0C5FC02FAE3" box="[1224,1394,1265,1291]" pageId="15" pageNumber="30" type="lectotype">LECTOTYPE</typeStatus>
,
<taxonomicName id="8EA39FD1FF80FFBF17C6A122FB2EFAD8" authority=", Burmeister, 1847" authorityName=", Burmeister" authorityYear="1847" box="[186,606,1302,1328]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF17C6A122F80FFAD8" box="[186,383,1302,1328]" italics="true" pageId="15" pageNumber="30">Inca burmeisteri</emphasis>
,
<bibRefCitation id="2D3299A3FF80FFBF16EDA122FB2EFAD8" author="Burmeister H." box="[401,606,1302,1328]" pageId="15" pageNumber="30" refId="ref9541" refString="Burmeister H. 1847. Handbuch der Entomologie, volume 5. Coleoptera Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin." type="book" year="1847">Burmeister, 1847</bibRefCitation>
</taxonomicName>
, des. Sousa &amp; Seidel 2020”;
<collectionCode id="2FB27C97FF80FFBF14CFA122FD7DFAD8" box="[947,1037,1302,1328]" collectionName="Germany, Halle a.S, Martin-Luther-Universitaet, Wissenschaftsbereich Zoologie" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="15" pageNumber="30">MLUH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="491CE452FF80FFBF1796A16DFBE8FA9B" blockId="15.[189,1379,1368,1432]" box="[234,664,1368,1395]" pageId="15" pageNumber="30">
<emphasis id="7BD73840FF80FFBF1796A16DFBE8FA9B" bold="true" box="[234,664,1368,1395]" pageId="15" pageNumber="30">
<typeStatus id="96185AF0FF80FFBF1796A16DF8D4FA9B" box="[234,420,1369,1395]" pageId="15" pageNumber="30" type="paralectotype">Paralectotypes</typeStatus>
of
<taxonomicName id="8EA39FD1FF80FFBF16B0A16DFBE8FA9B" authorityName=", Burmeister" authorityYear="1847" box="[460,664,1368,1395]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF16B0A16DFBE8FA9B" bold="true" box="[460,664,1368,1395]" italics="true" pageId="15" pageNumber="30">Inca burmeisteri</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A149FC13FA70" blockId="15.[189,1379,1368,1432]" box="[189,1379,1405,1432]" pageId="15" pageNumber="30">
<materialsCitation id="F9CBEE0FFF80FFBF17C1A149FC2FFA70" ID-GBIF-Occurrence="3097974327" box="[189,1375,1405,1432]" collectionCode="MLUH" country="Brazil" location="Rio de Janeiro" pageId="15" pageNumber="30" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
<collectingCountry id="31B4A4C2FF80FFBF17C1A149F856FA7F" box="[189,294,1405,1431]" name="Brazil" pageId="15" pageNumber="30">BRAZIL</collectingCountry>
–
<emphasis id="7BD73840FF80FFBF163FA149F88BFA7F" bold="true" box="[323,507,1405,1431]" pageId="15" pageNumber="30">
<collectingRegion id="8B672AB0FF80FFBF163FA149F88BFA7F" box="[323,507,1405,1431]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
</emphasis>
•
<specimenCount id="5FA52FDBFF80FFBF1569A149FB33FA70" box="[533,579,1405,1432]" pageId="15" pageNumber="30" type="male">1 ♂</specimenCount>
,
<specimenCount id="5FA52FDBFF80FFBF1533A149FBE2FA70" box="[591,658,1405,1432]" pageId="15" pageNumber="30" type="female">2 ♀♀</specimenCount>
; same collection data as for
<taxonomicName id="8EA39FD1FF80FFBF149DA14AFDF3FA7F" authorityName=", Burmeister" authorityYear="1847" box="[993,1155,1405,1431]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF149DA14AFDF3FA7F" box="[993,1155,1405,1431]" italics="true" pageId="15" pageNumber="30">I. burmeisteri</emphasis>
</taxonomicName>
<typeStatus id="96185AF0FF80FFBF13F6A149FD89FA7F" box="[1162,1273,1405,1431]" pageId="15" pageNumber="30" type="lectotype">lectotype</typeStatus>
;
<collectionCode id="2FB27C97FF80FFBF1279A14AFC2FFA70" box="[1285,1375,1406,1432]" collectionName="Germany, Halle a.S, Martin-Luther-Universitaet, Wissenschaftsbereich Zoologie" country="Germany" httpUri="http://grbio.org/cool/pvva-bewi" name="Martin Luther Universitaet" pageId="15" pageNumber="30">MLUH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A18BF8C7FA31" blockId="15.[189,439,1471,1497]" box="[189,439,1471,1497]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF17C1A18BF8C7FA31" bold="true" box="[189,439,1471,1497]" fontSize="11" level="3" pageId="15" pageNumber="30" reason="3">
<emphasis id="7BD73840FF80FFBF17C1A18BF8C7FA31" bold="true" box="[189,439,1471,1497]" pageId="15" pageNumber="30">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A1C4FBA7F9BB" blockId="15.[188,1398,1519,1619]" pageId="15" pageNumber="30">
<materialsCitation id="F9CBEE0FFF80FFBF17C1A1C4F8A1F9C7" ID-GBIF-Occurrence="3097974336" collectingDate="1999-04" collectionCode="UFRPE" collectorName="R. Rajs" country="Brazil" elevation="1200" location="Mury" municipality="Nova Friburgo" pageId="15" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro">
<collectingCountry id="31B4A4C2FF80FFBF17C1A1C4F856F9E2" box="[189,294,1520,1546]" name="Brazil" pageId="15" pageNumber="30">BRAZIL</collectingCountry>
–
<emphasis id="7BD73840FF80FFBF1643A1DBF884F9E1" bold="true" box="[319,500,1519,1546]" pageId="15" pageNumber="30">
<collectingRegion id="8B672AB0FF80FFBF1643A1DBF884F9E1" box="[319,500,1519,1546]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
</emphasis>
•
<specimenCount id="5FA52FDBFF80FFBF1577A1C4FB47F9E2" box="[523,567,1520,1546]" pageId="15" pageNumber="30" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="A9787E28FF80FFBF1538A1C4FB82F9E2" box="[580,754,1520,1546]" pageId="15" pageNumber="30">Nova Friburgo</collectingMunicipality>
,
<location id="4C7CB289FF80FFBF1581A1C4FA32F9E2" LSID="urn:lsid:plazi:treatment:C10A5544FF80FFA1151CA5B3FC2AFC8A:4C7CB289FF80FFBF1581A1C4FA32F9E2" box="[765,834,1520,1546]" country="Brazil" municipality="Nova Friburgo" name="Mury" pageId="15" pageNumber="30" stateProvince="Rio de Janeiro">Mury</location>
;
<elevation id="C28E0361FF80FFBF1430A1C4FA92F9E2" box="[844,994,1520,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="30" unit="m" value="1200.0">
<quantity id="8E5B49B7FF80FFBF1430A1C4FAD3F9E2" box="[844,931,1520,1546]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="30" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<date id="3D1DC292FF80FFBF1490A1C4FD13F9E2" box="[1004,1123,1520,1546]" pageId="15" pageNumber="30" value="1999-04">
<collectingDate id="2D593B7AFF80FFBF1490A1C4FD13F9E2" box="[1004,1123,1520,1546]" pageId="15" pageNumber="30" value="1999-04">Apr. 1999</collectingDate>
</date>
;
<collectorName id="E4568184FF80FFBF1312A1C4FDB3F9E2" box="[1134,1219,1520,1546]" pageId="15" pageNumber="30">R. Rajs</collectorName>
coll., “Coleção E &amp; P. Grossi”;
<collectionCode id="2FB27C97FF80FFBF160AA221F8A1F9C7" box="[374,465,1557,1583]" pageId="15" pageNumber="30">UFRPE</collectionCode>
</materialsCitation>
•
<materialsCitation id="F9CBEE0FFF80FFBF1695A220FBA3F9BB" ID-GBIF-Occurrence="3097974338" collectionCode="BMNH" country="Brazil" location="Rio de Janeiro" pageId="15" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="5FA52FDBFF80FFBF1695A220FB67F9C7" box="[489,535,1556,1583]" pageId="15" pageNumber="30" type="male">1 ♂</specimenCount>
; “Rio. Jan. [
<collectingRegion id="8B672AB0FF80FFBF15D8A220FA26F9C7" box="[676,854,1556,1583]" inferredBySuffix="true" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
], Fry,
<taxonomicName id="8EA39FD1FF80FFBF14E0A221FDC3F9C7" authority="Burm" authorityName="Burm" box="[924,1203,1556,1583]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF80FFBF14E0A221FD17F9C6" box="[924,1127,1556,1582]" italics="true" pageId="15" pageNumber="30">Ynca burmeisteri</emphasis>
Burm
</taxonomicName>
,
<taxonomicName id="8EA39FD1FF80FFBF13C1A221F873F9BB" authority="(Oliv) Burm" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF80FFBF13C1A221FC57F9C6" box="[1213,1319,1557,1582]" italics="true" pageId="15" pageNumber="30">irroratus</emphasis>
(Oliv) Burm
</taxonomicName>
; Brasilia, Fry Coll. 1905-100”;
<collectionCode id="2FB27C97FF80FFBF150BA20DFBA3F9BB" box="[631,723,1593,1619]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="15" pageNumber="30">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A248F82EF97D" blockId="15.[189,350,1659,1686]" box="[189,350,1659,1686]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF17C1A248F82EF97D" bold="true" box="[189,350,1659,1686]" fontSize="11" level="3" pageId="15" pageNumber="30" reason="3">
<emphasis id="7BD73840FF80FFBF17C1A248F82EF97D" bold="true" box="[189,350,1659,1686]" pageId="15" pageNumber="30">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A29FF8B5F92D" blockId="15.[189,453,1707,1733]" box="[189,453,1707,1733]" pageId="15" pageNumber="30">
<collectingRegion id="8B672AB0FF80FFBF17C1A29FF81AF92D" box="[189,362,1707,1733]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="30">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="31B4A4C2FF80FFBF1609A29FF8B0F92D" box="[373,448,1707,1733]" name="Brazil" pageId="15" pageNumber="30">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF80FFA017C1A2DAFD7FFC6C" lastPageId="16" lastPageNumber="31" pageId="15" pageNumber="30" type="description">
<paragraph id="491CE452FF80FFBF17C1A2DAF81DF8E0" blockId="15.[189,365,1774,1800]" box="[189,365,1774,1800]" pageId="15" pageNumber="30">
<heading id="1254533EFF80FFBF17C1A2DAF81DF8E0" bold="true" box="[189,365,1774,1800]" fontSize="11" level="3" pageId="15" pageNumber="30" reason="3">
<emphasis id="7BD73840FF80FFBF17C1A2DAF81DF8E0" bold="true" box="[189,365,1774,1800]" pageId="15" pageNumber="30">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF80FFBF1796A32AF85AF8D0" blockId="15.[189,1398,1822,1922]" box="[234,298,1822,1848]" pageId="15" pageNumber="30">
<emphasis id="7BD73840FF80FFBF1796A32AF85AF8D0" bold="true" box="[234,298,1822,1848]" pageId="15" pageNumber="30">Male</emphasis>
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A377FB09F86A" blockId="15.[189,1398,1822,1922]" pageId="15" pageNumber="30">
BODY (
<figureCitation id="D198F8D7FF80FFBF1668A376F8EFF8B5" box="[276,415,1858,1885]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="15" pageNumber="30">Fig. 1K–M</figureCitation>
). Total length including clypeal horns
<quantity id="8E5B49B7FF80FFBF1412A377FA9FF8B5" box="[878,1007,1859,1885]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="4.7" metricValueMin="4.3" pageId="15" pageNumber="30" unit="mm" value="45.0" valueMax="47.0" valueMin="43.0">43–47 mm</quantity>
(
<typeStatus id="96185AF0FF80FFBF137DA377FD14F8B5" box="[1025,1124,1859,1885]" pageId="15" pageNumber="30" type="neotype">neotype</typeStatus>
:
<quantity id="8E5B49B7FF80FFBF130DA377FDBBF8B5" box="[1137,1227,1859,1885]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="15" pageNumber="30" unit="mm" value="44.0">44 mm</quantity>
); width across humeri
<quantity id="8E5B49B7FF80FFBF1666A353F8E8F86A" box="[282,408,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="15" pageNumber="30" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
(
<typeStatus id="96185AF0FF80FFBF16DBA353FB7AF869" box="[423,522,1895,1921]" pageId="15" pageNumber="30" type="neotype">neotype</typeStatus>
:
<quantity id="8E5B49B7FF80FFBF1569A353FB1DF86A" box="[533,621,1895,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="30" unit="mm" value="18.0">18 mm</quantity>
).
</paragraph>
<paragraph id="491CE452FF80FFBF17C1A39EFBBCF801" blockId="15.[189,1398,1962,2025]" pageId="15" pageNumber="30">
COLOUR. Reddish brown; dorsal surface with metallic reflections; legs and meso- and metathoraX with reddish brown bright colour (
<figureCitation id="D198F8D7FF80FFBF1569A3FAFBCEF801" box="[533,702,1998,2025]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="15" pageNumber="30">Fig. 1K–M, R</figureCitation>
).
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A53DFDABFEAE" blockId="16.[188,1398,265,327]" pageId="16" pageNumber="31">
HEAD. Surface of frons with reddish brown waxy secretion; clypeal horns with inner dorsal carina weak, anterior area gradually interrupted at apex; dorsolateral tooth short and rounded (
<figureCitation id="D198F8D7FF9FFFA0130EA518FDBEFEAE" box="[1138,1230,300,326]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="16" pageNumber="31">Fig. 1R</figureCitation>
).
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A55AFBBEFDFD" blockId="16.[188,1399,366,533]" pageId="16" pageNumber="31">
THORAX. Lateral margin of pronotum slightly sinuous and deeply crenulated; lateromedial area with grooved fovea; longitudinal groove shallow; posterior angles rounded (
<figureCitation id="D198F8D7FF9FFFA0137DA5A5FDFAFE43" box="[1025,1162,401,427]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="16" pageNumber="31">Fig. 1K–M</figureCitation>
); anterior prosternal process evenly rounded, projected and densely setose in median area. Scutellar shield punctate in anterolateral area. Elytra with light yellow waxy maculae covering all surface; epipleuron with irregular and rounded marks on median area (
<figureCitation id="D198F8D7FF9FFFA01514A5CEFBB1FDFD" box="[616,705,506,533]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, G–N. I. axeli sp. nov. (A, D: ♀, G–N ♂). – B, E. I. neglectus sp. nov. – C, F. I. irroratus Chevrolat, 1833. A–C. Protibia, lateral view. D–F. Details of elytral epipleuron. G–N. Mouthparts. G–I. Right mandibles (dorsal, inner and ventral view). J–K. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="16" pageNumber="31">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A608FDAFFDBE" blockId="16.[189,1247,572,598]" box="[189,1247,572,598]" pageId="16" pageNumber="31">
LEGS. Posterior tooth of protibia short and rounded (
<figureCitation id="D198F8D7FF9FFFA0145FA608FA0FFDBE" box="[803,895,572,598]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, G–N. I. axeli sp. nov. (A, D: ♀, G–N ♂). – B, E. I. neglectus sp. nov. – C, F. I. irroratus Chevrolat, 1833. A–C. Protibia, lateral view. D–F. Details of elytral epipleuron. G–N. Mouthparts. G–I. Right mandibles (dorsal, inner and ventral view). J–K. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="16" pageNumber="31">Fig. 3C</figureCitation>
). Mesempodium with 2 setae.
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A649F8A2FD53" blockId="16.[189,1398,637,699]" pageId="16" pageNumber="31">ABDOMEN. Fovea of sternite VII slightly marked. Disc of pygidium densely punctate, lateral area with well-defined punctures.</paragraph>
<paragraph id="491CE452FF9FFFA017C1A6D6FC49FD14" blockId="16.[189,1337,738,764]" box="[189,1337,738,764]" pageId="16" pageNumber="31">
TERMINALIA. Aedeagus: outer distal process of parameres short and slightly rounded (
<figureCitation id="D198F8D7FF9FFFA013D5A6D6FC5BFD14" box="[1193,1323,738,764]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="16" pageNumber="31">Fig. 1N–O</figureCitation>
).
</paragraph>
<paragraph id="491CE452FF9FFFA01796A717F836FCD5" blockId="16.[189,1398,803,900]" box="[234,326,803,829]" pageId="16" pageNumber="31">
<emphasis id="7BD73840FF9FFFA01796A717F836FCD5" bold="true" box="[234,326,803,829]" pageId="16" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A773FD7FFC6C" blockId="16.[189,1398,803,900]" pageId="16" pageNumber="31">
BODY (
<figureCitation id="D198F8D7FF9FFFA01668A772F8FFFC89" box="[276,399,838,865]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. A–D, M. Inca axeli sp. nov. – E–H, N. Inca neglectus sp. nov. –I–L, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. M–O. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="16" pageNumber="31">Fig. 2I–K</figureCitation>
). Length 44.0–
<quantity id="8E5B49B7FF9FFFA0153AA772FBC3FC89" box="[582,691,838,865]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="16" pageNumber="31" unit="mm" value="48.5">48.5 mm</quantity>
; width across humeri 18.0–
<quantity id="8E5B49B7FF9FFFA0137EA772FD1FFC89" box="[1026,1135,838,865]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="16" pageNumber="31" unit="mm" value="19.5">19.5 mm</quantity>
. Disc of clypeus with large and rounded fovea below process of posteromedian area (
<figureCitation id="D198F8D7FF9FFFA014DFA75EFD72FC6C" box="[931,1026,874,900]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. A–D, M. Inca axeli sp. nov. – E–H, N. Inca neglectus sp. nov. –I–L, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. M–O. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="16" pageNumber="31">Fig. 2O</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF9FFFA017C1A79FFA5FFC1C" box="[189,815,939,1013]" pageId="16" pageNumber="31" type="distribution">
<paragraph id="491CE452FF9FFFA017C1A79FF828FC2D" blockId="16.[189,344,939,965]" box="[189,344,939,965]" pageId="16" pageNumber="31">
<heading id="1254533EFF9FFFA017C1A79FF828FC2D" bold="true" box="[189,344,939,965]" fontSize="11" level="3" pageId="16" pageNumber="31" reason="3">
<emphasis id="7BD73840FF9FFFA017C1A79FF828FC2D" bold="true" box="[189,344,939,965]" pageId="16" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF9FFFA017C1A7EEFA5FFC1C" blockId="16.[189,815,985,1013]" box="[189,815,985,1013]" pageId="16" pageNumber="31">
<collectingCountry id="31B4A4C2FF9FFFA017C1A7EEF879FC1C" box="[189,265,986,1012]" name="Brazil" pageId="16" pageNumber="31">Brazil</collectingCountry>
:
<collectingRegion id="8B672AB0FF9FFFA01668A7EEFB70FC1C" box="[276,512,986,1012]" country="Brazil" name="Rio de Janeiro" pageId="16" pageNumber="31">Rio de Janeiro State</collectingRegion>
(Nova Friburgo) (
<figureCitation id="D198F8D7FF9FFFA015ABA7EEFA6FFC1C" box="[727,799,985,1012]" captionStart="Fig" captionStartId="16.[190,233,1998,2024]" captionTargetBox="[189,1399,1195,1959]" captionTargetId="figure-187@16.[189,1399,1195,1959]" captionTargetPageId="16" captionText="Fig. 5. Distribution map of species of the Inca irroratus group." figureDoi="http://doi.org/10.5281/zenodo.4736145" httpUri="https://zenodo.org/record/4736145/files/figure.png" pageId="16" pageNumber="31">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1DDCB4DAFF9FFFA017C2A3FAFAD4F801" ID-DOI="http://doi.org/10.5281/zenodo.4736145" ID-Zenodo-Dep="4736145" box="[190,932,1998,2025]" httpUri="https://zenodo.org/record/4736145/files/figure.png" pageId="16" pageNumber="31" startId="16.[190,233,1998,2024]" targetBox="[189,1399,1195,1959]" targetPageId="16">
<paragraph id="491CE452FF9FFFA017C2A3FAFAD4F801" blockId="16.[190,932,1998,2025]" box="[190,932,1998,2025]" pageId="16" pageNumber="31">
<emphasis id="7BD73840FF9FFFA017C2A3FAF87DF800" bold="true" box="[190,269,1998,2024]" pageId="16" pageNumber="31">Fig. 5.</emphasis>
Distribution map of species of the
<taxonomicName id="8EA39FD1FF9FFFA015D6A3FBFA20F800" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[682,848,1999,2024]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="31" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9FFFA015D6A3FBFA20F800" box="[682,848,1999,2024]" italics="true" pageId="16" pageNumber="31">Inca irroratus</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<subSubSection id="01B9B7D9FF9EFFA117C1A53DFC2AFC8A" pageId="17" pageNumber="32" type="discussion">
<paragraph id="491CE452FF9EFFA117C1A53DF840FECB" blockId="17.[189,304,265,291]" box="[189,304,265,291]" pageId="17" pageNumber="32">
<heading id="1254533EFF9EFFA117C1A53DF840FECB" bold="true" box="[189,304,265,291]" fontSize="11" level="3" pageId="17" pageNumber="32" reason="3">
<emphasis id="7BD73840FF9EFFA117C1A53DF840FECB" bold="true" box="[189,304,265,291]" pageId="17" pageNumber="32">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF9EFFA117C1A503FAD0FD5E" blockId="17.[188,1400,311,694]" pageId="17" pageNumber="32">
The
<typeStatus id="96185AF0FF9EFFA1178DA503F828FEB9" box="[241,344,311,337]" pageId="17" pageNumber="32" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8EA39FD1FF9EFFA11602A50CFB70FEB9" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[382,512,312,337]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9EFFA11602A50CFB70FEB9" box="[382,512,312,337]" italics="true" pageId="17" pageNumber="32">I. irroratus</emphasis>
</taxonomicName>
cannot be found in the Muséum national d’histoire naturelle (Paris,
<collectingCountry id="31B4A4C2FF9EFFA11261A50CFC1FFEBA" box="[1309,1391,312,338]" name="France" pageId="17" pageNumber="32">France</collectingCountry>
) or in the Natural History Museum (BMNH, London,
<collectingCountry id="31B4A4C2FF9EFFA11439A56FFD60FE9E" box="[837,1040,347,374]" name="United Kingdom" pageId="17" pageNumber="32">United Kingdom</collectingCountry>
) and we therefore consider it lost. The description by Chevrolat is rather general and does not allow to precisely identify the species. According to Chevrolat, the species is “bearing a very short median horn preceded by the transverse carina”, a combination of characters none of the two species in question possess. Females exhibit an elevated carina on the head only in one species (
<figureCitation id="D198F8D7FF9EFFA1147FA5DDFA12FDEB" box="[771,866,489,515]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. A–D, M. Inca axeli sp. nov. – E–H, N. Inca neglectus sp. nov. –I–L, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. M–O. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="17" pageNumber="32">Fig. 2O</figureCitation>
), whereas the other (
<figureCitation id="D198F8D7FF9EFFA11321A5DDFDCAFDEB" box="[1117,1210,489,515]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. A–D, M. Inca axeli sp. nov. – E–H, N. Inca neglectus sp. nov. –I–L, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. M–O. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="17" pageNumber="32">Fig. 2N</figureCitation>
) only possesses a cone-shaped tubercle. None of the species have a horn. Additionally, based on the illustration of the type of
<taxonomicName id="8EA39FD1FF9EFFA11669A605F8E7FDA2" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[277,407,561,586]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9EFFA11669A605F8E7FDA2" box="[277,407,561,586]" italics="true" pageId="17" pageNumber="32">I. irroratus</emphasis>
</taxonomicName>
by
<bibRefCitation id="2D3299A3FF9EFFA116BFA604FB81FDA3" author="Gory H. &amp; Percheron A." box="[451,753,560,587]" pageId="17" pageNumber="32" refId="ref9872" refString="Gory H. &amp; Percheron A. 1833. Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris." type="book" year="1833">Gory &amp; Percheron (1833)</bibRefCitation>
, which lacks carina and horn, the female has a strongly crenulated pronotal margin and a medial tooth in the clypeal margin. To fiX the identity of
<taxonomicName id="8EA39FD1FF9EFFA11397A660FC1FFD85" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[1259,1391,596,621]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9EFFA11397A660FC1FFD85" box="[1259,1391,596,621]" italics="true" pageId="17" pageNumber="32">I. irroratus</emphasis>
</taxonomicName>
, we designate a male
<typeStatus id="96185AF0FF9EFFA116CCA64CFB7DFD7A" box="[432,525,632,658]" pageId="17" pageNumber="32" type="neotype">neotype</typeStatus>
(conspecific with females possessing the aforementioned characters) from BMNH (
<figureCitation id="D198F8D7FF9EFFA11659A6AFF8B7FD5D" box="[293,455,667,693]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. A–E, P. Inca axeli sp. nov., holotype (MZSP 22036). – F–J, Q. Inca neglectus sp. nov., paratype (MZSP). – K–O, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. P–R. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="17" pageNumber="32">Fig. 1K–O, R</figureCitation>
) from the type locality (Rio de Janeiro).
</paragraph>
<paragraph id="491CE452FF9EFFA117C1A6E9FC2AFC8A" blockId="17.[189,1400,732,866]" pageId="17" pageNumber="32">
Not knowing the exact number of
<typeStatus id="96185AF0FF9EFFA11513A6E9FBA6FD1F" box="[623,726,733,759]" pageId="17" pageNumber="32" type="syntype">syntypes</typeStatus>
and their depositories and to guarantee nomenclatural stability, we designate a male
<typeStatus id="96185AF0FF9EFFA11563A734FBFBFCF2" box="[543,651,768,794]" pageId="17" pageNumber="32" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="8EA39FD1FF9EFFA115C1A735FA10FCF2" box="[701,864,768,794]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF9EFFA115C1A735FA10FCF2" box="[701,864,768,794]" italics="true" pageId="17" pageNumber="32">I. burmeisteri</emphasis>
</taxonomicName>
from the
<typeStatus id="96185AF0FF9EFFA114A9A735FD40FCF3" box="[981,1072,769,795]" pageId="17" pageNumber="32" type="syntype">syntype</typeStatus>
series deposited in MLUH. The
<typeStatus id="96185AF0FF9EFFA1178FA710F82FFCD6" box="[243,351,804,830]" pageId="17" pageNumber="32" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="8EA39FD1FF9EFFA116F4A710FB5BFCD6" box="[392,555,804,830]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF9EFFA116F4A710FB5BFCD6" box="[392,555,804,830]" italics="true" pageId="17" pageNumber="32">I. burmeisteri</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="8EA39FD1FF9EFFA1146BA710FAEBFCD5" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[791,923,804,829]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9EFFA1146BA710FAEBFCD5" box="[791,923,804,829]" italics="true" pageId="17" pageNumber="32">I. irroratus</emphasis>
</taxonomicName>
, and therefore
<taxonomicName id="8EA39FD1FF9EFFA1132CA710FD83FCD6" box="[1104,1267,804,830]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="7BD73840FF9EFFA1132CA710FD83FCD6" box="[1104,1267,804,830]" italics="true" pageId="17" pageNumber="32">I. burmeisteri</emphasis>
</taxonomicName>
becomes a junior synonym. For
<taxonomicName id="8EA39FD1FF9EFFA116C9A77CFA11FC8A" authority="sensu Burmeister (1847)" authorityName="sensu Burmeister" authorityYear="1847" baseAuthorityName="Oliv" box="[437,865,839,866]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF9EFFA116C9A77CFB48FC89" box="[437,568,840,865]" italics="true" pageId="17" pageNumber="32">I. irroratus</emphasis>
sensu
<bibRefCitation id="2D3299A3FF9EFFA115F4A773FA11FC8A" author="Burmeister H." box="[648,865,839,866]" pageId="17" pageNumber="32" refId="ref9541" refString="Burmeister H. 1847. Handbuch der Entomologie, volume 5. Coleoptera Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin." type="book" year="1847">Burmeister (1847)</bibRefCitation>
</taxonomicName>
, see the description of
<taxonomicName id="8EA39FD1FF9EFFA1130CA77CFD88FC89" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1136,1272,839,865]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF9EFFA1130CA77CFD88FC89" box="[1136,1272,839,865]" italics="true" pageId="17" pageNumber="32">I. neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF9EFFA1127CA77CFC2AFC8A" box="[1280,1370,840,866]" pageId="17" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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