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<mods:title id="6A23074C326ABE912971D55406781113">A genus-level taxonomic review of primitively segmented spiders (Mesothelae, Liphistiidae)</mods:title>
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<mods:namePart id="86BDAB55ADAB6C0A69C9EC13FCF97967">Xu, Xin</mods:namePart>
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<mods:namePart id="DBD17253040990C3EBBEE1406533B3EF">Ono, Hirotsugu</mods:namePart>
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<paragraph id="793B95BACEA90502A073DAF8749C1A1F" pageId="5" pageNumber="126">Taxon classification Animalia Araneae Liphistiidae</paragraph>
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<subSubSection id="8C6273549B0E9ECB0D6D608648DEF534" pageId="5" pageNumber="126" type="nomenclature">
<paragraph id="7393A5382721D273F9E90B3C5E67DF1A" pageId="5" pageNumber="126">
Genus
<taxonomicName id="8F85B6FAB8F6651C0F3F80A9A3A4D25F" ID-CoL="4M6G" ID-ENA="1649362" LSID="http://zoobank.org/F70E24D5-C13B-4195-825C-A69A684AB893" authority="Xu &amp; Kuntner" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela" order="Araneae" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">Ganthela Xu &amp; Kuntner</taxonomicName>
<taxonomicNameLabel id="3D17C78B794475D49B46F0FA3D3FCE24" pageId="5" pageNumber="126">gen. n.</taxonomicNameLabel>
Figures 10-14
</paragraph>
</subSubSection>
<subSubSection id="C3B679860F0AD850570A757565975F6F" pageId="6" pageNumber="127" type="reference_group">
<paragraph id="B1DD82BE0CEDF6BD99E648FDDA3B9465" pageId="6" pageNumber="127">
<taxonomicName id="49A6EE0FAFF20F343861DB1302427E9A" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">
<pageBreakToken id="C587E70334BEB814150F5AF80CBBA771" pageId="6" pageNumber="127" start="start">Liphistius</pageBreakToken>
</taxonomicName>
:
<bibRefCitation id="0839C0B16A9FFBE2982517FA7A992D9E" author="Wang, JF" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="22" pageNumber="143" pagination="30 - 32" title="A new species of spider of the genus Liphistius from south China (Araneae: Liphistiidae)." volume="14" year="1989">Wang 1989</bibRefCitation>
, P. 30, description of
<taxonomicName id="F3FE1DB3533C39DD51D06520A24948CA" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Liphistius cipingensis</taxonomicName>
(=
<taxonomicName id="4CAA7BF9BFD579BF06DC91647D1E21F0" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Ganthela cipingensis</taxonomicName>
).
</paragraph>
<paragraph id="B905BD93A17DD25B8FB5D2D95A4AF358" pageId="6" pageNumber="127">
<taxonomicName id="BD044F73F0B4ED3B5C008F0C079EA9A7" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Songthela</taxonomicName>
:
<bibRefCitation id="B9F3FACB1B5FE1DF184AE2E16F8ADD3E" author="Ono, H" journalOrPublisher="Memoirs of the National Science Museum" pageId="20" pageNumber="141" pagination="145 - 151" title="Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae)." volume="33" year="2000">Ono 2000</bibRefCitation>
, P. 150, transferred
<taxonomicName id="44320613308556C71D68CBFB34A164A2" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Liphistius cipingensis</taxonomicName>
to
<taxonomicName id="AA489BF371CA390E4A7843A9DABE5C3F" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Songthela cipingensis</taxonomicName>
(=
<taxonomicName id="4F05F8E9FBEBBA3AC3DE177327F8A4C2" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Ganthela cipingensis</taxonomicName>
).
</paragraph>
<paragraph id="C13F7CCB46323E7017BA393E330CAE30" pageId="6" pageNumber="127">
<taxonomicName id="05165FEC041AB1773BB6704044EDEA08" class="Arachnida" family="Heptathelidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Heptathela</taxonomicName>
:
<bibRefCitation id="D53345995D29B121B7FE6363B3BADD43" author="Platnick, NI" journalOrPublisher="The New York Entomological Society, New York" pageId="20" pageNumber="141" title="Advances in spider taxonomy 1988 - 1991, with synonymies and transfers 1940 - 1980." year="1993">Platnick 1993</bibRefCitation>
, P. 77, transferred
<taxonomicName id="48D7CDECB087A4329881484CA435214D" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Liphistius cipingensis</taxonomicName>
to
<taxonomicName id="A6A15AF2BFACEC141B3B047F299DB6BE" class="Arachnida" family="Heptathelidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela cipingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipingensis">Heptathela cipingensis</taxonomicName>
(=
<taxonomicName id="66998C806A48F9B4874827DD256E5AA1" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela cipengensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="cipengensis">Ganthela cipengensis</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="54F7F4E0EA792B574289F61BF1583831" pageId="6" pageNumber="127" type="type species">
<paragraph id="418D4551B7665D6D83A8AA41182AD581" pageId="6" pageNumber="127">Type species.</paragraph>
<paragraph id="766891BC8D48E25A34722997658AF2FF" pageId="6" pageNumber="127">
<taxonomicName id="0C7C975AAC05890A9969BF1A65270EDE" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela yundingensis" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="yundingensis">Ganthela yundingensis</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection id="52884A1B937A25AF32F4AA86E8CE1AF3" pageId="6" pageNumber="127" type="etymology">
<paragraph id="53D31ADE4DA5C0A214CCFBE5422A83B5" pageId="6" pageNumber="127">Etymology.</paragraph>
<paragraph id="DFB6FD966ACBAC4C1BA6288AB1423C4F" pageId="6" pageNumber="127">
The genera of heptathelines contain in their name the word
<normalizedToken id="528D3B3BB401921885CB059EE4D72705" originalValue="thela">'thela'</normalizedToken>
referring to spinnerets as the Greek word thele means nipple-like protuberance (
<bibRefCitation id="EBBC54D4331F5409FC1022B8DACFDCBD" author="Ono, H" journalOrPublisher="Memoirs of the National Science Museum" pageId="20" pageNumber="141" pagination="145 - 151" title="Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae)." volume="33" year="2000">Ono 2000</bibRefCitation>
). We continue this tradition, but name the genus to start with Gan-, which refers to Jiangxi Province.
</paragraph>
</subSubSection>
<subSubSection id="DE2FFAD873327E785E0AF4A321369390" pageId="6" pageNumber="127" type="diagnosis">
<paragraph id="C74DC32D7224E18B4A77F1CE98845C6E" pageId="6" pageNumber="127">Diagnosis.</paragraph>
<paragraph id="9032CB0B04BD1EAB10BD3DC141EB0D22" pageId="6" pageNumber="127">
Males of
<taxonomicName id="F471F596E27E53F819A0A51053AA063B" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela" order="Araneae" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Ganthela</taxonomicName>
differs from all other
<taxonomicName id="E6466122FF3AB1ED6A76C3B101284EAA" lsidName="" pageId="6" pageNumber="127" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
genera by a smooth conductor with a distal spiniform apex (Figures 10-11), a flat opening embolus and scale-like contrategulum (Figures 10-11), females can be identified by a single pair of similar receptacular clusters (Figures 13-14).
</paragraph>
</subSubSection>
<subSubSection id="3A622FA5769048D089B7C0C09599EB8C" pageId="7" pageNumber="128" type="description">
<paragraph id="AAFCBB0F1F04A6C75A96B109053869FC" pageId="7" pageNumber="128">
<pageBreakToken id="EB21C410F6D5DCC38184F0CAC56E493B" pageId="7" pageNumber="128" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph id="81E0597CB491A8A7773FB62E8DAC5DC8" pageId="7" pageNumber="128">Total length (excluding chelicerae) = 8-15 mm (N = 35); male palpal conductor smooth, wide, leaf-shaped, with a spiniform apex (Figures 10-11); spinose paracymbium relative short (Figures 10, 12); embolus with a flat opening (Figures 10-11); contrategulum scale-like with a smooth margin (Figures 10-11); female genitalia with paired receptacular clusters of similar size, situated at the anterior margin of the bursa copulatrix with tubular stems (Figures 13-14).</paragraph>
</subSubSection>
<subSubSection id="6EEE73C8B85B5DA817E1546E221D24D5" pageId="7" pageNumber="128" type="species composition">
<paragraph id="F5A094BF7391DC3A9D8EB2D2ECA47E23" pageId="7" pageNumber="128">Species composition.</paragraph>
<paragraph id="448BF5034463A112DA4BB7F9144DCB6E" pageId="7" pageNumber="128">
<taxonomicName id="8ED95C68AA61D83CE61A6683983B0268" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela cipingensis" order="Araneae" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="cipingensis">Ganthela cipingensis</taxonomicName>
(Wang, 1989), comb. n. (7♀), male is unknown,
<taxonomicName id="4EB4B202CA09C12D9EA5BAFC77A57C40" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela yundingensis" order="Araneae" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="yundingensis">Ganthela yundingensis</taxonomicName>
Xu, sp. n. (1♂1♀), one undescribed species from Jiangxi Province, China (11♀), and four undescribed species from Fujian Province, China (1♀, 1♂11♀, 1♀, and 3♀, respectively).
</paragraph>
</subSubSection>
<subSubSection id="3E63492A1BD590CDA15069EB58D48286" pageId="7" pageNumber="128" type="distribution">
<paragraph id="4C96294BADECB1F4B8DBE1B5897501A6" pageId="7" pageNumber="128">Distribution.</paragraph>
<paragraph id="006DC9084817B9FF91F8C20AF9FC41CC" pageId="7" pageNumber="128">China (Fujian, Jiangxi).</paragraph>
</subSubSection>
<subSubSection id="4EFBEE33B368FDC1741D9035219F52CC" pageId="7" pageNumber="128" type="remarks">
<paragraph id="690BC0D290448C89FC9850E2269D963A" pageId="7" pageNumber="128">Remarks.</paragraph>
<paragraph id="A131F496E7690A14F4E57D64E92117C9" pageId="7" pageNumber="128">
<bibRefCitation id="3F9979244FEE289599290645CD98F43D" author="Wang, JF" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="22" pageNumber="143" pagination="30 - 32" title="A new species of spider of the genus Liphistius from south China (Araneae: Liphistiidae)." volume="14" year="1989">Wang (1989)</bibRefCitation>
placed
<taxonomicName id="187E0E2FCF36EFCBC3EC602A0B7D9BEA" class="Arachnida" family="Heptathelidae" genus="Ganthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganthela cipingensis" order="Araneae" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="cipingensis">Ganthela cipingensis</taxonomicName>
in
<taxonomicName id="593A43C6AB3761D997E1A4644DB82C3D" class="Arachnida" family="Liphistiidae" genus="Liphistius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liphistius" order="Araneae" pageId="7" pageNumber="128" phylum="Arthropoda" rank="genus">Liphistius</taxonomicName>
based on the presence of eight spinnerets. Our collections from the type locality contain specimens with seven spinnerets. The number of spinnerets thus varies intraspecifically.
</paragraph>
</subSubSection>
</treatment>
</document>