treatments-xml/data/BE/A5/0F/BEA50FCA88F98E51ECA3890F6D7F632E.xml

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<mods:title id="7BC0807D1C76BB742B154FDE1EAAC9EC">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="44391A799FA34E98F9987F7112483372">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="F6E0306AC649935921CFA0721FCB3DBC">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<treatment id="BEA50FCA88F98E51ECA3890F6D7F632E" ID-GBIF-Taxon="182223711" LSID="urn:lsid:zoobank.org:act:6839CC5D-ED40-4D10-BE41-1717E4A57D8E" httpUri="http://treatment.plazi.org/id/BEA50FCA88F98E51ECA3890F6D7F632E" lastPageId="43" lastPageNumber="254" pageId="41" pageNumber="252">
<subSubSection id="6200AD02F61086620C9952BC80A60557" pageId="41" pageNumber="252" type="nomenclature">
<paragraph id="D4FF9AA34D86095BA205AA12165B94BE" pageId="41" pageNumber="252">
21. 
<taxonomicName id="63BC406F83644E4844F6A67CAFABE205" ID-CoL="6C8Z3" LSID="http://zoobank.org/6839CC5D-ED40-4D10-BE41-1717E4A57D8E" authority="Liebherr, 2016" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus subulatus" order="Coleoptera" pageId="41" pageNumber="252" phylum="Arthropoda" rank="species" species="subulatus" status="sp. n.">Cyphocoleus subulatus</taxonomicName>
<taxonomicNameLabel id="053F8F09A37F8F39551882589DFA54CF" pageId="41" pageNumber="252">sp. n.</taxonomicNameLabel>
<figureCitation id="0FC21F2DE9DF93093DE9F02560EA8700" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cyphocoleus subulatus sp. n. male paratype dorsal view: NEW CALEDONIA, Ciu, near Mt. Canala, 300 m el., 1 - 3 - i- 1955, rainforest, E. O. Wilson (MCZ)." figureDoi="10.3897/dez.63.10241.figure1" httpUri="https://binary.pensoft.net/fig/111537" pageId="41" pageNumber="252">Figures 1</figureCitation>
<figureCitation id="C55ADBD219FFE31754D929BB2DC29DA3" captionStart="Figures 26–27" captionStartId="F4" captionText="Figures 26 - 27. Right maxillary stipes, lacinia, galea, and palp, Cyphocoleus subulatus: 26, dorsal view; 27, ventral view. Single-segmented galea of Cyphocoleus and other Homethina lies laterad the lacinia, with a rabbeted groove on the dorsomedial surface of the galea accommodating the lateral margin of the lacinia, thereby supporting lacinia during alternating, rotary movements of the maxillae. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures26-27" httpUri="https://binary.pensoft.net/fig/111540" pageId="41" pageNumber="252">, 26</figureCitation>
<figureCitation id="6EBB3FA8638E1660503DFD52AD891681" captionStart="Figures 26–27" captionStartId="F4" captionText="Figures 26 - 27. Right maxillary stipes, lacinia, galea, and palp, Cyphocoleus subulatus: 26, dorsal view; 27, ventral view. Single-segmented galea of Cyphocoleus and other Homethina lies laterad the lacinia, with a rabbeted groove on the dorsomedial surface of the galea accommodating the lateral margin of the lacinia, thereby supporting lacinia during alternating, rotary movements of the maxillae. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures26-27" httpUri="https://binary.pensoft.net/fig/111540" pageId="41" pageNumber="252">, 27</figureCitation>
<figureCitation id="312D204FE0DDAB12DBBDBC41315CC070" captionStart="Figures 28–33" captionStartId="F5" captionText="Figures 28 - 33. 28 - 32 Male aedeagus with associated abdominal segments VIII and IX, dorsal view, Cyphocoleus spp.: 28, Cyphocoleus heterogenus; 29, Cyphocoleus angustatus; 30, Cyphocoleus mirabilis; 31, Cyphocoleus cordatus; 32, Cyphocoleus subulatus. 33. Abdominal lateroventrites VIII (left) and laterotergites VIII (right), showing membranous division of laterotergites, male Cyphocoleus latipennis. Dorsal gland reservoir is globose without dorsal lobe. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures28-33" httpUri="https://binary.pensoft.net/fig/111541" pageId="41" pageNumber="252">, 32</figureCitation>
<figureCitation id="4B41C2AD44799952428EB61E70E97162" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="41" pageNumber="252">, 112</figureCitation>
<figureCitation id="4C03CC751C549BFC85DAFF6D6A08589A" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="41" pageNumber="252">, 120</figureCitation>
<figureCitation id="F21C586D422618F7B1E3580973539636" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="41" pageNumber="252">, 155-158</figureCitation>
<figureCitation id="DC8DDA3587226BEE44B9A0B62DDCEE80" captionStart="Figures 161–164" captionStartId="F24" captionText="Figures 161 - 164. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 161, Cyphocoleus flavipes Yahoue; 162, Cyphocoleus bourailensis, Bourail; 163, Cyphocoleus subulatus, Mt. Koghi; 124, Cyphocoleus iledespinsensis, Ile des Pins." figureDoi="10.3897/dez.63.10241.figures161-164" httpUri="https://binary.pensoft.net/fig/111563" pageId="41" pageNumber="252">, 163</figureCitation>
<figureCitation id="FB048DA4A728179741981278A46304D6" captionStart="Figure 165" captionStartId="F25" captionText="Figure 165. Distribution map of Cyphocoleus subulatus and Cyphocoleus iledespinsensis." figureDoi="10.3897/dez.63.10241.figure165" httpUri="https://binary.pensoft.net/fig/111564" pageId="41" pageNumber="252">, 165</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7F2E50316060ACE1C3255AB90B18711" pageId="42" pageNumber="253" type="diagnosis">
<paragraph id="2AD5C89F4789B834FF86A9F571FB17A5" pageId="42" pageNumber="253">
<pageBreakToken id="3D6D819EA958381C140F8B8BD9699203" pageId="42" pageNumber="253" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph id="92CAD8F8A3DD0DE45160A563142879E6" pageId="42" pageNumber="253">
Among species with orbicular pronotum and broadly cordate elytra, this species (Fig. 
<figureCitation id="2B674B059DCB731A17766B5124A70392" captionStart="Figures 161–164" captionStartId="F24" captionText="Figures 161 - 164. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 161, Cyphocoleus flavipes Yahoue; 162, Cyphocoleus bourailensis, Bourail; 163, Cyphocoleus subulatus, Mt. Koghi; 124, Cyphocoleus iledespinsensis, Ile des Pins." figureDoi="10.3897/dez.63.10241.figures161-164" httpUri="https://binary.pensoft.net/fig/111563" pageId="42" pageNumber="253">163</figureCitation>
) shares the possession of the posterior supraorbital seta with only 
<taxonomicName id="3719599D4D51824ABB0F373055E2A6DA" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus latipennis" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="35BE2F072BED3C0044626A20CEE42831" italics="true" pageId="42" pageNumber="253">Cyphocoleus latipennis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="01C95BE960279F3E1E39D6F6655762EF" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="42" pageNumber="253">122</figureCitation>
) and 
<taxonomicName id="8029B9EC3F38FDC145C654DE9AF15419" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus iledespinsensis" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="iledespinsensis">
<emphasis id="B60AB57C95EC13A381682556ED7C0381" italics="true" pageId="42" pageNumber="253">Cyphocoleus iledespinsensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="ECDD57C536536AE75F3AA49E38244FC5" captionStart="Figures 161–164" captionStartId="F24" captionText="Figures 161 - 164. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 161, Cyphocoleus flavipes Yahoue; 162, Cyphocoleus bourailensis, Bourail; 163, Cyphocoleus subulatus, Mt. Koghi; 124, Cyphocoleus iledespinsensis, Ile des Pins." figureDoi="10.3897/dez.63.10241.figures161-164" httpUri="https://binary.pensoft.net/fig/111563" pageId="42" pageNumber="253">164</figureCitation>
). Beetles of 
<taxonomicName id="BF4A28FB54090EF78CD292E97D0EB9BE" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus subulatus" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="subulatus">
<emphasis id="6DADEFD64853617D5D438FB48F9D855C" italics="true" pageId="42" pageNumber="253">Cyphocoleus subulatus</emphasis>
</taxonomicName>
can be diagnosed from the former by presence of evident, transversely stretched isodiametric microsculpture on the head, not the shallow, difficult to trace transverse-line microsculpture of 
<taxonomicName id="05AD99BFE25B0BBBEF7EE6658AF32B65" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus latipennis" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="latipennis">
<emphasis id="466AC0465C67174710DD9C7E54275D6D" italics="true" pageId="42" pageNumber="253">Cyphocoleus latipennis</emphasis>
</taxonomicName>
. Conversely, this species can be diagnosed from the latter by the pale antennae, and femora with flavous ground color and dark basal and apical bands, versus the uniformly fuscous antennae and femora observed in 
<taxonomicName id="D103650AC44920CC0322E3781EA073F4" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus iledespinsensis" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="iledespinsensis">
<emphasis id="A8BBFF5F28039A5CEE0515CC3633F6C2" italics="true" pageId="42" pageNumber="253">Cyphocoleus iledespinsensis</emphasis>
</taxonomicName>
. The eyes of this species are more convex, ocular ratio 1.54-1.57 compared to those of 
<taxonomicName id="8CBD55EF9AA4C392E929C44AEE566FFE" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus iledespinsensis" order="Diptera" pageId="42" pageNumber="253" phylum="Arthropoda" rank="species" species="iledespinsensis">
<emphasis id="3C16042EA9C0C0AA1ABF38CD6044C784" italics="true" pageId="42" pageNumber="253">Cyphocoleus iledespinsensis</emphasis>
</taxonomicName>
, where the broad, little convex eyes result in a ratio of 1.49. If a male can be dissected, the differences in the aedeagal median lobes are distinctive (see male genitalia sections for the two species descriptions). Standardized body length 6.0-7.2 mm.
</paragraph>
</subSubSection>
<subSubSection id="DE3CD2F4AFC7F72AE3F52A366D5C0990" lastPageId="43" lastPageNumber="254" pageId="42" pageNumber="253" type="description">
<paragraph id="4511E34D0C9276DC3724E84E2BA5AD92" pageId="42" pageNumber="253">Description</paragraph>
<paragraph id="F12B90D404F61D968CD6C1627A4FF1B0" lastPageId="43" lastPageNumber="254" pageId="42" pageNumber="253">
(n = 5). Head capsule ovoid, neck moderately constricted with genae moderately convergent behind eyes in dorsal view; eyes moderately convex, outer profile extended beyond curvature of gena behind eye, 
<pageBreakToken id="6EB799830D92BBB446D5FE4698E24E44" pageId="43" pageNumber="254" start="start">16</pageBreakToken>
-18 ommatidia across horizontal diameter of eye; frons with well-developed median carina that is surrounded by transversely wrinkled median flat spot; frontal grooves narrow, sinuous anteriorly from between eyes onto clypeus, frontoclypeal suture obsolete; mandibles moderately elongate, length 1.9 
<normalizedToken id="850BA1C814C07A9F841909C15414A132" originalValue="×">x</normalizedToken>
distance from antennal articulatory socket to anterolateral margin of labrum; antennae moderately elongate, scape length 2.5 
<normalizedToken id="C9FD57E6054E1F8C03EBEEE2F6A4CF5F" originalValue="×">x</normalizedToken>
maximal breadth; gena setose, subgenal seta present. Pronotum with front angles tightly rounded, slightly protruded, nearly right; lateral marginal bead distinctly sinuate before depressed median base that is margined medially by narrow bead; median longitudinal impression narrowly and finely incised, extended from just inside basal marginal bead nearly to anterior marginal bead, transverse wrinkles on either side of impression not aligned; anterior transverse impression very shallow, irregularly impressed in association with shallow longitudinal wrinkles on anterior callosity; proepisternum not visible in dorsal view, smooth ventrally, with fine vertical wrinkles along dorsal margin; prosternal process slightly convex medially anterad procoxae, with shallow medial depression ventrally, broadly convex posteriorly. Elytra broadly cordate, basal margin slightly concave to base of fourth stria, then posteriorly curved across broad humeri; discal intervals convex, intervals 3-5 slightly more convex, broader basally; elytral striae deep, narrow, impunctate; parascutellar seta present, articulatory socket papillate; dorsal elytral setae absent; lateral elytral setae arranged as 6 + 7, the articulatory sockets papillate; both subapical and apical elytral setae present, the articulatory socket for apical seta smaller, not papillate; subapical sinuation distinctly concave, lateral portion of concavity joined to depressed lateral margin in broad curve, medial margin convex to suture; elytral apices conjoined, margin not or only slightly invaginated at fused suture. Mesepisternum impunctate; metepisternal dorsal length subequal to diagonal width; metasternum with broad, shallow median fossa at base of metasternal process. Abdomen with apical margin of apical ventrite shallowly concave medially in both males and females; males with one seta each side of apical margin of apical ventrite, females with two seta each side. Legs moderately elongate; metacoxae bisetose; mt1 length/tibial length = 0.25; mt4 length to apex of outer lobe 1.6 
<normalizedToken id="605E23AEE70A5B7ADB9EDBB6E4F1DD3B" originalValue="×">x</normalizedToken>
median tarsomere length, 3-4 ventrolateral setae each side. Microsculpture well developed on pronotal disc, a combination of upraised isodiametric and transverse sculpticells, the surface matte; elytral microsculpture an irregularly present transverse mesh, sculpticell borders shallow, difficult to trace in glossier portions of surface; pelage little developed, head apparently glabrous, pronotal and elytral disc with sparsely distributed, extremely short microsetae, visible only in lateral view; prosternum with only a few very short microsetae, pterothoracic and abdominal ventrites and legs apparently glabrous except for macrosetae. Coloration of body dorsum and venter, including elytral epipleuron, uniformly piceous; legs paler, rufobrunneous with piceous cast except for flavous median femoral band (a minority of specimens lack this pale band and so have concolorous, rufobrunneous leg segments).
</paragraph>
<paragraph id="517134723E4FB33ADCC9F83075A0A76C" pageId="43" pageNumber="254">
<emphasis id="2966265FBC4E4D7EEE53696AC2523E6E" italics="true" pageId="43" pageNumber="254">Male genitalia</emphasis>
(n = 7). Male aedeagal median lobe elongate, heavily sclerotized, parallel sided from base to ostial opening; dorsoventral breadth at midlength 0.13 
<normalizedToken id="0A7753FE71D4BAC9652A6C7309C4F1BB" originalValue="×">x</normalizedToken>
distance from tip to base of closed basal bulb (Figs 
<figureCitation id="56692F6FEC75B17B67B7ED503C6BEA83" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="43" pageNumber="254">155-158</figureCitation>
); elongate lobe apex extended beyond ostial opening in a broad dorsally directed curve, tip tightly rounded, dorsoventral breadth at midlength of apical extension 0.18 
<normalizedToken id="C845E01A26C181C2E7F70743B9334F03" originalValue="×">x</normalizedToken>
distance from distal margin of ostial opening to tip; ostial opening on ventral surface of median lobe shaft (Fig. 
<figureCitation id="EDFFC11B96894F9535E280F68E90A189" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="43" pageNumber="254">157</figureCitation>
); sac covered with microtrichia, but without any heavily sclerotized structures, length at least 1.6 
<normalizedToken id="0144DC1E47E385B2633C6EBA0C620CDE" originalValue="×">x</normalizedToken>
breadth.
</paragraph>
<paragraph id="EBAA3A305F1E98F80EE7D93D13F52E43" pageId="43" pageNumber="254">
<emphasis id="2FCCDE3FC4F0C1EBA4418E24B0342F3A" italics="true" pageId="43" pageNumber="254">Female reproductive tract</emphasis>
(n = 3). Bursa copulatrix columnar, distance from base of gonocoxites to spermathecal basal sclerite 4 
<normalizedToken id="F5742B36E36FC746A87EE6C865786ADA" originalValue="×">x</normalizedToken>
maximum breadth (dissection compressed under cover slip) (Fig. 
<figureCitation id="658D5CFE1C471FA02C6B6E8CE0701E61" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="43" pageNumber="254">112</figureCitation>
); bursal walls thin, translucent, lumenal surface smooth except for thin band of very fine setae at midlength; basal gonocoxite with apical fringe of five to six setae (Fig. 
<figureCitation id="C476056FD9856B7291A0CADC24F32F0C" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="43" pageNumber="254">120</figureCitation>
); apical gonocoxite broadly subtriangular with pointed apex and two lateral ensiform setae.
</paragraph>
</subSubSection>
<subSubSection id="C6BEAFC9DA13D69EBBD72828E4AB44E0" pageId="43" pageNumber="254" type="types">
<paragraph id="596A0076412B62CD5C5098C8F61A23C6" pageId="43" pageNumber="254">Types.</paragraph>
<paragraph id="BFC4E5B0EC50651B5937EA8B0BA93E11" pageId="43" pageNumber="254">
Holotype male (ANIC deposited in MNHN): NEW CALEDONIA / Col 
<normalizedToken id="857AA5C7F0775C33A33F38BD7C1FCFBB" originalValue="d’Amieu">d'Amieu</normalizedToken>
, 1800-2000 / 12-13 Dec., 1973 / G.B. &amp;S. R. Monteith // HOLOTYPE / 
<taxonomicName id="F19CF933569C34D52F4F9CE8FD1427A5" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="43" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="5FCC18817960725E7843F5E95B783354" italics="true" pageId="43" pageNumber="254">Cyphocoleus</emphasis>
</taxonomicName>
/ 
<taxonomicName id="A551FBF00B9405E5401634F574B7A9C6" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subulatus" order="Diptera" pageId="43" pageNumber="254" phylum="Arthropoda" rank="species" species="subulatus">
<emphasis id="3C4A8E19F42EAD842E179F457D9B37D1" italics="true" pageId="43" pageNumber="254">subulatus</emphasis>
</taxonomicName>
/ J.K. Liebherr 2016 (black-margined red label).
</paragraph>
<paragraph id="540DA86F64D0A84FF5FCB9E9703F4772" pageId="43" pageNumber="254">
Paratypes (239 specimens). NEW CALEDONIA: 
<normalizedToken id="AE14E0BC41C11F937B3B0E603D43C947" originalValue="Aoupinié">Aoupinie</normalizedToken>
, top camp, 850 m el., pyrethrum trees &amp; logs, 
<geoCoordinate id="A157D52350CABB91FD0EF8DE6C572BEB" degrees="21" direction="south" minutes="11" orientation="latitude" precision="925" value="-21.183332">21°11'S</geoCoordinate>
, 
<geoCoordinate id="AC2433DB8C5F7AD5E61B380AA54A9091" degrees="165" direction="east" minutes="18" orientation="longitude" precision="925" value="165.3">165°18'E</geoCoordinate>
, 02-03xi2001, Burwell &amp; Monteith (QMB, 2; lot no. 8716); La Crouen, 
<geoCoordinate id="DBA5F72BCD5F743B59534E2C086EE0EA" degrees="21" direction="south" minutes="33" orientation="latitude" precision="925" value="-21.55">
21°33 
<normalizedToken id="D48CD76CF43D73EEEEDD7A78B70FD0C2" originalValue="S´">S'</normalizedToken>
</geoCoordinate>
<geoCoordinate id="2D4FF9447917E0BB1EDBCB4F7256CDA8" degrees="165" direction="east" minutes="55" orientation="longitude" precision="925" value="165.91667">165°55'E</geoCoordinate>
, 16iii1961, Sedlacek (BPBM, 1); Col 
<normalizedToken id="E68A30715FFB351756A5ABA0CE497235" originalValue="d’Amieu">d'Amieu</normalizedToken>
, 545-610 m el., 
<geoCoordinate id="12056B721587CCAC213CE5371F8C9E4F" degrees="21" direction="south" minutes="33" orientation="latitude" precision="925" value="-21.55">21°33'S</geoCoordinate>
, 
<geoCoordinate id="B47AF88168C141341B925B14B20B7D42" degrees="165" direction="east" minutes="48" orientation="longitude" precision="925" value="165.8">165°48'E</geoCoordinate>
, 12-13xii1973, G. &amp; S. Monteith (ANIC, 13), 18-19xii1973, G. &amp; S. Monteith (ANIC, 2); Col 
<normalizedToken id="4C2F34B11E340872FF548F33172C4B2B" originalValue="d’Amieu">d'Amieu</normalizedToken>
, 6 km NNE, 300 m el., 
<geoCoordinate id="E2B5842A52C6B18CF5B757338AECB151" degrees="21" direction="south" minutes="33.5" orientation="latitude" precision="92" value="-21.558332">21°33.5'S</geoCoordinate>
, 
<geoCoordinate id="EC7E3C37D7F5B6F1494B46050C75636E" degrees="165" direction="east" minutes="51" orientation="longitude" precision="925" value="165.85">165°51'E</geoCoordinate>
, 13xi2000, Bouchard, Burwell &amp; Monteith (QMB, 3; lot no. 9924; QMB, 2; lot no. 9923), pyrethrum trees &amp; logs, 11xi2001, Burwell (QMB, 2; lot no. 8678), 390 m el., ex nr. 
<emphasis id="18A501F30B77B2181082BCE01EAC1F9C" italics="true" pageId="43" pageNumber="254">Mirasmius</emphasis>
, 
<geoCoordinate id="8C9BBF87D033E75552925D47FAEF241A" degrees="21" direction="south" minutes="33.5" orientation="latitude" precision="92" value="-21.558332">21°33.5'S</geoCoordinate>
, 
<geoCoordinate id="FF0DD626356F3C3120907910DCD561A1" degrees="165" direction="east" minutes="45.6" orientation="longitude" precision="92" value="165.76">165°45.6'E</geoCoordinate>
, 16iii2007, Leschen (EMEC, 1; NZAC, 1); 440 m el., fogging logs, 
<geoCoordinate id="BAC495081A2210172FE5080AED546550" degrees="21" direction="south" minutes="33.5" orientation="latitude" precision="92" value="-21.558332">21°33.5'S</geoCoordinate>
, 
<geoCoordinate id="E183CF36E6EA32CDBC4AFB38EE2B3B24" degrees="165" direction="east" minutes="45.3" orientation="longitude" precision="92" value="165.755">165°45.3'E</geoCoordinate>
, 16iii2007, Leschen (NZAC, 1); Mt. Kanala, 
<geoCoordinate id="5CD465729C71F51CFCB110EFE8591CAC" degrees="21" direction="south" minutes="34" orientation="latitude" precision="925" value="-21.566668">21°34'S</geoCoordinate>
, 
<geoCoordinate id="CB4D5CA17D2356A42E695DE9B52F9518" degrees="165" direction="east" minutes="56" orientation="longitude" precision="925" value="165.93333">165°56'E</geoCoordinate>
, 03xi1911, Sarasin &amp; Roux (SMTD, 1); Ciu nr. Mt. Canala, rainforest, 
<geoCoordinate id="D591A280B07DB152CBCBAAB0FBBC99AA" degrees="21" direction="south" minutes="34" orientation="latitude" precision="925" value="-21.566668">21°34'S</geoCoordinate>
, 
<geoCoordinate id="3EC40F6927F657518B4D16AF289D3F0F" degrees="165" direction="east" minutes="57" orientation="longitude" precision="925" value="165.95">165°57'E</geoCoordinate>
, 01-03i1955 Wilson (MCZ, 2); Table Unio road, 500 m el., 
<geoCoordinate id="085D83494BB69993CC0775A6C8A5894B" degrees="21" direction="south" minutes="34" orientation="latitude" precision="925" value="-21.566668">21°34'S</geoCoordinate>
, 
<geoCoordinate id="4D93F2EDBBB09641E4F7076C98F6113A" degrees="165" direction="east" minutes="46" orientation="longitude" precision="925" value="165.76666">165°46'E</geoCoordinate>
, 14xi2000, Bouchard, Burwell &amp; Monteith (QMB, 2; lot no. 9946), 800 m el., 
<geoCoordinate id="0EC5DA57BFD7FFBE64A97D16916FDFE7" degrees="21" direction="south" minutes="34" orientation="latitude" precision="925" value="-21.566668">21°34'S</geoCoordinate>
, 
<geoCoordinate id="01A5CFC1D871C60CAEF8BA032896ED21" degrees="165" direction="east" minutes="46" orientation="longitude" precision="925" value="165.76666">165°46'E</geoCoordinate>
, 14xi2000, Bouchard, Burwell &amp; Monteith (QMB, 1; lot no. 9934); Canala, 4 km S Mia, 380 m el., 
<geoCoordinate id="9F07E2753DCA2DA48F1A626E1256241B" degrees="21" direction="south" minutes="34.7" orientation="latitude" precision="92" value="-21.578333">21°34.7'S</geoCoordinate>
, 
<geoCoordinate id="138D07B204C66835442C146CC53BAE01" degrees="165" direction="east" minutes="58.3" orientation="longitude" precision="92" value="165.97166">165°58.3'E</geoCoordinate>
, river valley, rainforest, Wanat &amp; Ruta (MNHW, 1); Mt. Rembai 650 m el., 
<geoCoordinate id="6CA5172D0C2234C646E31B84240D87C7" degrees="21" direction="south" minutes="35" orientation="latitude" precision="925" value="-21.583334">21°35'S</geoCoordinate>
, 
<geoCoordinate id="577DBFFFCC52662F87308A4CC3E715AA" degrees="165" direction="east" minutes="51" orientation="longitude" precision="925" value="165.85">165°51'E</geoCoordinate>
, 09v1984 Monteith &amp; Cook (QMB, 3), 700-900 m el., rainforest litter, 
<geoCoordinate id="0BF86DFDA24A6FED28E4E8B669347C1B" degrees="21" direction="south" minutes="35" orientation="latitude" precision="925" value="-21.583334">21°35'S</geoCoordinate>
, 
<geoCoordinate id="89B64ECD33B242BFD6A510F0802A0305" degrees="165" direction="east" minutes="51" orientation="longitude" precision="925" value="165.85">165°51'E</geoCoordinate>
, 09v1984, Monteith &amp; Cook (QMB, 9); Col 
<normalizedToken id="E1C96648C583D9A99533A458FC862ED1" originalValue="d’Amieu">d'Amieu</normalizedToken>
Forest Station, 400 m el., 
<geoCoordinate id="21C585411947D3929AD7680B87B1ADC2" degrees="21" direction="south" minutes="35" orientation="latitude" precision="925" value="-21.583334">21°35'S</geoCoordinate>
, 
<geoCoordinate id="638720C1126EA0ACCE2E5AC99FCD7F1A" degrees="165" direction="east" minutes="50" orientation="longitude" precision="925" value="165.83333">165°50'E</geoCoordinate>
, 08-09v1984, Monteith &amp; Cook (QMB, 4); Mt. Rembai top junction, 780 m el., hand collecting, rainforest, 
<geoCoordinate id="E7DE9DCCA32C6A1A1418EA45AEF2961A" degrees="21" direction="south" minutes="35" orientation="latitude" precision="925" value="-21.583334">21°35'S</geoCoordinate>
, 
<geoCoordinate id="21C48BDE24211BC98D3C1F58DBE76DB9" degrees="165" direction="east" minutes="51" orientation="longitude" precision="925" value="165.85">165°51'E</geoCoordinate>
, 30xii2004, Monteith (QMB, 1; lot no. 12007); Plateau de Dogny trail, 870 m el., litter, 
<geoCoordinate id="4FBB743EFA2898E660AA6EE76C0F0E80" degrees="21" direction="south" minutes="37.2" orientation="latitude" precision="92" value="-21.62">21°37.2'S</geoCoordinate>
, 
<geoCoordinate id="E5D0448AE1BAB14E7C91748A704DEEB3" degrees="165" direction="east" minutes="52.5" orientation="longitude" precision="92" value="165.875">165°52.5'E</geoCoordinate>
, 15iii2007, Leschen &amp; Will (EMEC, 1); Plateau de Dogny trail, 245 m el., flood debris, 
<geoCoordinate id="4C224F00F83F0B3DF782EC2A0CD092B5" degrees="21" direction="south" minutes="37.8" orientation="latitude" precision="92" value="-21.63">21°37.8'S</geoCoordinate>
, 
<geoCoordinate id="9A1B69D073E33486CB977E8D9272BCA2" degrees="165" direction="east" minutes="52" orientation="longitude" precision="925" value="165.86667">165°52'E</geoCoordinate>
, 15iii2007, Leschen (NZAC, 1), Will (EMEC, 12); Col 
<normalizedToken id="B9C2A60E653364ADABC301CA931853EC" originalValue="d’Amieu">d'Amieu</normalizedToken>
litter, 
<geoCoordinate id="0ACED605C8A15FFCBD1BC7EF12981F52" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="432F5DCF4F8F8CD41E6EFB19909C164C" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 06ix1990, Raven (QMB, 1); Farino, 5 km N, 400 m el., night collecting, 
<geoCoordinate id="2718893063ABF72BCB4CA69F7638D952" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="E63FDF624DA9BA0625C16D287A8BFF93" degrees="165" direction="east" minutes="46" orientation="longitude" precision="925" value="165.76666">165°46'E</geoCoordinate>
, 14xi2002, Burwell &amp; Monteith (QMB, 2; lot no. 11186); Col 
<normalizedToken id="951F2300E9577EA7BE0388A3848AEF41" originalValue="d’Amieu">d'Amieu</normalizedToken>
, west slope, 470 m el., pyrethrum logs, 
<geoCoordinate id="1EAAEA7646897C5A175AF4FDBD9BEBAE" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="6E3419FC4EEBDC071965EE28747BE702" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 17xi2002, Monteith (QMB, 5; lot no. 11180), hand collecting, 
<geoCoordinate id="9131B0B697780AD50F5AD1F524664813" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="3848A0B2BCCEBAD54EB3E99C158FFCE6" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 25xi2003, Monteith (QMB, 1; lot no. 11426), pyrethrum trees &amp; logs, 
<geoCoordinate id="1413983B27B708B087D03D55586A37A5" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="311A898378B28BDA63B04835729EC8FB" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 25xi2003, Monteith (QMB, 8; lot no. 11427), pyrethrum log w/ epiphytes, 
<geoCoordinate id="A32EA5FE040E15A19296B0818B09E8DA" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="AB511DC80A12F191B7EECF5EB662CBB6" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 27i2004, Monteith (QMB, 1; lot no. 11517), pyrethrum log w/o epiphytes, 
<geoCoordinate id="F795F2918503AAB117FE8615D8766764" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="58B8D4E41CF272802C39BA3A680D3817" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 27xi2004, Monteith (QMB, 14; lot no. 11516), pyrethrum trees &amp; logs, 
<geoCoordinate id="4D5451CA0FB247C44B45D58245E0AB3A" degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
, 
<geoCoordinate id="0EEA8E1EA160E4886206D115C147131A" degrees="165" direction="east" minutes="49" orientation="longitude" precision="925" value="165.81667">165°49'E</geoCoordinate>
, 29ix2004 Monteith (QMB, 1; lot no. 11667); 
<normalizedToken id="5508A8E8B46D89A7BB24043F0455F592" originalValue="Sarraméa">Sarramea</normalizedToken>
, 2 km NE, 280 m el., 
<geoCoordinate id="61376C13F8464BEC5341A799D778ABA6" degrees="21" direction="south" minutes="38" orientation="latitude" precision="925" value="-21.633333">21°38'S</geoCoordinate>
, 
<geoCoordinate id="40763EB61D238988F3A68451B48445B1" degrees="165" direction="east" minutes="52" orientation="longitude" precision="925" value="165.86667">165°52'E</geoCoordinate>
, 30xi2009, Schuh (NMHW, 2); Table Unio, 700-1000 m el., 
<geoCoordinate id="AF981E7E3712D59CC057A90D7F64A377" degrees="21" direction="south" minutes="43" orientation="latitude" precision="925" value="-21.716667">21°43'S</geoCoordinate>
, 
<geoCoordinate id="538F892B008DDA505E3BCFBD84CA6E5E" degrees="165" direction="east" minutes="47" orientation="longitude" precision="925" value="165.78334">165°47'E</geoCoordinate>
, 10v1984, Monteith &amp; Cook (QMB, 1); 
<normalizedToken id="133C8859489EA753F5C844984A7F3FC7" originalValue="Sarraméa">Sarramea</normalizedToken>
, Plateau de Dogny, Bois mort, 
<geoCoordinate id="3D2C3682E15D47A6A8B1BFE9E68EF335" degrees="21" direction="south" minutes="46" orientation="latitude" precision="925" value="-21.766666">21°46'S</geoCoordinate>
, 
<geoCoordinate id="3F3CF2B30F6B3CF49673EB650FD57F56" degrees="165" direction="east" minutes="55" orientation="longitude" precision="925" value="165.91667">165°55'E</geoCoordinate>
, 23vi1971 (MNHN, 5); Mt. Mou, 380 m el., small stream, 
<geoCoordinate id="72D8DE6D96BE500D83BC28F850614E7C" degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
, 
<geoCoordinate id="1812ABED2F1F236081060ED62B9F763E" degrees="166" direction="east" minutes="20" orientation="longitude" precision="925" value="166.33333">166°20'E</geoCoordinate>
, 23xi2009, 
<normalizedToken id="F4B85F11CE5939E5E2CAEC3F9062A83D" originalValue="Jäch">Jaech</normalizedToken>
(NMHW, 1); Col de la Pirogue, 
<geoCoordinate id="D5874F91542D8DB41DEA02221A5F215E" degrees="22" direction="south" minutes="05" orientation="latitude" precision="925" value="-22.083334">22°05'S</geoCoordinate>
, 
<geoCoordinate id="756721BF2ADD4F56C7FBCBD6B84EBA7D" degrees="166" direction="east" minutes="19" orientation="longitude" precision="925" value="166.31667">166°19'E</geoCoordinate>
, 23i1962, Krauss (BPBM, 2); 
<normalizedToken id="A7FFF7DB56DEBF03622C6AC1E914CACC" originalValue="Rivière">Riviere</normalizedToken>
Bleue, Kaori Geant, 
<geoCoordinate id="A3079F1E5DAE31D73DB266C198A15ADA" degrees="22" direction="south" minutes="06" orientation="latitude" precision="925" value="-22.1">22°06'S</geoCoordinate>
, 
<geoCoordinate id="D814CB733F0A38ACC5ACC58C4784EF1E" degrees="166" direction="east" minutes="39" orientation="longitude" precision="925" value="166.65">166°39'E</geoCoordinate>
, 11xi2000, Bouchard, Burwell &amp; Monteith (QMB, 9; lot no. 9957), pyrethrum trunks &amp; logs, 
<geoCoordinate id="9CEABAB02FA33A59D2E082E0503B096F" degrees="22" direction="south" minutes="06" orientation="latitude" precision="925" value="-22.1">22°06'S</geoCoordinate>
, 
<geoCoordinate id="7BB7EAF77CBFD2EB7E92C9247A154546" degrees="166" direction="east" minutes="39" orientation="longitude" precision="925" value="166.65">166°39'E</geoCoordinate>
, 11xi2000, Monteith (QMB, 14; lot no. 9954), 160 m el., pyrethrum trees &amp; logs, 
<geoCoordinate id="8A65E999A8C321F1C91A427AD08676BB" degrees="22" direction="south" minutes="06" orientation="latitude" precision="925" value="-22.1">22°06'S</geoCoordinate>
, 
<geoCoordinate id="3B74346756742F3EE2D18CE075219C45" degrees="166" direction="east" minutes="39" orientation="longitude" precision="925" value="166.65">166°39'E</geoCoordinate>
, 19xi2002, Monteith (QMB, 2; lot no. 11202); 11-21xi2002, Monteith &amp; Wright (QMB, 1; lot no. 11203), Pont Germain 160 m el., 
<geoCoordinate id="3A3C67D4DDB52319D6B581A5AC43AF31" degrees="22" direction="south" minutes="06" orientation="latitude" precision="925" value="-22.1">22°06'S</geoCoordinate>
, 
<geoCoordinate id="734491B6E6C8B7B1D0909A9D0F8FFC31" degrees="166" direction="east" minutes="39" orientation="longitude" precision="925" value="166.65">166°39'E</geoCoordinate>
, 11xi2000, Bouchard, Burwell &amp; Monteith (QMB, 9; lot no. 9950), 
<normalizedToken id="B05B50283E560B749F6B9FE3FDBAEE3C" originalValue="L’houp">L'houp</normalizedToken>
Geant Trail, 350 m el., 
<geoCoordinate id="0DABEFB2474B6AE5E0C8E2B453B2A2E2" degrees="22" direction="south" minutes="09" orientation="latitude" precision="925" value="-22.15">22°09'S</geoCoordinate>
, 
<geoCoordinate id="06C07798C27700CE900A2AD498F0381B" degrees="166" direction="east" minutes="41.2" orientation="longitude" precision="92" value="166.68666">166°41.2'E</geoCoordinate>
, 26xi2009, 
<normalizedToken id="20DABD14AFC90CDE828F490B12CA43A5" originalValue="Jäch">Jaech</normalizedToken>
(NMHW, 1); Mt. Koghi, 
<geoCoordinate id="5557473E3C67D899CD85AF49D0814C5E" degrees="22" direction="south" minutes="10" orientation="latitude" precision="925" value="-22.166666">22°10'S</geoCoordinate>
, 
<geoCoordinate id="F67180A066666E83C26BF5035AC1C4D8" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
, 26-30i1963, Kuschel (BPBM, 1), 480 m el., 24-x-2008, fogged log, Wanat (MNHW, 4), 500-800 m el., 23-27x1967, J. &amp; M. Sedlacek (BPBM, 3), 500-750 m el., m, 25-26x1967, J. &amp; M. Sedlacek (BPBM, 2), 610-670 m el., 20-23xi1973, G. &amp; S. Monteith (ANIC, 8), 12-13ii1977, J. Balogh (RScC, 2), 500 m el., 26viii1978, S. &amp; J. Peck (CNC, 2), 400 m el., 26v1984, Monteith &amp; Cook (ANIC, 3), 400 m el., 26v1984, Monteith &amp; Cook (ANIC, 2), rainforest 500 m el., sifted litter, 23v1987, Raven (QMB, 2), pyrethrum trunks &amp; logs, 27xi2000, Monteith (QMB, 1; lot no. 9942), 750 m el., pyrethrum trunks &amp; logs, 29xi2000, Monteith (QMB, 8; lot no. 9944); 500 m el., hand collecting, 27i2002, Monteith (QMB, 1; lot no. 8917), night collecting, 02-03xi2002, Burwell, Monteith &amp; Wright (QMB, 2; lot no. 11088), 500 m el., 22xi2000, Bouchard, Burwell &amp; Monteith (QMB, 2; lot no. 9932), dung pitfalls, 22xi2000, Monteith (QMB, 3; lot no. 9933), pyrethrum trunks &amp; logs, 22xi2000, Monteith (QMB, 12; lot no. 9931), 700 m el., 
<geoCoordinate id="0941264FE87F2727DDD76966A4A58AA3" degrees="22" direction="south" minutes="10.5" orientation="latitude" precision="92" value="-22.175">22°10.5'S</geoCoordinate>
, 
<geoCoordinate id="2CA9438592E22962FC6E3A953518115B" degrees="166" direction="east" minutes="30.8" orientation="longitude" precision="92" value="166.51334">166°30.8'E</geoCoordinate>
, 12iii2007, Will (EMEC, 1), 550 m el., small streams, 
<geoCoordinate id="A4BD0A4E2FCA9D74E73FF1604D149B22" degrees="22" direction="south" minutes="10.5" orientation="latitude" precision="92" value="-22.175">22°10.5'S</geoCoordinate>
, 
<geoCoordinate id="6ADA99AE24FB213485C09137F7AEEDA7" degrees="166" direction="east" minutes="30.3" orientation="longitude" precision="92" value="166.505">166°30.3'E</geoCoordinate>
, 25xi2009, 
<normalizedToken id="BA1E20235AF6560F9E63AEC202D29E8D" originalValue="Jäch">Jaech</normalizedToken>
(NMHW, 2), 485 m el., beating fungi &amp; dead branches, 
<geoCoordinate id="2F6ECCB7320AA27306FA65D9296360DC" degrees="22" direction="south" minutes="10.7" orientation="latitude" precision="92" value="-22.178333">22°10.7'S</geoCoordinate>
, 
<geoCoordinate id="8075CAE43EFF19E0F62831A35853CD53" degrees="166" direction="east" minutes="30.4" orientation="longitude" precision="92" value="166.50667">166°30.4'E</geoCoordinate>
, 12iii2007, Leschen (EMEC, 2; NZAC, 1); Mt. Chapeau de Gendarme [= Mt. Malaoui], forest brook, under rocks, 
<geoCoordinate id="BC4711445D02D219A76E22A2C94CE58D" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
, 
<geoCoordinate id="344A5339D125FFA7A96DCFE2A7C0F29D" degrees="166" direction="east" minutes="30" orientation="longitude" precision="925" value="166.5">166°30'E</geoCoordinate>
, 22x1944, Herron (MSUC, 5), low pasture, 22x1944, Herron (MSUC, 1), under logs, 22x1944, Herron (MSUC, 1); 
<normalizedToken id="86DEBC2F4A4F8F933FB347DCCC07F733" originalValue="Forêt">Foret</normalizedToken>
de Thi to Hamma, 700-800 m el., 
<geoCoordinate id="E78CE0D73F877021F0CE8394E111C1EF" degrees="22" direction="south" minutes="13" orientation="latitude" precision="925" value="-22.216667">22°13'S</geoCoordinate>
, 
<geoCoordinate id="896F382ED34FFFB0430B41FDA5D0CF1D" degrees="166" direction="east" minutes="32" orientation="longitude" precision="925" value="166.53334">166°32'E</geoCoordinate>
, 16vii1958, Malkin &amp; Rageau (BPBM, 2); 
<normalizedToken id="A16881BC746B071685E7C0386E489E76" originalValue="Forêt">Foret</normalizedToken>
de Thi, 100-300 m el., 
<geoCoordinate id="1CDB060E8E4155A5FE13505C65CD5A12" degrees="22" direction="south" minutes="13" orientation="latitude" precision="925" value="-22.216667">22°13'S</geoCoordinate>
, 
<geoCoordinate id="48A921FFC0FA21EDBA9E7A23162226FE" degrees="166" direction="east" minutes="32" orientation="longitude" precision="925" value="166.53334">166°32'E</geoCoordinate>
, 25iii1961, Sedlacek (BPBM, 2), 05viii1966, Oesterr. NC Exped. (RScC, 1), 30x1967, J. &amp; M. Sedlacek (BPBM, 1), 100-300 m el., 07ix1979, Nishida (BPBM, 1), 150 m el., 21v1984, Monteith &amp; Cook (ANIC, 2; QMB, 3); Pic du Grand Kaori, site 2, rainforest, 250 m el., hand collecting, 
<geoCoordinate id="AE3800900B03B6876F2CF640929C1AC0" degrees="22" direction="south" minutes="17" orientation="latitude" precision="925" value="-22.283333">22°17'S</geoCoordinate>
, 
<geoCoordinate id="F75BAE2C284E614FAA209949F614C3D7" degrees="166" direction="east" minutes="53" orientation="longitude" precision="925" value="166.88333">166°53'E</geoCoordinate>
, 22-24xi2004, QMB Party (QMB, 2; lot no. 11772), pyrethrum, rainforest, 22-24xi2004, Monteith &amp; Burwell (QMB, 1; lot no. 11773); 
<normalizedToken id="F12EBD0645BC9043F061CB88C9BA978D" originalValue="Rivière">Riviere</normalizedToken>
des Pirogues Headwaters, 350-400 m el., 
<geoCoordinate id="051BA4113FD25BB93175F4D2D7B32501" degrees="22" direction="south" minutes="19" orientation="latitude" precision="925" value="-22.316668">22°19'S</geoCoordinate>
, 
<geoCoordinate id="A4F3B0315D507F57D6E01F35752BC410" degrees="166" direction="east" minutes="44" orientation="longitude" precision="925" value="166.73334">166°44'E</geoCoordinate>
, 22v1984, Monteith &amp; Cook (ANIC, 2; QMB, 4); 
<normalizedToken id="CE9D73D682B944CB99B460E576418305" originalValue="Forêt">Foret</normalizedToken>
Nord, site 2, rainforest, 200 m el., day hand collecting, 
<geoCoordinate id="7685B8003E842F4B977A2299F45614A8" degrees="22" direction="south" minutes="19" orientation="latitude" precision="925" value="-22.316668">22°19'S</geoCoordinate>
, 
<geoCoordinate id="37E75579A23A0FED2E3E7D0210DBD3DF" degrees="166" direction="east" minutes="55" orientation="longitude" precision="925" value="166.91667">166°55'E</geoCoordinate>
, 02-03xii2004, QMB Party (QMB, 3; lot no. 11829).
</paragraph>
</subSubSection>
<subSubSection id="141135DB743FDB81289DA444372C6F76" pageId="43" pageNumber="254" type="etymology">
<paragraph id="004EECF343BBE757F5D45E88D445B216" pageId="43" pageNumber="254">Etymology.</paragraph>
<paragraph id="9BB08AB7FB66DA27F66315790572CB5D" pageId="43" pageNumber="254">
The Latin adjectival 
<taxonomicName id="056E8A51B7940CE6B1D1A77E541D1EBD" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="subulatus" order="Diptera" pageId="43" pageNumber="254" phylum="Arthropoda" rank="species" species="subulatus">subulatus</taxonomicName>
, awl-shaped or pointed (
<bibRefCitation id="4B99217F92C6B4AB7BEF3FE05A042452" author="Brown, RW" journalOrPublisher="Smithsonian Institution Press, Washington, DC" pageId="48" pageNumber="259" refId="B8" refString="Brown, RW, 1956. Composition of Scientific Words. Smithsonian Institution Press, Washington, DC" title="Composition of Scientific Words." year="1956">Brown 1956</bibRefCitation>
), forms the basis this 
<normalizedToken id="2674183CB8B73BFDBC9338207A1374D1" originalValue="species’">species'</normalizedToken>
epithet based on the distinctively pointed male aedeagal median lobe (Figs 
<figureCitation id="3B2481CAB1B260758B557717E3394901" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="43" pageNumber="254">155-158</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9AD2074B622D646E1BE90F651C6C6E82" pageId="43" pageNumber="254" type="distribution">
<paragraph id="FED45C2C4FCE7386D960A83E7A5BF959" pageId="43" pageNumber="254">Distribution and habitat.</paragraph>
<paragraph id="80D28EB499CA75C4FAAF58302BC64FF1" pageId="43" pageNumber="254">
This species is known from the southern half of Grande Terre, with a northerly outlying locality, based on current specimens, at 
<normalizedToken id="4F3BACFBE2BFE8516068C832E98243F9" originalValue="Aoupinié">Aoupinie</normalizedToken>
(Fig. 
<figureCitation id="6522275568CE1ADF47B706C3D9669D95" captionStart="Figure 165" captionStartId="F25" captionText="Figure 165. Distribution map of Cyphocoleus subulatus and Cyphocoleus iledespinsensis." figureDoi="10.3897/dez.63.10241.figure165" httpUri="https://binary.pensoft.net/fig/111564" pageId="43" pageNumber="254">165</figureCitation>
). Beetles have been abundantly collected along the 
<normalizedToken id="E9BF6C752CAE83BCEF757412DCFAE369" originalValue="SarraméaCanala">SarrameaCanala</normalizedToken>
Road, and also in the mountains outside 
<normalizedToken id="396DB7D2F4E013148746BFB0654DD97D" originalValue="Nouméa">Noumea</normalizedToken>
. Localities range in elevation from 100-900 m. Collecting situations are quite diverse, including dead branches with fungi, pyrethrin fogged logs with and without epiphytes, the margins of small streams, and within flood debris.
</paragraph>
<caption id="B3318D35EA402906524745818C286F88" ID-Zenodo-Dep="985986" doi="10.3897/dez.63.10241.figure165" httpUri="https://binary.pensoft.net/fig/111564" pageId="43" pageNumber="254" start="Figure 165" startId="F25">
<paragraph id="9D7E5CA25EF9B4991A6329E6F1E38C9F" pageId="43" pageNumber="254">
<emphasis id="2EDC5ED387C6F6104217F72B14C2CDE6" bold="true" pageId="43" pageNumber="254">Figure 165.</emphasis>
Distribution map of 
<taxonomicName id="C58A9B03606D64B7D650D29D5DB94D0C" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus subulatus" order="Diptera" pageId="43" pageNumber="254" phylum="Arthropoda" rank="species" species="subulatus">
<emphasis id="6862C15094CD50D57684EB112D182F84" italics="true" pageId="43" pageNumber="254">Cyphocoleus subulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="C0BEFC6475020550BE06885689D5DEF9" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus iledespinsensis" order="Diptera" pageId="43" pageNumber="254" phylum="Arthropoda" rank="species" species="iledespinsensis">
<emphasis id="1F7CC47F509BEDC2C562D623B4E9B5C7" italics="true" pageId="43" pageNumber="254">Cyphocoleus iledespinsensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>