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<document id="A61BD7858C81B970A7E45C49F50696D8" ID-DOI="10.11646/phytotaxa.437.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13873878" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727815621804" checkinUser="felipe" docAuthor="Thomas, Daniel C. &amp; Ardi, Wisnu H." docDate="2020" docId="BC25F3022009FFC1FF15FC6FE11A938F" docLanguage="en" docName="phytotaxa.437.2.4.pdf" docOrigin="Phytotaxa 437 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.437.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia hooveriana Wiriadinata 2013" docType="treatment" docVersion="2" lastPageNumber="86" masterDocId="401C8B7A2003FFCCFF9DFFB5E012921F" masterDocTitle="Synopsis of Begonia (Begoniaceae) of southwest Sulawesi and the Selayar Islands, Indonesia, including one new species" masterLastPageNumber="96" masterPageNumber="73" pageNumber="83" updateTime="1727817875856" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="829D076E45EC2E84D0F2D7DBEDC7DD05">Synopsis of Begonia (Begoniaceae) of southwest Sulawesi and the Selayar Islands, Indonesia, including one new species</mods:title>
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<mods:namePart id="E0DAD6D60E325F830DCC3193CD12931B">Thomas, Daniel C.</mods:namePart>
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<mods:namePart id="EADBD32003FF97971B039B65C9FB8593">Ardi, Wisnu H.</mods:namePart>
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<mods:title id="05E208C9CEB5F232813E2D69A4D8B892">Phytotaxa</mods:title>
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<treatment id="BC25F3022009FFC1FF15FC6FE11A938F" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F" httpUri="http://treatment.plazi.org/id/BC25F3022009FFC1FF15FC6FE11A938F" lastPageId="13" lastPageNumber="86" pageId="10" pageNumber="83">
<subSubSection id="7C96119F2009FFC6FF15FC6FE33391EB" box="[136,801,985,1013]" pageId="10" pageNumber="83" type="nomenclature">
<paragraph id="343342142009FFC6FF15FC6FE33391EB" blockId="10.[136,801,985,1048]" box="[136,801,985,1013]" pageId="10" pageNumber="83">
<heading id="6F7BF5782009FFC6FF15FC6FE33391EB" box="[136,801,985,1013]" fontSize="11" level="2" pageId="10" pageNumber="83" reason="4">
<emphasis id="06F89E062009FFC6FF15FC6FE08F91EB" bold="true" box="[136,157,986,1012]" pageId="10" pageNumber="83">7.</emphasis>
<taxonomicName id="F38C39972009FFC6FF39FC6FE2B091EA" ID-CoL="87YMT" authority="Wiriadinata (2013: 445)" authorityName="Wiriadinata" authorityPageNumber="445" authorityYear="2013" box="[164,674,985,1013]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="10" pageNumber="83" phylum="Tracheophyta" rank="species" species="hooveriana">
<emphasis id="06F89E062009FFC6FF39FC6FE19D91EB" bold="true" box="[164,399,985,1012]" italics="true" pageId="10" pageNumber="83">Begonia hooveriana</emphasis>
<bibRefCitation id="501D3FE52009FFC6FE08FC6FE2B091EA" author="Wiriadinata, H." box="[405,674,986,1013]" pageId="10" pageNumber="83" pagination="445 - 448" refId="ref20383" refString="Wiriadinata, H. (2013) A new species of Begonia (Begoniaceae) from South Sulawesi, Indonesia. Reinwardtia 13: 445 - 448." type="journal article" year="2013">Wiriadinata (2013: 445)</bibRefCitation>
</taxonomicName>
(
<figureCitation id="ACB75E912009FFC6FD2FFC6FE2E291EB" box="[690,752,986,1012]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[179,1408,188,1926]" captionTargetId="figure-22@2.[179,1408,187,1926]" captionTargetPageId="2" captionText="FIGURE 1. Distribution maps of Begonia species occurring on the southwestern arm of Sulawesi or the Selayar Islands (georeferenced specimen data from major collections [B, BO, E, K, KRB, L, SING], the Begonia Resource Center [Hughes et al. 2015a], as well as some additional confirmed observations). Grey lines indicate 1° graticules." figureDoi="http://doi.org/10.5281/zenodo.13873880" httpUri="https://zenodo.org/record/13873880/files/figure.png" pageId="10" pageNumber="83">Fig. 1</figureCitation>
&amp;
<figureCitation id="ACB75E912009FFC6FC95FC6FE30A91EB" box="[776,792,986,1012]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="10" pageNumber="83">4</figureCitation>
)
</heading>
</paragraph>
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<paragraph id="343342142009FFC6FF15FC4BE1459607" blockId="10.[136,801,985,1048]" box="[136,343,1022,1048]" pageId="10" pageNumber="83">
Sect.
<taxonomicName id="F38C39972009FFC6FF5BFC4AE1459607" authorityName="de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" box="[198,343,1023,1048]" pageId="10" pageNumber="80" rank="section" section="Petermannia">
<emphasis id="06F89E062009FFC6FF5BFC4AE1459607" box="[198,343,1023,1048]" italics="true" pageId="10" pageNumber="83">Petermannia</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7C96119F2009FFC6FF15FBFCE58096EF" pageId="10" pageNumber="83" type="materials_examined">
<paragraph id="343342142009FFC6FF15FBFCE14096B8" blockId="10.[136,1452,1097,1191]" pageId="10" pageNumber="83">
<emphasis id="06F89E062009FFC6FF15FBFCE0C89641" bold="true" box="[136,218,1097,1118]" pageId="10" pageNumber="83">Type:—</emphasis>
<materialsCitation id="84E448492009FFC6FF47FBFCE15C96B8" collectingDate="1998-03-09" collectionCode="BO" collectorName="W. S. Hoover" country="Indonesia" elevation="900" location="Central Park Makele" municipality="Tanah Toraja" pageId="10" pageNumber="83" specimenCode="BO1454232, BO1454237" specimenCount="2" stateProvince="Sulawesi Utara" typeStatus="lectotype">
<collectingCountry id="4C9B02842009FFC6FF47FBFCE1729641" box="[218,352,1097,1118]" name="Indonesia" pageId="10" pageNumber="83">INDONESIA</collectingCountry>
.
<collectingRegion id="F6488CF62009FFC6FEF3FBFCE1D59640" box="[366,455,1097,1119]" country="Indonesia" name="Sulawesi Utara" pageId="10" pageNumber="83">Sulawesi</collectingRegion>
,
<collectingRegion id="F6488CF62009FFC6FE4BFBFCE2609640" box="[470,626,1097,1119]" country="Indonesia" name="Sulawesi Selatan" pageId="10" pageNumber="83">South Sulawesi</collectingRegion>
,
<collectingMunicipality id="D457D86E2009FFC6FD1DFBFCE3179640" box="[640,773,1097,1119]" pageId="10" pageNumber="83">Tanah Toraja</collectingMunicipality>
,
<location id="315314CF2009FFC6FC8EFBFCE3F79640" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF2009FFC6FC8EFBFCE3F79640" box="[787,997,1097,1119]" country="Indonesia" municipality="Tanah Toraja" name="Central Park Makele" pageId="10" pageNumber="83" stateProvince="Sulawesi Utara">Central Park Makele</location>
, alt.
<quantity id="F374EFF12009FFC6FB80FBFCE4739640" box="[1053,1121,1097,1119]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="83" unit="m" value="900.0">
<elevation id="BFA1A5272009FFC6FB80FBFCE4739640" box="[1053,1121,1097,1119]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="83" unit="m" value="900.0">900 m</elevation>
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,
<date id="403264D42009FFC6FBEDFBFCE4E39641" box="[1136,1265,1097,1119]" pageId="10" pageNumber="83" value="1998-03-09">
<collectingDate id="50769D3C2009FFC6FBEDFBFCE4E39641" box="[1136,1265,1097,1119]" pageId="10" pageNumber="83" value="1998-03-09">9 Mar. 1998</collectingDate>
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,
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<collectorName id="997927C22009FFC6FA9DFBFFE56E9640" box="[1280,1404,1098,1119]" pageId="10" pageNumber="83">W.S. Hoover</collectorName>
889
</emphasis>
(
<collectionCode id="529DDAD12009FFC6FF59FBD8E0FA969C" box="[196,232,1133,1155]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="10" pageNumber="83" type="Herbarium">BO</collectionCode>
[1454234]!,
<typeStatus id="EB37FCB62009FFC6FEFEFBD8E1A9969C" box="[355,443,1133,1155]" pageId="10" pageNumber="83" type="lectotype">lectotype</typeStatus>
designated here;
<typeStatus id="EB37FCB62009FFC6FDECFBD8E2F7969C" box="[625,741,1133,1155]" pageId="10" pageNumber="83" type="isolectotype">isolectotype</typeStatus>
BO[5 sheets:
<specimenCode id="642AEA6F2009FFC6FCE1FBD8E3EB969C" box="[892,1017,1133,1155]" collectionCode="BO" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="10" pageNumber="83" type="Herbarium">BO1454232</specimenCode>
!; BO1454233!; BO1454235!; BO1454236!;
<specimenCode id="642AEA6F2009FFC6FF20FB24E12F96B8" box="[189,317,1169,1191]" collectionCode="BO" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="10" pageNumber="83" type="Herbarium">BO1454237</specimenCode>
!])
</materialsCitation>
.
</paragraph>
<paragraph id="343342142009FFC6FF15FB60E58096EF" blockId="10.[136,1452,1237,2093]" box="[136,1426,1237,1265]" pageId="10" pageNumber="83">
<emphasis id="06F89E062009FFC6FF15FB60E1B796F0" bold="true" box="[136,421,1237,1263]" pageId="10" pageNumber="83">Additional literature:—</emphasis>
Tebbit (2005: 100) [drawing];
<bibRefCitation id="501D3FE52009FFC6FD67FB63E40F96EE" author="Thomas, D. C. &amp; Ardi, W. H. &amp; Hughes, M." box="[762,1053,1238,1265]" pageId="10" pageNumber="83" pagination="225 - 255" refId="ref20207" refString="Thomas, D. C., Ardi, W. H. &amp; Hughes, M. (2011) Nine new species of Begonia (Begoniaceae) from South and West Sulawesi, Indonesia. Edinburgh Journal of Botany 68: 225 - 255. https: // doi. org / 10.1017 / S 0960428611000072" type="journal article" year="2011">
Thomas
<emphasis id="06F89E062009FFC6FCC6FB62E38696EF" box="[859,916,1238,1264]" italics="true" pageId="10" pageNumber="83">et al.</emphasis>
(2011: 252)
</bibRefCitation>
[as
<emphasis id="06F89E062009FFC6FBC8FB62E48696EF" box="[1109,1172,1239,1264]" italics="true" pageId="10" pageNumber="83">B. sp.</emphasis>
I; identification key].
</paragraph>
</subSubSection>
<subSubSection id="7C96119F2009FFC0FF20FB4CE596931F" lastPageId="12" lastPageNumber="85" pageId="10" pageNumber="83" type="description">
<paragraph id="343342142009FFC0FF20FB4CE596931F" blockId="10.[136,1452,1237,2093]" lastBlockId="12.[136,1452,158,2093]" lastPageId="12" lastPageNumber="85" pageId="10" pageNumber="83">
<emphasis id="06F89E062009FFC6FF20FB4CE17F970C" bold="true" box="[189,365,1273,1299]" pageId="10" pageNumber="83">Description:—</emphasis>
Perennial, erect, monoecious herb to ca.
<quantity id="F374EFF12009FFC6FCABFB4FE39B970B" box="[822,905,1274,1300]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="83" unit="cm" value="140.0">140 cm</quantity>
tall.
<emphasis id="06F89E062009FFC6FC5FFB4FE3EE970B" bold="true" box="[962,1020,1274,1300]" italics="true" pageId="10" pageNumber="83">Stem</emphasis>
branched; internodes
<quantity id="F374EFF12009FFC6FB68FB4FE579970B" box="[1269,1387,1274,1301]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="15.0" metricValueMin="2.5" pageId="10" pageNumber="83" unit="cm" value="8.75" valueMax="15.0" valueMin="2.5">2.515 cm</quantity>
long, slightly swollen at the nodes, pale green or reddish, with white spots, glabrous except for microscopic glandular hairs.
<emphasis id="06F89E062009FFC6FF15FAF7E0C59743" bold="true" box="[136,215,1346,1372]" italics="true" pageId="10" pageNumber="83">Leaves</emphasis>
alternate;
<emphasis id="06F89E062009FFC6FED1FAF7E1B09743" box="[332,418,1346,1372]" italics="true" pageId="10" pageNumber="83">stipules</emphasis>
semi-persistent, ca. 1.54.7 ×
<quantity id="F374EFF12009FFC6FD69FAF7E3619743" box="[756,883,1346,1372]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.1" metricValueMin="1.2" pageId="10" pageNumber="83" unit="cm" value="1.65" valueMax="2.1" valueMin="1.2">1.22.1 cm</quantity>
, ovate, with an abaxially prominent midrib, margin translucent, keeled, apex narrowed into bristle up to
<quantity id="F374EFF12009FFC6FD4DFAD3E303979F" box="[720,785,1382,1408]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="83" unit="mm" value="3.0">3 mm</quantity>
long, glabrous, green;
<emphasis id="06F89E062009FFC6FB8DFAD3E475979F" box="[1040,1127,1382,1408]" italics="true" pageId="10" pageNumber="83">petioles</emphasis>
<quantity id="F374EFF12009FFC6FBF3FAD3E4DC979F" box="[1134,1230,1382,1409]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="10" pageNumber="83" unit="cm" value="12.5" valueMax="20.0" valueMin="5.0">520 cm</quantity>
long, concolourous with the stem, glabrous;
<emphasis id="06F89E062009FFC6FE07FA3FE1FA97BB" box="[410,488,1418,1444]" italics="true" pageId="10" pageNumber="83">lamina</emphasis>
basifixed, 828 ×
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, ovate to elliptic, asymmetric, base cordate and lobes not overlapping to overlapping, apex acuminate, margin broadly dentate, denticulate between the larger teeth, teeth bristle-pointed, adaxial surface green or reddish green, smooth and shiny, iridescent with pale green veins or sometimes variegated with white bands between the veins, glabrous; abaxial surface pale green, reddish-greenish or maroon in the central part of the leaves, glabrous or very sparsely hairy with bristly hairs on the veins; venation palmate-pinnate, primary veins (5)7, actinodromous, secondary veins craspedodromous.
<emphasis id="06F89E062009FFC6FB8DF988E4D39447" bold="true" box="[1040,1217,1597,1624]" italics="true" pageId="10" pageNumber="83">Inflorescences:</emphasis>
protogynous;
<emphasis id="06F89E062009FFC6FAFCF98BE1359463" italics="true" pageId="10" pageNumber="83">female inflorescences</emphasis>
2-flowered, 1
<specimenCount id="228A899D2009FFC6FE46F9D7E2519463" box="[475,579,1634,1660]" count="4" pageId="10" pageNumber="83" type="female">4 female</specimenCount>
inflorescences basal to male inflorescence part, peduncles (3)
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long, bracts caducous, up to 15 ×
<quantity id="F374EFF12009FFC6FD99F933E25A94BF" box="[516,584,1670,1696]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="83" unit="mm" value="9.0">9 mm</quantity>
, broadly ovate, pale green, glabrous;
<emphasis id="06F89E062009FFC6FC73F933E4D994BF" box="[1006,1227,1670,1696]" italics="true" pageId="10" pageNumber="83">male inflorescences</emphasis>
paniculate-cymose, composed of up to 7 subumbellate partial inflorescences, each with up to 12 flowers, peduncle of male inflorescence
<quantity id="F374EFF12009FFC6FF15F97BE0E794F7" box="[136,245,1742,1768]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="4.0" metricValueMin="0.1" pageId="10" pageNumber="83" unit="mm" value="20.5" valueMax="40.0" valueMin="1.0">140 mm</quantity>
long, pale green, glabrous, bracts caducous, 1015 ×
<quantity id="F374EFF12009FFC6FCD0F97BE3A894F7" box="[845,954,1742,1769]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="10" pageNumber="83" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">915 mm</quantity>
, broadly ovate to suborbicular, pale green or reddish-green, glabrous.
<emphasis id="06F89E062009FFC6FE00F944E28C9513" box="[413,670,1777,1804]" italics="true" pageId="10" pageNumber="83">
<emphasis id="06F89E062009FFC6FE00F944E22E9513" bold="true" box="[413,572,1777,1804]" italics="true" pageId="10" pageNumber="83">Male flowers:</emphasis>
pedicels
</emphasis>
<quantity id="F374EFF12009FFC6FD38F947E31D9513" box="[677,783,1778,1804]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="4.0" metricValueMin="0.8" pageId="10" pageNumber="83" unit="mm" value="24.0" valueMax="40.0" valueMin="8.0">840 mm</quantity>
long, pale green or reddish-green, glabrous;
<emphasis id="06F89E062009FFC6FA9CF947E5569513" box="[1281,1348,1778,1804]" italics="true" pageId="10" pageNumber="83">tepals</emphasis>
2, white, white tinged pink, or greenish-whitish, 1223 ×
<quantity id="F374EFF12009FFC6FD2BF8A3E321952F" box="[694,819,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="10" pageNumber="83" unit="mm" value="21.5" valueMax="28.0" valueMin="15.0">1528 mm</quantity>
, broadly ovate to suborbicular, base slightly cordate to truncate, apex rounded, outer surface glabrous;
<emphasis id="06F89E062009FFC6FD01F88FE333954B" box="[668,801,1850,1876]" italics="true" pageId="10" pageNumber="83">androecium</emphasis>
of ca. 7288 stamens, yellow, filaments up to ca.
<quantity id="F374EFF12009FFC6FAD2F88FE5BE954B" box="[1359,1452,1850,1877]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="83" unit="mm" value="1.5">1.5 mm</quantity>
long, slightly fused at the very base, anthers up to ca.
<quantity id="F374EFF12009FFC6FD42F8EBE3269567" box="[735,820,1886,1912]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="83" unit="mm" value="1.2">1.2 mm</quantity>
long, obovate, dehiscing through unilaterally positioned slits that are ca. ½ as long as the anthers.
<emphasis id="06F89E062009FFC6FDE3F834E3B69583" box="[638,932,1921,1948]" italics="true" pageId="10" pageNumber="83">
<emphasis id="06F89E062009FFC6FDE3F834E32F9583" bold="true" box="[638,829,1921,1948]" italics="true" pageId="10" pageNumber="83">Female flowers:</emphasis>
pedicels
</emphasis>
(10)
<quantity id="F374EFF12009FFC6FC76F837E4799583" box="[1003,1131,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="10" pageNumber="83" unit="mm" value="22.5" valueMax="30.0" valueMin="15.0">1530 mm</quantity>
long, pale green, glabrous;
<emphasis id="06F89E062009FFC6FF15F813E0D995DF" box="[136,203,1958,1984]" italics="true" pageId="10" pageNumber="83">tepals</emphasis>
5(6), white, unequal, larger 2028 ×
<quantity id="F374EFF12009FFC6FD17F813E31595DF" box="[650,775,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="10" pageNumber="83" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">1824 mm</quantity>
, ovate to elliptic, smallest one 1824 ×
<quantity id="F374EFF12009FFC6FB48F813E55795DF" box="[1237,1349,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="10" pageNumber="83" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">914 mm</quantity>
, obovate or elliptic, outer surface glabrous;
<emphasis id="06F89E062009FFC6FD93F87EE25C95FB" box="[526,590,1995,2020]" italics="true" pageId="10" pageNumber="83">ovary</emphasis>
1025 ×
<quantity id="F374EFF12009FFC6FD26F87FE30995FB" box="[699,795,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="10" pageNumber="83" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
, cylindrical or ellipsoid, wings 3, subequal, base rounded, apex rounded to truncate, widest point up to
<quantity id="F374EFF12009FFC6FD1FF85BE2C09A17" box="[642,722,2030,2056]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="83" unit="mm" value="12.0">12 mm</quantity>
(apically or subapically), glabrous, locules 3, placentation axile, placentae bilamellate; style ca.
<quantity id="F374EFF12009FFC6FE6EF7A7E25E9A33" box="[499,588,2066,2093]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="83" unit="mm" value="6.5">6.5 mm</quantity>
long, basally fused, 3branched, each stylodium bifurcate in the stigmatic region, stigmatic surface a spirally twisted papillose band, orange.
<emphasis id="06F89E06200FFFC0FC10FF2BE3C192A7" bold="true" box="[909,979,158,184]" italics="true" pageId="12" pageNumber="85">Fruit:</emphasis>
dry capsule;
<emphasis id="06F89E06200FFFC0FBECFF2BE4F392A7" box="[1137,1249,158,184]" italics="true" pageId="12" pageNumber="85">peduncles</emphasis>
<quantity id="F374EFF1200FFFC0FB76FF2BE57A92A6" box="[1259,1384,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="7.5" metricValueMin="1.5" pageId="12" pageNumber="85" unit="mm" value="45.0" valueMax="75.0" valueMin="15.0">1575 mm</quantity>
long;
<emphasis id="06F89E06200FFFC0FF15FF77E0F692C3" box="[136,228,194,220]" italics="true" pageId="12" pageNumber="85">pedicels</emphasis>
<quantity id="F374EFF1200FFFC0FF77FF77E17192C2" box="[234,355,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="12" pageNumber="85" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">1030 mm</quantity>
long, pale green, glabrous;
<emphasis id="06F89E06200FFFC0FD0AFF77E34C92C3" box="[663,862,194,220]" italics="true" pageId="12" pageNumber="85">seed-bearing part</emphasis>
(8)1025 ×
<quantity id="F374EFF1200FFFC0FC64FF77E47792C2" box="[1017,1125,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="12" pageNumber="85" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">610 mm</quantity>
(excluding the wings), ovoid to ellipsoid, wing shape as for ovary, pale green or reddish green, glabrous.
<emphasis id="06F89E06200FFFC0FC45FF53E40B931E" bold="true" box="[984,1049,230,257]" italics="true" pageId="12" pageNumber="85">Seeds</emphasis>
barrel-shaped, ca.
<quantity id="F374EFF1200FFFC0FB73FF53E556931E" box="[1262,1348,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="85" unit="mm" value="0.3">0.3 mm</quantity>
long.
</paragraph>
</subSubSection>
<caption id="60F3129C2008FFC7FF15F8E3E0A99A3F" ID-DOI="http://doi.org/10.5281/zenodo.13873888" ID-Zenodo-Dep="13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="11" pageNumber="84" startId="11.[136,229,1878,1900]" targetBox="[151,1435,190,1853]" targetPageId="11" targetType="figure">
<paragraph id="343342142008FFC7FF15F8E3E0A99A3F" blockId="11.[136,1452,1878,2080]" pageId="11" pageNumber="84">
<emphasis id="06F89E062008FFC7FF15F8E3E0EC9574" bold="true" box="[136,254,1878,1900]" pageId="11" pageNumber="84">FIGURE 4.</emphasis>
<taxonomicName id="F38C39972008FFC7FE98F8E3E1DB9574" authorityName="Wiriadinata" authorityYear="2013" box="[261,457,1878,1899]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="84" phylum="Tracheophyta" rank="species" species="hooveriana">
<emphasis id="06F89E062008FFC7FE98F8E3E1DB9574" box="[261,457,1878,1899]" italics="true" pageId="11" pageNumber="84">Begonia hooveriana</emphasis>
</taxonomicName>
. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I:
<emphasis id="06F89E062008FFC7FF03F853E1F695E4" box="[158,484,2022,2043]" italics="true" pageId="11" pageNumber="84">D.C. Thomas &amp; W.H. Ardi 08-82;</emphasis>
B, D, G, H:
<emphasis id="06F89E062008FFC7FDFFF853E3BF95E4" box="[610,941,2022,2043]" italics="true" pageId="11" pageNumber="84">D.C. Thomas &amp; W.H. Ardi 09-100</emphasis>
; C, E:
<emphasis id="06F89E062008FFC7FC6EF853E46F95E4" box="[1011,1149,2022,2043]" italics="true" pageId="11" pageNumber="84">W.H. Ardi 323</emphasis>
. Photos: D.C. Thomas &amp; W.H. Ardi.
</paragraph>
</caption>
<subSubSection id="7C96119F200FFFC1FF20FEBFE11A938F" lastPageId="13" lastPageNumber="86" pageId="12" pageNumber="85" type="distribution">
<paragraph id="34334214200FFFC0FF20FEBFE483933B" blockId="12.[136,1452,158,2093]" box="[189,1169,266,293]" pageId="12" pageNumber="85">
<emphasis id="06F89E06200FFFC0FF20FEBFE167933B" bold="true" box="[189,373,266,292]" pageId="12" pageNumber="85">Distribution:—</emphasis>
<collectingCountry id="4C9B0284200FFFC0FEE8FEBFE1F7933B" box="[373,485,266,292]" name="Indonesia" pageId="12" pageNumber="85">Indonesia</collectingCountry>
, endemic to
<collectingRegion id="F6488CF6200FFFC0FDEFFEBFE2C9933B" box="[626,731,266,292]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="85">Sulawesi</collectingRegion>
: South and
<collectingRegion id="F6488CF6200FFFC0FCC3FEBFE40F933B" box="[862,1053,266,292]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
(
<figureCitation id="ACB75E91200FFFC0FBB1FEBFE491933B" box="[1068,1155,266,292]" captionStart="FIGURE 1" captionStartId="2.[136,229,1953,1975]" captionTargetBox="[179,1408,188,1926]" captionTargetId="figure-22@2.[179,1408,187,1926]" captionTargetPageId="2" captionText="FIGURE 1. Distribution maps of Begonia species occurring on the southwestern arm of Sulawesi or the Selayar Islands (georeferenced specimen data from major collections [B, BO, E, K, KRB, L, SING], the Begonia Resource Center [Hughes et al. 2015a], as well as some additional confirmed observations). Grey lines indicate 1° graticules." figureDoi="http://doi.org/10.5281/zenodo.13873880" httpUri="https://zenodo.org/record/13873880/files/figure.png" pageId="12" pageNumber="85">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="34334214200FFFC0FF20FE9BE4E29373" blockId="12.[136,1452,158,2093]" pageId="12" pageNumber="85">
<emphasis id="06F89E06200FFFC0FF20FE9BE12C9357" bold="true" box="[189,318,302,328]" pageId="12" pageNumber="85">Habitat:—</emphasis>
Lowland to hill rain forest; growing in crevices of limestone rock, terrestrially at the base of limestone cliffs and hills, or on mossy granite rocks; partially shaded to open areas at ca.
<quantity id="F374EFF1200FFFC0FC61FEE7E4699372" box="[1020,1147,338,365]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="12" pageNumber="85" unit="m" value="550.0" valueMax="900.0" valueMin="200.0">200900 m</quantity>
elevation.
</paragraph>
<paragraph id="34334214200FFFC0FF20FEC3E21B90AF" blockId="12.[136,1452,158,2093]" pageId="12" pageNumber="85">
<emphasis id="06F89E06200FFFC0FF20FEC3E134938F" bold="true" box="[189,294,374,400]" pageId="12" pageNumber="85">Notes:—</emphasis>
In the original description,
<emphasis id="06F89E06200FFFC0FDC6FEC2E308938F" box="[603,794,374,401]" italics="true" pageId="12" pageNumber="85">W.S. Hoover 885</emphasis>
(BO) was designated as
<typeStatus id="EB37FCB6200FFFC0FBA5FEC3E488938F" box="[1080,1178,374,400]" pageId="12" pageNumber="85" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="501D3FE5200FFFC0FB34FEC3E58C938E" author="Wiriadinata, H." box="[1193,1438,374,401]" pageId="12" pageNumber="85" pagination="445 - 448" refId="ref20383" refString="Wiriadinata, H. (2013) A new species of Begonia (Begoniaceae) from South Sulawesi, Indonesia. Reinwardtia 13: 445 - 448." type="journal article" year="2013">Wiriadinata 2013: 45</bibRefCitation>
), and no
<typeStatus id="EB37FCB6200FFFC0FF47FE2FE12493AB" box="[218,310,410,436]" pageId="12" pageNumber="85" type="isotype">isotypes</typeStatus>
or other additional material were indicated. Specimens with the specified collection number could not be located in BO, but six sheets indicating W.S. Hoover, H. Wiriadinata, T. &amp; M. Hoover as collectors, 889 as the collection number, and the same locality and collection date as specified in the original description were present. The collection number 885 in the original description was probably a simple clerical error, but there are six herbarium sheets present in BO that are not labelled with sheet numbers, and there is no additional information on barcodes identifying the
<typeStatus id="EB37FCB6200FFFC0FEAFFDFBE1869077" box="[306,404,590,616]" pageId="12" pageNumber="85" type="holotype">holotype</typeStatus>
in the original description. Consequently, we designate one of the six sheets of
<emphasis id="06F89E06200FFFC0FA9BFDFAE0C49093" italics="true" pageId="12" pageNumber="85">W.S. Hoover et al. 889</emphasis>
as a
<typeStatus id="EB37FCB6200FFFC0FE93FDC7E1679093" box="[270,373,626,652]" pageId="12" pageNumber="85" type="lectotype">lectotype</typeStatus>
here. All sheets have flowering material, but the sheet stamped BO-1454234 is the only one that has both male and female flowers.
</paragraph>
<paragraph id="34334214200FFFC0FF20FD0EE2E397BB" blockId="12.[136,1452,158,2093]" pageId="12" pageNumber="85">
<taxonomicName id="F38C3997200FFFC0FF20FD0EE1B090CB" authorityName="Wiriadinata" authorityYear="2013" box="[189,418,698,724]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="85" phylum="Tracheophyta" rank="species" species="hooveriana">
<emphasis id="06F89E06200FFFC0FF20FD0EE1B090CB" box="[189,418,698,724]" italics="true" pageId="12" pageNumber="85">Begonia hooveriana</emphasis>
</taxonomicName>
was originally described from a single collection from a heavily disturbed limestone habitat in the centre of the city of Makale (
<bibRefCitation id="501D3FE5200FFFC0FDB5FD6BE2E690E7" author="Wiriadinata, H." box="[552,756,734,760]" pageId="12" pageNumber="85" pagination="445 - 448" refId="ref20383" refString="Wiriadinata, H. (2013) A new species of Begonia (Begoniaceae) from South Sulawesi, Indonesia. Reinwardtia 13: 445 - 448." type="journal article" year="2013">Wiriadinata 2013</bibRefCitation>
). Several other collections from South and
<collectingRegion id="F6488CF6200FFFC0FB77FD6BE5B990E7" box="[1258,1451,734,760]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
show characters that link them to
<taxonomicName id="F38C3997200FFFC0FE66FCB6E28B9103" authorityName="Wiriadinata" authorityYear="2013" box="[507,665,770,796]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="85" phylum="Tracheophyta" rank="species" species="hooveriana">
<emphasis id="06F89E06200FFFC0FE66FCB6E28B9103" box="[507,665,770,796]" italics="true" pageId="12" pageNumber="85">B. hooveriana</emphasis>
</taxonomicName>
: a similar erect or ascending growth habit and well-developed greyishbrownish periderm on older stems (
<figureCitation id="ACB75E91200FFFC0FD81FC93E28B915F" box="[540,665,806,832]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4A, C</figureCitation>
); relatively large, glabrous stipules with relatively short apical bristle (
<figureCitation id="ACB75E91200FFFC0FF0DFCFFE0F9917B" box="[144,235,842,868]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4D</figureCitation>
); relatively large male flowers in subumbellate partial inflorescences (
<figureCitation id="ACB75E91200FFFC0FC66FCFFE42C917B" box="[1019,1086,842,868]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4</figureCitation>
E-G); and a ovary and fruit wing attachment frequently conspicuously below the base of the seed-bearing part (
<figureCitation id="ACB75E91200FFFC0FC41FCDBE4519197" box="[988,1091,878,904]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4E, I</figureCitation>
). However, there is considerable morphological heterogeneity among these collections, e.g. leaf laminas are sometimes succulent and relatively small in plants growing in limestone rock crevices and sun-exposed habitats (e.g.,
<emphasis id="06F89E06200FFFC0FC7DFC03E4F191CF" box="[992,1251,950,977]" italics="true" pageId="12" pageNumber="85">Thomas &amp; Ardi 09-101</emphasis>
), but much larger and thinner leaf laminas can be found in other populations growing in soil or more shaded habitats (e.g.,
<emphasis id="06F89E06200FFFC0FAA8FC6FE11D9607" italics="true" pageId="12" pageNumber="85">Thomas &amp; Ardi 09-100</emphasis>
); male partial inflorescences can be sometimes subsessile (
<figureCitation id="ACB75E91200FFFC0FC3DFC4BE3E69607" box="[928,1012,1022,1048]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4F</figureCitation>
), but in most individuals they are borne on peduncles up to
<quantity id="F374EFF1200FFFC0FEFFFB97E1889622" box="[354,410,1058,1085]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="85" unit="cm" value="4.0">4 cm</quantity>
long (
<figureCitation id="ACB75E91200FFFC0FE7FFB97E2289623" box="[482,570,1058,1084]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4E</figureCitation>
); material from the
<typeStatus id="EB37FCB6200FFFC0FC85FB96E35A9622" box="[792,840,1059,1085]" pageId="12" pageNumber="85">type</typeStatus>
locality has ovaries that are relatively long and have a cylindric seed-bearing part, the wing attachment is considerably below the base of the seed-bearing part, and stigma branches are rather compact (
<figureCitation id="ACB75E91200FFFC0FE40FBDFE23C969B" box="[477,558,1130,1156]" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4I</figureCitation>
), while in other collections the ovary is much shorter and ellipsoid, the wing attachment is just below the seed-bearing part, and the stylodia are more elongated (e.g.,
<emphasis id="06F89E06200FFFC0FBF8FB3BE57796B7" box="[1125,1381,1166,1193]" italics="true" pageId="12" pageNumber="85">Thomas &amp; Ardi 09-100</emphasis>
) (
<figureCitation id="ACB75E91200FFFC0FAE3FB3BE0BC96D3" captionStart="FIGURE 4" captionStartId="11.[136,229,1878,1900]" captionTargetBox="[151,1435,190,1853]" captionTargetId="figure-17@11.[151,1435,190,1853]" captionTargetPageId="11" captionText="FIGURE 4. Begonia hooveriana. A: Habit. B: Habit, in partially shaded habitat. C: Habit, growing in crevice of limestone rock in sunexposed habitat. D: Stipule; scale bar = 1 cm. E: Inflorescence and young infructescence; scale bar = 3 cm. F: Subsessile male partial inflorescences, scale bar = 5 cm. G: Male flower, front view; scale bar = 2 cm. H: Female inflorescence; scale bar = 2 cm. I: Female inflorescence (material from type locality); scale bar = 2 cm. J: Ovary cross-section, bilamellate placentation, scale bar = 3 mm. A, F, I: D.C. Thomas &amp; W.H. Ardi 08-82; B, D, G, H: D.C. Thomas &amp; W.H. Ardi 09-100; C, E: W.H. Ardi 323. Photos: D.C. Thomas &amp; W.H. Ardi." figureDoi="http://doi.org/10.5281/zenodo.13873888" httpUri="https://zenodo.org/record/13873888/files/figure.png" pageId="12" pageNumber="85">Fig. 4H</figureCitation>
). Most of this variability seems to be in a morphological continuum, however, and further investigations are needed to evaluate whether meaningful segregates can be identified. Moreover, some of the observed variability may be the result of introgression with sympatrically occurring species (e.g.,
<taxonomicName id="F38C3997200FFFC0FCF2FB4EE4CC970B" authority="Ardi &amp; D. C. Thomas" authorityName="Ardi &amp; D. C. Thomas" box="[879,1246,1274,1300]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="85" phylum="Tracheophyta" rank="species" species="siregarii">
<emphasis id="06F89E06200FFFC0FCF2FB4EE3FC970B" box="[879,1006,1275,1300]" italics="true" pageId="12" pageNumber="85">B. siregarii</emphasis>
Ardi &amp; D.C.Thomas
</taxonomicName>
[Ardi
<emphasis id="06F89E06200FFFC0FAB6FB4EE576970B" box="[1323,1380,1274,1300]" italics="true" pageId="12" pageNumber="85">et al.</emphasis>
2014: 263]), and this aspect requires further investigation ideally employing a population genetic approach. Plants cultivated at Bogor Botanic Gardens from material collected at the
<typeStatus id="EB37FCB6200FFFC0FC9BFAF6E3249742" box="[774,822,1347,1373]" pageId="12" pageNumber="85">type</typeStatus>
locality regularly flowered and developed mature-sized male flowers that, however, were always abscised before opening. This dropping of male flowers before opening may be an indicator of a hybrid origin (Dewitte
<emphasis id="06F89E06200FFFC0FDF7FA3EE2B097BB" box="[618,674,1418,1444]" italics="true" pageId="12" pageNumber="85">et al.</emphasis>
2011).
</paragraph>
<paragraph id="34334214200FFFC0FF20FA1AE58497D7" blockId="12.[136,1452,158,2093]" box="[189,1430,1454,1481]" pageId="12" pageNumber="85">
<taxonomicName id="F38C3997200FFFC0FF20FA1AE1B097D7" authorityName="Wiriadinata" authorityYear="2013" box="[189,418,1454,1480]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="85" phylum="Tracheophyta" rank="species" species="hooveriana">
<emphasis id="06F89E06200FFFC0FF20FA1AE1B097D7" box="[189,418,1454,1480]" italics="true" pageId="12" pageNumber="85">Begonia hooveriana</emphasis>
</taxonomicName>
has been observed to be locally cultivated as outdoor ornamental plant in
<collectingRegion id="F6488CF6200FFFC0FB7CFA1BE58097D7" box="[1249,1426,1454,1480]" country="Indonesia" name="Sulawesi Selatan" pageId="12" pageNumber="85">South Sulawesi</collectingRegion>
.
</paragraph>
<paragraph id="34334214200FFFC0FF20FA67E1E994F7" blockId="12.[136,1452,158,2093]" pageId="12" pageNumber="85">
<emphasis id="06F89E06200FFFC0FF20FA67E2E697F3" bold="true" box="[189,756,1490,1516]" pageId="12" pageNumber="85">Provisional IUCN conservation assessment:—</emphasis>
Vulnerable (VU) B1ab(iii),B2ab(iii). This species has been collected at several localities in South and
<collectingRegion id="F6488CF6200FFFC0FDE1FA43E353940F" box="[636,833,1526,1552]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
. The collections from
<collectingRegion id="F6488CF6200FFFC0FBDAFA43E51B940F" box="[1095,1289,1526,1552]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
are in legally protected areas (Wera Forest Reserve and Lore Lindu National Park), but the majority of collections are from
<collectingRegion id="F6488CF6200FFFC0FAF7F9AFE0FF9447" country="Indonesia" name="Sulawesi Selatan" pageId="12" pageNumber="85">South Sulawesi</collectingRegion>
from localities that are within or in close proximity to settlements and agricultural land. Some of the collection localities in the
<taxonomicName id="F38C3997200FFFC0FEA3F9D7E1BB9463" box="[318,425,1634,1660]" pageId="12" pageNumber="90" rank="subPhylum" subPhylum="Enrekang">Enrekang</taxonomicName>
Karst and Tanah Toraja were heavily disturbed. This species is able to cope with a certain degree of habitat disturbance, but because of its fragmented distribution, the reduction of lowland and hill forest habitats in South and
<collectingRegion id="F6488CF6200FFFC0FEE1F91FE22994DB" box="[380,571,1706,1732]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
(Cannon
<emphasis id="06F89E06200FFFC0FD37F91EE2F194DB" box="[682,739,1706,1732]" italics="true" pageId="12" pageNumber="85">et al.</emphasis>
2005, 2007), and the small EOO (ca.
<quantity id="F374EFF1200FFFC0FB13F91FE4EB94DB" box="[1166,1273,1706,1732]" metricMagnitude="6" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="85" unit="km" value="3100.0">3,100 km</quantity>
<superScript id="C3F9EF5C200FFFC0FB64F91FE51394A7" attach="left" box="[1273,1281,1706,1720]" fontSize="6" pageId="12" pageNumber="85">2</superScript>
) and AOO, we assess this species as Vulnerable.
</paragraph>
<paragraph id="34334214200FFFC0FF20F947E5BE9A33" blockId="12.[136,1452,158,2093]" pageId="12" pageNumber="85">
<emphasis id="06F89E06200FFFC0FF20F947E2689513" bold="true" box="[189,634,1778,1804]" pageId="12" pageNumber="85">Additional specimens examined:—</emphasis>
<materialsCitation id="84E44849200FFFC0FDE7F947E323952F" collectingDate="1979-06-01" collectorName="M. M. J. v. Balgooy" country="Indonesia" location="Parampadende" municipality="Wera Waterfall" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Utara">
<collectingCountry id="4C9B0284200FFFC0FDE7F947E3049513" box="[634,790,1778,1804]" name="Indonesia" pageId="12" pageNumber="85">INDONESIA</collectingCountry>
.
<collectingRegion id="F6488CF6200FFFC0FCB2F947E38A9513" box="[815,920,1778,1804]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="85">Sulawesi</collectingRegion>
.
<emphasis id="06F89E06200FFFC0FC2DF947E4819513" bold="true" box="[944,1171,1778,1804]" pageId="12" pageNumber="85">
<collectingRegion id="F6488CF6200FFFC0FC2DF947E49C9513" box="[944,1166,1778,1804]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="85">Central Sulawesi</collectingRegion>
:
</emphasis>
<collectingMunicipality id="D457D86E200FFFC0FB35F946E5709513" box="[1192,1378,1778,1805]" pageId="12" pageNumber="85">Wera Waterfall</collectingMunicipality>
, near
<location id="315314CF200FFFC0FF15F8A3E122952F" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FF15F8A3E122952F" box="[136,304,1814,1840]" country="Indonesia" municipality="Wera Waterfall" name="Parampadende" pageId="12" pageNumber="85" stateProvince="Sulawesi Utara">Parampadende</location>
,
<date id="403264D4200FFFC0FEA1F8A3E1DA952F" box="[316,456,1814,1841]" pageId="12" pageNumber="85" value="1979-06-01">
<collectingDate id="50769D3C200FFFC0FEA1F8A3E1DA952F" box="[316,456,1814,1841]" pageId="12" pageNumber="85" value="1979-06-01">1 June 1979</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FE49F8A2E2C9952F" box="[468,731,1814,1840]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FE49F8A2E289952F" box="[468,667,1814,1840]" pageId="12" pageNumber="85">M.M.J.v. Balgooy</collectorName>
3619
</emphasis>
(A, K)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FCA0F8A3E17E954B" collectingDate="2002-05-25" collectionCode="CEB" collectorName="Ramadanil" country="Indonesia" county="Pakuli" location="Donggala" municipality="Biromaru" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingCounty id="DD523A98200FFFC0FCA0F8A3E395952F" box="[829,903,1814,1840]" pageId="12" pageNumber="85">Pakuli</collectingCounty>
,
<collectingMunicipality id="D457D86E200FFFC0FC09F8A3E410952F" box="[916,1026,1814,1840]" pageId="12" pageNumber="85">Biromaru</collectingMunicipality>
,
<location id="315314CF200FFFC0FB93F8A3E46C952F" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FB93F8A3E46C952F" box="[1038,1150,1814,1840]" country="Indonesia" county="Pakuli" municipality="Biromaru" name="Donggala" pageId="12" pageNumber="85" stateProvince="Sulawesi Tengah">Donggala</location>
,
<date id="403264D4200FFFC0FB17F8A3E536952F" box="[1162,1316,1814,1841]" pageId="12" pageNumber="85" value="2002-05-25">
<collectingDate id="50769D3C200FFFC0FB17F8A3E536952F" box="[1162,1316,1814,1841]" pageId="12" pageNumber="85" value="2002-05-25">25 May 2002</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FAADF8A3E10B954B" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FAADF8A3E5B9952F" box="[1328,1451,1814,1840]" pageId="12" pageNumber="85">Ramadanil</collectorName>
et al. RP 805
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FEB5F88FE176954B" box="[296,356,1850,1876]" country="Indonesia" lsid="urn:lsid:biocol.org:col:14168" name="Tadulako University" pageId="12" pageNumber="85" type="Herbarium">CEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FEEAF88FE1DE9567" collectingDate="2007-05-11" collectionCode="CEB" collectorName="Ramadanil Pitopang &amp; Sylva PC" country="Indonesia" county="Pakuli" location="Gumbasa River" municipality="Maleo" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingCounty id="DD523A98200FFFC0FEEAF88FE1D3954B" box="[375,449,1850,1876]" pageId="12" pageNumber="85">Pakuli</collectingCounty>
,
<collectingMunicipality id="D457D86E200FFFC0FE50F88FE207954B" box="[461,533,1850,1876]" pageId="12" pageNumber="85">Maleo</collectingMunicipality>
nesting ground, near
<location id="315314CF200FFFC0FC9BF88FE3A5954B" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FC9BF88FE3A5954B" box="[774,951,1850,1876]" country="Indonesia" county="Pakuli" municipality="Maleo" name="Gumbasa River" pageId="12" pageNumber="85" stateProvince="Sulawesi Tengah">Gumbasa River</location>
,
<date id="403264D4200FFFC0FC5FF88FE44A954B" box="[962,1112,1850,1877]" pageId="12" pageNumber="85" value="2007-05-11">
<collectingDate id="50769D3C200FFFC0FC5FF88FE44A954B" box="[962,1112,1850,1877]" pageId="12" pageNumber="85" value="2007-05-11">11 May 2007</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FBFEF88FE1689567" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FBFEF88FE55E954B" box="[1123,1356,1850,1876]" pageId="12" pageNumber="85">Ramadanil Pitopang</collectorName>
&amp;
<collectorName id="997927C2200FFFC0FAF2F88FE0BE9566" pageId="12" pageNumber="85">Sylva PC</collectorName>
UNTAD, RP 2046
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FE15F8EBE1D69567" box="[392,452,1886,1912]" country="Indonesia" lsid="urn:lsid:biocol.org:col:14168" name="Tadulako University" pageId="12" pageNumber="85" type="Herbarium">CEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FE4AF8EBE5B59567" box="[471,1447,1886,1912]" collectingDate="2018-07-29" collectionCode="BO, SING, KRB" collectorName="W. H. Ardi" country="Indonesia" location="Maleo" municipality="Saluki" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingMunicipality id="D457D86E200FFFC0FE4AF8EBE2339567" box="[471,545,1886,1912]" pageId="12" pageNumber="85">Saluki</collectingMunicipality>
,
<location id="315314CF200FFFC0FDB1F8EBE2669567" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FDB1F8EBE2669567" box="[556,628,1886,1912]" country="Indonesia" municipality="Saluki" name="Maleo" pageId="12" pageNumber="85" stateProvince="Sulawesi Tengah">Maleo</location>
nesting ground,
<date id="403264D4200FFFC0FCB3F8EBE3D39567" box="[814,961,1886,1912]" pageId="12" pageNumber="85" value="2018-07-29">
<collectingDate id="50769D3C200FFFC0FCB3F8EBE3D39567" box="[814,961,1886,1912]" pageId="12" pageNumber="85" value="2018-07-29">29 July 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FC56F8EAE4DC9567" box="[971,1230,1886,1912]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FC56F8EAE42B9567" box="[971,1081,1886,1912]" pageId="12" pageNumber="85">W.H. Ardi</collectorName>
et al. WI 249
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FB40F8EBE5159567" box="[1245,1287,1886,1912]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="12" pageNumber="85" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200FFFC0FA8FF8EBE5449567" box="[1298,1366,1886,1912]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="12" pageNumber="85" type="Herbarium">SING</collectionCode>
,
<collectionCode id="529DDAD1200FFFC0FAFCF8EBE58D9567" box="[1377,1439,1886,1912]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="12" pageNumber="85" type="Herbarium">KRB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FF15F837E4719583" box="[136,1123,1922,1949]" collectingDate="2018-08-10" collectionCode="BO, FIPIA, KRB, SING" collectorName="W. H. Ardi" country="Indonesia" location="Wera" municipality="Sigi" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingMunicipality id="D457D86E200FFFC0FF15F837E0AA9583" box="[136,184,1922,1948]" pageId="12" pageNumber="85">Sigi</collectingMunicipality>
,
<location id="315314CF200FFFC0FF59F836E0EC9582" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FF59F836E0EC9582" box="[196,254,1923,1949]" country="Indonesia" municipality="Sigi" name="Wera" pageId="12" pageNumber="85" stateProvince="Sulawesi Tengah">Wera</location>
waterfall,
<date id="403264D4200FFFC0FEE4F837E2059583" box="[377,535,1922,1948]" pageId="12" pageNumber="85" value="2018-08-10">
<collectingDate id="50769D3C200FFFC0FEE4F837E2059583" box="[377,535,1922,1948]" pageId="12" pageNumber="85" value="2018-08-10">10 Aug. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FDBEF836E33D9583" box="[547,815,1922,1948]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FDBEF836E2819583" box="[547,659,1922,1948]" pageId="12" pageNumber="85">W.H. Ardi</collectorName>
et al. WI 317
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FCDDF837E3789583" box="[832,874,1922,1948]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="12" pageNumber="85" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200FFFC0FCEAF837E3D29583" box="[887,960,1922,1948]" country="Indonesia" lsid="urn:lsid:biocol.org:col:13104" name="Institut Teknologi Bandung" pageId="12" pageNumber="85" type="Herbarium">FIPIA</collectionCode>
,
<collectionCode id="529DDAD1200FFFC0FC51F836E41A9582" box="[972,1032,1923,1949]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="12" pageNumber="85" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200FFFC0FB88F837E4499583" box="[1045,1115,1922,1948]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="12" pageNumber="85" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="84E44849200FFFC0FBF2F837E23C95DF" collectingDate="1929-06-17" collectionCode="BO" collectorName="G. K. Kjellberg" country="Indonesia" location="Pasoei" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Selatan">
<emphasis id="06F89E06200FFFC0FBF2F837E53D9583" bold="true" box="[1135,1327,1922,1948]" pageId="12" pageNumber="85">
<collectingRegion id="F6488CF6200FFFC0FBF2F837E5389583" box="[1135,1322,1922,1948]" country="Indonesia" name="Sulawesi Selatan" pageId="12" pageNumber="85">South Sulawesi</collectingRegion>
:
</emphasis>
<location id="315314CF200FFFC0FAAAF837E5919583" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FAAAF837E5919583" box="[1335,1411,1922,1948]" country="Indonesia" name="Pasoei" pageId="12" pageNumber="85" stateProvince="Sulawesi Selatan">Pasoei</location>
,
<date id="403264D4200FFFC0FA12F837E0E895DF" pageId="12" pageNumber="85" value="1929-06-17">
<collectingDate id="50769D3C200FFFC0FA12F837E0E895DF" pageId="12" pageNumber="85" value="1929-06-17">17 june 1929</collectingDate>
</date>
.
<emphasis id="06F89E06200FFFC0FE9BF812E1F895DF" box="[262,490,1958,1985]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FE9BF812E1B995DF" box="[262,427,1958,1985]" pageId="12" pageNumber="85">G.K. Kjellberg</collectorName>
1396
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FE64F813E23795DF" box="[505,549,1958,1984]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="12" pageNumber="85" type="Herbarium">BO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FDA4F812E5B595DF" box="[569,1447,1958,1985]" collectingDate="1998-09-03" collectionCode="BO" collectorName="W. S. Hoover" country="Indonesia" location="Central Park Makele" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Selatan">
near
<location id="315314CF200FFFC0FDECF813E34E95DF" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FDECF813E34E95DF" box="[625,860,1958,1984]" country="Indonesia" name="Central Park Makele" pageId="12" pageNumber="85" stateProvince="Sulawesi Selatan">Central Park Makele</location>
,
<date id="403264D4200FFFC0FCFAF813E3FD95DF" box="[871,1007,1958,1984]" pageId="12" pageNumber="85" value="1998-09-03">
<collectingDate id="50769D3C200FFFC0FCFAF813E3FD95DF" box="[871,1007,1958,1984]" pageId="12" pageNumber="85" value="1998-09-03">3 Sep. 1998</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FC67F812E4E595DF" box="[1018,1271,1958,1985]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FC67F812E49495DF" box="[1018,1158,1959,1985]" pageId="12" pageNumber="85">W.S. Hoover</collectorName>
et al. 889
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FA9BF813E52295DF" box="[1286,1328,1958,1984]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="12" pageNumber="85" type="Herbarium">BO</collectionCode>
[6 sheets])
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FF15F87EE3BE95FB" box="[136,940,1994,2021]" collectingDate="2003-11-15" collectionCode="SING" collectorName="J. Vermeulen" country="Indonesia" location="Kpg Lokkok" municipality="Buntu area" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Selatan">
<collectingMunicipality id="D457D86E200FFFC0FF15F87EE11495FA" box="[136,262,1995,2021]" pageId="12" pageNumber="85">Buntu area</collectingMunicipality>
,
<location id="315314CF200FFFC0FE8FF87EE1B795FB" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FE8FF87EE1B795FB" box="[274,421,1994,2021]" country="Indonesia" municipality="Buntu area" name="Kpg Lokkok" pageId="12" pageNumber="85" stateProvince="Sulawesi Selatan">Kpg Lokkok</location>
,
<date id="403264D4200FFFC0FE2CF87FE25C95FB" box="[433,590,1994,2021]" pageId="12" pageNumber="85" value="2003-11-15">
<collectingDate id="50769D3C200FFFC0FE2CF87FE25C95FB" box="[433,590,1994,2021]" pageId="12" pageNumber="85" value="2003-11-15">15 Nov. 2003</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FDC7F87EE33E95FB" box="[602,812,1994,2020]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FDC7F87EE2F995FB" box="[602,747,1994,2020]" pageId="12" pageNumber="85">J. Vermeulen</collectorName>
2299
</emphasis>
(L,
<collectionCode id="529DDAD1200FFFC0FCC0F87FE3B195FB" box="[861,931,1994,2020]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="12" pageNumber="85" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FC25F87FE45F9A17" collectingDate="2008-05-16" collectorName="D. C. Thomas &amp; W. H. Ardi" country="Indonesia" county="Botanic Gardens" location="Makale" municipality="Tanah Toraja" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Bali">
cultivated at
<collectingRegion id="F6488CF6200FFFC0FBD6F87FE46895FB" box="[1099,1146,1994,2020]" country="Indonesia" name="Bali" pageId="12" pageNumber="85">Bali</collectingRegion>
<collectingCounty id="DD523A98200FFFC0FB1FF87FE52C95FB" box="[1154,1342,1994,2020]" pageId="12" pageNumber="85">Botanic Gardens</collectingCounty>
(material collected in
<collectingMunicipality id="D457D86E200FFFC0FE93F85BE1B09A17" box="[270,418,2030,2056]" pageId="12" pageNumber="85">Tanah Toraja</collectingMunicipality>
,
<location id="315314CF200FFFC0FE31F85BE2169A17" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FE31F85BE2169A17" box="[428,516,2030,2056]" country="Indonesia" county="Botanic Gardens" municipality="Tanah Toraja" name="Makale" pageId="12" pageNumber="85" stateProvince="Bali">Makale</location>
),
<date id="403264D4200FFFC0FD8BF85BE2BF9A17" box="[534,685,2030,2057]" pageId="12" pageNumber="85" value="2008-05-16">
<collectingDate id="50769D3C200FFFC0FD8BF85BE2BF9A17" box="[534,685,2030,2057]" pageId="12" pageNumber="85" value="2008-05-16">16 May 2008</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FD2AF85AE4329A17" box="[695,1056,2030,2057]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FD2AF85AE3589A17" box="[695,842,2030,2057]" pageId="12" pageNumber="85">D.C. Thomas</collectorName>
&amp;
<collectorName id="997927C2200FFFC0FCF1F85AE3CB9A17" box="[876,985,2030,2056]" pageId="12" pageNumber="85">W.H. Ardi</collectorName>
08-82
</emphasis>
(E)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FBCAF85BE3189A33" collectingDate="2009-04-29" collectionCode="BO" collectorName="D. C. Thomas &amp; W. H. Ardi" country="Indonesia" location="Makale" municipality="Tanah Toraja" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Selatan">
<collectingMunicipality id="D457D86E200FFFC0FBCAF85BE4F99A17" box="[1111,1259,2030,2056]" pageId="12" pageNumber="85">Tanah Toraja</collectingMunicipality>
, close to
<location id="315314CF200FFFC0FACDF85BE5B59A17" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FACDF85BE5B59A17" box="[1360,1447,2030,2056]" country="Indonesia" municipality="Tanah Toraja" name="Makale" pageId="12" pageNumber="85" stateProvince="Sulawesi Selatan">Makale</location>
,
<date id="403264D4200FFFC0FF15F7A7E10F9A33" box="[136,285,2066,2092]" pageId="12" pageNumber="85" value="2009-04-29">
<collectingDate id="50769D3C200FFFC0FF15F7A7E10F9A33" box="[136,285,2066,2092]" pageId="12" pageNumber="85" value="2009-04-29">29 Apr. 2009</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FEB5F7A6E2B59A33" box="[296,679,2066,2093]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FEB5F7A6E1AC9A33" box="[296,446,2066,2093]" pageId="12" pageNumber="85">D.C. Thomas</collectorName>
&amp;
<collectorName id="997927C2200FFFC0FE7FF7A6E2429A33" box="[482,592,2066,2092]" pageId="12" pageNumber="85">W.H. Ardi</collectorName>
09-100
</emphasis>
(
<collectionCode id="529DDAD1200FFFC0FD2BF7A7E2F29A33" box="[694,736,2066,2092]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="12" pageNumber="85" type="Herbarium">BO</collectionCode>
, E)
</materialsCitation>
;
<materialsCitation id="84E44849200FFFC0FC8BF7A7E5BE9A33" box="[790,1452,2066,2093]" collectingDate="2009-04-29" collectorName="D. C. Thomas" country="Indonesia" location="Makale" municipality="Tanah Toraja" pageId="12" pageNumber="85" specimenCount="1" stateProvince="Sulawesi Selatan">
<collectingMunicipality id="D457D86E200FFFC0FC8BF7A7E3B89A33" box="[790,938,2066,2092]" pageId="12" pageNumber="85">Tanah Toraja</collectingMunicipality>
, close to
<location id="315314CF200FFFC0FB89F7A7E4799A33" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200FFFC0FB89F7A7E4799A33" box="[1044,1131,2066,2092]" country="Indonesia" municipality="Tanah Toraja" name="Makale" pageId="12" pageNumber="85" stateProvince="Sulawesi Selatan">Makale</location>
,
<date id="403264D4200FFFC0FBEBF7A7E5199A33" box="[1142,1291,2066,2092]" pageId="12" pageNumber="85" value="2009-04-29">
<collectingDate id="50769D3C200FFFC0FBEBF7A7E5199A33" box="[1142,1291,2066,2092]" pageId="12" pageNumber="85" value="2009-04-29">29 Apr. 2009</collectingDate>
</date>
,
<emphasis id="06F89E06200FFFC0FA8BF7A6E5BE9A33" box="[1302,1452,2066,2093]" italics="true" pageId="12" pageNumber="85">
<collectorName id="997927C2200FFFC0FA8BF7A6E5BE9A33" box="[1302,1452,2066,2093]" pageId="12" pageNumber="85">D.C. Thomas</collectorName>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="34334214200EFFC1FF15FF2AE11A938F" blockId="13.[136,1452,158,400]" pageId="13" pageNumber="86">
<emphasis id="06F89E06200EFFC1FF15FF2AE17592A7" box="[136,359,158,185]" italics="true" pageId="13" pageNumber="86">&amp; W.H. Ardi 09-101</emphasis>
(BO, E);
<materialsCitation id="84E44849200EFFC1FE4EFF2BE4D192A7" box="[467,1219,158,185]" collectingDate="2018-11-03" collectionCode="BO, KRB, SING" collectorName="Bambapuang Karst &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FE4EFF2BE2BE92A6" box="[467,684,158,185]" pageId="13" pageNumber="86">Bambapuang Karst</collectorName>
,
<date id="403264D4200EFFC1FD2AFF2BE35092A7" box="[695,834,158,185]" pageId="13" pageNumber="86" value="2018-11-03">
<collectingDate id="50769D3C200EFFC1FD2AFF2BE35092A7" box="[695,834,158,185]" pageId="13" pageNumber="86" value="2018-11-03">3 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FCD0FF2AE3F892A7" box="[845,1002,158,184]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FC14FF2BE3A892A7" box="[905,954,158,184]" pageId="13" pageNumber="86">Ardi</collectorName>
323
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FC64FF2BE43192A7" box="[1017,1059,158,184]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FBB3FF2AE47892A6" box="[1070,1130,159,185]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FBE8FF2BE4A992A7" box="[1141,1211,158,184]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FB53FF2BE2BE92C3" collectingDate="2018-11-03" collectionCode="BO, KRB, SING" collectorName="Bambapuang Karst &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FB53FF2BE5B592A6" box="[1230,1447,158,185]" pageId="13" pageNumber="86">Bambapuang Karst</collectorName>
,
<date id="403264D4200EFFC1FF15FF77E10B92C3" box="[136,281,194,221]" pageId="13" pageNumber="86" value="2018-11-03">
<collectingDate id="50769D3C200EFFC1FF15FF77E10B92C3" box="[136,281,194,221]" pageId="13" pageNumber="86" value="2018-11-03">3 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FEBAFF76E1D892C3" box="[295,458,194,220]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FEFBFF77E18592C3" box="[358,407,194,220]" pageId="13" pageNumber="86">Ardi</collectorName>
325
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FE41FF77E21492C3" box="[476,518,194,220]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FD89FF76E24292C2" box="[532,592,195,221]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FDC3FF77E2B692C3" box="[606,676,194,220]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FD27FF76E0C4931F" collectingDate="2018-11-05" collectionCode="BO, KRB, SING" collectorName="Karst Baroko &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FD27FF76E34792C3" box="[698,853,194,221]" pageId="13" pageNumber="86">Karst Baroko</collectorName>
,
<taxonomicName id="F38C3997200EFFC1FCFEFF77E3C392C3" box="[867,977,194,220]" pageId="13" pageNumber="90" rank="subPhylum" subPhylum="Enrekang">Enrekang</taxonomicName>
,
<date id="403264D4200EFFC1FC43FF77E46292C3" box="[990,1136,194,221]" pageId="13" pageNumber="86" value="2018-11-05">
<collectingDate id="50769D3C200EFFC1FC43FF77E46292C3" box="[990,1136,194,221]" pageId="13" pageNumber="86" value="2018-11-05">5 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FBE0FF76E53392C3" box="[1149,1313,194,220]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FB20FF77E4FC92C3" box="[1213,1262,194,220]" pageId="13" pageNumber="86">Ardi</collectorName>
337
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FAAEFF77E54F92C3" box="[1331,1373,194,220]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FAF6FF76E5B592C2" box="[1387,1447,195,221]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FF15FF53E0DC931F" box="[136,206,230,256]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FF7FFF52E440931F" box="[226,1106,230,257]" collectingDate="2018-11-05" collectionCode="BO, KRB, SING" collectorName="Karst Benteng Alla &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FF7FFF52E1A9931F" box="[226,443,230,257]" pageId="13" pageNumber="86">Karst Benteng Alla</collectorName>
,
<taxonomicName id="F38C3997200EFFC1FE5BFF53E226931F" box="[454,564,230,256]" pageId="13" pageNumber="90" rank="subPhylum" subPhylum="Enrekang">Enrekang</taxonomicName>
,
<date id="403264D4200EFFC1FDDDFF53E2DE931F" box="[576,716,230,257]" pageId="13" pageNumber="86" value="2018-11-05">
<collectingDate id="50769D3C200EFFC1FDDDFF53E2DE931F" box="[576,716,230,257]" pageId="13" pageNumber="86" value="2018-11-05">5 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FD4AFF52E365931F" box="[727,887,230,256]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FC88FF53E354931F" box="[789,838,230,256]" pageId="13" pageNumber="86">Ardi</collectorName>
338
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FC1BFF53E3A2931F" box="[902,944,230,256]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FC21FF52E3EA931E" box="[956,1016,231,257]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FB99FF53E458931F" box="[1028,1098,230,256]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FBC0FF53E253933B" collectingDate="2018-11-05" collectionCode="BO, KRB, SING" collectorName="Gandang Batu &amp; Toraja &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FBC0FF53E513931E" box="[1117,1281,230,257]" pageId="13" pageNumber="86">Gandang Batu</collectorName>
,
<collectorName id="997927C2200EFFC1FA91FF53E544931F" box="[1292,1366,230,256]" pageId="13" pageNumber="86">Toraja</collectorName>
,
<date id="403264D4200EFFC1FAFCFF53E0D1933B" pageId="13" pageNumber="86" value="2018-11-05">
<collectingDate id="50769D3C200EFFC1FAFCFF53E0D1933B" pageId="13" pageNumber="86" value="2018-11-05">5 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FF50FEBEE178933B" box="[205,362,266,292]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FE94FEBFE128933B" box="[265,314,266,292]" pageId="13" pageNumber="86">Ardi</collectorName>
339
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FEE5FEBFE1B0933B" box="[376,418,266,292]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FE30FEBEE1FB933A" box="[429,489,267,293]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FE6EFEBFE22B933B" box="[499,569,266,292]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FDD6FEBFE541933B" box="[587,1363,266,293]" collectingDate="2018-11-08" collectionCode="BO, KRB, SING" collectorName="Pemandian Tilangnga &amp; Ardi" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FDD6FEBFE352933B" box="[587,832,266,292]" pageId="13" pageNumber="86">Pemandian Tilangnga</collectorName>
,
<date id="403264D4200EFFC1FCD7FEBFE3C6933B" box="[842,980,266,293]" pageId="13" pageNumber="86" value="2018-11-08">
<collectingDate id="50769D3C200EFFC1FCD7FEBFE3C6933B" box="[842,980,266,293]" pageId="13" pageNumber="86" value="2018-11-08">8 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FC43FEBEE469933B" box="[990,1147,266,292]" italics="true" pageId="13" pageNumber="86">
W.H.
<collectorName id="997927C2200EFFC1FB86FEBFE45E933B" box="[1051,1100,266,292]" pageId="13" pageNumber="86">Ardi</collectorName>
348
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FB14FEBFE4A1933B" box="[1161,1203,266,292]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FB23FEBEE4E8933A" box="[1214,1274,267,293]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FA99FEBFE558933B" box="[1284,1354,266,292]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FAC0FEBFE2B49373" collectingDate="2004-04-22" collectingDateMax="2018-11-08" collectingDateMin="2004-04-22" collectionCode="BO, KRB, SING, BKL" collectorName="Londa &amp; Torajan Tomb &amp; W. H. Ardi &amp; P. Harwood" location="Cultivated" municipality="Brooklyn Botanic Gardens" pageId="13" pageNumber="86" specimenCount="1">
<collectorName id="997927C2200EFFC1FAC0FEBFE5B5933B" box="[1373,1447,266,292]" pageId="13" pageNumber="86">Londa</collectorName>
,
<collectorName id="997927C2200EFFC1FF15FE9BE13A9357" box="[136,296,302,328]" pageId="13" pageNumber="86">Torajan Tomb</collectorName>
,
<date id="403264D4200EFFC1FEA9FE9BE1D39357" box="[308,449,302,329]" pageId="13" pageNumber="86" value="2018-11-08">
<collectingDate id="50769D3C200EFFC1FEA9FE9BE1D39357" box="[308,449,302,329]" pageId="13" pageNumber="86" value="2018-11-08">8 Nov. 2018</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FE50FE9AE27C9357" box="[461,622,302,328]" italics="true" pageId="13" pageNumber="86">
<collectorName id="997927C2200EFFC1FE50FE9AE22E9357" box="[461,572,302,328]" pageId="13" pageNumber="86">W.H. Ardi</collectorName>
350
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FDE0FE9BE2B59357" box="[637,679,302,328]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="86" type="Herbarium">BO</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FD29FE9AE2E29356" box="[692,752,303,329]" country="Indonesia" httpUri="http://grbio.org/cool/c6c6-q10t" name="Kebun Raya Bogor" pageId="13" pageNumber="86" type="Herbarium">KRB</collectionCode>
,
<collectionCode id="529DDAD1200EFFC1FD61FE9BE3509357" box="[764,834,302,328]" country="Singapore" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="13" pageNumber="86" type="Herbarium">SING</collectionCode>
).
<emphasis id="06F89E06200EFFC1FCC8FE9BE45B9357" bold="true" box="[853,1097,302,328]" pageId="13" pageNumber="86">
<location id="315314CF200EFFC1FCC8FE9BE3C19357" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200EFFC1FCC8FE9BE3C19357" box="[853,979,302,328]" municipality="Brooklyn Botanic Gardens" name="Cultivated" pageId="13" pageNumber="86">Cultivated</location>
material:
</emphasis>
cultivated at
<collectingMunicipality id="D457D86E200EFFC1FB7EFE9BE0FA9373" pageId="13" pageNumber="86">Brooklyn Botanic Gardens</collectingMunicipality>
,
<date id="403264D4200EFFC1FF68FEE7E1999373" box="[245,395,338,364]" pageId="13" pageNumber="86" value="2004-04-22">
<collectingDate id="50769D3C200EFFC1FF68FEE7E1999373" box="[245,395,338,364]" pageId="13" pageNumber="86" value="2004-04-22">22 Apr. 2004</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FE05FEE6E2429373" box="[408,592,338,364]" italics="true" pageId="13" pageNumber="86">
<collectorName id="997927C2200EFFC1FE05FEE6E20C9373" box="[408,542,338,364]" pageId="13" pageNumber="86">P. Harwood</collectorName>
126
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FDFCFEE7E28D9373" box="[609,671,338,364]" country="USA" lsid="urn:lsid:biocol.org:col:15693" name="Brooklyn Botanic Garden" pageId="13" pageNumber="86" type="Herbarium">BKL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E44849200EFFC1FD2EFEE7E116938F" collectingDate="2007-06-26" collectionCode="BKL" collectorName="M. C. Tebbitt" location="Brooklyn Botanic Gardens" pageId="13" pageNumber="86" specimenCount="1">
cultivated at
<location id="315314CF200EFFC1FCDBFEE7E4659373" LSID="urn:lsid:plazi:treatment:BC25F3022009FFC1FF15FC6FE11A938F:315314CF200EFFC1FCDBFEE7E4659373" box="[838,1143,338,364]" name="Brooklyn Botanic Gardens" pageId="13" pageNumber="86">Brooklyn Botanic Gardens</location>
,
<date id="403264D4200EFFC1FB19FEE7E5009373" box="[1156,1298,338,365]" pageId="13" pageNumber="86" value="2007-06-26">
<collectingDate id="50769D3C200EFFC1FB19FEE7E5009373" box="[1156,1298,338,365]" pageId="13" pageNumber="86" value="2007-06-26">26 Jun 2007</collectingDate>
</date>
,
<emphasis id="06F89E06200EFFC1FA83FEE6E0BD938F" italics="true" pageId="13" pageNumber="86">
<collectorName id="997927C2200EFFC1FA83FEE6E5BE9373" box="[1310,1452,338,365]" pageId="13" pageNumber="86">M.C. Tebbitt</collectorName>
s.n.
</emphasis>
(
<collectionCode id="529DDAD1200EFFC1FF23FEC3E0EE938F" box="[190,252,374,400]" country="USA" lsid="urn:lsid:biocol.org:col:15693" name="Brooklyn Botanic Garden" pageId="13" pageNumber="86" type="Herbarium">BKL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>