205 lines
19 KiB
XML
205 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt & Liew, Thor-Seng" docDate="2017" docId="B7DFEA90BA3825A78EDE0664169FD0C8" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus alticola Foon & Liew, 2017, sp. n." docType="treatment" docUuid="E88B3EDB-0DEE-417F-BDEA-613A67ADA99B" docUuidSource="ZooBank" docVersion="6" lastPageNumber="9" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="8" updateTime="1668164728433" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Foon, Junn Kitt</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>692</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>81</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.692.14706</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.692.14706</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-692-1</mods:identifier>
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<mods:identifier type="ZBK">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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<mods:identifier type="ZooBank">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132786246" LSID="urn:lsid:zoobank.org:act:E88B3EDB-0DEE-417F-BDEA-613A67ADA99B" httpUri="http://treatment.plazi.org/id/B7DFEA90BA3825A78EDE0664169FD0C8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph pageId="7" pageNumber="8">
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<taxonomicName LSID="http://zoobank.org/E88B3EDB-0DEE-417F-BDEA-613A67ADA99B" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus alticola" order="Architaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="alticola">Alycaeus alticola</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
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Figures 7J, 8, 31G
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="type locality">
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<paragraph pageId="7" pageNumber="8">Type locality.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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PHG 77 Bukit Mengapur, Pahang (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="3.7450001">3°44'42"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="102.83778">102°50'16"E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="type material">
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<paragraph pageId="7" pageNumber="8">Type material.</paragraph>
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<paragraph pageId="7" pageNumber="8">Holotype. PHG 77 Bukit Mengapur, Pahang: BOR/MOL 12977. Paratypes. PHG 77 Bukit Mengapur, Pahang: BOR/MOL 6888/1, BOR/MOL 8398/26.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="materials_examined">
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<paragraph pageId="7" pageNumber="8">Other examined materials.</paragraph>
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<paragraph pageId="7" pageNumber="8">PHG 77 Bukit Mengapur, Pahang: BOR/MOL 8397(11).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="etymology">
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<paragraph pageId="7" pageNumber="8">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Latin for "dweller at heights". Named after the
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<normalizedToken originalValue="species’">species'</normalizedToken>
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apparent ecological preference for habitats at higher and drier parts of limestone hills.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
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<paragraph pageId="7" pageNumber="8">Description.</paragraph>
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<paragraph pageId="7" pageNumber="8">Protoconch. Smooth.</paragraph>
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<paragraph pageId="7" pageNumber="8">Shell shape. Conical. Shell height: 5.08-6.04 mm. Shell width: 5.04-5.87 mm.</paragraph>
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<paragraph pageId="7" pageNumber="8">Spire. Spire height: 1.64-2.23 mm. Spire width: 2.32-2.71 mm. Number of whorls: up to 5. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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Whorl constriction. At about 4
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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whorls posterior of protoconch.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<pageBreakToken pageId="8" pageNumber="9" start="start">Breathing</pageBreakToken>
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tube. Length: 0.42-0.58 mm.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">Aperture and peristome. Aperture circular, moderately expanded. Aperture height: 2.40-2.70 mm. Aperture width: 2.32-2.72 mm. Peristome double, not thickened, slightly notched at suture. Peristome orientation 35-38° oblique with respect to the coiling axis.</paragraph>
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<paragraph pageId="8" pageNumber="9">Spiral lines. Indistinct, regularly spaced. Approximately 22-44 lines per 1 mm.</paragraph>
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<paragraph pageId="8" pageNumber="9">Radial ribs running anterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 10-14 ribs per 1 mm.</paragraph>
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<paragraph pageId="8" pageNumber="9">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced, white and thicker than those anterior of breathing tube, evenly spaced. Approximately 9-15 ribs per 1 mm.</paragraph>
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<paragraph pageId="8" pageNumber="9">Radial ribs running posterior of breathing tube. Radial ribs absent immediately posterior of constriction, becoming pronounced and regularly spaced at the middle until prior to aperture. Approximately 7-15 ribs per 1 mm.</paragraph>
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<paragraph pageId="8" pageNumber="9">Operculum. Concave, rounded. Exterior covered with very thin calcareous layer, with indistinct appressed multilamellae. Interior covered with proteinaceous coating, smooth, with indistinct appressed multilamellae and indistinct mamilla.</paragraph>
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<paragraph pageId="8" pageNumber="9">Shell colour. Whorls yellow, fading to white posterior of constriction. Peristome white.</paragraph>
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<paragraph pageId="8" pageNumber="9">Living animal. Body cream-white with fine brown blotches. Head brown. Tentacles brown.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="habitat">
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<paragraph pageId="8" pageNumber="9">Habitat and ecology.</paragraph>
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<paragraph pageId="8" pageNumber="9">Lives on rock surfaces covered with powdery lichen at higher and drier areas of the limestone hills, often at open places exposed to ambient sunlight. They never appear on wet and mossy sections of limestone rocks and are rarely seen in lower parts of limestone hills that are shaded by forest cover.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="distribution">
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<paragraph pageId="8" pageNumber="9">Distribution range.</paragraph>
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<paragraph pageId="8" pageNumber="9">Restricted to Bukit Mengapur, Pahang.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="differential diagnosis">
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<paragraph pageId="8" pageNumber="9">Differential diagnosis.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus alticola" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="alticola">Alycaeus alticola</taxonomicName>
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sp. n. varies in the degree of ultimate whorl expansion within population but has a consistently more expanded and globose whorl compared to
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<taxonomicName lsidName="A. costacrassa" pageId="8" pageNumber="9" rank="species" species="costacrassa">A. costacrassa</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="A. selangoriensis" pageId="8" pageNumber="9" rank="species" species="selangoriensis">A. selangoriensis</taxonomicName>
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sp. n. and
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<taxonomicName lsidName="A. kapayanensis" pageId="8" pageNumber="9" rank="species" species="kapayanensis">A. kapayanensis</taxonomicName>
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.
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus alticola" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="alticola">Alycaeus alticola</taxonomicName>
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sp. n. shares similar shell shape with
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus charasensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="charasensis">Alycaeus charasensis</taxonomicName>
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sp. n. but differs in the presence of pronounced radial ribs and spiral lines, as well as less expanded ultimate whorl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="discussion">
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<paragraph pageId="8" pageNumber="9">Discussion.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName lsidName="A. alticola" pageId="8" pageNumber="9" rank="species" species="alticola">A. alticola</taxonomicName>
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sp. n. is among the larger yellow, conical shelled
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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in Peninsular Malaysia.
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</paragraph>
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<caption pageId="8" pageNumber="9">
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<paragraph pageId="8" pageNumber="9">
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Figure 7. Photographs of 18 living
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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species.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus balingensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="balingensis">Alycaeus balingensis</taxonomicName>
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Tomlin, 1948, BOR/MOL 8356 C
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus liratulus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="liratulus">Alycaeus liratulus</taxonomicName>
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(Preston, 1907), BOR/MOL 8334 D
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti</taxonomicName>
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Morgan, 1885b, BOR/MOL 6835 E
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus conformis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="conformis">Alycaeus conformis</taxonomicName>
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Fulton, 1902, BOR/MOL 6809 F
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus gibbosulus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gibbosulus">Alycaeus gibbosulus</taxonomicName>
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Stoliczka, 1872, BOR/MOL 6850 G
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus gibbosulus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="gibbosulus">Alycaeus gibbosulus</taxonomicName>
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Stoliczka, 1872, BOR/MOL 8526 H
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
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Morgan, 1885a, BOR/MOL 8341 I
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
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Morgan, 1885a, BOR/MOL 8336 J
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus alticola" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="alticola">Alycaeus alticola</taxonomicName>
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sp. n., BOR/MOL 8398 K
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus charasensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="charasensis">Alycaeus charasensis</taxonomicName>
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sp. n., BOR/MOL 8399 L
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus clementsi" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="clementsi">Alycaeus clementsi</taxonomicName>
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sp. n., BOR/MOL 8364 M
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus costacrassa" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="costacrassa">Alycaeus costacrassa</taxonomicName>
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sp. n., BOR/MOL 6811 N
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus expansus" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="expansus">Alycaeus expansus</taxonomicName>
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sp. n., BOR/MOL 6367 O
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kapayanensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="kapayanensis">Alycaeus kapayanensis</taxonomicName>
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Morgan, 1885b, BOR/MOL 13005 P
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kelantanensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="kelantanensis">Alycaeus kelantanensis</taxonomicName>
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Sykes, 1902, BOR/MOL 8325 Q
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kelantanensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="kelantanensis">Alycaeus kelantanensis</taxonomicName>
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Sykes, 1902, BOR/MOL 6200 R
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kurauensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="kurauensis">Alycaeus kurauensis</taxonomicName>
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sp. n., BOR/MOL 6851 S
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
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Crosse, 1879a, BOR/MOL 6852 T
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus regalis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="regalis">Alycaeus regalis</taxonomicName>
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sp. n., BOR/MOL 6881 U
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus selangoriensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="selangoriensis">Alycaeus selangoriensis</taxonomicName>
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sp. n., BOR/MOL 6371
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<normalizedToken originalValue="V–W">V-W</normalizedToken>
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus senyumensis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="senyumensis">Alycaeus senyumensis</taxonomicName>
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sp. n., BOR/MOL 6249. All photographs by Junn Kitt Foon.
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</paragraph>
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</caption>
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<caption pageId="8" pageNumber="9">
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<paragraph pageId="8" pageNumber="9">
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Figure 8.
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus alticola" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="alticola">Alycaeus alticola</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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Shell of holotype, BOR/MOL 8398
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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Close up of shell of holotype, BOR/MOL 8398
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<normalizedToken originalValue="I–L">I-L</normalizedToken>
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Operculum of paratype, BOR/MOL 8398
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<normalizedToken originalValue="M–Q">M-Q</normalizedToken>
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Shell of paratype, BOR/MOL 8398. Scale bars:
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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,
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<normalizedToken originalValue="I–Q">I-Q</normalizedToken>
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1 mm; H 0.5 mm. All photographs by Junn Kitt Foon.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |