treatments-xml/data/B7/53/87/B75387965F46BF3CFCB5FA7BFDCBF7A3.xml
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<document ID-DOI="10.37520/aemnp.2020.008" ID-ISSN="1804-6487" ID-Zenodo-Dep="3879304" ID-ZooBank="16B1C03F-93E7-4BFF-B895-F846EA4CD029" approvalRequired="4" approvalRequired_for_taxonomicNames="2" approvalRequired_for_treatments="2" checkinTime="1591380478429" checkinUser="tatiana" docAuthor="Liu, Hsing-Che, Hu, Fang-Shuo &amp; Fikáček, Martin" docDate="2020" docId="B75387965F46BF3CFCB5FA7BFDCBF7A3" docLanguage="en" docName="ActaEntMusNatPra.60.1.155-162.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@751e664e" docStyleName="actaentmusnatpra.2018to2019.journal_article" docTitle="Coelostoma (Lachnocoelostoma) phallicum Orchymont 1940" docType="treatment" docVersion="2" lastPageId="4" lastPageNumber="159" masterDocId="4B6AFFEE5F45BF38FFCDFFFFFFB5FFBE" masterDocTitle="Review of the genus Coelostoma of Taiwan with description of a new species (Coleoptera: Hydrophilidae)" masterLastPageNumber="162" masterPageNumber="155" pageId="3" pageNumber="158" updateTime="1591380803713" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the genus Coelostoma of Taiwan with description of a new species (Coleoptera: Hydrophilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Hsing-Che</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hu, Fang-Shuo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fikáček, Martin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-03-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>60</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>155</mods:start>
<mods:end>162</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.37520/aemnp.2020.008</mods:identifier>
<mods:identifier type="ISSN">1804-6487</mods:identifier>
<mods:identifier type="Zenodo-Dep">3879304</mods:identifier>
<mods:identifier type="ZooBank">16B1C03F-93E7-4BFF-B895-F846EA4CD029</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:B75387965F46BF3CFCB5FA7BFDCBF7A3" httpUri="http://treatment.plazi.org/id/B75387965F46BF3CFCB5FA7BFDCBF7A3" lastPageId="4" lastPageNumber="159" pageId="3" pageNumber="158">
<subSubSection pageId="3" pageNumber="158" type="nomenclature">
<paragraph blockId="3.[828,1377,1412,1527]" pageId="3" pageNumber="158">
<heading centered="true" fontSize="10" level="2" pageId="3" pageNumber="158" reason="3">
<taxonomicName authority="Orchymont, 1940" authorityName="Orchymont" authorityYear="1940" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="phallicum" subGenus="Lachnocoelostoma">
<emphasis bold="true" pageId="3" pageNumber="158">
<emphasis bold="true" box="[888,1020,1412,1435]" italics="true" pageId="3" pageNumber="158">Coelostoma</emphasis>
(
<emphasis bold="true" box="[1035,1247,1412,1435]" italics="true" pageId="3" pageNumber="158">Lachnocoelostoma</emphasis>
)
<emphasis bold="true" box="[1263,1377,1412,1435]" italics="true" pageId="3" pageNumber="158">phallicum</emphasis>
<bibRefCitation author="ORCHYMONT A. d'" box="[1031,1233,1444,1467]" pageId="3" pageNumber="158" pagination="157 - 197" refId="ref7487" refString="ORCHYMONT A. d' 1940: Contribution a l'etude des Palpicornia XIV. Bulletin et Annales de la Societe Entomologique de Belgique 80: 157 - 197." type="journal article" year="1940">Orchymont, 1940</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1073,1192,1474,1493]" pageId="3" pageNumber="158" type="description">
<paragraph blockId="3.[828,1377,1412,1527]" box="[1073,1192,1474,1493]" pageId="3" pageNumber="158">
(
<figureCitation box="[1079,1151,1474,1493]" captionStart="Fig" captionStartId="4.[151,182,1301,1320]" captionTargetBox="[178,1413,195,1286]" captionTargetId="figure@4.[169,1419,189,1286]" captionTargetPageId="4" captionText="Fig. 2. male genitalia and front view of the head of Taiwanese Coelostoma species.AF genitals in dorsal view:A C. fallaciosum Orchymont, 1936; B C. vitalisi Orchymont, 1923; C C. phallicum Orchymont, 1940; D C. wui Orchymont, 1940; E C. bhutanicum Jayaswal, 1972; F C. stultum (Walker, 1858), specimen from Japan: Iriomote Island; GH front view of head with labrum: G C. taiwanense sp. nov., paratype; H C. fallaciosum." figureDoi="http://doi.org/10.5281/zenodo.3879306" httpUri="https://zenodo.org/record/3879306/files/figure.png" pageId="3" pageNumber="158">Figs 2C</figureCitation>
,
<figureCitation box="[1160,1185,1474,1493]" captionStart="Fig" captionStartId="5.[152,183,1195,1214]" captionTargetBox="[170,1418,189,1180]" captionTargetId="figure@5.[170,1418,189,1180]" captionTargetPageId="5" captionText="Fig. 3. Habitat of Coelostoma species.A Hsinchu County, Zhudong: habitat of C. stultum (Walker, 1858); B Hsinchu County, Hengshan Township: habitat of C. wui Orchymont, 1940; C Taichung City, Wufeng District:habitat of C. taiwanense sp.nov.and C. bhutanicum Jayaswal, 1972; D Hsinchu County, Beipu Township: habitat of C. fallaciosum Orchymont, 1936; E Yilan County, Jiaoxi Township: habitat of C. phallicum Orchymont, 1940; F Taipei city, Neihu District: habitat of C. vitalisi Orchymont, 1923. Photos by H.-C. Liu (AC, F), R. Matsushima (D) and F.-S. Hu (E)." figureDoi="http://doi.org/10.5281/zenodo.3879308" httpUri="https://zenodo.org/record/3879308/files/figure.png" pageId="3" pageNumber="158">3E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[828,1235,1508,1527]" pageId="3" pageNumber="158" type="reference_group">
<paragraph blockId="3.[828,1377,1412,1527]" box="[828,1235,1508,1527]" pageId="3" pageNumber="158">
<treatmentCitationGroup box="[828,1235,1508,1527]" pageId="3" pageNumber="158">
<taxonomicName authority="Orchymont, 1940: 158" authorityName="Orchymont" authorityPageNumber="158" authorityYear="1940" box="[828,1230,1508,1527]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="phallicum">
<treatmentCitation author="ORCHYMONT A. d'" box="[828,1230,1508,1527]" page="158" pageId="3" pageNumber="158" year="1940">
<emphasis box="[828,1024,1508,1527]" italics="true" pageId="3" pageNumber="158">Coelostoma phallicum</emphasis>
<bibRefCitation author="ORCHYMONT A. d'" box="[1030,1230,1508,1527]" pageId="3" pageNumber="158" pagination="157 - 197" refId="ref7487" refString="ORCHYMONT A. d' 1940: Contribution a l'etude des Palpicornia XIV. Bulletin et Annales de la Societe Entomologique de Belgique 80: 157 - 197." type="journal article" year="1940">Orchymont, 1940: 158</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="158" type="materials_examined">
<paragraph blockId="3.[828,1438,1550,1862]" pageId="3" pageNumber="158">
<emphasis bold="true" box="[828,1133,1550,1569]" pageId="3" pageNumber="158">
Material examined.
<collectingCountry box="[1017,1107,1550,1569]" name="Taiwan" pageId="3" pageNumber="158">TAIWAN</collectingCountry>
:
<collectingRegion box="[1119,1133,1550,1569]" country="Taiwan" name="Taitung" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1133,1193,1554,1567]" pageId="3" pageNumber="158">TAITUNG</emphasis>
</collectingRegion>
</emphasis>
<emphasis bold="true" box="[1193,1200,1550,1569]" pageId="3" pageNumber="158">:</emphasis>
1 (
<collectionCode box="[1257,1311,1550,1569]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="158">TARI</collectionCode>
): Hsiaotianchi (
<emphasis bold="true" box="[835,900,1575,1595]" pageId="3" pageNumber="158">
<collectionCode box="[835,900,1575,1595]" pageId="3" pageNumber="158">ṬXṮ</collectionCode>
</emphasis>
),
<date box="[922,1006,1577,1596]" pageId="3" pageNumber="158" value="2016-04-02">2.iv.2016</date>
, B.-X. Guo leg.
<collectingRegion box="[1161,1216,1577,1596]" country="Taiwan" name="Yilan" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1161,1176,1577,1596]" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1177,1216,1581,1594]" pageId="3" pageNumber="158">
<subScript attach="right" box="[1177,1216,1581,1594]" fontSize="6" pageId="3" pageNumber="158">YILAN</subScript>
</emphasis>
</emphasis>
</collectingRegion>
<emphasis bold="true" box="[1216,1223,1577,1596]" pageId="3" pageNumber="158">:</emphasis>
2 (
<collectionCode box="[1296,1369,1577,1596]" pageId="3" pageNumber="158">KMNH</collectionCode>
): Yijie (
<emphasis bold="true" box="[835,876,1600,1621]" pageId="3" pageNumber="158">áỡ</emphasis>
)ʼ [incorrect spelling, correct name of the locality is Aijie (
<emphasis bold="true" box="[1375,1417,1600,1621]" pageId="3" pageNumber="158">Ṟỡ</emphasis>
)], Yunshan Township [
<emphasis bold="true" box="[995,1058,1628,1649]" pageId="3" pageNumber="158">DƜǿ</emphasis>
],
<date box="[1074,1195,1630,1649]" pageId="3" pageNumber="158" value="2000-09-12" valueMax="2000-09-14" valueMin="2000-09-12">1214.ix.2000</date>
, Takashi Shimada leg.; 2,
<specimenCount box="[828,876,1653,1676]" pageId="3" pageNumber="158" type="female">6 ♀♀</specimenCount>
(
<collectionCode box="[889,955,1657,1676]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="158">NMNS</collectionCode>
): Jiaoxi Township [
<emphasis bold="true" box="[1122,1185,1655,1676]" pageId="3" pageNumber="158">ḌṰǿ</emphasis>
], Tangwei Brook Park [
<emphasis bold="true" pageId="3" pageNumber="158">ȁā ặảợċă</emphasis>
],
<geoCoordinate box="[954,1070,1683,1702]" degrees="24" direction="north" minutes="49" orientation="latitude" pageId="3" pageNumber="158" precision="1" seconds="41.9" value="24.828306">24°49'41.9&quot;N</geoCoordinate>
<geoCoordinate box="[1078,1204,1683,1702]" degrees="121" direction="east" minutes="46" orientation="longitude" pageId="3" pageNumber="158" precision="1" seconds="12.4" value="121.77011">121°46'12.4&quot;E</geoCoordinate>
,
<date box="[1215,1306,1683,1702]" pageId="3" pageNumber="158" value="2018-04-22">22.iv.2018</date>
, F. S. Hu leg., hand collecting; 5 (
<collectionCode box="[1022,1083,1710,1729]" pageId="3" pageNumber="158">HCLC</collectionCode>
): Jiaoxi Township [
<emphasis bold="true" box="[1244,1307,1708,1729]" pageId="3" pageNumber="158">ḌṰǿ</emphasis>
],
<geoCoordinate box="[1324,1436,1710,1729]" degrees="24.824385" direction="north" orientation="latitude" pageId="3" pageNumber="158" precision="1" value="24.824385">24.824385°N</geoCoordinate>
<geoCoordinate box="[828,950,1737,1756]" degrees="121.769800" direction="east" orientation="longitude" pageId="3" pageNumber="158" precision="1" value="121.7698">121.769800°E</geoCoordinate>
,
<date box="[961,1047,1737,1756]" pageId="3" pageNumber="158" value="2019-07-02">2.vii.2019</date>
, Hsing-Che Liu leg;
<specimenCount box="[1233,1305,1737,1756]" pageId="3" pageNumber="158" type="generic">26 spec.</specimenCount>
(
<collectionCode box="[1318,1379,1737,1756]" pageId="3" pageNumber="158">HCLC</collectionCode>
): Luodong sports park (
<emphasis bold="true" box="[986,1112,1760,1781]" pageId="3" pageNumber="158">ůǥḋḙċă</emphasis>
), Luodong To. [
<emphasis bold="true" box="[1256,1320,1760,1781]" pageId="3" pageNumber="158">ůǥḃ</emphasis>
],
<geoCoordinate box="[1339,1433,1763,1782]" degrees="24.6841" direction="north" orientation="latitude" pageId="3" pageNumber="158" precision="5" value="24.6841">24.6841°N</geoCoordinate>
,
<geoCoordinate box="[828,929,1790,1809]" degrees="121.7517" direction="east" orientation="longitude" pageId="3" pageNumber="158" precision="5" value="121.7517">121.7517°E</geoCoordinate>
,
<date box="[937,1017,1790,1809]" pageId="3" pageNumber="158" value="2020-01-19">19.i.2020</date>
, leg. F. S. Hu (near pond).
<collectingRegion box="[1240,1313,1790,1809]" country="Taiwan" name="Hsinchu" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1240,1256,1790,1809]" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1256,1313,1794,1807]" pageId="3" pageNumber="158">HSINCHU</emphasis>
</emphasis>
</collectingRegion>
<emphasis bold="true" box="[1313,1320,1790,1809]" pageId="3" pageNumber="158">:</emphasis>
1 (
<collectionCode box="[1366,1427,1790,1809]" pageId="3" pageNumber="158">HCLC</collectionCode>
): Emei Township [
<emphasis bold="true" box="[964,1026,1814,1835]" pageId="3" pageNumber="158">ŔḎǿ</emphasis>
],
<date box="[1041,1129,1817,1836]" pageId="3" pageNumber="158" value="2018-09-11">11.ix.2018</date>
,Hsing-Che Liu leg;
<collectingRegion box="[1292,1345,1817,1836]" country="Taiwan" name="Taipei" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1292,1306,1817,1836]" pageId="3" pageNumber="158">
<emphasis bold="true" box="[1306,1345,1821,1834]" pageId="3" pageNumber="158">
<subScript attach="left" box="[1306,1345,1821,1834]" fontSize="6" pageId="3" pageNumber="158">TAIPEI</subScript>
</emphasis>
</emphasis>
</collectingRegion>
: 1,
<specimenCount box="[1391,1436,1813,1836]" pageId="3" pageNumber="158" type="female">1 ♀♀</specimenCount>
(
<collectionCode box="[834,900,1843,1862]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="158">NMNS</collectionCode>
): NTU Farm [
<emphasis bold="true" box="[1017,1143,1840,1861]" pageId="3" pageNumber="158">ȒŘRḤōö</emphasis>
],
<date box="[1160,1251,1843,1862]" pageId="3" pageNumber="158" value="2011-09-22">22.ix.2011</date>
, Y. T. Wang (
<emphasis bold="true" box="[1361,1424,1840,1861]" pageId="3" pageNumber="158">ŦṪŽ</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="159" pageId="3" pageNumber="158" type="diagnosis">
<paragraph blockId="3.[828,1437,1887,2070]" pageId="3" pageNumber="158">
<emphasis bold="true" box="[828,948,1887,1910]" pageId="3" pageNumber="158">Diagnosis.</emphasis>
Body length
<quantity box="[1107,1246,1887,1910]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="3" pageNumber="158" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.25.5 mm</quantity>
, maximum body width
<quantity box="[904,1049,1919,1942]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="3" pageNumber="158" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.23.3 mm</quantity>
. Body black. Labrum yellowish. Maxillary and labial palpi yellowish. Elytra densely punctate, without traces of series, sutural stria reaching basal half of elytra. Ventral face of pro- and mesofemora densely pubescent.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3879306" ID-Zenodo-Dep="3879306" httpUri="https://zenodo.org/record/3879306/files/figure.png" pageId="4" pageNumber="159" startId="4.[151,182,1301,1320]" targetBox="[178,1413,195,1286]" targetPageId="4">
<paragraph blockId="4.[151,1437,1301,1374]" pageId="4" pageNumber="159">
Fig. 2. male genitalia and front view of the head of Taiwanese
<taxonomicName box="[670,772,1301,1320]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="subGenus" subGenus="Coelostoma">
<emphasis box="[670,772,1301,1320]" italics="true" pageId="4" pageNumber="159">Coelostoma</emphasis>
</taxonomicName>
species.AF genitals in dorsal view:A
<taxonomicName authority="Orchymont, 1936" authorityName="Orchymont" authorityYear="1936" box="[1154,1433,1301,1320]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="fallaciosum">
<emphasis box="[1154,1168,1301,1320]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<taxonomicName authority="Orchymont" authorityName="Orchymont" authorityYear="1936" box="[1177,1433,1301,1320]" pageId="4" pageNumber="159" rank="species" species="fallaciosum">
<taxonomicName box="[1177,1277,1301,1320]" pageId="4" pageNumber="159" rank="species" species="fallaciosum">
<emphasis box="[1177,1277,1301,1320]" italics="true" pageId="4" pageNumber="159">fallaciosum</emphasis>
</taxonomicName>
<bibRefCitation author="ORCHYMONT A. d'" box="[1281,1433,1301,1320]" pageId="4" pageNumber="159" pagination="1 - 38" refId="ref7450" refString="ORCHYMONT A. d' 1936: Revision des Coelostoma (s. str.) non americains. Memoires du Musee Royal d ' Historire Naturelle de Belgique 7 (2): 1 - 38." type="journal article" year="1936">Orchymont, 1936</bibRefCitation>
</taxonomicName>
</taxonomicName>
; B
<taxonomicName authority="Orchymont, 1923" authorityName="Orchymont" authorityYear="1923" box="[187,428,1327,1347]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[187,201,1328,1347]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<taxonomicName authority="Orchymont" authorityName="Orchymont" authorityYear="1923" box="[211,428,1327,1346]" pageId="4" pageNumber="159" rank="species" species="vitalisi">
<taxonomicName box="[211,269,1327,1346]" pageId="4" pageNumber="159" rank="species" species="vitalisi">
<emphasis box="[211,269,1327,1346]" italics="true" pageId="4" pageNumber="159">vitalisi</emphasis>
</taxonomicName>
<bibRefCitation author="ORCHYMONT A. d'" box="[275,428,1327,1346]" pageId="4" pageNumber="159" pagination="416 - 421" refId="ref7399" refString="ORCHYMONT A. d' 1923: Neue order interessante Sphaeridiinen und Hydrophilinen der Malayischen Region. Treubia 3: 416 - 421." type="journal article" year="1923">Orchymont, 1923</bibRefCitation>
</taxonomicName>
</taxonomicName>
; C
<taxonomicName authority="Orchymont" authorityName="Orchymont" authorityYear="1940" box="[472,740,1327,1347]" class="Insecta" family="Hydrophilidae" genus="C." kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="phallicum">
<emphasis box="[472,486,1328,1347]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<taxonomicName authority="Orchymont" authorityName="Orchymont" authorityYear="1940" box="[497,740,1327,1346]" pageId="4" pageNumber="159" rank="species" species="phallicum">
<taxonomicName box="[497,582,1327,1346]" pageId="4" pageNumber="159" rank="species" species="phallicum">
<emphasis box="[497,582,1327,1346]" italics="true" pageId="4" pageNumber="159">phallicum</emphasis>
</taxonomicName>
<bibRefCitation author="ORCHYMONT A. d'" box="[588,740,1327,1346]" pageId="4" pageNumber="159" pagination="157 - 197" refId="ref7487" refString="ORCHYMONT A. d' 1940: Contribution a l'etude des Palpicornia XIV. Bulletin et Annales de la Societe Entomologique de Belgique 80: 157 - 197." type="journal article" year="1940">Orchymont, 1940</bibRefCitation>
</taxonomicName>
</taxonomicName>
; D
<taxonomicName authority="Orchymont, 1940" authorityName="Orchymont" authorityYear="1940" box="[786,1000,1327,1347]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="wui">
<emphasis box="[786,800,1328,1347]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<taxonomicName authority="Orchymont" authorityName="Orchymont" authorityYear="1940" box="[811,1000,1327,1346]" pageId="4" pageNumber="159" rank="species" species="wui">
<emphasis box="[811,842,1328,1346]" italics="true" pageId="4" pageNumber="159">wui</emphasis>
<bibRefCitation author="ORCHYMONT A. d'" box="[847,1000,1327,1346]" pageId="4" pageNumber="159" pagination="157 - 197" refId="ref7487" refString="ORCHYMONT A. d' 1940: Contribution a l'etude des Palpicornia XIV. Bulletin et Annales de la Societe Entomologique de Belgique 80: 157 - 197." type="journal article" year="1940">Orchymont, 1940</bibRefCitation>
</taxonomicName>
</taxonomicName>
; E
<taxonomicName authority="Jayaswal, 1972" authorityName="Jayaswal" authorityYear="1972" box="[1043,1306,1327,1347]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="bhutanicum">
<taxonomicName authority="" authorityName="" box="[1043,1169,1327,1347]" class="Centricae" family="Chironomidae" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="159" phylum="Diatomeae" rank="species" species="bhutanicum">
<emphasis box="[1043,1057,1328,1347]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<emphasis box="[1068,1169,1327,1346]" italics="true" pageId="4" pageNumber="159">bhutanicum</emphasis>
</taxonomicName>
<bibRefCitation author="JAYASWAL K. P." box="[1174,1306,1327,1346]" pageId="4" pageNumber="159" pagination="409 - 412" refId="ref6933" refString="JAYASWAL K. P. 1972: On the two new species of the hydrophilid beetles (Coleoptera: Hydrophilidae). Zoologischer Anzeiger 189: 409 - 412." type="journal article" year="1972">Jayaswal, 1972</bibRefCitation>
</taxonomicName>
; F
<taxonomicName authority="(Walker, 1858)" baseAuthorityName="Walker" baseAuthorityYear="1858" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="stultum">
<emphasis box="[1348,1362,1328,1347]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<emphasis box="[1373,1436,1327,1346]" italics="true" pageId="4" pageNumber="159">stultum</emphasis>
(
<bibRefCitation author="WALKER F." box="[157,273,1354,1373]" pageId="4" pageNumber="159" pagination="202 - 209" refId="ref7631" refString="WALKER F. 1858: Characters of some apparently undescribed Ceylon Insects. Annals and Magazine of Natural History, Series 3 2: 202 - 209." type="journal article" year="1858">Walker, 1858</bibRefCitation>
)
</taxonomicName>
, specimen from Japan: Iriomote Island; GH front view of head with labrum: G
<taxonomicName authority="Liu &amp; Hu &amp; Fikáček, 2020" authorityName="Liu &amp; Hu &amp; Fikáček" authorityYear="2020" box="[990,1110,1354,1374]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="taiwanense" status="sp. nov.">
<emphasis box="[990,1004,1355,1374]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<emphasis box="[1014,1110,1355,1373]" italics="true" pageId="4" pageNumber="159">taiwanense</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1114,1182,1354,1373]" pageId="4" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
, paratype; H
<taxonomicName authorityName="Orchymont" authorityYear="1936" box="[1307,1431,1354,1374]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="fallaciosum">
<emphasis box="[1307,1321,1355,1374]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<taxonomicName box="[1331,1431,1354,1373]" pageId="4" pageNumber="159" rank="species" species="fallaciosum">
<emphasis box="[1331,1431,1354,1373]" italics="true" pageId="4" pageNumber="159">fallaciosum</emphasis>
</taxonomicName>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="4.[151,761,1478,2077]" pageId="4" pageNumber="159">
<emphasis bold="true" box="[181,351,1478,1501]" italics="true" pageId="4" pageNumber="159">Male genitalia</emphasis>
(
<figureCitation box="[371,463,1478,1501]" captionStart="Fig" captionStartId="4.[151,182,1301,1320]" captionTargetBox="[178,1413,195,1286]" captionTargetId="figure@4.[169,1419,189,1286]" captionTargetPageId="4" captionText="Fig. 2. male genitalia and front view of the head of Taiwanese Coelostoma species.AF genitals in dorsal view:A C. fallaciosum Orchymont, 1936; B C. vitalisi Orchymont, 1923; C C. phallicum Orchymont, 1940; D C. wui Orchymont, 1940; E C. bhutanicum Jayaswal, 1972; F C. stultum (Walker, 1858), specimen from Japan: Iriomote Island; GH front view of head with labrum: G C. taiwanense sp. nov., paratype; H C. fallaciosum." figureDoi="http://doi.org/10.5281/zenodo.3879306" httpUri="https://zenodo.org/record/3879306/files/figure.png" pageId="4" pageNumber="159">Fig. 2C</figureCitation>
). Aedeagus
<quantity box="[605,693,1478,1501]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="159" unit="mm" value="1.5">1.5 mm</quantity>
long, apical part of the median lobe trilobed apically, much shorter than parameres; gonopore semicircular; apical third of the paramere bends inwards.
</paragraph>
<paragraph blockId="4.[151,761,1478,2077]" pageId="4" pageNumber="159">
<emphasis bold="true" box="[151,406,1606,1629]" pageId="4" pageNumber="159">Differential diagnosis.</emphasis>
<taxonomicName authorityName="Orchymont" authorityYear="1940" box="[411,656,1606,1629]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="phallicum">
<emphasis box="[411,656,1606,1629]" italics="true" pageId="4" pageNumber="159">Coelostoma phallicum</emphasis>
</taxonomicName>
is unique among Taiwanese
<taxonomicName box="[349,479,1638,1661]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="subGenus" subGenus="Coelostoma">
<emphasis box="[349,479,1638,1661]" italics="true" pageId="4" pageNumber="159">Coelostoma</emphasis>
</taxonomicName>
in having the median lobe trilobed apically.
</paragraph>
<paragraph blockId="4.[151,761,1478,2077]" pageId="4" pageNumber="159">
<emphasis bold="true" box="[151,273,1702,1725]" pageId="4" pageNumber="159">Bionomics</emphasis>
(
<figureCitation box="[292,380,1702,1725]" captionStart="Fig" captionStartId="5.[152,183,1195,1214]" captionTargetBox="[170,1418,189,1180]" captionTargetId="figure@5.[170,1418,189,1180]" captionTargetPageId="5" captionText="Fig. 3. Habitat of Coelostoma species.A Hsinchu County, Zhudong: habitat of C. stultum (Walker, 1858); B Hsinchu County, Hengshan Township: habitat of C. wui Orchymont, 1940; C Taichung City, Wufeng District:habitat of C. taiwanense sp.nov.and C. bhutanicum Jayaswal, 1972; D Hsinchu County, Beipu Township: habitat of C. fallaciosum Orchymont, 1936; E Yilan County, Jiaoxi Township: habitat of C. phallicum Orchymont, 1940; F Taipei city, Neihu District: habitat of C. vitalisi Orchymont, 1923. Photos by H.-C. Liu (AC, F), R. Matsushima (D) and F.-S. Hu (E)." figureDoi="http://doi.org/10.5281/zenodo.3879308" httpUri="https://zenodo.org/record/3879308/files/figure.png" pageId="4" pageNumber="159">Fig. 3E</figureCitation>
). The specimens of
<taxonomicName authorityName="Orchymont" authorityYear="1940" box="[614,760,1702,1726]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="phallicum">
<emphasis box="[614,632,1703,1726]" italics="true" pageId="4" pageNumber="159">C</emphasis>
.
<emphasis box="[650,760,1702,1725]" italics="true" pageId="4" pageNumber="159">phallicum</emphasis>
</taxonomicName>
from Yilan were found in mud with abundant
<emphasis box="[664,759,1736,1759]" italics="true" pageId="4" pageNumber="159">Brassica</emphasis>
plants and from accumulations of rotten leaves in a small, artificial brook draining the water from a hot spring hotel. They were found either submerged in water or on the border of the shore zone of the stream. The specimen from Hsinchu was collected from the surface of a wet stone near the stream. The beetles were observed active at night.
<emphasis bold="true" box="[151,299,1958,1981]" pageId="4" pageNumber="159">Distribution.</emphasis>
This species is known from
<collectingCountry box="[600,664,1958,1981]" name="China" pageId="4" pageNumber="159">China</collectingCountry>
(
<collectingRegion country="China" name="Guangdong" pageId="4" pageNumber="159">Guangdong</collectingRegion>
,
<collectingRegion box="[216,296,1990,2013]" country="China" name="Hainan" pageId="4" pageNumber="159">Hainan</collectingRegion>
,
<collectingRegion box="[305,401,1990,2013]" country="China" name="Guangxi" pageId="4" pageNumber="159">Guangxi</collectingRegion>
),
<collectingCountry box="[418,531,1990,2013]" name="Cambodia" pageId="4" pageNumber="159">Cambodia</collectingCountry>
,
<collectingCountry box="[539,646,1990,2013]" name="Indonesia" pageId="4" pageNumber="159">Indonesia</collectingCountry>
,
<collectingCountry box="[655,756,1990,2013]" name="Malaysia" pageId="4" pageNumber="159">Malaysia</collectingCountry>
,
<collectingCountry box="[151,204,2022,2045]" name="Laos" pageId="4" pageNumber="159">Laos</collectingCountry>
and
<collectingCountry box="[257,349,2022,2045]" name="Vietnam" pageId="4" pageNumber="159">Vietnam</collectingCountry>
(
<bibRefCitation author="ORCHYMONT A. d'" box="[365,559,2022,2045]" pageId="4" pageNumber="159" pagination="157 - 197" refId="ref7487" refString="ORCHYMONT A. d' 1940: Contribution a l'etude des Palpicornia XIV. Bulletin et Annales de la Societe Entomologique de Belgique 80: 157 - 197." type="journal article" year="1940">
<subScript attach="left" box="[385,494,2028,2044]" fontSize="7" pageId="4" pageNumber="159">ORCHYMONT</subScript>
1940
</bibRefCitation>
,
<bibRefCitation author="HANSEN M." box="[570,717,2022,2045]" pageId="4" pageNumber="159" refId="ref6868" refString="HANSEN M. 1999: World Catalogue of Insects 2: Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, 416 pp." type="book" year="1999">
<subScript attach="left" box="[590,654,2028,2044]" fontSize="7" pageId="4" pageNumber="159">HANSEN</subScript>
1999
</bibRefCitation>
,
<bibRefCitation author="JIA F. - L. &amp; ASTON P. &amp; FIKACEK M." pageId="4" pageNumber="159" pagination="354 - 376" refId="ref7132" refString="JIA F. - L., ASTON P. &amp; FIKACEK M. 2014: Review of the Chinese species of the genus Coelostoma Brulle, 1835 (Coleoptera: Hydrophilidae: Sphaeridiinae). Zootaxa 3887 (3): 354 - 376." type="journal article" year="2014">
<subScript attach="left" box="[740,760,2028,2044]" fontSize="7" pageId="4" pageNumber="159">JIA</subScript>
et al. 2014
</bibRefCitation>
). New record for
<collectingCountry box="[462,542,2054,2077]" name="Taiwan" pageId="4" pageNumber="159">Taiwan</collectingCountry>
(
<figureCitation box="[557,624,2054,2077]" captionStart="Fig" captionStartId="7.[151,182,970,989]" captionTargetBox="[154,757,192,945]" captionTargetId="figure@7.[148,762,186,964]" captionTargetPageId="7" captionText="Fig. 4. Map of Taiwan showing distribution of species of the Coelostoma species." figureDoi="http://doi.org/10.5281/zenodo.3879310" httpUri="https://zenodo.org/record/3879310/files/figure.png" pageId="4" pageNumber="159">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>