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<mods:title id="2CB8B121CB8062F101E838629CF9F779">Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<treatment id="B7275C4E7519A9577EE86C6A80ED05E4" ID-GBIF-Taxon="146754233" LSID="urn:lsid:plazi:treatment:B7275C4E7519A9577EE86C6A80ED05E4" httpUri="http://treatment.plazi.org/id/B7275C4E7519A9577EE86C6A80ED05E4" lastPageId="9" lastPageNumber="126" pageId="7" pageNumber="124">
<subSubSection id="0ECB1C6C53143B19D2624337D9165ECB" pageId="7" pageNumber="124" type="nomenclature">
<paragraph id="44C9C4E19AE886280B410F3381CF2A5B" pageId="7" pageNumber="124">
<taxonomicName id="5FC4F4F8BB335D3A9484A73346D57BF0" ID-CoL="4R2NN" authority="Luze, 1904" authorityName="Luze" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Microsaurus) fusicornis" order="Coleoptera" pageId="7" pageNumber="124" phylum="Arthropoda" rank="species" species="fusicornis" subGenus="Microsaurus">Quedius (Microsaurus) fusicornis Luze, 1904</taxonomicName>
Fig. 8
</paragraph>
</subSubSection>
<subSubSection id="114ED0BE0397732F3600ED498A94BCDD" pageId="8" pageNumber="125" type="reference_group">
<paragraph id="21200D89E1130D5914CD4847BD236668" pageId="8" pageNumber="125">
<taxonomicName id="CE217138BF10962F858E085F208DBBC5" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fusicornis" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="fusicornis">
<pageBreakToken id="A5CBB6E358B9EFE63984DEF63F5B824F" pageId="8" pageNumber="125" start="start">Quedius</pageBreakToken>
fusicornis
</taxonomicName>
Luze, 1904, 28 (original description);
<bibRefCitation id="CF0DD2FD65686A176A72C77FF569ED70" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli 1924</bibRefCitation>
, 69 (characters, notes)
</paragraph>
</subSubSection>
<subSubSection id="AA2CAB6B5BFD30104D13E369A3FBB30E" pageId="8" pageNumber="125" type="materials_examined">
<paragraph id="7643456BA366B4E4D10FAE07751F0AC4" pageId="8" pageNumber="125">Material examined.</paragraph>
<paragraph id="A8C23F0ED40E5E05521EEA3E53EA8BB0" pageId="8" pageNumber="125">
Type material: Tajikistan or Uzbekistan: Lectotype (here designated): ♂, &quot;♂/ Seravchan Putchin Pass. Glasunov 1892 [printed]/ Type fusicornis Luze [handwritten]/ ex. coll. Luze [printed]/ ex. coll. Scheerpeltz [printed]/ Typus Quedius fusicornis Luze [pre-printed]
<normalizedToken id="8A50C9CC8DA14F58DA712D4194834EBD" originalValue="">''</normalizedToken>
; Paralectotypes: 1 ♀, &quot;Seravchan Putchin Pass. Glasunov 1892 [printed]/ Type fusicornis Luze [handwritten]/ Quedius fusicornis Luze [handwritten]/ [square orange piece of paper]
<normalizedToken id="B3063FF02F777477E67F3A5E02BE2004" originalValue="">''</normalizedToken>
; 1 ♀, &quot;Seravchan Boschara Glasunow 1892 [printed]/ Type fusicornis Luze [handwritten]/ Quedius fusicornis Luze [pre-printed]
<normalizedToken id="F270F380647C8344A6538F2DE918DC6D" originalValue="">''</normalizedToken>
(Fig. 8F, G) (NMW).
</paragraph>
</subSubSection>
<subSubSection id="42E7B3061608F045B9F5A2CF42C9BE3D" pageId="8" pageNumber="125" type="materials_examined">
<paragraph id="E89DC346428D19F1245A927A6B5199DE" pageId="8" pageNumber="125">Additional material.</paragraph>
<paragraph id="C6124CED9A56F671026819DFFB7028D4" pageId="8" pageNumber="125">
Uzbekistan: 1 ♂, Samarqand Region, Aman Kutan, 04.VII.1932, V.V. Gussakovsky leg.; Kyrgyzstan: 1 ♂, Kyrgyz-Alatoo Mts, 09.VII.2010, 72°28'
<geoCoordinate id="46E4D2308CDEE75D1E7DF4E4880A0124" direction="north" orientation="latitude" precision="5555" value="38.6">38.6N</geoCoordinate>
, 42°48'
<geoCoordinate id="A98F6E6704DBAE51472654CBDE8A23A7" direction="east" orientation="longitude" precision="5555" value="49.2">49.2E</geoCoordinate>
, V.A. Kastcheev leg. (ZIN).
</paragraph>
</subSubSection>
<subSubSection id="51FE48DD74ACC75A9E155B9234C65040" pageId="8" pageNumber="125" type="type material">
<paragraph id="F50865E7E00E4DD22DCBAE68A5C3A517" pageId="8" pageNumber="125">Comments on taxonomy and type material.</paragraph>
<paragraph id="BA8BEA7D575B951725D5A62EF60087A2" pageId="8" pageNumber="125">
In the original description,
<bibRefCitation id="11B4E033719C6355BAA4C14779B1A0D3" pageId="8" pageNumber="125">Luze (1904)</bibRefCitation>
provided no information on the type material, but he indicated 7.7-8.5 mm body size range for the species. This suggests that he must have had more than one specimen to base a description on. He also indicated &quot;Seravschan: Putschin-Pass,
<normalizedToken id="87C30B4852CB46658CB23EA34F66E78B" originalValue="Boschara">Boschara''</normalizedToken>
as a locality that his material was from. Finally, we know from the introduction in
<normalizedToken id="C8B425334452094182DFC14CE09570C6" originalValue="Luzes">Luze's</normalizedToken>
paper that the material he examined was collected by Glasunov. Therefore, a single male (NMW) and two females (ZIN) that we examined and that match the original description morphologically and in the label data, are syntypes.
<bibRefCitation id="7EB99DB2624CF15AEF6F807149BC193B" pageId="8" pageNumber="125">Luze (1904)</bibRefCitation>
compared
<taxonomicName id="82E2577DDDF1BCCC0F76790F5AA442DF" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fusicornis" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="fusicornis">Quedius fusicornis</taxonomicName>
with his
<taxonomicName id="45D3C692615771EF9D7FAFBFFE8608DC" lsidName="Q. solskyi" pageId="8" pageNumber="125" rank="species" species="solskyi">Q. solskyi</taxonomicName>
and the widespread
<taxonomicName id="885DF133E1F9B0C5678FD3121DE2B9C5" lsidName="Q. cruentus" pageId="8" pageNumber="125" rank="species" species="cruentus">Q. cruentus</taxonomicName>
Ol.
<bibRefCitation id="6BD3CD1B5CE0984231F242243355CB58" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli (1924)</bibRefCitation>
apparently based his short notes about this species exclusively on
<normalizedToken id="C1272F0F6E8AD9184CEA156650315544" originalValue="Luzes">Luze's</normalizedToken>
description, without seeing any material. Similarly to other species of
<taxonomicName id="474AE1961D3826596E31E571FDE7F91C" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
described by
<bibRefCitation id="A31CAFE985D92CC0F7D38ADB6052E8DB" pageId="8" pageNumber="125">Luze (1904)</bibRefCitation>
,
<taxonomicName id="D826B67E873568C72303B97A633AE234" lsidName="Q. fusicornis" pageId="8" pageNumber="125" rank="species" species="fusicornis">Q. fusicornis</taxonomicName>
is missing in the monograph by
<bibRefCitation id="050379B14E2968D85D4F813DD77799D5" pageId="8" pageNumber="125">Coiffait (1978)</bibRefCitation>
who apparently overlooked
<normalizedToken id="BE844F12E0862FF99268E739227786CE" originalValue="Luzes">Luze's</normalizedToken>
publication. Here we provide a redescription and first illustrations of this poorly known species, including its aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="29EB999CDC0213D735A9D2A69496DC76" pageId="8" pageNumber="125" type="redescription">
<paragraph id="9253AB3547B2EAB77C14A6C8832B9D3D" pageId="8" pageNumber="125">Redescription.</paragraph>
<paragraph id="7060B5313E50F6AE98C9D5AD4B0F0CBC" pageId="8" pageNumber="125">Measurements and ratios (range, arithmetic mean; n = 6): HL: 1.0-1.2 (1.1); HW: 1.0-1.4 (1.1); PL: 1.1-1.5 (1.3); PW: 1.3-1.6 (1.4); EL: 1.5-1.7 (1.6); EW: 1.3-1.6 (1.5); FB: 3.7-4.4 (4.0); TL: 6.0-8.6 (7.3); HL/HW: 0.9-1.1 (1.0); PL/PW: 0.8-0.9 (0.9); EL/EW: 1.00-1.2 (1.1).</paragraph>
<paragraph id="789F76EFEF23728D2241F5FAF8480B67" pageId="8" pageNumber="125">Body length: 6.0-8.6 (7.3); head, scutellum and abdomen blackish, pronotum and hind margins of abdominal tergites slightly paler; elytra light red or orange; palpi, antennae and legs brown; body glossy (Fig. 8A, B).</paragraph>
<paragraph id="177B45CD20F49ED75E603A48252ABBCC" pageId="8" pageNumber="125">Head approximately as wide as long HL/HW: 0.9-1.1 (1.0); eyes small, not convex; temples slightly longer or as long as longitudinal diameter of eye; posterior frontal puncture closer to posterior margin of head than to anterior frontal puncture; temporal puncture closer to posterior margin of head than to posterior margin of eye; two vertical punctures behind posterior frontal puncture arranged as slightly oblique line between posterior margin of eye and dorsal part of neck; microsculpture of entire surface of head with transverse waves.</paragraph>
<paragraph id="426728C37941F44CAFD1913F2EDC3FBF" pageId="8" pageNumber="125">
Antennae moderately long, antennal segments: 3rd longer than 2nd, 4
<normalizedToken id="D63B141FF0CE0AD058952FD502FED44F" originalValue="th">th-</normalizedToken>
10th gradually widening towards apex of antenna.
</paragraph>
<paragraph id="159C4B3A36515DFE40774172D5426996" pageId="8" pageNumber="125">Pronotum slightly wider than long PL/PW: 0.8-0.9 (0.9), widest at about posterior third, gradually narrowing anteriad; hind angles rounded but distinct; dorsal and sublateral rows each with three punctures; microsculpture with transverse waves similar to that on posterior part of head. Scutellum impunctate with microsculpture as on pronotum. Elytra parallel-sided, slightly longer than wide, as long as or slightly longer than pronotum and narrower than maximum width of pronotum; punctation dense; setation gray; interspaces shiny, with distinct minute irregularities.</paragraph>
<paragraph id="AF8B2033DD3263F17A0C7A02E7486A61" pageId="8" pageNumber="125">Abdomen: punctation fine and moderately dense; interspaces with vaguely distinct minute irregularities; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="FAE3CEA89C976F301DE19108DC95C62F" pageId="8" pageNumber="125">
Male: protarsi with tarsomeres 1-4 dilated stronger than in females. Aedeagus (Fig. 8
<normalizedToken id="45C653CC1FAD494D53DEF40447BFC4E4" originalValue="CE">C-E</normalizedToken>
: median lobe parallel-sided along most of its length with broad and obtuse apex and tooth located near apex (Fig. 8C). Paramere parallel-sided, narrowing only in rhomboid apical portion; its apex almost reaching apex of median lobe, with two pairs of apical setae and two pairs of lateral setae below apex, with 6 peg setae arranged in two regular longitudinal rows apically extending basad over pairs of lateral setae (Fig. 8D).
</paragraph>
</subSubSection>
<subSubSection id="584BC035659C90760CBFE8BA7D9F174B" pageId="8" pageNumber="125" type="comparisons">
<paragraph id="58B06B04FB6C72510FF7B45FFB266989" pageId="8" pageNumber="125">Comparison.</paragraph>
<paragraph id="F948AFFF7ACD5FC7348415FD160C368F" pageId="8" pageNumber="125">
<taxonomicName id="122866441A8D888D89624DD4389C4C23" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fusicornis" order="Coleoptera" pageId="8" pageNumber="125" phylum="Arthropoda" rank="species" species="fusicornis">Quedius fusicornis</taxonomicName>
is similar to
<taxonomicName id="337D2D04CCB7FFCB76285C8B3F7B38F9" lsidName="Q. capitalis" pageId="8" pageNumber="125" rank="species" species="capitalis">Q. capitalis</taxonomicName>
. For comparison, see the latter species above. From other similar species such as
<taxonomicName id="2513427A3F910E395984FD09A037CF38" lsidName="Q. solskyi" pageId="8" pageNumber="125" rank="species" species="solskyi">Q. solskyi</taxonomicName>
,
<taxonomicName id="71EAEBE8C093BDCB56429091B0F59092" lsidName="Q. cruentus" pageId="8" pageNumber="125" rank="species" species="cruentus">Q. cruentus</taxonomicName>
and
<taxonomicName id="9BF92B4ABC9410B908BEAB61FF3BCA1A" lsidName="Q. ochripennis" pageId="8" pageNumber="125" rank="species" species="ochripennis">Q. ochripennis</taxonomicName>
, it can be easily distinguished by the structure of the apical part of the paramere with two medially situated short rows of peg setae (3 in each row) extending basad the pairs of lateral setae.
</paragraph>
</subSubSection>
<subSubSection id="36EE446A7770BDB3E6937EB0E5E5DDC0" lastPageId="9" lastPageNumber="126" pageId="8" pageNumber="125" type="distribution">
<paragraph id="86D72B96330A52F8B30C52247EC05141" pageId="8" pageNumber="125">Distribution.</paragraph>
<paragraph id="44E27332D6E28872DA3FC679A25595B3" lastPageId="9" lastPageNumber="126" pageId="8" pageNumber="125">
We were not able to locate the type locality &quot;Putchin Pass&quot; situated somewhere along Zeravchan River that is extended from eastern Uzbekistan to western Tajikistan. Additional material was studied from eastern Uzbekistan (near Aman-Kutan) and north-west
<pageBreakToken id="A3F374ABF0F024FEDBDFA5DA34EA7D34" pageId="9" pageNumber="126" start="start">ern</pageBreakToken>
Kyrgyzstan (Kyrgyz-Alatoo). Finally, one specimen was from
<normalizedToken id="8B18295D23248634ED8053521C2C8C42" originalValue="Tangi-Gharuh">'Tangi-Gharuh'</normalizedToken>
, a toponym in Afghanistan that we could not locate.
</paragraph>
<caption id="CF4513C61CAD4D7F80466FEE7E35F6E7" pageId="9" pageNumber="126">
<paragraph id="7FD5555C87513959C796AE9FB0741F8D" pageId="9" pageNumber="126">
Figure 8.
<taxonomicName id="0977706641F5417220FD49A495DBF92C" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fusicornis" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="fusicornis">Quedius fusicornis</taxonomicName>
, types. A, lectotype, male, habitus; B, paralectotype, female, habitus.
<normalizedToken id="1D14D410E16AA17C00BF76C5699EEEFD" originalValue="CE">C-E</normalizedToken>
,
<normalizedToken id="CD8E28605F4247E1755B2E53FD1E6FB1" originalValue="аedeagus">aedeagus</normalizedToken>
of the lectotype: C, lateral view; D, paramere, underside; E, median lobe, ventral view. F (lectotype), G, (paralectotype), labels. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="61FC78F3FCE9986283CACCB29828080C" pageId="9" pageNumber="126" type="bionomics">
<paragraph id="B832510D08D976B6147F1513008D3B72" pageId="9" pageNumber="126">Bionomics.</paragraph>
<paragraph id="3B70161799A61FB98B10E2F2EB0E4D71" pageId="9" pageNumber="126">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>