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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.86.75801" ID-GBIF-Dataset="a816417f-204c-4403-9407-01cc6152be6f" ID-PMC="PMC8760234" ID-Pensoft-Pub="1314-4049-86-1" ID-Pensoft-UUID="A65492DEA8E55FC9B9050FC172C2748B" ID-PubMed="35095303" ModsDocID="1314-4049-86-1" checkinTime="1641658256192" checkinUser="pensoft" docAuthor="Dissanayake, Lakmali S., Marasinghe, Diana S., Samarakoon, Milan C., Maharachchikumbura, Sajeewa S. N., Mortimer, Peter E., Hyde, Kevin D., Kuo, Chang-Hsin &amp; Kang, Ji-Chuan" docDate="2022" docId="B66F83E054D95E0F91BD1AA0A819845C" docLanguage="en" docName="MycoKeys 86: 1-17" docOrigin="MycoKeys 86" docPubDate="2022-01-07" docSource="http://dx.doi.org/10.3897/mycokeys.86.75801" docTitle="Iodosphaeria chiayiensis Marasinghe, C. H. Kuo &amp; K. D. Hyde, sp. nov." docType="treatment" docVersion="4" id="A65492DEA8E55FC9B9050FC172C2748B" lastPageNumber="1" masterDocId="A65492DEA8E55FC9B9050FC172C2748B" masterDocTitle="Three new species of Iodosphaeria (Xylariomycetidae): I. chiayiensis, I. jinghongensis and I. thailandica" masterLastPageNumber="17" masterPageNumber="1" pageNumber="1" updateTime="1668136780210" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new species of Iodosphaeria (Xylariomycetidae): I. chiayiensis, I. jinghongensis and I. thailandica</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dissanayake, Lakmali S.</mods:namePart>
<mods:affiliation>Engineering and Research Center for Southwest Bio-Pharmaceutical Resources of National Education Ministry of China, Guizhou University, Guiyang 550025, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marasinghe, Diana S.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Samarakoon, Milan C.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maharachchikumbura, Sajeewa S. N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9127-0783</mods:nameIdentifier>
<mods:affiliation>School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu 611731, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mortimer, Peter E.</mods:namePart>
<mods:affiliation>Centre for Mountain Futures (CMF), Kunming Institute of Botany, Honghe County 654400, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hyde, Kevin D.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Innovative Institute for Plant Health, Zhongkai University of Agriculture and Engineering, Guangzhou 510225, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kuo, Chang-Hsin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9011-6530</mods:nameIdentifier>
<mods:affiliation>Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kang, Ji-Chuan</mods:namePart>
<mods:affiliation>Engineering and Research Center for Southwest Bio-Pharmaceutical Resources of National Education Ministry of China, Guizhou University, Guiyang 550025, China</mods:affiliation>
<mods:nameIdentifier type="email">jckang@gzu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>86</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>17</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.86.75801</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.86.75801</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-86-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">A65492DEA8E55FC9B9050FC172C2748B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="191943043" LSID="urn:lsid:plazi:treatment:B66F83E054D95E0F91BD1AA0A819845C" httpUri="http://treatment.plazi.org/id/B66F83E054D95E0F91BD1AA0A819845C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="B66F83E0-54D9-5E0F-91BD-1AA0A819845C" authority="Marasinghe, C. H. Kuo &amp; K. D. Hyde, sp. nov." authorityName="Marasinghe, C. H. Kuo &amp; K. D. Hyde, sp. nov." class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria chiayiensis" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="chiayiensis">Iodosphaeria chiayiensis Marasinghe, C.H. Kuo &amp; K.D. Hyde, sp. nov.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Iodosphaeria chiayiensis (MFLU 21 - 0042, holotype) a substrate b ascomata on the host surface c section of ascoma d appearance of setae on peridium e peridium f, g asci h J + apical ring (in Melzer's reagent) i paraphyses j-m ascospores (m stained in lactophenol cotton blue). Scale bars: 50 μm (c, e); 5 μm (d); 20 μm (f-g); 5 μm (h, i); 10 μm (j-m)." figureDoi="10.3897/mycokeys.86.75801.figure2" httpUri="https://binary.pensoft.net/fig/631569" pageId="0" pageNumber="1">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria chiayiensis" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria chiayiensis</emphasis>
</taxonomicName>
refers to the city name where the fungus was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype">
<paragraph pageId="0" pageNumber="1">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="1">MFLU 21-0042.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Saprobic</emphasis>
on dead leaves of an unidentified host.
<emphasis bold="true" pageId="0" pageNumber="1">Sexual morph</emphasis>
:
<emphasis italics="true" pageId="0" pageNumber="1">Ascomata</emphasis>
150-190
<normalizedToken originalValue="×">x</normalizedToken>
160-200
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 170
<normalizedToken originalValue="×">x</normalizedToken>
180
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 10), globose to subglobose, superficial, black, solitary to gregarious, consisting of numerous long, flexuous setae.
<emphasis italics="true" pageId="0" pageNumber="1">Setae</emphasis>
3-5
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, arising from cells at the peridium surface, brown, unbranched, septate, apex flattened.
<emphasis italics="true" pageId="0" pageNumber="1">Ostiole</emphasis>
periphysate, apapillate.
<emphasis italics="true" pageId="0" pageNumber="1">Peridium</emphasis>
50-55
<normalizedToken originalValue="μm">μm</normalizedToken>
wide (x̅ = 53.4
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 10), comprises two layers of
<emphasis italics="true" pageId="0" pageNumber="1">textura angularis</emphasis>
cells, outer layer of dark brown to black thick-walled cells, and an inner layer of flattened, light brown.
<emphasis italics="true" pageId="0" pageNumber="1">Paraphyses</emphasis>
2-4
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, shorter than asci, hyaline, embedded in a gelatinous matrix.
<emphasis italics="true" pageId="0" pageNumber="1">Asci</emphasis>
60-90
<normalizedToken originalValue="×">x</normalizedToken>
8-10
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 72.9
<normalizedToken originalValue="×">x</normalizedToken>
9.2
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 30), 8-spored, unitunicate, cylindrical, shortly pedicellate, apex rounded, with a J+ apical ring.
<emphasis italics="true" pageId="0" pageNumber="1">Ascospores</emphasis>
15-20
<normalizedToken originalValue="×">x</normalizedToken>
4-6
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 17.2
<normalizedToken originalValue="×">x</normalizedToken>
5.2
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 30), overlapping uni-seriate, ellipsoidal to allantoid, aseptate, hyaline, guttulate.
<emphasis bold="true" pageId="0" pageNumber="1">Asexual morph</emphasis>
: Undetermined
</paragraph>
<caption doi="10.3897/mycokeys.86.75801.figure2" httpUri="https://binary.pensoft.net/fig/631569" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria chiayiensis" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria chiayiensis</emphasis>
</taxonomicName>
(MFLU 21-0042, holotype)
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
substrate
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
ascomata on the host surface
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
section of ascoma
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
appearance of setae on peridium
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
peridium
<emphasis bold="true" pageId="0" pageNumber="1">f, g</emphasis>
asci
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
J+ apical ring (in
<normalizedToken originalValue="Melzers">Melzer's</normalizedToken>
reagent)
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
paraphyses
<emphasis bold="true" pageId="0" pageNumber="1">j-m</emphasis>
ascospores (
<emphasis bold="true" pageId="0" pageNumber="1">m</emphasis>
stained in lactophenol cotton blue). Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">c, e</emphasis>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
); 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">f-g</emphasis>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">h, i</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">j-m</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2019-09-10" collectorName="D. S Marasinghe" country="Taiwan" location="Fanlu Township area" specimenCount="1" stateProvince="Chiayi" typeStatus="holotype">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
,
<collectingRegion country="Taiwan" name="Chiayi">Chiayi</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B66F83E054D95E0F91BD1AA0A819845C:F9C6B1995CBDEB2D6E1B2E672AC406A5" country="Taiwan" name="Fanlu Township area" stateProvince="Chiayi">Fanlu Township area</location>
, on dead leaves of an undetermined species,
<collectingDate value="2019-09-10">10 September 2019</collectingDate>
,
<collectorName>D.S Marasinghe</collectorName>
, DTF018 (MFLU 21-0042,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>holotype</typeStatus>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria chiayiensis" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria chiayiensis</emphasis>
</taxonomicName>
resembles
<taxonomicName lsidName="I. polygoni" pageId="0" pageNumber="1" rank="species" species="polygoni">
<emphasis italics="true" pageId="0" pageNumber="1">I. polygoni</emphasis>
</taxonomicName>
which has globose to sub globose, superficial, solitary to gregarious ascomata, cylindrical, short pedicellate asci with J+, apical rings and ellipsoidal to allantoid, aseptate, guttulate ascospores. However,
<taxonomicName lsidName="I. chiayiensis" pageId="0" pageNumber="1" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. chiayiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="I. polygoni" pageId="0" pageNumber="1" rank="species" species="polygoni">
<emphasis italics="true" pageId="0" pageNumber="1">I. polygoni</emphasis>
</taxonomicName>
in having smaller ascomata (150-190
<normalizedToken originalValue="×">x</normalizedToken>
160-200
<normalizedToken originalValue="μm">μm</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">vs.</emphasis>
270-475
<normalizedToken originalValue="×">x</normalizedToken>
250-500
<normalizedToken originalValue="μm">μm</normalizedToken>
) and shorter asci (60-90
<normalizedToken originalValue="×">x</normalizedToken>
8-10
<normalizedToken originalValue="μm">μm</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">vs.</emphasis>
150-180
<normalizedToken originalValue="×">x</normalizedToken>
10-13
<normalizedToken originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation DOI="https://doi.org/10.1017/S095375629600319X" author="Hsieh, WH" journalOrPublisher="Mycological Research" pageId="0" pageNumber="1" pagination="841 - 842" refId="B12" refString="Hsieh, WH, Chen, CY, Sivanesan, A, 1997. Iodosphaeria polygoni sp. nov. a new pyrenomycete from Taiwan. Mycological Research 101: 841 - 842, DOI: https://doi.org/10.1017/S095375629600319X" title="Iodosphaeria polygoni sp. nov. a new pyrenomycete from Taiwan." url="https://doi.org/10.1017/S095375629600319X" volume="101" year="1997">Hsieh et al. 1997</bibRefCitation>
). In the multi-gene phylogenetic analyses (Figure
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. RAxML tree based on a combined dataset of partial LSU and ITS sequence analyses. The tree is rooted to Delonicicola siamense (MFLUCC 15 - 0670) and Furfurella luteostiolata (CBS 143620). Type strains are in bold, and the newly generated strains are in red." figureDoi="10.3897/mycokeys.86.75801.figure1" httpUri="https://binary.pensoft.net/fig/631568" pageId="0" pageNumber="1">1</figureCitation>
), our collection (
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria chiayiensis" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria chiayiensis</emphasis>
</taxonomicName>
, MFLU 21-0042) has close affinity to
<taxonomicName lsidName="I. thailandica." pageId="0" pageNumber="1" rank="species" species="thailandica.">
<emphasis italics="true" pageId="0" pageNumber="1">I. thailandica.</emphasis>
</taxonomicName>
However, it was not possible to compare
<taxonomicName lsidName="I. chiayiensis" pageId="0" pageNumber="1" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. chiayiensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="I. jinghongensis" pageId="0" pageNumber="1" rank="species" species="jinghongensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. jinghongensis</emphasis>
</taxonomicName>
as they occur as different morphs.
</paragraph>
</subSubSection>
</treatment>
</document>