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<document id="D88346C97269F941F37EAA5642F8B626" ID-DOI="10.11646/zootaxa.4748.1.3" ID-GBIF-Dataset="f267c3f7-48d8-45f7-a480-b1660dc6b391" ID-ISSN="1175-5326" ID-Zenodo-Dep="3697578" ID-ZooBank="6F6367D1-5F1B-439D-B737-FEF761155D5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583398626205" checkinUser="plazi" docAuthor="Evans, Arthur V. &amp; Smith, Andrew B. T." docDate="2020" docId="B63C87C2F37BFFB52FA8FF67FE8DFC3D" docLanguage="en" docName="zootaxa.4748.1.3.pdf" docOrigin="Zootaxa 4748 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acoma nonglabrata Evans &amp; Smith 2020, new species" docType="treatment" docUuid="8EBF5164-E12F-4C2B-BE80-21ACF3429289" docUuidSource="ZooBank" docVersion="8" lastPageNumber="61" masterDocId="4A05FFBAF372FFBF2F3FFFFEFFD0FFF6" masterDocTitle="On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889" masterLastPageNumber="77" masterPageNumber="51" pageNumber="60" updateTime="1698724959701" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="38128B406626E96791BAAE4516FDDDD2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D25B9F5B1FF60EB0907514AA165CE0B3">On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889</mods:title>
</mods:titleInfo>
<mods:name id="62446546C179FF13B0AECA9E8D24B2FC" type="personal">
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<mods:namePart id="EF7B440CC0FED86D7D0B4969242FDFB5">Evans, Arthur V.</mods:namePart>
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<mods:namePart id="EB511791F08FE4042841E5DD31CA4207">Smith, Andrew B. T.</mods:namePart>
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<mods:title id="1460DFE3A6C89EDC46009310B6007932">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0B69B2DE4B581E4D6AB36FDD998824E5">
<mods:date id="9CDD919F801FE6F949269273F5BC42E1">2020</mods:date>
<mods:detail id="1F2F09FC9459C8F61569166F33D06EAD" type="pubDate">
<mods:number id="96BA1E85E85D51D7131E40A736CB698E">2020-03-05</mods:number>
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<mods:detail id="F69F3128F3F5C27E37892F533A4121C1" type="volume">
<mods:number id="E37D4272A2F1790F3228AA76942C5037">4748</mods:number>
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<mods:classification id="E47359CE96B54749A84D509962ED764D">journal article</mods:classification>
<mods:identifier id="A156BA0A31ED57F904E43A535B011AB4" type="DOI">10.11646/zootaxa.4748.1.3</mods:identifier>
<mods:identifier id="8B068F0CC93A201E4FC9FB68DFA28B28" type="GBIF-Dataset">f267c3f7-48d8-45f7-a480-b1660dc6b391</mods:identifier>
<mods:identifier id="C2545040EBCEE87B3C7E719428393044" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="56C77BA600593FA7075940CD992A1E5F" type="Zenodo-Dep">3697578</mods:identifier>
<mods:identifier id="3A2548A587CCDE89EBA627C025A21189" type="ZooBank">6F6367D1-5F1B-439D-B737-FEF761155D5C</mods:identifier>
</mods:mods>
<treatment id="B63C87C2F37BFFB52FA8FF67FE8DFC3D" ID-DOI="http://doi.org/10.5281/zenodo.3704226" ID-GBIF-Taxon="162443301" ID-Zenodo-Dep="3704226" LSID="urn:lsid:zoobank.org:act:8EBF5164-E12F-4C2B-BE80-21ACF3429289" httpUri="http://treatment.plazi.org/id/B63C87C2F37BFFB52FA8FF67FE8DFC3D" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
<subSubSection id="768F655FF37BFFB62FA8FF67FD3CFF45" box="[151,748,153,180]" pageId="9" pageNumber="60" type="nomenclature">
<paragraph id="3E2A36D4F37BFFB62FA8FF67FD3CFF45" blockId="9.[151,922,153,244]" box="[151,748,153,180]" pageId="9" pageNumber="60">
<heading id="656281B8F37BFFB62FA8FF67FD3CFF45" bold="true" box="[151,748,153,180]" fontSize="11" level="1" pageId="9" pageNumber="60" reason="1">
<emphasis id="0CE1EAC6F37BFFB62FA8FF67FD3CFF45" bold="true" box="[151,748,153,180]" pageId="9" pageNumber="60">
<taxonomicName id="F9954D57F37BFFB62FA8FF67FD80FF45" authority="Evans &amp; Smith" authorityName="Evans &amp; Smith" authorityYear="2020" box="[151,592,153,180]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="nonglabrata" status="sp. nov.">
<emphasis id="0CE1EAC6F37BFFB62FA8FF67FE57FF42" bold="true" box="[151,391,153,180]" italics="true" pageId="9" pageNumber="60">Acoma nonglabrata</emphasis>
Evans &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="17D257BDF37BFFB62D63FF64FD3CFF45" box="[604,748,153,180]" pageId="9" pageNumber="60" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF37BFFB62FA8FF43FC4AFF02" pageId="9" pageNumber="60" type="description">
<paragraph id="3E2A36D4F37BFFB62FA8FF43FECBFF21" blockId="9.[151,922,153,244]" box="[151,283,189,215]" pageId="9" pageNumber="60">
<figureCitation id="A6AE2A51F37BFFB62FA8FF43FECBFF21" box="[151,283,189,215]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1980,2005]" captionTargetBox="[151,1435,734,1953]" captionTargetId="figure@9.[151,1435,734,1954]" captionTargetPageId="9" captionText="FIGURES 17–22. Acoma nonglabrata Evans &amp; Smith, new species, male holotype. 17, Dorsal habitus; 18, oblique habitus; 19, ventral habitus; 20, head; 21, dorsal parameres; 22, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697586" httpUri="https://zenodo.org/record/3697586/files/figure.png" pageId="9" pageNumber="60">Figs. 17–22</figureCitation>
</paragraph>
<paragraph id="3E2A36D4F37BFFB62FA8FF22FC4AFF02" blockId="9.[151,922,153,244]" box="[151,922,220,244]" pageId="9" pageNumber="60">
<uri id="4A043AD6F37BFFB62FA8FF22FC4AFF02" box="[151,922,220,244]" pageId="9" pageNumber="60">
urn:lsid:zoobank.org:act:
<uuid id="4A330C01F37BFFB62EA5FF22FC4AFF02" box="[410,922,220,244]" pageId="9" pageNumber="60">8EBF5164-E12F-4C2B-BE80-21ACF3429289</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF37BFFB62FA8FEDDFB65FDAB" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="3E2A36D4F37BFFB62FA8FEDDFE7BFE5F" blockId="9.[151,1437,290,713]" pageId="9" pageNumber="60">
<emphasis id="0CE1EAC6F37BFFB62FA8FEDDFE91FECB" bold="true" box="[151,321,290,317]" pageId="9" pageNumber="60">Type material</emphasis>
.
<materialsCitation id="8EFD3C89F37BFFB62E6CFEDDFE7BFE5F" ID-GBIF-Occurrence="2573655802" collectingDate="1973-06-18" collectorName="K. B. Blair" country="Mexico" determinerName="W. B. Warner" location="SierraJuarez" pageId="9" pageNumber="60" specimenCount="2" typeStatus="holotype">
<typeStatus id="E12E8876F37BFFB62E6CFEDDFE6BFECB" box="[339,443,291,317]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
male (CASCTYPE 19748), labeled “
<collectingCountry id="46827644F37BFFB62C50FEDDFC35FECB" box="[879,997,291,317]" name="Mexico" pageId="9" pageNumber="60">MEXICO</collectingCountry>
: BajaCal. Norte / Guadalupe Canyon, E. / side of
<location id="3B4A600FF37BFFB62E1FFEB9FE7AFE97" LSID="urn:lsid:plazi:treatment:B63C87C2F37BFFB52FA8FF67FE8DFC3D:3B4A600FF37BFFB62E1FFEB9FE7AFE97" box="[288,426,327,353]" country="Mexico" name="SierraJuarez" pageId="9" pageNumber="60">SierraJuarez</location>
/
<date id="4A2B1014F37BFFB62EFBFEB9FD98FE97" box="[452,584,327,353]" pageId="9" pageNumber="60" value="1973-06-18">
<collectingDate id="5A6FE9FCF37BFFB62EFBFEB9FD98FE97" box="[452,584,327,353]" pageId="9" pageNumber="60" value="1973-06-18">18-VI-1973</collectingDate>
</date>
S.C. / Williams &amp;
<collectorName id="93605302F37BFFB62C16FEB9FC4FFE97" box="[809,927,327,353]" pageId="9" pageNumber="60">K.B. Blair</collectorName>
/ SCW # 311 (2) //
<taxonomicName id="F9954D57F37BFFB62BB7FEB9FB3FFE97" authorityName="Casey" authorityYear="1889" box="[1160,1263,327,353]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">ACOMA</taxonomicName>
/ OCHLERA / HOWDEN / OR N.SP. NEAR / det.
<determinerName id="CBF6DA10F37BFFB62D1CFE95FD60FE73" box="[547,688,363,389]" pageId="9" pageNumber="60">W.B.Warner</determinerName>
2015 // CASENT / 8424501 //
<typeStatus id="E12E8876F37BFFB62B34FE95FB4FFE73" box="[1035,1183,363,389]" pageId="9" pageNumber="60" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="F9954D57F37BFFB62B8DFE95FAD3FE73" authorityName="Casey" authorityYear="1889" box="[1202,1283,363,389]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">Acoma</taxonomicName>
/ nonglabrata / Evans &amp; Smith, 2019”.
</materialsCitation>
</paragraph>
<paragraph id="3E2A36D4F37BFFB62FF8FE4DFA53FDE3" blockId="9.[151,1437,290,713]" pageId="9" pageNumber="60">
<typeStatus id="E12E8876F37BFFB62FF8FE4DFEE1FE3B" box="[199,305,435,461]" pageId="9" pageNumber="60" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="2893FD5DF37BFFB62E7EFE4DFE4BFE38" box="[321,411,435,462]" pageId="9" pageNumber="60" type="male">7 males</specimenCount>
): 5,
<materialsCitation id="8EFD3C89F37BFFB62EF7FE4DFAAFFDE3" ID-GBIF-Occurrence="2573655807" collectingDate="1973-06-18" collectionCode="CASC, CMNC" collectorName="E. L. Sleeper" country="Mexico" determinerName="W. B. Warner" location="Bajia de Los" pageId="9" pageNumber="60" specimenCount="1" typeStatus="paratype">
same data as holotype, except no determination label,
<collectionCode id="5884AE11F37BFFB62B27FE4DFBB0FE3B" box="[1048,1120,435,461]" pageId="9" pageNumber="60">CASC</collectionCode>
database numbers CASENT 8424502–8424506; 2, “
<collectingCountry id="46827644F37BFFB62EA8FE29FE0AFE07" box="[407,474,471,497]" name="Mexico" pageId="9" pageNumber="60">MEX</collectingCountry>
. BajaCalif.Norte /
<quantity id="F96D9B31F37BFFB62D9BFE28FD39FE07" box="[676,745,470,497]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="60" unit="km" value="5.3">5.3km</quantity>
S
<location id="3B4A600FF37BFFB62C3BFE29FC5DFE07" LSID="urn:lsid:plazi:treatment:B63C87C2F37BFFB52FA8FF67FE8DFC3D:3B4A600FF37BFFB62C3BFE29FC5DFE07" box="[772,909,471,497]" country="Mexico" name="Bajia de Los" pageId="9" pageNumber="60">Bajia de Los</location>
/ Angeles VII.9–10.78 / 59029.6 /
<collectorName id="93605302F37BFFB62A3CFE29FA43FE07" box="[1283,1427,471,497]" pageId="9" pageNumber="60">E. L. Sleeper</collectorName>
” (
<collectionCode id="5884AE11F37BFFB62FA0FE05FF23FDE3" box="[159,243,507,533]" pageId="9" pageNumber="60">CMNC</collectionCode>
) and “BCN-
<quantity id="F96D9B31F37BFFB62E41FE04FE19FDE3" box="[382,457,506,533]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="60" unit="km" value="5.3">5.3 km</quantity>
S / BAJIA DE LOS / ANGELES / VII-9/10-1978 // 59029.6 /
<collectorName id="93605302F37BFFB62B48FE05FAD8FDE3" box="[1143,1288,507,533]" pageId="9" pageNumber="60">E. L. Sleeper</collectorName>
” (
<collectionCode id="5884AE11F37BFFB62A1FFE05FAA8FDE3" box="[1312,1400,507,533]" pageId="9" pageNumber="60">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3E2A36D4F37BFFB62FF8FDE1FB65FDAB" blockId="9.[151,1437,290,713]" pageId="9" pageNumber="60">
<materialsCitation id="8EFD3C89F37BFFB62FF8FDE1FB61FDAB" ID-GBIF-Occurrence="2573655803" collectionCode="AVEC, CASC, CMNC, UNAM" pageId="9" pageNumber="60" specimenCount="5" typeStatus="paratype">
<typeStatus id="E12E8876F37BFFB62FF8FDE1FEE4FDCF" box="[199,308,543,569]" pageId="9" pageNumber="60" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="5884AE11F37BFFB62EC7FDE1FD91FDCF" box="[504,577,543,569]" pageId="9" pageNumber="60">AVEC</collectionCode>
(
<specimenCount id="2893FD5DF37BFFB62D6BFDE1FD1DFDCF" box="[596,717,543,569]" pageId="9" pageNumber="60" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424502),
<collectionCode id="5884AE11F37BFFB62CF5FDE1FBC3FDCF" box="[970,1043,543,569]" pageId="9" pageNumber="60">CASC</collectionCode>
(
<specimenCount id="2893FD5DF37BFFB62B1BFDE1FB7BFDCF" box="[1060,1195,543,569]" pageId="9" pageNumber="60" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
; CASENT 8424505– 8424506),
<collectionCode id="5884AE11F37BFFB62E2EFDBDFEB3FDAB" box="[273,355,579,605]" pageId="9" pageNumber="60">CMNC</collectionCode>
(
<specimenCount id="2893FD5DF37BFFB62E4AFDBDFE3CFDAB" box="[373,492,579,605]" pageId="9" pageNumber="60" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424503),
<collectionCode id="5884AE11F37BFFB62DDBFDBDFCEAFDAB" box="[740,826,579,605]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="9" pageNumber="60">UNAM</collectionCode>
(
<specimenCount id="2893FD5DF37BFFB62C73FDBDFC13FDAB" box="[844,963,579,605]" pageId="9" pageNumber="60" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424504)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF37BFFB62FF8FD98FE7EFD3F" pageId="9" pageNumber="60" type="diagnosis">
<paragraph id="3E2A36D4F37BFFB62FF8FD98FE7EFD3F" blockId="9.[151,1437,290,713]" pageId="9" pageNumber="60">
<emphasis id="0CE1EAC6F37BFFB62FF8FD98FE92FD77" bold="true" box="[199,322,614,641]" pageId="9" pageNumber="60">Diagnosis.</emphasis>
Light reddish tan dorsally with head and pronotum darker; anterior clypeal margin emarginate medially; antennae composed of 10 antennomeres, antennomeres 8–10 lamellate; protibia distinctly tridentate; elytral disc with scattered setae.
</paragraph>
</subSubSection>
<caption id="6AEA665CF37BFFB62FA8F842FC98F80F" ID-DOI="http://doi.org/10.5281/zenodo.3697586" ID-Zenodo-Dep="3697586" httpUri="https://zenodo.org/record/3697586/files/figure.png" pageId="9" pageNumber="60" startId="9.[151,264,1980,2005]" targetBox="[151,1435,734,1953]" targetPageId="9">
<paragraph id="3E2A36D4F37BFFB62FA8F842FC98F80F" blockId="9.[151,1436,1980,2041]" pageId="9" pageNumber="60">
<emphasis id="0CE1EAC6F37BFFB62FA8F842FE9DF823" bold="true" box="[151,333,1980,2005]" pageId="9" pageNumber="60">FIGURES 17–22</emphasis>
.
<taxonomicName id="F9954D57F37BFFB62E67F843FD13F823" authority="Evans &amp; Smith" authorityName="Evans &amp; Smith" authorityYear="2020" box="[344,707,1980,2005]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="nonglabrata" status="sp. nov.">
<emphasis id="0CE1EAC6F37BFFB62E67F843FDF1F823" box="[344,545,1981,2005]" italics="true" pageId="9" pageNumber="60">Acoma nonglabrata</emphasis>
Evans &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="17D257BDF37BFFB62DF3F843FC98F823" box="[716,840,1980,2005]" pageId="9" pageNumber="60" rank="species">new species</taxonomicNameLabel>
, male holotype.
<emphasis id="0CE1EAC6F37BFFB62CD3F842FBD5F823" bold="true" box="[1004,1029,1980,2005]" pageId="9" pageNumber="60">17</emphasis>
, Dorsal habitus;
<emphasis id="0CE1EAC6F37BFFB62B8FF842FB19F823" bold="true" box="[1200,1225,1980,2005]" pageId="9" pageNumber="60">18</emphasis>
, oblique habitus;
<emphasis id="0CE1EAC6F37BFFB62A42F842FA46F823" bold="true" box="[1405,1430,1980,2005]" pageId="9" pageNumber="60">19</emphasis>
, ventral habitus;
<emphasis id="0CE1EAC6F37BFFB62E05F81EFE83F80F" bold="true" box="[314,339,2016,2041]" pageId="9" pageNumber="60">20</emphasis>
, head;
<emphasis id="0CE1EAC6F37BFFB62EA1F81EFE67F80F" bold="true" box="[414,439,2016,2041]" pageId="9" pageNumber="60">21</emphasis>
, dorsal parameres;
<emphasis id="0CE1EAC6F37BFFB62D42F81EFD46F80F" bold="true" box="[637,662,2016,2041]" pageId="9" pageNumber="60">22</emphasis>
, lateral genitalia.
</paragraph>
</caption>
<subSubSection id="768F655FF378FFB52FF8FF66FB17FCE1" pageId="10" pageNumber="61" type="description">
<paragraph id="3E2A36D4F378FFB52FF8FF66FC93FD39" blockId="10.[151,1437,152,971]" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FF66FE89FF45" bold="true" box="[199,345,152,179]" pageId="10" pageNumber="61">Description.</emphasis>
<typeStatus id="E12E8876F378FFB52E5FFF67FE18FF45" box="[352,456,153,179]" pageId="10" pageNumber="61" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="A6AE2A51F378FFB52D2AFF67FD4EFF45" box="[533,670,153,179]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1980,2005]" captionTargetBox="[151,1435,734,1953]" captionTargetId="figure@9.[151,1435,734,1954]" captionTargetPageId="9" captionText="FIGURES 17–22. Acoma nonglabrata Evans &amp; Smith, new species, male holotype. 17, Dorsal habitus; 18, oblique habitus; 19, ventral habitus; 20, head; 21, dorsal parameres; 22, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697586" httpUri="https://zenodo.org/record/3697586/files/figure.png" pageId="10" pageNumber="61">Figs. 17–22</figureCitation>
), length:
<quantity id="F96D9B31F378FFB52C38FF66FC8FFF45" box="[775,863,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="61" unit="mm" value="6.5">6.5 mm</quantity>
, width:
<quantity id="F96D9B31F378FFB52C85FF66FBC3FF45" box="[954,1043,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="61" unit="mm" value="3.5">3.5 mm</quantity>
. BODY testaceous, elongate, moderately convex. HEAD (
<figureCitation id="A6AE2A51F378FFB52E92FF43FDD1FF21" box="[429,513,189,215]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1980,2005]" captionTargetBox="[151,1435,734,1953]" captionTargetId="figure@9.[151,1435,734,1954]" captionTargetPageId="9" captionText="FIGURES 17–22. Acoma nonglabrata Evans &amp; Smith, new species, male holotype. 17, Dorsal habitus; 18, oblique habitus; 19, ventral habitus; 20, head; 21, dorsal parameres; 22, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697586" httpUri="https://zenodo.org/record/3697586/files/figure.png" pageId="10" pageNumber="61">Fig. 20</figureCitation>
) with impunctate vertex shiny and slightly depressed, frons and clypeus rugose, clypeus with large shallow punctures evident and separated by their own diameters; supraorbital carina obsolete at base of head and clypeus. Clypeus subtrapezoidal, lateral margins moderately reflexed, anterior margin broadly and shallowly concave with angles not prominent. Ocular canthus about one-quarter length of eye when viewed laterally, setose with setae longer than canthus. PRONOTUM slightly wider than long, broadest medially, convex, about one-third length of elytron; disc convex, glabrous, moderately punctate, most punctures separated by 1–2 diameters, with longitudinal impunctate area medially; lateral and posterior margins with narrow bead; anterior margin moderately bisinuate between obtuse anterior angles, with broad bead; anterolateral margin slightly sinuate and converging anteriorly, posterolateral margin straight and converging slightly posteriorly, with posterior angles broadly angulate. SCUTELLUM parabolic, about one-quarter width of elytron, disc mostly finely shagreened. ELYTRON convex, shiny, finely shagreened, disc punctostriate with punctures becoming confused on apical and lateral declivities, setigerous punctures with long suberect setae. LEGS with tridentate protibia moderately emarginate between teeth. PYGIDIUM parabolic, shiny, finely shagreened, with scattered setigerous punctures, setae long. GENITALIA with parameres symmetrical, slender and slightly sinuate when viewed laterally, with membranous flange extending to apical fourth, and not as long as phallobasis (
<figureCitation id="A6AE2A51F378FFB52D92FD4BFCE6FD39" box="[685,822,693,719]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1980,2005]" captionTargetBox="[151,1435,734,1953]" captionTargetId="figure@9.[151,1435,734,1954]" captionTargetPageId="9" captionText="FIGURES 17–22. Acoma nonglabrata Evans &amp; Smith, new species, male holotype. 17, Dorsal habitus; 18, oblique habitus; 19, ventral habitus; 20, head; 21, dorsal parameres; 22, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697586" httpUri="https://zenodo.org/record/3697586/files/figure.png" pageId="10" pageNumber="61">Figs. 21–22</figureCitation>
).
</paragraph>
<paragraph id="3E2A36D4F378FFB52FF8FD26FB17FCE1" blockId="10.[151,1437,152,971]" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FD26FEEFFD05" bold="true" box="[199,319,728,755]" pageId="10" pageNumber="61">Variation.</emphasis>
The
<typeStatus id="E12E8876F378FFB52E46FD27FE79FD05" box="[377,425,729,755]" pageId="10" pageNumber="61">type</typeStatus>
series is consistent in length, color, and surface sculpturing, but does exhibit minor variation in the degree of concavity of the anterior clypeal margin and prominence of clypeal angles.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB52FF8FCDFFEB8FCA9" pageId="10" pageNumber="61" type="etymology">
<paragraph id="3E2A36D4F378FFB52FF8FCDFFEB8FCA9" blockId="10.[151,1437,152,971]" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FCDFFE9EFCCD" bold="true" box="[199,334,801,827]" pageId="10" pageNumber="61">Etymology.</emphasis>
This specific epithet is in reference to the superficial resemblance to
<taxonomicName id="F9954D57F378FFB52B75FCDFFB18FCCD" authorityName="Cazier" authorityYear="1953" box="[1098,1224,801,827]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="glabrata">
<emphasis id="0CE1EAC6F378FFB52B75FCDFFB8BFCCD" box="[1098,1115,801,827]" italics="true" pageId="10" pageNumber="61">A</emphasis>
.
<emphasis id="0CE1EAC6F378FFB52B57FCDFFB18FCCD" box="[1128,1224,801,827]" italics="true" pageId="10" pageNumber="61">glabrata</emphasis>
</taxonomicName>
, which also occurs in
<collectingRegion id="FC51F836F378FFB52F8BFCBBFEB3FCA9" box="[180,355,837,863]" country="Mexico" name="Baja California" pageId="10" pageNumber="61">Baja California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB52FF8FC97FE8DFC3D" pageId="10" pageNumber="61" type="discussion">
<paragraph id="3E2A36D4F378FFB52FF8FC97FE8DFC3D" blockId="10.[151,1437,152,971]" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FC97FEEBFC75" bold="true" box="[199,315,873,899]" pageId="10" pageNumber="61">Remarks.</emphasis>
With 10 antennomeres and the club consisting of three antennomeres,
<taxonomicName id="F9954D57F378FFB52B60FC97FAD9FC75" authority="Evans &amp; Smith, 2020" authorityName="Evans &amp; Smith" authorityYear="2020" box="[1119,1289,873,899]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="nonglabrata" status="sp. nov.">
<emphasis id="0CE1EAC6F378FFB52B60FC97FBA0FC75" box="[1119,1136,873,899]" italics="true" pageId="10" pageNumber="61">A</emphasis>
.
<emphasis id="0CE1EAC6F378FFB52B40FC97FAD9FC75" box="[1151,1289,873,899]" italics="true" pageId="10" pageNumber="61">nonglabrata</emphasis>
</taxonomicName>
<emphasis id="0CE1EAC6F378FFB52A2DFC97FA4CFC75" bold="true" box="[1298,1436,872,899]" pageId="10" pageNumber="61">
<taxonomicNameLabel id="17D257BDF378FFB52A2DFC97FA4CFC75" box="[1298,1436,872,899]" pageId="10" pageNumber="61" rank="species">new species</taxonomicNameLabel>
</emphasis>
superficially resembles
<taxonomicName id="F9954D57F378FFB52E9EFC73FDF1FC51" authorityName="Cazier" authorityYear="1953" box="[417,545,909,935]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="glabrata">
<emphasis id="0CE1EAC6F378FFB52E9EFC73FE62FC51" box="[417,434,909,935]" italics="true" pageId="10" pageNumber="61">A</emphasis>
.
<emphasis id="0CE1EAC6F378FFB52EFEFC73FDF1FC51" box="[449,545,909,935]" italics="true" pageId="10" pageNumber="61">glabrata</emphasis>
</taxonomicName>
, but is readily distinguished by the setose elytral disc and more angulate lateral pronotal margins.
</paragraph>
</subSubSection>
</treatment>
</document>
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