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<document id="EFE8531431B6E38B1E50C6DC6FC90B48" ID-DOI="10.11646/zootaxa.4748.1.3" ID-GBIF-Dataset="f267c3f7-48d8-45f7-a480-b1660dc6b391" ID-ISSN="1175-5326" ID-Zenodo-Dep="3697578" ID-ZooBank="6F6367D1-5F1B-439D-B737-FEF761155D5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583398626205" checkinUser="plazi" docAuthor="Evans, Arthur V. &amp; Smith, Andrew B. T." docDate="2020" docId="B63C87C2F378FFB42FA8FBE3FAF6FDED" docLanguage="en" docName="zootaxa.4748.1.3.pdf" docOrigin="Zootaxa 4748 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acoma pararobusta Evans &amp; Smith 2020, new species" docType="treatment" docUuid="9342C833-0F57-4CCB-9819-C38902BAE444" docUuidSource="ZooBank" docVersion="8" lastPageNumber="62" masterDocId="4A05FFBAF372FFBF2F3FFFFEFFD0FFF6" masterDocTitle="On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889" masterLastPageNumber="77" masterPageNumber="51" pageNumber="61" updateTime="1698724959701" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="C47C15917FB40341E72D06BF585C6CA3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0D2733E9A67F31B30715F17E15A2B05D">On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889</mods:title>
</mods:titleInfo>
<mods:name id="E40CA2DCC2F86A2D7A3493F813F9FD09" type="personal">
<mods:role id="F99C7477D343C8077E5AA760C10E68CE">
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<mods:namePart id="F378B0212ADD7FC15CFEFC67E4BBCD63">Evans, Arthur V.</mods:namePart>
</mods:name>
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<mods:role id="1CCD419F6813784D34C228F8A8587CF4">
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</mods:role>
<mods:namePart id="982C68CB8610CA92DC193EA5D5964942">Smith, Andrew B. T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B85CE9FD4F0E4928CB9B2DD87CDC5470">text</mods:typeOfResource>
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<mods:title id="7AF0A48E3CE4D10C7BFB85BDA7EF048F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F2E861A73F353EEEED93012911BE2551">
<mods:date id="7854B373D673631526CA764385B1E4E4">2020</mods:date>
<mods:detail id="D44E25EFC1B3EF890E3FA553BB961004" type="pubDate">
<mods:number id="A4F5BAEDFA3955CA136E9F2C3EFB257D">2020-03-05</mods:number>
</mods:detail>
<mods:detail id="5CAF1A11A01A562C2F90AA94CA3BD533" type="volume">
<mods:number id="51DAC25D7FCE21F7B0F14332BE9CFA87">4748</mods:number>
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<mods:number id="0C1F0929DF3B63C8843E796C7B3AB5AC">1</mods:number>
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<mods:end id="81AA4D6901A51D8110876E477D84F188">77</mods:end>
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<mods:classification id="D3290023789ED41E93FDEDF7095690FB">journal article</mods:classification>
<mods:identifier id="F1E7DB7079F91A44DDFD861CFB3E34DC" type="DOI">10.11646/zootaxa.4748.1.3</mods:identifier>
<mods:identifier id="8E76F06AAF769A3B2EAEEFF50C5C5E9C" type="GBIF-Dataset">f267c3f7-48d8-45f7-a480-b1660dc6b391</mods:identifier>
<mods:identifier id="87A6EBE5EC7947ADCBB74AD6E2B09113" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E6D5CB3ED8D994A9F1D2B444A5FC2557" type="Zenodo-Dep">3697578</mods:identifier>
<mods:identifier id="1DCE09ACDF64A05404B229EE8A5ECA2A" type="ZooBank">6F6367D1-5F1B-439D-B737-FEF761155D5C</mods:identifier>
</mods:mods>
<treatment id="B63C87C2F378FFB42FA8FBE3FAF6FDED" ID-DOI="http://doi.org/10.5281/zenodo.3704224" ID-GBIF-Taxon="162443302" ID-Zenodo-Dep="3704224" LSID="urn:lsid:zoobank.org:act:9342C833-0F57-4CCB-9819-C38902BAE444" httpUri="http://treatment.plazi.org/id/B63C87C2F378FFB42FA8FBE3FAF6FDED" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61">
<subSubSection id="768F655FF378FFB52FA8FBE3FD3AFBC1" box="[151,746,1053,1080]" pageId="10" pageNumber="61" type="nomenclature">
<paragraph id="3E2A36D4F378FFB52FA8FBE3FD3AFBC1" blockId="10.[151,917,1053,1144]" box="[151,746,1053,1080]" pageId="10" pageNumber="61">
<heading id="656281B8F378FFB52FA8FBE3FD3AFBC1" bold="true" box="[151,746,1053,1080]" fontSize="11" level="1" pageId="10" pageNumber="61" reason="1">
<emphasis id="0CE1EAC6F378FFB52FA8FBE3FD3AFBC1" bold="true" box="[151,746,1053,1080]" pageId="10" pageNumber="61">
<taxonomicName id="F9954D57F378FFB52FA8FBE3FD9EFBC1" authority="Evans &amp; Smith" authorityName="Evans &amp; Smith" authorityYear="2020" box="[151,590,1053,1080]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="pararobusta" status="sp. nov.">
<emphasis id="0CE1EAC6F378FFB52FA8FBE3FE56FBCE" bold="true" box="[151,390,1053,1080]" italics="true" pageId="10" pageNumber="61">Acoma pararobusta</emphasis>
Evans &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="17D257BDF378FFB52D64FBE3FD3AFBC1" box="[603,746,1053,1079]" pageId="10" pageNumber="61" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB52FA8FBBFFC45FB8E" pageId="10" pageNumber="61" type="description">
<paragraph id="3E2A36D4F378FFB52FA8FBBFFECCFBAD" blockId="10.[151,917,1053,1144]" box="[151,284,1089,1115]" pageId="10" pageNumber="61">
<figureCitation id="A6AE2A51F378FFB52FA8FBBFFECCFBAD" box="[151,284,1089,1115]" captionStart="FIGURES 23–28" captionStartId="11.[151,264,1893,1918]" captionTargetBox="[151,1435,561,1867]" captionTargetId="figure@11.[151,1435,561,1869]" captionTargetPageId="11" captionText="FIGURES 23–28. Acoma pararobusta Evans &amp; Smith, new species, male holotype. 23, Dorsal habitus; 24, oblique habitus; 25, ventral habitus; 26, head; 27, dorsal parameres; 28, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697588" httpUri="https://zenodo.org/record/3697588/files/figure.png" pageId="10" pageNumber="61">Figs. 23–28</figureCitation>
</paragraph>
<paragraph id="3E2A36D4F378FFB52FA8FB9EFC45FB8E" blockId="10.[151,917,1053,1144]" box="[151,917,1120,1144]" pageId="10" pageNumber="61">
<uri id="4A043AD6F378FFB52FA8FB9EFC45FB8E" box="[151,917,1120,1144]" pageId="10" pageNumber="61">
urn:lsid:zoobank.org:act:
<uuid id="4A330C01F378FFB52EA5FB9EFC45FB8E" box="[410,917,1120,1144]" pageId="10" pageNumber="61">9342C833-0F57-4CCB-9819-C38902BAE444</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB52FA8FB58FA5BFA82" pageId="10" pageNumber="61" type="materials_examined">
<paragraph id="3E2A36D4F378FFB52FA8FB58FB95FAFE" blockId="10.[151,1437,1189,2045]" pageId="10" pageNumber="61">
<materialsCitation id="8EFD3C89F378FFB52FA8FB58FB95FAFE" ID-GBIF-Occurrence="2575869366" collectingDate="1971-07-29" collectorName="A. R. Hardy" country="Mexico" determinerName="W. B. Warner" location="At U. V. light / H. G. Real &amp; R. E. Main" municipality="Baja Calif. Sur / El Sargento" pageId="10" pageNumber="61" specimenCount="3" specimenCount-male="1" typeStatus="holotype">
<emphasis id="0CE1EAC6F378FFB52FA8FB58FEEDFB36" bold="true" box="[151,317,1189,1216]" pageId="10" pageNumber="61">Type material</emphasis>
.
<typeStatus id="E12E8876F378FFB52E75FB58FE62FB36" box="[330,434,1190,1216]" pageId="10" pageNumber="61" type="holotype">Holotype</typeStatus>
<specimenCount id="2893FD5DF378FFB52E87FB58FE3EFB36" box="[440,494,1190,1216]" pageId="10" pageNumber="61" type="male">male</specimenCount>
(CASTYPE 19749), labeled “
<collectingCountry id="46827644F378FFB52C02FB58FC55FB36" box="[829,901,1190,1216]" name="Mexico" pageId="10" pageNumber="61">MEX</collectingCountry>
.:
<collectingMunicipality id="DE4EACAEF378FFB52CADFB58FB1FFB36" box="[914,1231,1190,1216]" pageId="10" pageNumber="61">Baja Calif. Sur / El Sargento</collectingMunicipality>
,
<date id="4A2B1014F378FFB52BE6FB58FAA8FB36" box="[1241,1400,1190,1216]" pageId="10" pageNumber="61" value="1971-07-29">
<collectingDate id="5A6FE9FCF378FFB52BE6FB58FAA8FB36" box="[1241,1400,1190,1216]" pageId="10" pageNumber="61" value="1971-07-29">29.VII. / 1971</collectingDate>
</date>
.
<location id="3B4A600FF378FFB52ABFFB58FDCFFB12" LSID="urn:lsid:plazi:treatment:B63C87C2F378FFB42FA8FBE3FAF6FDED:3B4A600FF378FFB52ABFFB58FDCFFB12" country="Mexico" municipality="Baja Calif. Sur / El Sargento" name="At U. V. light / H. G. Real &amp; R. E. Main" pageId="10" pageNumber="61">At U.V. light / H.G. Real &amp; R.E. Main</location>
//
<emphasis id="0CE1EAC6F378FFB52D03FB34FDABFB12" bold="true" box="[572,635,1226,1252]" pageId="10" pageNumber="61">
n.
<specimenCount id="2893FD5DF378FFB52D66FB34FDABFB12" box="[601,635,1226,1252]" pageId="10" pageNumber="61" type="generic">sp.</specimenCount>
</emphasis>
? / DET. /
<collectorName id="93605302F378FFB52DD4FB34FCBCFB12" box="[747,876,1226,1252]" pageId="10" pageNumber="61">A.R. Hardy</collectorName>
1979 //
<taxonomicName id="F9954D57F378FFB52CF9FB34FBC7FB12" authorityName="Casey" authorityYear="1889" box="[966,1047,1226,1252]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">
<location id="3B4A600FF378FFB52CF9FB34FBC7FB12" LSID="urn:lsid:plazi:treatment:B63C87C2F378FFB42FA8FBE3FAF6FDED:3B4A600FF378FFB52CF9FB34FBC7FB12" box="[966,1047,1226,1252]" country="Mexico" municipality="Baja Calif. Sur / El Sargento" name="Acoma" pageId="10" pageNumber="61">Acoma</location>
</taxonomicName>
/
<emphasis id="0CE1EAC6F378FFB52B13FB34FBBAFB12" bold="true" box="[1068,1130,1226,1252]" pageId="10" pageNumber="61">
n.
<specimenCount id="2893FD5DF378FFB52B76FB34FBBAFB12" box="[1097,1130,1226,1252]" pageId="10" pageNumber="61" type="generic">sp.</specimenCount>
</emphasis>
3 / det.
<determinerName id="CBF6DA10F378FFB52B84FB34FA98FB12" box="[1211,1352,1226,1252]" pageId="10" pageNumber="61">W.B.Warner</determinerName>
2015 // CASENT / 8424521 // HOLOTYPE /
<taxonomicName id="F9954D57F378FFB52D7EFB10FD42FAFE" authorityName="Casey" authorityYear="1889" box="[577,658,1262,1288]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="genus">
<location id="3B4A600FF378FFB52D7EFB10FD42FAFE" LSID="urn:lsid:plazi:treatment:B63C87C2F378FFB42FA8FBE3FAF6FDED:3B4A600FF378FFB52D7EFB10FD42FAFE" box="[577,658,1262,1288]" country="Mexico" municipality="Baja Calif. Sur / El Sargento" name="Acoma" pageId="10" pageNumber="61">Acoma</location>
</taxonomicName>
/ pararobusta /
<location id="3B4A600FF378FFB52C7FFB11FC3EFAFE" LSID="urn:lsid:plazi:treatment:B63C87C2F378FFB42FA8FBE3FAF6FDED:3B4A600FF378FFB52C7FFB11FC3EFAFE" box="[832,1006,1262,1289]" country="Mexico" municipality="Baja Calif. Sur / El Sargento" name="Evans &amp; Smith" pageId="10" pageNumber="61">Evans &amp; Smith</location>
, 2019”.
</materialsCitation>
</paragraph>
<paragraph id="3E2A36D4F378FFB52FF8FAEDFEA1FAA6" blockId="10.[151,1437,1189,2045]" pageId="10" pageNumber="61">
<materialsCitation id="8EFD3C89F378FFB52FF8FAEDFEBDFAA6" ID-GBIF-Occurrence="2575869367" collectionCode="CASC" pageId="10" pageNumber="61" specimenCount="1" typeStatus="paratype">
<typeStatus id="E12E8876F378FFB52FF8FAEDFEE7FADB" box="[199,311,1299,1325]" pageId="10" pageNumber="61" type="paratype">Paratypes</typeStatus>
, (
<specimenCount id="2893FD5DF378FFB52E76FAECFE75FADB" box="[329,421,1298,1325]" pageId="10" pageNumber="61" type="male">2 males</specimenCount>
): same data as holotype, except no determination labels,
<collectionCode id="5884AE11F378FFB52B2AFAECFB8EFADA" box="[1045,1118,1298,1324]" pageId="10" pageNumber="61">CASC</collectionCode>
database numbers CASENT 8424522–8424523
</materialsCitation>
.
</paragraph>
<paragraph id="3E2A36D4F378FFB52FF8FAA5FA5BFA82" blockId="10.[151,1437,1189,2045]" box="[199,1419,1369,1397]" pageId="10" pageNumber="61">
<materialsCitation id="8EFD3C89F378FFB52FF8FAA5FA57FA82" ID-GBIF-Occurrence="2573655805" box="[199,1415,1369,1397]" collectionCode="CASC, CMNC" pageId="10" pageNumber="61" specimenCount="2" typeStatus="paratype">
<typeStatus id="E12E8876F378FFB52FF8FAA5FEE4FA83" box="[199,308,1371,1397]" pageId="10" pageNumber="61" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="5884AE11F378FFB52ECCFAA4FDECFA82" box="[499,572,1370,1396]" pageId="10" pageNumber="61">CASC</collectionCode>
(
<specimenCount id="2893FD5DF378FFB52D70FAA4FD16FA83" box="[591,710,1370,1397]" pageId="10" pageNumber="61" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424523),
<collectionCode id="5884AE11F378FFB52C81FAA4FBC0FA82" box="[958,1040,1370,1396]" pageId="10" pageNumber="61">CMNC</collectionCode>
(
<specimenCount id="2893FD5DF378FFB52B1DFAA4FB49FA83" box="[1058,1177,1370,1397]" pageId="10" pageNumber="61" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424522)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB52FF8FA83FC21FA17" pageId="10" pageNumber="61" type="diagnosis">
<paragraph id="3E2A36D4F378FFB52FF8FA83FC21FA17" blockId="10.[151,1437,1189,2045]" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FA83FE92FA6E" bold="true" box="[199,322,1405,1432]" pageId="10" pageNumber="61">Diagnosis.</emphasis>
Rufopiceous or rufobrunneus dorsally; anterior clypeal margin shallowly emarginate or rounded medially; frontoclypeal suture tumid; antennae composed of 9 antennomeres, antennomeres 5–9 lamellate, subequal in length; protibia distinctly tridentate; elytral disc with scattered, pale setae.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF378FFB42FF8FA17FCD6FE91" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61" type="description">
<paragraph id="3E2A36D4F378FFB42FF8FA17FDC2FEE9" blockId="10.[151,1437,1189,2045]" lastBlockId="11.[151,1437,153,539]" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61">
<emphasis id="0CE1EAC6F378FFB52FF8FA17FE89F9F2" bold="true" box="[199,345,1513,1540]" pageId="10" pageNumber="61">Description.</emphasis>
<typeStatus id="E12E8876F378FFB52E5EFA14FE19F9F2" box="[353,457,1514,1540]" pageId="10" pageNumber="61" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="A6AE2A51F378FFB52D27FA14FD72F9F2" box="[536,674,1514,1540]" captionStart="FIGURES 23–28" captionStartId="11.[151,264,1893,1918]" captionTargetBox="[151,1435,561,1867]" captionTargetId="figure@11.[151,1435,561,1869]" captionTargetPageId="11" captionText="FIGURES 23–28. Acoma pararobusta Evans &amp; Smith, new species, male holotype. 23, Dorsal habitus; 24, oblique habitus; 25, ventral habitus; 26, head; 27, dorsal parameres; 28, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697588" httpUri="https://zenodo.org/record/3697588/files/figure.png" pageId="10" pageNumber="61">Figs. 23–28</figureCitation>
), length: 9.0 mm, width: 4.0 mm. BODY dark brown, elongate, robust, moderately convex. HEAD (
<figureCitation id="A6AE2A51F378FFB52D2BF9F0FDB6F9DF" box="[532,614,1550,1577]" captionStart="FIGURES 23–28" captionStartId="11.[151,264,1893,1918]" captionTargetBox="[151,1435,561,1867]" captionTargetId="figure@11.[151,1435,561,1869]" captionTargetPageId="11" captionText="FIGURES 23–28. Acoma pararobusta Evans &amp; Smith, new species, male holotype. 23, Dorsal habitus; 24, oblique habitus; 25, ventral habitus; 26, head; 27, dorsal parameres; 28, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697588" httpUri="https://zenodo.org/record/3697588/files/figure.png" pageId="10" pageNumber="61">Fig. 26</figureCitation>
) with narrowly raised transverse impunctate area, frons and base of clypeus deeply and rugosely punctate, reflexed clypeal margins smoother, with deep punctures mostly separated by their own diameters; short and irregular supraorbital carina obsolete at bases of head and clypeus. Clypeus somewhat semicircular, lateral margins narrowly reflexed, apical margin more broadly reflexed and slightly concave. Ocular canthus thick and broadly rounded apically, about one-third length of eye when viewed laterally, densely setose with setae somewhat longer than canthus. PRONOTUM about half length of elytron, as long as wide; disc glabrous with moderately dense punctures usually separated by twice their diameters sometimes briefly confluent on either side of narrow longitudinal impunctate area medially, punctures become denser laterally, even more so anteriorly; anterolateral margins sinuate, abruptly converging anteriorly from middle, posterolateral margins subparallel, shallowly concave medially; lateral and posterior margins with narrow bead, lateral margins distinctly setose with setae long; anterior margin mostly concave between acute and produced anterior angles, slightly protruding medially, with broad bead; posterior angles broadly obtuse. SCUTELLUM broadly elliptical, basal half sparsely and setigerously punctate, setae coarse. ELYTRON shiny and setose, with deep, narrow striae on disc with setigerous punctures bearing recumbent setae becoming short and irregular on declivities. LEGS with tridentate protibia deeply emarginate between apical and middle teeth, shallowly emarginate between middle and basal teeth. PYGIDIUM broadly parabolic, rugosely punctate along sides, and setose with setae long, erect. GENITALIA with parameres symmetrical, slender and slightly sinuate when viewed laterally, with membranous flange extending to apical fourth, and not as long as phallobasis (
<figureCitation id="A6AE2A51F379FFB42E43FEFBFDD5FEE9" box="[380,517,261,287]" captionStart="FIGURES 23–28" captionStartId="11.[151,264,1893,1918]" captionTargetBox="[151,1435,561,1867]" captionTargetId="figure@11.[151,1435,561,1869]" captionTargetPageId="11" captionText="FIGURES 23–28. Acoma pararobusta Evans &amp; Smith, new species, male holotype. 23, Dorsal habitus; 24, oblique habitus; 25, ventral habitus; 26, head; 27, dorsal parameres; 28, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697588" httpUri="https://zenodo.org/record/3697588/files/figure.png" pageId="11" pageNumber="62">Figs. 27–28</figureCitation>
).
</paragraph>
<paragraph id="3E2A36D4F379FFB42FF8FED6FCD6FE91" blockId="11.[151,1437,153,539]" pageId="11" pageNumber="62">
<emphasis id="0CE1EAC6F379FFB42FF8FED6FEEFFEB5" bold="true" box="[199,319,296,323]" pageId="11" pageNumber="62">Variation.</emphasis>
Length: 7.5–9.0 mm, width: 3.2–4.0 mm. Color varies from light to dark brown. Anterior clypeal margin ranges from slightly concave to nearly rounded.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF379FFB42FF8FE8FFB06FE7D" box="[199,1238,369,395]" pageId="11" pageNumber="62" type="etymology">
<paragraph id="3E2A36D4F379FFB42FF8FE8FFB06FE7D" blockId="11.[151,1437,153,539]" box="[199,1238,369,395]" pageId="11" pageNumber="62">
<emphasis id="0CE1EAC6F379FFB42FF8FE8FFE9EFE7D" bold="true" box="[199,334,369,395]" pageId="11" pageNumber="62">Etymology.</emphasis>
This is named in recognition of the similarities with
<taxonomicName id="F9954D57F379FFB42CA0FE8FFB02FE7D" authority="Van Dyke, 1928" authorityName="Van Dyke" authorityYear="1928" box="[927,1234,369,395]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="0CE1EAC6F379FFB42CA0FE8FFC60FE7D" box="[927,944,369,395]" italics="true" pageId="11" pageNumber="62">A</emphasis>
.
<emphasis id="0CE1EAC6F379FFB42C81FE8FFBC3FE7D" box="[958,1043,369,395]" italics="true" pageId="11" pageNumber="62">robusta</emphasis>
<bibRefCitation id="5A044B25F379FFB42B26FE8FFB02FE7D" author="Van Dyke, E. C." box="[1049,1234,369,395]" pageId="11" pageNumber="62" pagination="151 - 162" refId="ref15726" refString="Van Dyke, E. C. (1928) Notes and descriptions of new species of Scarabaeidae from western North America. The Pan-Pacific Entomologist, 4, 151 - 162." type="journal article" year="1928">Van Dyke, 1928</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF379FFB42FF8FE6BFAF6FDED" pageId="11" pageNumber="62" type="discussion">
<paragraph id="3E2A36D4F379FFB42FF8FE6BFAF6FDED" blockId="11.[151,1437,153,539]" pageId="11" pageNumber="62">
<emphasis id="0CE1EAC6F379FFB42FF8FE6BFEEBFE59" bold="true" box="[199,315,405,431]" pageId="11" pageNumber="62">Remarks.</emphasis>
<taxonomicName id="F9954D57F379FFB42E7EFE6BFDCDFE59" authority="Evans &amp; Smith, 2020" authorityName="Evans &amp; Smith" authorityYear="2020" box="[321,541,405,431]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="pararobusta" status="sp. nov.">
<emphasis id="0CE1EAC6F379FFB42E7EFE6BFDCDFE59" box="[321,541,405,431]" italics="true" pageId="11" pageNumber="62">Acoma pararobusta</emphasis>
</taxonomicName>
<emphasis id="0CE1EAC6F379FFB42D1CFE6BFD7BFE59" bold="true" box="[547,683,404,431]" pageId="11" pageNumber="62">
<taxonomicNameLabel id="17D257BDF379FFB42D1CFE6BFD7BFE59" box="[547,683,404,431]" pageId="11" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
is distinguished from all other species in the genus by the characters given in the diagnosis. In addition to the tumid frontoclypeal suture, it is distinguished from
<taxonomicName id="F9954D57F379FFB42BB3FE47FB2EFE25" authorityName="Van Dyke" authorityYear="1928" box="[1164,1278,441,467]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="0CE1EAC6F379FFB42BB3FE47FB4DFE25" box="[1164,1181,441,467]" italics="true" pageId="11" pageNumber="62">A</emphasis>
.
<emphasis id="0CE1EAC6F379FFB42B94FE47FB2EFE25" box="[1195,1278,441,467]" italics="true" pageId="11" pageNumber="62">robusta</emphasis>
</taxonomicName>
by the coarse, rugose sculpturing on the frons and base of the clypeus, occasional confluent punctures on pronotal disc, and the somewhat semicircular clypeal margin with bases that are more or less continuous with the supraorbital carinae.
</paragraph>
</subSubSection>
</treatment>
</document>
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