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<document id="E842AB4CAD8DE455E6D8A45E55E98A92" ID-DOI="10.11646/zootaxa.4748.1.3" ID-GBIF-Dataset="f267c3f7-48d8-45f7-a480-b1660dc6b391" ID-ISSN="1175-5326" ID-Zenodo-Dep="3697578" ID-ZooBank="6F6367D1-5F1B-439D-B737-FEF761155D5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583398626205" checkinUser="plazi" docAuthor="Evans, Arthur V. &amp; Smith, Andrew B. T." docDate="2020" docId="B63C87C2F377FFB82FA8F954FB5CFC3D" docLanguage="en" docName="zootaxa.4748.1.3.pdf" docOrigin="Zootaxa 4748 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acoma chihuahuaensis Evans &amp; Smith 2020, new species" docType="treatment" docUuid="BD4FB72A-5BD1-4671-886A-C0E05D1BE391" docUuidSource="ZooBank" docVersion="8" lastPageNumber="58" masterDocId="4A05FFBAF372FFBF2F3FFFFEFFD0FFF6" masterDocTitle="On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889" masterLastPageNumber="77" masterPageNumber="51" pageNumber="56" updateTime="1698724959701" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9E764D4F567EE871DBEBE9EA7E7EF46F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="834E82A08072D0FC1ACAD06AA0BC3892">On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889</mods:title>
</mods:titleInfo>
<mods:name id="7BE986BF829F74540CDF2338AC1C2015" type="personal">
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<mods:namePart id="1F232DD4667139AA8663A144C22DC375">Evans, Arthur V.</mods:namePart>
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<mods:namePart id="2B0E60E0CA387AF29879FEA5BBBFFC1B">Smith, Andrew B. T.</mods:namePart>
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<mods:title id="7F0FDFD7C924F9219C3B7A9687D7962B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5530438E8166C9DDA9F9328F7F890622">
<mods:date id="5DBD9D0814BA4C5EA6336C05E50F0836">2020</mods:date>
<mods:detail id="05217FCC9E5042F8EA948A933451EF09" type="pubDate">
<mods:number id="023FBDF70EB8CB030A9B7A691D5B56B4">2020-03-05</mods:number>
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<mods:number id="F3884E485CDBD543CB5958896AB9C414">4748</mods:number>
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<mods:classification id="DDFAB6D218EF0EDAD3999A872EFDAA7B">journal article</mods:classification>
<mods:identifier id="485D7B678FAD01D30B34686A4FAABFB1" type="DOI">10.11646/zootaxa.4748.1.3</mods:identifier>
<mods:identifier id="A19A5AF9A707A2DDA9E7D29733AF966D" type="GBIF-Dataset">f267c3f7-48d8-45f7-a480-b1660dc6b391</mods:identifier>
<mods:identifier id="49EB0A1B894DE7039929571BE0874C20" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E59EC4D90E52423A3811C3FE812B2854" type="Zenodo-Dep">3697578</mods:identifier>
<mods:identifier id="0C9E6A06995B359A3506ACD3253FAA4E" type="ZooBank">6F6367D1-5F1B-439D-B737-FEF761155D5C</mods:identifier>
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<treatment id="B63C87C2F377FFB82FA8F954FB5CFC3D" ID-DOI="http://doi.org/10.5281/zenodo.3704220" ID-GBIF-Taxon="162443303" ID-Zenodo-Dep="3704220" LSID="urn:lsid:zoobank.org:act:BD4FB72A-5BD1-4671-886A-C0E05D1BE391" httpUri="http://treatment.plazi.org/id/B63C87C2F377FFB82FA8F954FB5CFC3D" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56">
<subSubSection id="768F655FF377FFBA2FA8F954FCC8F932" box="[151,792,1706,1733]" pageId="5" pageNumber="56" type="nomenclature">
<paragraph id="3E2A36D4F377FFBA2FA8F954FCC8F932" blockId="5.[151,940,1706,1797]" box="[151,792,1706,1733]" pageId="5" pageNumber="56">
<heading id="656281B8F377FFBA2FA8F954FCC8F932" bold="true" box="[151,792,1706,1733]" fontSize="11" level="1" pageId="5" pageNumber="56" reason="1">
<emphasis id="0CE1EAC6F377FFBA2FA8F954FCC8F932" bold="true" box="[151,792,1706,1733]" pageId="5" pageNumber="56">
<taxonomicName id="F9954D57F377FFBA2FA8F954FDACF932" authority="Evans &amp; Smith" authorityName="Evans &amp; Smith" authorityYear="2020" box="[151,636,1706,1733]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="chihuahuaensis" status="sp. nov.">
<emphasis id="0CE1EAC6F377FFBA2FA8F954FE63F933" bold="true" box="[151,435,1706,1733]" italics="true" pageId="5" pageNumber="56">Acoma chihuahuaensis</emphasis>
Evans &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="17D257BDF377FFBA2DB7F955FCC8F932" box="[648,792,1706,1733]" pageId="5" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF377FFBA2FA8F930FC7CF8F3" pageId="5" pageNumber="56" type="description">
<paragraph id="3E2A36D4F377FFBA2FA8F930FEDDF91E" blockId="5.[151,940,1706,1797]" box="[151,269,1741,1768]" pageId="5" pageNumber="56">
<figureCitation id="A6AE2A51F377FFBA2FA8F930FEDDF91E" box="[151,269,1741,1768]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1817,1842]" captionTargetBox="[151,1435,345,1792]" captionTargetId="figure@6.[151,1435,345,1792]" captionTargetPageId="6" captionText="FIGURES 5–10. Acoma chihuahuaensis Evans &amp; Smith, new species, male holotype. 5, Dorsal habitus; 6, oblique habitus; 7, ventral habitus; 8, head; 9, dorsal parameres; 10, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697582" httpUri="https://zenodo.org/record/3697582/files/figure.png" pageId="5" pageNumber="56">Figs. 5–10</figureCitation>
</paragraph>
<paragraph id="3E2A36D4F377FFBA2FA8F913FC7CF8F3" blockId="5.[151,940,1706,1797]" box="[151,940,1773,1797]" pageId="5" pageNumber="56">
<uri id="4A043AD6F377FFBA2FA8F913FC7CF8F3" box="[151,940,1773,1797]" pageId="5" pageNumber="56">
urn:lsid:zoobank.org:act:
<uuid id="4A330C01F377FFBA2E9FF913FC7CF8F3" box="[416,940,1773,1797]" pageId="5" pageNumber="56">BD4FB72A-5BD1-4671-886A-C0E05D1BE391</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="768F655FF377FFB92FA8F8CAFB65FF21" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56" type="materials_examined">
<paragraph id="3E2A36D4F377FFBA2FA8F8CAFB54F860" blockId="5.[151,1437,1843,2014]" pageId="5" pageNumber="56">
<emphasis id="0CE1EAC6F377FFBA2FA8F8CAFE95F8B8" bold="true" box="[151,325,1843,1870]" pageId="5" pageNumber="56">Type material.</emphasis>
<materialsCitation id="8EFD3C89F377FFBA2E72F8CDFB54F860" ID-GBIF-Occurrence="2573655800" collectingDate="1982-07-23" collectionCode="CASC" collectorName="Fred G. Andrews &amp; Dr. Alan R. Hardy" country="Mexico" location="Type" pageId="5" pageNumber="56" specimenCount="2" stateProvince="Chihuahua" typeStatus="holotype">
<typeStatus id="E12E8876F377FFBA2E72F8CDFE65F8BB" box="[333,437,1843,1869]" pageId="5" pageNumber="56" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="5884AE11F377FFBA2D3DF8CAFD9DF8B8" box="[514,589,1844,1870]" pageId="5" pageNumber="56">CASC</collectionCode>
<location id="3B4A600FF377FFBA2D6CF8CAFD5BF8B8" LSID="urn:lsid:plazi:treatment:B63C87C2F377FFB82FA8F954FB5CFC3D:3B4A600FF377FFBA2D6CF8CAFD5BF8B8" box="[595,651,1844,1870]" country="Mexico" name="Type" pageId="5" pageNumber="56" stateProvince="Chihuahua">Type</location>
19747), labeled “
<collectingCountry id="46827644F377FFBA2C6DF8CAFC18F8B8" box="[850,968,1844,1870]" name="Mexico" pageId="5" pageNumber="56">MEXICO</collectingCountry>
:
<collectingRegion id="FC51F836F377FFBA2CEDF8CDFB9BF8BB" box="[978,1099,1843,1869]" country="Mexico" name="Chihuahua" pageId="5" pageNumber="56">Chihuahua</collectingRegion>
/
<locationDeviation id="2F982FF6F377FFBA2B5DF8CAFAE0F8BB" box="[1122,1328,1843,1870]" location="Concho" pageId="5" pageNumber="56">
<quantity id="F96D9B31F377FFBA2B5DF8CAFB62F8BB" box="[1122,1202,1843,1870]" metricMagnitude="4" metricUnit="m" metricValue="1.14263424" pageId="5" pageNumber="56" unit="mi" value="7.1">7.1 mi.</quantity>
W Concho
</locationDeviation>
/
<date id="4A2B1014F377FFBA2A78F8CAFF1FF884" pageId="5" pageNumber="56" value="1982-07-23">
<collectingDate id="5A6FE9FCF377FFBA2A78F8CAFF1FF884" pageId="5" pageNumber="56" value="1982-07-23">VII-23- 1982</collectingDate>
</date>
/
<collectorName id="93605302F377FFBA2FDAF8A9FE74F887" box="[229,420,1879,1906]" pageId="5" pageNumber="56">Fred G. Andrews</collectorName>
/ coll. blacklight //
<collectorName id="93605302F377FFBA2D40F8A6FC9BF887" box="[639,843,1879,1906]" pageId="5" pageNumber="56">Dr. Alan R. Hardy</collectorName>
/ Donation to the / California Academy / of Sciences // CASENT / 8424526 //
<typeStatus id="E12E8876F377FFBA2E90F882FD93F860" box="[431,579,1916,1942]" pageId="5" pageNumber="56" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="F9954D57F377FFBA2D68F882FD78F860" authorityName="Casey" authorityYear="1889" box="[599,680,1916,1942]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">Acoma</taxonomicName>
/ chihuahuaensis / Evans &amp; Smith, 2019”.
</materialsCitation>
</paragraph>
<paragraph id="3E2A36D4F377FFBA2FF8F85EFED1F828" blockId="5.[151,1437,1843,2014]" pageId="5" pageNumber="56">
<typeStatus id="E12E8876F377FFBA2FF8F85EFEE7F84C" box="[199,311,1952,1978]" pageId="5" pageNumber="56" type="paratype">Paratypes</typeStatus>
, (
<specimenCount id="2893FD5DF377FFBA2E6EF85EFE64F84C" box="[337,436,1951,1978]" pageId="5" pageNumber="56" type="male">9 males</specimenCount>
):
<materialsCitation id="8EFD3C89F377FFBA2EF2F85EFF2DF828" ID-GBIF-Occurrence="2573655804" collectingDate="1982-07-23" collectionCode="CASC" collectorName="Fred G. Andrews &amp; Dr. Alan R. Hardy" country="Mexico" location="Type" pageId="5" pageNumber="56" specimenCount="1" stateProvince="Chihuahua" typeStatus="paratype">
same data as holotype, except with
<collectionCode id="5884AE11F377FFBA2CBFF85EFC19F84C" box="[896,969,1952,1978]" pageId="5" pageNumber="56">CASC</collectionCode>
database numbers CASENT 8424527– 8424535
</materialsCitation>
.
</paragraph>
<paragraph id="3E2A36D4F374FFB92FF8FF67FB65FF21" blockId="6.[151,1436,152,323]" pageId="6" pageNumber="57">
<materialsCitation id="8EFD3C89F374FFB92FF8FF67FB61FF21" ID-GBIF-Occurrence="2573655806" collectionCode="AVEC, CASC, CMNC, UNAM" pageId="6" pageNumber="57" specimenCount="9" typeStatus="paratype">
<typeStatus id="E12E8876F374FFB92FF8FF67FEE4FF45" box="[199,308,153,179]" pageId="6" pageNumber="57" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="5884AE11F374FFB92EC7FF67FD91FF45" box="[504,577,153,179]" pageId="6" pageNumber="57">AVEC</collectionCode>
(
<specimenCount id="2893FD5DF374FFB92D6BFF67FD1DFF45" box="[596,717,153,179]" pageId="6" pageNumber="57" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424534),
<collectionCode id="5884AE11F374FFB92CF5FF67FBC3FF45" box="[970,1043,153,179]" pageId="6" pageNumber="57">CASC</collectionCode>
(
<specimenCount id="2893FD5DF374FFB92B1BFF67FB7BFF45" box="[1060,1195,153,179]" pageId="6" pageNumber="57" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
; CASENT 8424527– 8424532),
<collectionCode id="5884AE11F374FFB92E2EFF43FEB3FF21" box="[273,355,189,215]" pageId="6" pageNumber="57">CMNC</collectionCode>
(
<specimenCount id="2893FD5DF374FFB92E4AFF43FE3CFF21" box="[373,492,189,215]" pageId="6" pageNumber="57" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424535),
<collectionCode id="5884AE11F374FFB92DDBFF43FCEAFF21" box="[740,826,189,215]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="6" pageNumber="57">UNAM</collectionCode>
(
<specimenCount id="2893FD5DF374FFB92C73FF43FC13FF21" box="[844,963,189,215]" pageId="6" pageNumber="57" type="generic" typeStatus="paratype">1 paratype</specimenCount>
; CASENT 8424533)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF374FFB92FF8FF1EFEEEFEB5" pageId="6" pageNumber="57" type="diagnosis">
<paragraph id="3E2A36D4F374FFB92FF8FF1EFEEEFEB5" blockId="6.[151,1436,152,323]" pageId="6" pageNumber="57">
<emphasis id="0CE1EAC6F374FFB92FF8FF1EFE92FF0D" bold="true" box="[199,322,224,251]" pageId="6" pageNumber="57">Diagnosis.</emphasis>
Brown dorsally; anterior clypeal margin emarginate medially; antennae composed of 10 antennomeres, antennomeres 8–10 lamellate; protibia distinctly tridentate; elytra with disc glabrous and scattered setae along margins.
</paragraph>
</subSubSection>
<caption id="6AEA665CF374FFB92FA8F8E7FCFEF8A0" ID-DOI="http://doi.org/10.5281/zenodo.3697582" ID-Zenodo-Dep="3697582" httpUri="https://zenodo.org/record/3697582/files/figure.png" pageId="6" pageNumber="57" startId="6.[151,264,1817,1842]" targetBox="[151,1435,345,1792]" targetPageId="6">
<paragraph id="3E2A36D4F374FFB92FA8F8E7FCFEF8A0" blockId="6.[151,1436,1817,1878]" pageId="6" pageNumber="57">
<emphasis id="0CE1EAC6F374FFB92FA8F8E7FE91F8C4" bold="true" box="[151,321,1817,1842]" pageId="6" pageNumber="57">FIGURES 5–10</emphasis>
.
<taxonomicName id="F9954D57F374FFB92E71F8E4FD0DF8C4" authority="Evans &amp; Smith" authorityName="Evans &amp; Smith" authorityYear="2020" box="[334,733,1817,1842]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chihuahuaensis" status="sp. nov.">
<emphasis id="0CE1EAC6F374FFB92E71F8E4FDE9F8C4" box="[334,569,1818,1842]" italics="true" pageId="6" pageNumber="57">Acoma chihuahuaensis</emphasis>
Evans &amp; Smith
</taxonomicName>
,
<taxonomicNameLabel id="17D257BDF374FFB92DD8F8E4FCB4F8C4" box="[743,868,1817,1842]" pageId="6" pageNumber="57" rank="species">new species</taxonomicNameLabel>
, male holotype.
<emphasis id="0CE1EAC6F374FFB92B33F8E4FBC9F8C4" bold="true" box="[1036,1049,1818,1842]" pageId="6" pageNumber="57">5</emphasis>
, Dorsal habitus;
<emphasis id="0CE1EAC6F374FFB92BF9F8E7FB03F8C4" bold="true" box="[1222,1235,1817,1842]" pageId="6" pageNumber="57">6</emphasis>
, oblique habitus;
<emphasis id="0CE1EAC6F374FFB92AB6F8E4FA46F8C4" bold="true" box="[1417,1430,1818,1842]" pageId="6" pageNumber="57">7</emphasis>
, ventral habitus;
<emphasis id="0CE1EAC6F374FFB92E05F8C3FE97F8A0" bold="true" box="[314,327,1853,1878]" pageId="6" pageNumber="57">8</emphasis>
, head;
<emphasis id="0CE1EAC6F374FFB92EAEF8C3FE4EF8A0" bold="true" box="[401,414,1853,1878]" pageId="6" pageNumber="57">9</emphasis>
, dorsal parameres;
<emphasis id="0CE1EAC6F374FFB92D5BF8C3FDADF8A0" bold="true" box="[612,637,1853,1878]" pageId="6" pageNumber="57">10</emphasis>
, lateral genitalia.
</paragraph>
</caption>
<subSubSection id="768F655FF374FFB82FF8F87DFB4BFCE1" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57" type="description">
<paragraph id="3E2A36D4F374FFB82FF8F87DFDD3FD5D" blockId="6.[151,1436,1923,2022]" lastBlockId="7.[151,1437,153,971]" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
<emphasis id="0CE1EAC6F374FFB92FF8F87DFE89F868" bold="true" box="[199,345,1923,1950]" pageId="6" pageNumber="57">Description.</emphasis>
<typeStatus id="E12E8876F374FFB92E61F87AFE16F868" box="[350,454,1924,1950]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="A6AE2A51F374FFB92D2EF87AFD5AF868" box="[529,650,1923,1950]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1817,1842]" captionTargetBox="[151,1435,345,1792]" captionTargetId="figure@6.[151,1435,345,1792]" captionTargetPageId="6" captionText="FIGURES 5–10. Acoma chihuahuaensis Evans &amp; Smith, new species, male holotype. 5, Dorsal habitus; 6, oblique habitus; 7, ventral habitus; 8, head; 9, dorsal parameres; 10, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697582" httpUri="https://zenodo.org/record/3697582/files/figure.png" pageId="6" pageNumber="57">Figs. 5–10</figureCitation>
), length:
<quantity id="F96D9B31F374FFB92DCEF87AFC98F868" box="[753,840,1924,1950]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="6" pageNumber="57" unit="mm" value="6.9">6.9 mm</quantity>
, width:
<quantity id="F96D9B31F374FFB92C9FF87AFC27F868" box="[928,1015,1924,1950]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="57" unit="mm" value="2.6">2.6 mm</quantity>
. BODY elongate, moderately convex, and uniformly brown dorsally, underside and appendages slightly lighter. HEAD (
<figureCitation id="A6AE2A51F374FFB92B11F856FBA1F834" box="[1070,1137,1960,1986]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1817,1842]" captionTargetBox="[151,1435,345,1792]" captionTargetId="figure@6.[151,1435,345,1792]" captionTargetPageId="6" captionText="FIGURES 5–10. Acoma chihuahuaensis Evans &amp; Smith, new species, male holotype. 5, Dorsal habitus; 6, oblique habitus; 7, ventral habitus; 8, head; 9, dorsal parameres; 10, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697582" httpUri="https://zenodo.org/record/3697582/files/figure.png" pageId="6" pageNumber="57">Fig. 8</figureCitation>
) rugosely punctate; supraorbital carinae becoming obsolete at base of clypeus. Clypeus subtrapezoidal with margins strongly reflexed, lateral margins arcuate basally, becoming straight before broadly concave anterior margin flanked by prominent obtuse angles. Ocular canthus slightly less than one-third of eye length in lateral view, with scattered pale setae each not quite as long as canthus. PRONOTUM slightly wider than long, broadest at middle, and less than half length of elytron; disc convex, glabrous, shiny, sparsely punctate with punctures separated by four or more of their diameters, with a narrow longitudinal impunctate line medially; margins with distinct bead, anterior margin bisinuate between obtuse anterior angles, with marginal bead relatively broad and raised; anterolateral margins straight, converging anteriorly, posterolateral margins slightly converging posteriorly with posterolateral angles broadly rounded. SCUTELLUM small, parabolic, about one-fifth basal width of elytron, finely and setigerously punctate basally, and glabrous and becoming impunctate apically. ELYTRON convex, with disc shiny, glabrous, and distinctly punctostriate with deep punctures separated by 1.5–3.0 X their diameters, punctures becoming confused at apical declivity; margins setigerous, setae on lateral margins becoming shorter apically. LEGS with tridentate protibia moderately emarginate between apical and middle teeth, and shallowly emarginate between middle and basal teeth. PYGIDIUM parabolic, shiny, finely shagreened, with scattered setigerous punctures, setae long. GENITALIA with parameres symmetrical, slender and slightly sinuate when viewed laterally, with membranous flange extending to apical fifth, and about as long as phallobasis (
<figureCitation id="A6AE2A51F375FFB82E43FD6FFE27FD5D" box="[380,503,657,683]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1817,1842]" captionTargetBox="[151,1435,345,1792]" captionTargetId="figure@6.[151,1435,345,1792]" captionTargetPageId="6" captionText="FIGURES 5–10. Acoma chihuahuaensis Evans &amp; Smith, new species, male holotype. 5, Dorsal habitus; 6, oblique habitus; 7, ventral habitus; 8, head; 9, dorsal parameres; 10, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697582" httpUri="https://zenodo.org/record/3697582/files/figure.png" pageId="7" pageNumber="58">Figs. 9–10</figureCitation>
).
</paragraph>
<paragraph id="3E2A36D4F375FFB82FF8FD4AFB4BFCE1" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="58">
<emphasis id="0CE1EAC6F375FFB82FF8FD4AFEEFFD39" bold="true" box="[199,319,692,719]" pageId="7" pageNumber="58">Variation.</emphasis>
Length:
<quantity id="F96D9B31F375FFB82E9DFD4AFDFAFD39" box="[418,554,692,719]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.9" metricValueMin="5.9" pageId="7" pageNumber="58" unit="mm" value="6.4" valueMax="6.9" valueMin="5.9">5.9–6.9 mm</quantity>
, width:
<quantity id="F96D9B31F375FFB82DBDFD4BFCDCFD39" box="[642,780,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="7" pageNumber="58" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2–2.6 mm</quantity>
. Color varies from light brown to brown, occasionally with elytra lighter, while the head and pronotum are slightly darker. Clypeus varies from subtrapezoidal to nearly parabolic with narrowly to broadly concave anterior margin flanked by prominent angles or not.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF375FFB82FF8FCDFFA5FFCCD" box="[199,1423,801,827]" pageId="7" pageNumber="58" type="etymology">
<paragraph id="3E2A36D4F375FFB82FF8FCDFFA5FFCCD" blockId="7.[151,1437,153,971]" box="[199,1423,801,827]" pageId="7" pageNumber="58">
<emphasis id="0CE1EAC6F375FFB82FF8FCDFFE9EFCCD" bold="true" box="[199,334,801,827]" pageId="7" pageNumber="58">Etymology.</emphasis>
This species is named after the Mexican state of
<collectingRegion id="FC51F836F375FFB82C4AFCDFFC21FCCD" box="[885,1009,801,827]" country="Mexico" name="Chihuahua" pageId="7" pageNumber="58">Chihuahua</collectingRegion>
, where the
<typeStatus id="E12E8876F375FFB82B4EFCDFFB71FCCD" box="[1137,1185,801,827]" pageId="7" pageNumber="58">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="768F655FF375FFB82FF8FCBBFB5CFC3D" pageId="7" pageNumber="58" type="discussion">
<paragraph id="3E2A36D4F375FFB82FF8FCBBFB5CFC3D" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="58">
<emphasis id="0CE1EAC6F375FFB82FF8FCBBFEEBFCA9" bold="true" box="[199,315,837,863]" pageId="7" pageNumber="58">Remarks.</emphasis>
<taxonomicName id="F9954D57F375FFB82E7CFCBBFD42FCA9" authority="Cazier, 1953" authorityName="Cazier" authorityYear="1953" box="[323,658,836,863]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="glabrata">
<emphasis id="0CE1EAC6F375FFB82E7CFCBBFE28FCA9" box="[323,504,837,863]" italics="true" pageId="7" pageNumber="58">Acoma glabrata</emphasis>
<bibRefCitation id="5A044B25F375FFB82D3FFCBBFD42FCA9" author="Cazier, M. A." box="[512,658,836,863]" pageId="7" pageNumber="58" pagination="1 - 13" refId="ref13258" refString="Cazier, M. A. (1953) Review of the scarab genus Acoma. American Museum Novitates, 1624, 1 - 13." type="journal article" year="1953">Cazier, 1953</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F9954D57F375FFB82DA1FCBBFC97FCA9" authority="Evans &amp; Smith, 2020" authorityName="Evans &amp; Smith" authorityYear="2020" box="[670,839,837,863]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="nonglabrata" status="sp. nov.">
<emphasis id="0CE1EAC6F375FFB82DA1FCBBFD7FFCA9" box="[670,687,837,863]" italics="true" pageId="7" pageNumber="58">A</emphasis>
.
<emphasis id="0CE1EAC6F375FFB82D82FCBBFC97FCA9" box="[701,839,837,863]" italics="true" pageId="7" pageNumber="58">nonglabrata</emphasis>
</taxonomicName>
<emphasis id="0CE1EAC6F375FFB82C70FCBBFC09FCA9" bold="true" box="[847,985,836,863]" pageId="7" pageNumber="58">
<taxonomicNameLabel id="17D257BDF375FFB82C70FCBBFC09FCA9" box="[847,985,836,863]" pageId="7" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="F9954D57F375FFB82B26FCBBFB36FCA9" authority="Evans &amp; Smith, 2020" authorityName="Evans &amp; Smith" authorityYear="2020" box="[1049,1254,837,863]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="chihuahuaensis" status="sp. nov.">
<emphasis id="0CE1EAC6F375FFB82B26FCBBFBFAFCA9" box="[1049,1066,837,863]" italics="true" pageId="7" pageNumber="58">A</emphasis>
.
<emphasis id="0CE1EAC6F375FFB82B07FCBBFB36FCA9" box="[1080,1254,837,863]" italics="true" pageId="7" pageNumber="58">chihuahuaensis</emphasis>
</taxonomicName>
<emphasis id="0CE1EAC6F375FFB82BD1FCBBFAA8FCA9" bold="true" box="[1262,1400,836,863]" pageId="7" pageNumber="58">
<taxonomicNameLabel id="17D257BDF375FFB82BD1FCBBFAA8FCA9" box="[1262,1400,836,863]" pageId="7" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
all have antennae with 10 antennomeres, of which 8–10 form the lamellate club.
<taxonomicName id="F9954D57F375FFB82B39FC97FADAFC75" authority="Evans &amp; Smith, 2020" authorityName="Evans &amp; Smith" authorityYear="2020" box="[1030,1290,873,899]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="chihuahuaensis" status="sp. nov.">
<emphasis id="0CE1EAC6F375FFB82B39FC97FADAFC75" box="[1030,1290,873,899]" italics="true" pageId="7" pageNumber="58">Acoma chihuahuaensis</emphasis>
</taxonomicName>
<emphasis id="0CE1EAC6F375FFB82A2DFC97FA4CFC75" bold="true" box="[1298,1436,872,899]" pageId="7" pageNumber="58">
<taxonomicNameLabel id="17D257BDF375FFB82A2DFC97FA4CFC75" box="[1298,1436,872,899]" pageId="7" pageNumber="58" rank="species">new species</taxonomicNameLabel>
</emphasis>
is readily distinguished among these species by its darker color, distinct elytral punctostriae with deep punctures, more rounded and less pronounced protibial teeth, and its occurrence in mainland
<collectingCountry id="46827644F375FFB82B10FC4FFB58FC3D" box="[1071,1160,945,971]" name="Mexico" pageId="7" pageNumber="58">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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