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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.686.12206" ID-GBIF-Dataset="82b32106-bef0-417d-81d4-f47f22c33fa9" ID-PMC="PMC5672565" ID-Pensoft-Pub="1313-2970-686-15" ID-PubMed="29200915" ID-ZBK="976E23A1CFC74CB381705B59452825A6" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-686-15" ModsDocOrigin="ZooKeys 686" ModsDocTitle="Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)" checkinTime="1502744466696" checkinUser="pensoft" docAuthor="Sousa, Pedro, Arnedo, Miquel A. &amp; Harris, D. James" docDate="2017" docId="A6FABAEED2BDDA89B1E31C016EE280B4" docLanguage="en" docName="ZooKeys 686: 15-84" docOrigin="ZooKeys 686" docSource="http://dx.doi.org/10.3897/zookeys.686.12206" docTitle="Buthus tunetatus Herbst 1800" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FF8DFFA394365835FF81FFFFFF80FF91" masterDocTitle="Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)" masterLastPageNumber="84" masterPageNumber="15" pageNumber="37" updateTime="1668164638468" updateUser="ExternalLinkService">
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<mods:title>Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sousa, Pedro</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Arnedo, Miquel A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harris, D. James</mods:namePart>
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<mods:date>2017</mods:date>
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<mods:number>686</mods:number>
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<treatment ID-GBIF-Taxon="132648855" LSID="urn:lsid:plazi:treatment:A6FABAEED2BDDA89B1E31C016EE280B4" httpUri="http://treatment.plazi.org/id/A6FABAEED2BDDA89B1E31C016EE280B4" lastPageNumber="37" pageId="22" pageNumber="37">
<subSubSection pageId="22" pageNumber="37" type="nomenclature">
<paragraph pageId="22" pageNumber="37">
<pageBreakToken pageId="22" pageNumber="37" start="start">51</pageBreakToken>
.
<taxonomicName authority="Herbst, 1800" authorityName="Herbst" authorityYear="1800" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus tunetatus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetatus">Buthus tunetatus (Herbst, 1800)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="37" type="reference_group">
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Scorpio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpio tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetanus">Scorpio tunetanus</taxonomicName>
<bibRefCitation author="Herbst, JFW" journalOrPublisher="Gottlieb August Lange, Berlin" pageId="27" pageNumber="42" title="Natursystem der ungefluegelten Insekten. Naturgeschichte der Skorpionen." year="1800">Herbst 1800</bibRefCitation>
: 68-69, pl. III, fig. 3 (not pl. II, fig. 2, as listed in the text);
<bibRefCitation pageId="22" pageNumber="37">Latreille 1804</bibRefCitation>
: 122-124.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus (Leiurus) tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetanus" subGenus="Leiurus">Androctonus (Leiurus) tunetanus</taxonomicName>
: Ehrenberg in
<bibRefCitation author="Hemprich, FW" journalOrPublisher="Verhandungen der Gessellschaft naturforschende Freunde in Berlin" pageId="27" pageNumber="42" pagination="348 - 362" title="Vorlaeufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Scorpione und deren geographischen Verbreitung, nach den eigenen Beobachtungen von Dr. Hemprich und Dr. Ehrenberg." url="https://books.google.pt/books?id=8OoTAAAAQAAJ" volume="1" year="1829">Hemprich and Ehrenberg 1829</bibRefCitation>
: 354.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus (Leiurus) tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="subspecies" species="tunetanus" subGenus="Leiurus" subspecies="genuinus">Androctonus (Leiurus) tunetanus genuinus</taxonomicName>
: Ehrenberg in
<bibRefCitation author="Hemprich, FW" journalOrPublisher="Verhandungen der Gessellschaft naturforschende Freunde in Berlin" pageId="27" pageNumber="42" pagination="348 - 362" title="Vorlaeufige Uebersicht der in Nord-Afrika und West-Asien einheimischen Scorpione und deren geographischen Verbreitung, nach den eigenen Beobachtungen von Dr. Hemprich und Dr. Ehrenberg." url="https://books.google.pt/books?id=8OoTAAAAQAAJ" volume="1" year="1829">Hemprich and Ehrenberg 1829</bibRefCitation>
: 354.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus (Liurus) tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetanus" subGenus="Liurus">Androctonus (Liurus) tunetanus</taxonomicName>
: Ehrenberg in
<bibRefCitation author="Hemprich, FW" journalOrPublisher="Verhandungen der Gessellschaft naturforschende Freunde in Berlin" pageId="27" pageNumber="42" url="https://doi.org/10.5962/bhl.title.107403" year="1831">Hemprich and Ehrenberg 1831</bibRefCitation>
(pages unnumbered)
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus (Liurus) tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="subspecies" species="tunetanus" subGenus="Liurus" subspecies="genuinus">Androctonus (Liurus) tunetanus genuinus</taxonomicName>
: Ehrenberg in
<bibRefCitation author="Hemprich, FW" journalOrPublisher="Verhandungen der Gessellschaft naturforschende Freunde in Berlin" pageId="27" pageNumber="42" url="https://doi.org/10.5962/bhl.title.107403" year="1831">Hemprich and Ehrenberg 1831</bibRefCitation>
(pages unnumbered).
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Androctonus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Androctonus tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetanus">Androctonus tunetanus</taxonomicName>
: C. L.
<bibRefCitation author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" title="Die Arachniden. Getreu nach der Natur abgebildet und beschrieben. 12. C. H." url="http://www.biodiversitylibrary.org/item/55844" year="1845">Koch 1845</bibRefCitation>
: 15-19, pl. CCCCI (sic), fig. 968; C. L.
<bibRefCitation author="Koch, CL" journalOrPublisher="Zeh'schen Buchhandlung, Nuernberg" pageId="29" pageNumber="44" url="http://www.biodiversitylibrary.org/bibliography/43744" year="1850">Koch 1850</bibRefCitation>
: 90.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="occitanus">Buthus occitanus</taxonomicName>
(MIS):
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
and Whitman 2005 (part): 106.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="subspecies" species="occitanus" subspecies="tunetanus">Buthus occitanus tunetanus</taxonomicName>
:
<bibRefCitation author="Birula, AA" journalOrPublisher="Bulletin de l'Academie Imperiale des Sciences de St-Petersbourg [Izviestiia Imperatorskoi akademii nauk]" pageId="23" pageNumber="38" pagination="105 - 113" title="Bemerkungen ueber einige neue oder wenig bekaunte Skorpionnenformen Nord-Afrikas." volume="19" year="1903">Birula 1903</bibRefCitation>
: 107;
<bibRefCitation author="Birula, AA" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="23" pageNumber="38" pagination="115 - 192" title="Ueber Scorpio maurus Linne und seine Unterarten." url="https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf" volume="39" year="1910">Birula 1910</bibRefCitation>
: 118;
<bibRefCitation author="Borelli, A" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="24" pageNumber="39" pagination="148 - 159" title="Contributo allo studio della fauna Libica. Materiali raccolti nelle zone di Misurata e Homs (1912 - 13). Scorpioni." volume="6" year="1914 a">Borelli 1914a</bibRefCitation>
: 154-155;
<bibRefCitation author="Borelli, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano" pageId="24" pageNumber="39" pagination="456 - 464" title="Gli scorpioni del Museo Civico di Storia Naturale di Milano." volume="53" year="1914 b">Borelli 1914b</bibRefCitation>
: 461;
<bibRefCitation author="Borelli, A" journalOrPublisher="Universita di Torino" pageId="24" pageNumber="39" pagination="1 - 16" title="Missione zoologica del Dr. E. Festa in Cirenaica. XVI. Scorpioni e Solifughi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R." volume="39" year="1924">Borelli 1924</bibRefCitation>
: 4-5;
<bibRefCitation author="Borelli, A" journalOrPublisher="Annali del Museo civico di Storia Naturale Giacomo Doria" pageId="24" pageNumber="39" pagination="346 - 355" title="Risultati zoologici della Missione inviata dalla R. Societa Geografica Italiana per l'explorazione dell'oasi di Giarabud (1926 - 1927). Scorpioni e Solifughi." volume="52" year="1928">Borelli 1928</bibRefCitation>
: 351;
<bibRefCitation pageId="22" pageNumber="37">Giltay 1929</bibRefCitation>
: 196-197;
<bibRefCitation author="Werner, F" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien" pageId="37" pageNumber="52" pagination="1 - 34" title="Wissenschaftliche Ergebnisse einer zoologischen Forschungsreise nach Westalgerien und Marokko. III. Scorpiones." url="http://www.zobodat.at/publikation_articles.php?id=169321" volume="138" year="1929">Werner 1929</bibRefCitation>
: 30-31;
<bibRefCitation author="Caporiacco, L di" journalOrPublisher="Annali del Museo civico di Storia Naturale Giacomo Doria" pageId="24" pageNumber="39" pagination="395 - 408" title="Spedizione scientifica all'Oasi di Cufra (Marzo-Luglio 1931). Scorpioni e Solifugi." volume="55" year="1932">Caporiacco 1932</bibRefCitation>
: 395-396;
<bibRefCitation author="Schenkel, E" journalOrPublisher="Revue Suisse de Zoologie" pageId="34" pageNumber="49" pagination="375 - 396" title="Notizen ueeber einige Scorpione und Solifugen." volume="39" year="1932">Schenkel 1932</bibRefCitation>
: 379-380;
<bibRefCitation author="Werner, F" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien" pageId="37" pageNumber="52" pagination="285 - 306" title="Ergebnisse einer zoologischen Forschungsreise nach Marokko. VI. Skorpione." url="http://www.zobodat.at/publikation_articles.php?id=180394" volume="141" year="1932">Werner 1932</bibRefCitation>
: 300-305;
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de la Societe d'Histoire Naturelle d'Afrique du Nord" pageId="33" pageNumber="48" pagination="90 - 100" title="Scorpions du Sahara central. Mission du Hoggar III (Fevrier a Mai 1928)." volume="4" year="1934">Pallary 1934</bibRefCitation>
: 99;
<bibRefCitation pageId="22" pageNumber="37">Borelli 1934</bibRefCitation>
: 169;
<bibRefCitation author="Werner, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="37" pageNumber="52" pagination="82 - 88" title="Ueber skorpione aus Marokko." volume="45" year="1934 b">Werner 1934b</bibRefCitation>
: 84-85, fig. 4;
<bibRefCitation author="Werner, F" journalOrPublisher="Strand, Riga" pageId="37" pageNumber="52" pagination="171 - 193" title="Neu-Eingaenge von Skorpionen im Zoologischen Museum in Hamburg. Festschrift zum 60. Geburtstage von Professor Dr. Embrik." volume="2" year="1936">Werner 1936</bibRefCitation>
: 173;
<bibRefCitation author="Caporiacco, L di" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano" pageId="24" pageNumber="39" pagination="340 - 354" title="Risultati scientifici della Missione del Prof. G. Scortecci nel Fezzan e sui Tassili (1936). Scorpioni e Solifughi." volume="76" year="1937">Caporiacco 1937</bibRefCitation>
: 345;
<bibRefCitation author="Schenkel, E" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="34" pageNumber="49" pagination="186 - 201" title="Mitteilungen ueber Spinnentiere. Skorpione aus Marokko." volume="60" year="1949">Schenkel 1949</bibRefCitation>
: 186;
<bibRefCitation author="Vachon, M" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="35" pageNumber="50" pagination="334 - 396" title="Etudes sur les Scorpions. III (suite). Description des Scorpions du Nord de l'Afrique." volume="27" year="1949 c">Vachon 1949c</bibRefCitation>
: 344-353, fig. 381-393;
<bibRefCitation author="Vachon, M" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="35" pageNumber="50" pagination="46 - 104" title="Etudes sur les Scorpions. (suite et fin). Determination des Scorpions du Nord-Ouest de l'Afrique." volume="29" year="1951 a">Vachon 1951a</bibRefCitation>
: fig. 670;
<bibRefCitation author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon 1952a</bibRefCitation>
: 272-281, fig. 381-393, 670;
<bibRefCitation author="Vachon, M" journalOrPublisher="Toxicon" pageId="36" pageNumber="51" pagination="209 - 218" title="Liste des scorpions connus en Egypte, Arabie, Israeli, Liban, Syrie, Jordanie, Turquie, Irak, Iran." volume="4" year="1966">Vachon 1966</bibRefCitation>
: 211;
<normalizedToken originalValue="Peréz">Perez</normalizedToken>
1974: 22;
<bibRefCitation author="Levy, G" journalOrPublisher="The Israel Academy of Sciences an d Humanit es, Jerusalem" pageId="30" pageNumber="45" title="Fauna Palaestina. Arachnida I: Scorpiones." year="1980">Levy and Amitai 1980</bibRefCitation>
: 16;
<bibRefCitation author="El-Hennawy, HK" journalOrPublisher="Serket" pageId="25" pageNumber="40" pagination="95 - 153" title="A catalogue of the scorpions described from the Arab countries (1758 - 1990) (Arachnida: Scorpionida)." volume="2" year="1992">El-Hennawy 1992</bibRefCitation>
: 98, 121;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Klapalekiana" pageId="29" pageNumber="44" pagination="19 - 21" title="First report of Orthochirusinnesi (Scorpionida: Buthidae) from Morocco." volume="31" year="1995">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
1995
</bibRefCitation>
: 20;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="29" pageNumber="44" pagination="177 - 185" title="A check-list of scorpions (Arachnida) in the collection of the Hungarian Natural History Museum, Budapest." volume="89" year="1997">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
1997
</bibRefCitation>
: 179;
<bibRefCitation pageId="22" pageNumber="37">Gantenbein et al. 1998</bibRefCitation>
a: 51;
<bibRefCitation pageId="22" pageNumber="37">Gantenbein et al. 1998</bibRefCitation>
b: 33-39;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Madagaskar, Jihlava" pageId="29" pageNumber="44" title="Stiri." year="1998">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
1998
</bibRefCitation>
: 106;
<bibRefCitation pageId="22" pageNumber="37">Fet and Lowe 2000</bibRefCitation>
: 97;
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Biogeographica" pageId="30" pageNumber="45" pagination="109 - 127" title="Considerations sur les modeles de distribution et differentiation du genre Buthus Leach, 1815, avec la description d'une nouvelle espece des montagnes du Tassili des Ajjer, Algerie (Scorpiones, Buthidae)." volume="78" year="2002">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
2002
</bibRefCitation>
, p. 113, 115, fig. 8-9, 11, 13;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Serket" pageId="29" pageNumber="44" pagination="1 - 23" title="A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany." volume="8" year="2002">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
2002
</bibRefCitation>
: 6;
<bibRefCitation author="Gantenbein, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="26" pageNumber="41" pagination="119 - 130" title="The phylogeographic importance of the Strait of Gibraltar as a gene flow barrier in terrestrial arthropods: a case study with the scorpion Buthusoccitanus as model organism." url="https://doi.org/10.1016/S1055-7903(03)00031-9" volume="28" year="2003">
Gantenbein and
<normalizedToken originalValue="Largiadèr">Largiader</normalizedToken>
2003
</bibRefCitation>
: 120, 122;
<bibRefCitation author="Ben Othmen, A" journalOrPublisher="Biological Journal of the Linnean Society" pageId="23" pageNumber="38" pagination="255 - 265" title="Allozymic differentiation of Tunisian populations of Androctonus species and Buthusoccitanus (Scorpiones: Buthidae)." url="https://doi.org/10.1111/j.1095-8312.2003.00285.x" volume="81" year="2004">Ben Othmen et al. 2004</bibRefCitation>
: 257;
<bibRefCitation pageId="22" pageNumber="37">Touloun 2012</bibRefCitation>
: 37, 41.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus (Buthus) occitanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="subspecies" species="occitanus" subGenus="Buthus" subspecies="tunetanus">Buthus (Buthus) occitanus tunetanus</taxonomicName>
:
<bibRefCitation author="Birula, AA" journalOrPublisher="Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien" pageId="23" pageNumber="38" pagination="121 - 152" title="Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. F. Werner's nach dem Aegyptischen Sudan und Nord-Uganda. XIV. Scorpiones und Solifugae." url="http://www.zobodat.at/publikation_articles.php?id=148339" volume="117" year="1908">Birula 1908</bibRefCitation>
: 123-124;
<bibRefCitation author="Birula, AA" journalOrPublisher="Abteilung fuer Systematik, Geographie und Biologie der Tiere" pageId="23" pageNumber="38" pagination="505 - 522" title="Scorpione und Solifugen von Tripolis und Barka. Nach der Sammlung von Dr. Bruno Klaptocz im jahre 19 - 06. Zoologische Jahrbuecher." volume="28" year="1909">Birula 1909</bibRefCitation>
: 507-508, fig. B;
<bibRefCitation author="Birula, AA" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="23" pageNumber="38" pagination="115 - 192" title="Ueber Scorpio maurus Linne und seine Unterarten." url="https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf" volume="39" year="1910">Birula 1910</bibRefCitation>
: 156-157;
<bibRefCitation author="Birula, AA" journalOrPublisher="Zapiski Kavkazskogo Muzeya, Seriia A [Memoires du Musee du Caucase, Se ́ rie A]" pageId="23" pageNumber="38" pagination="1 - 253" title="[authored as Byalynitskii-Birulya, A. A.] ArachnoideaArthrogastra Caucasica. Pars I. Scorpiones." volume="5" year="1917 a">Birula 1917a</bibRefCitation>
: 223;
<bibRefCitation author="Roewer, CF" journalOrPublisher="Senckenbergiana" pageId="34" pageNumber="49" pagination="205 - 244" title="Ueber eine neuerworbene Sammlung von Skorpionen des Natur-Museums Senckenberg." volume="26" year="1943">Roewer 1943</bibRefCitation>
: 206.
</paragraph>
<paragraph pageId="22" pageNumber="37">
<taxonomicName class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus tunetanus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="species" species="tunetanus">Buthus tunetanus</taxonomicName>
:
<bibRefCitation pageId="22" pageNumber="37">Simon 1872</bibRefCitation>
: 251-252;
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Revue Suisse de Zoologie" pageId="30" pageNumber="45" pagination="875 - 912" title="Complements a la faune de scorpions (Arachnida) de l'Afrique du Nord, avec des considerations sur le genre Buthus Leach, 1815." volume="110" year="2003">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
2003
</bibRefCitation>
: 897-899, fig. 57-61;
<bibRefCitation author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="29" pageNumber="44" pagination="1 - 16" title="Review of Tunisian Species of the Genus Buthus with Descriptions of Two New Species and a Discussion of Ehrenberg's Types (Scorpiones: Buthidae)." url="http://www.science.marshall.edu/fet/euscorpius/p2006_34.pdf" volume="34" year="2006">
<normalizedToken originalValue="Kovařík">Kovarik</normalizedToken>
2006
</bibRefCitation>
: 2, 8, 10, 15, fig. 16-19.;
<bibRefCitation author="Sadine, SE" journalOrPublisher="Arachnides" pageId="34" pageNumber="49" pagination="1 - 10" title="Premieres donnees sur la diversite scorpionique dans la region du Souf (Algerie)." volume="61" year="2011">Sadine et al. 2011</bibRefCitation>
: 6;
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="31" pageNumber="46" pagination="11 - 18" title="A new species of Buthus Leach, 1815 from Egypt (Scorpiones, Buthidae)." volume="16" year="2012">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
and Cloudsley-Thompson 2012
</bibRefCitation>
: 13-16, fig. 8;
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Serket" pageId="31" pageNumber="46" pagination="8 - 15" title="Confirmation of a new species of Buthus Leach, 1815 from Alexandria, Egypt (Scorpiones, Buthidae)." volume="13" year="2012">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
and Simon 2012
</bibRefCitation>
: 12;
<bibRefCitation author="Lourenco, WR" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="31" pageNumber="46" pagination="63 - 68" title="A new species of Buthus Leach, 1815 from Algeria (Scorpiones, Buthidae)." volume="16" year="2013">
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
2013
</bibRefCitation>
: 65-66; Rossi, Tropea and
<normalizedToken originalValue="Yağmur">Yagmur</normalizedToken>
2013: 4-5, 7
</paragraph>
<paragraph pageId="22" pageNumber="37">
Scorpion Tunetanus (ISS):
<bibRefCitation pageId="22" pageNumber="37">Latreille 1817</bibRefCitation>
: 106.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="37" type="type material">
<paragraph pageId="22" pageNumber="37">Type material.</paragraph>
<paragraph pageId="22" pageNumber="37">
Types lost according to
<bibRefCitation pageId="22" pageNumber="37">Fet and Lowe (2000)</bibRefCitation>
, Tunisia.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="37" type="distribution">
<paragraph pageId="22" pageNumber="37">Distribution.</paragraph>
<paragraph pageId="22" pageNumber="37">The species is currently distributed across Algeria, Libya, Morocco and Tunisia, and doubtfully in the island of Malta.</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="37" type="remarks">
<paragraph pageId="22" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="22" pageNumber="37">
Because of the description of new species from Tunisia, and the lack of both type specimen and locality (beyond the country), a neotype for
<taxonomicName lsidName="B. tunetanus" pageId="22" pageNumber="37" rank="species" species="tunetanus">B. tunetanus</taxonomicName>
is necessary to stabilize the taxonomy of Tunisian
<taxonomicName class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus" order="Scorpiones" pageId="22" pageNumber="37" phylum="Arthropoda" rank="genus">Buthus</taxonomicName>
. As explained for
<taxonomicName lsidName="B. paris" pageId="22" pageNumber="37" rank="species" species="paris">B. paris</taxonomicName>
, this is further complicated by recent diagnoses of
<taxonomicName lsidName="B. tunetanus" pageId="22" pageNumber="37" rank="species" species="tunetanus">B. tunetanus</taxonomicName>
that differ from those offered by
<bibRefCitation author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon (1952a)</bibRefCitation>
, and as such if
<normalizedToken originalValue="Vachons">Vachon's</normalizedToken>
<taxonomicName lsidName="B. tuntetanus" pageId="22" pageNumber="37" rank="species" species="tuntetanus">B. tuntetanus</taxonomicName>
material is found in the MNHN it should be given priority in the future designation of a neotype.
<bibRefCitation author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon (1952a)</bibRefCitation>
did not formally described any variety of
<taxonomicName lsidName="B. tunetanus" pageId="22" pageNumber="37" rank="species" species="tunetanus">B. tunetanus</taxonomicName>
, but he again split the specimens that compose the species into four regions: 1) the typical region, corresponding to north and central Tunisia; 2) the southern montane region of Algeria, specimens from which have subsequently been described as
<taxonomicName lsidName="B. tassili" pageId="22" pageNumber="37" rank="species" species="tassili">B. tassili</taxonomicName>
; 3) the Algerian Saharan Atlas and the southern region of the High Plateau; and 4) the disjunct desert regions of southern Tunisia, western central Algeria and eastern central Morocco. It is unclear whether region 3 or 4 might either correspond to
<taxonomicName lsidName="B. dunlopi" pageId="22" pageNumber="37" rank="species" species="dunlopi">B. dunlopi</taxonomicName>
or
<taxonomicName lsidName="B. saharicus" pageId="22" pageNumber="37" rank="species" species="saharicus">B. saharicus</taxonomicName>
. As explained in
<bibRefCitation pageId="22" pageNumber="37">Fet and Lowe (2000)</bibRefCitation>
, the name
<taxonomicName lsidName="A. (Leiurus) t." pageId="22" pageNumber="37" rank="subspecies" species="t." subGenus="Leiurus" subspecies="genuinus">A. (Leiurus) t. genuinus</taxonomicName>
refers to the nominotypical form of the species and as such the adjective &quot;
<taxonomicName genus="Buthoidea" lsidName="Buthoidea genuinus" pageId="22" pageNumber="37" rank="species" species="genuinus">genuinus</taxonomicName>
&quot; is not an available subspecific name.
</paragraph>
</subSubSection>
</treatment>
</document>