Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)
Author
Sousa, Pedro
Author
Arnedo, Miquel A.
Author
Harris, D. James
text
ZooKeys
2017
686
15
84
http://dx.doi.org/10.3897/zookeys.686.12206
journal article
http://dx.doi.org/10.3897/zookeys.686.12206
1313-2970-686-15
976E23A1CFC74CB381705B59452825A6
976E23A1CFC74CB381705B59452825A6
51
.
Buthus tunetatus (Herbst, 1800)
Scorpio tunetanus
Herbst 1800
: 68-69, pl. III, fig. 3 (not pl. II, fig. 2, as listed in the text);
Latreille 1804
: 122-124.
Androctonus (Leiurus) tunetanus
: Ehrenberg in
Hemprich and Ehrenberg 1829
: 354.
Androctonus (Leiurus) tunetanus genuinus
: Ehrenberg in
Hemprich and Ehrenberg 1829
: 354.
Androctonus (Liurus) tunetanus
: Ehrenberg in
Hemprich and Ehrenberg 1831
(pages unnumbered)
Androctonus (Liurus) tunetanus genuinus
: Ehrenberg in
Hemprich and Ehrenberg 1831
(pages unnumbered).
Androctonus tunetanus
: C. L.
Koch 1845
: 15-19, pl. CCCCI (sic), fig. 968; C. L.
Koch 1850
: 90.
Buthus occitanus
(MIS):
Kovarik
and Whitman 2005 (part): 106.
Buthus occitanus tunetanus
:
Birula 1903
: 107;
Birula 1910
: 118;
Borelli 1914a
: 154-155;
Borelli 1914b
: 461;
Borelli 1924
: 4-5;
Borelli 1928
: 351;
Giltay 1929
: 196-197;
Werner 1929
: 30-31;
Caporiacco 1932
: 395-396;
Schenkel 1932
: 379-380;
Werner 1932
: 300-305;
Pallary 1934
: 99;
Borelli 1934
: 169;
Werner 1934b
: 84-85, fig. 4;
Werner 1936
: 173;
Caporiacco 1937
: 345;
Schenkel 1949
: 186;
Vachon 1949c
: 344-353, fig. 381-393;
Vachon 1951a
: fig. 670;
Vachon 1952a
: 272-281, fig. 381-393, 670;
Vachon 1966
: 211;
Perez
1974: 22;
Levy and Amitai 1980
: 16;
El-Hennawy 1992
: 98, 121;
Kovarik
1995
: 20;
Kovarik
1997
: 179;
Gantenbein et al. 1998
a: 51;
Gantenbein et al. 1998
b: 33-39;
Kovarik
1998
: 106;
Fet and Lowe 2000
: 97;
Lourenco
2002
, p. 113, 115, fig. 8-9, 11, 13;
Kovarik
2002
: 6;
Gantenbein and
Largiader
2003
: 120, 122;
Ben Othmen et al. 2004
: 257;
Touloun 2012
: 37, 41.
Buthus (Buthus) occitanus tunetanus
:
Birula 1908
: 123-124;
Birula 1909
: 507-508, fig. B;
Birula 1910
: 156-157;
Birula 1917a
: 223;
Roewer 1943
: 206.
Buthus tunetanus
:
Simon 1872
: 251-252;
Lourenco
2003
: 897-899, fig. 57-61;
Kovarik
2006
: 2, 8, 10, 15, fig. 16-19.;
Sadine et al. 2011
: 6;
Lourenco
and Cloudsley-Thompson 2012
: 13-16, fig. 8;
Lourenco
and Simon 2012
: 12;
Lourenco
2013
: 65-66; Rossi, Tropea and
Yagmur
2013: 4-5, 7
Scorpion Tunetanus (ISS):
Latreille 1817
: 106.
Type material.
Types lost according to
Fet and Lowe (2000)
, Tunisia.
Distribution.
The species is currently distributed across Algeria, Libya, Morocco and Tunisia, and doubtfully in the island of Malta.
Remarks.
Because of the description of new species from Tunisia, and the lack of both type specimen and locality (beyond the country), a neotype for
B. tunetanus
is necessary to stabilize the taxonomy of Tunisian
Buthus
. As explained for
B. paris
, this is further complicated by recent diagnoses of
B. tunetanus
that differ from those offered by
Vachon (1952a)
, and as such if
Vachon's
B. tuntetanus
material is found in the MNHN it should be given priority in the future designation of a neotype.
Vachon (1952a)
did not formally described any variety of
B. tunetanus
, but he again split the specimens that compose the species into four regions: 1) the typical region, corresponding to north and central Tunisia; 2) the southern montane region of Algeria, specimens from which have subsequently been described as
B. tassili
; 3) the Algerian Saharan Atlas and the southern region of the High Plateau; and 4) the disjunct desert regions of southern Tunisia, western central Algeria and eastern central Morocco. It is unclear whether region 3 or 4 might either correspond to
B. dunlopi
or
B. saharicus
. As explained in
Fet and Lowe (2000)
, the name
A. (Leiurus) t. genuinus
refers to the nominotypical form of the species and as such the adjective "
genuinus
" is not an available subspecific name.