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<document id="D9586D2450F568DDE5EEA3840D5DC3B8" ID-CLB-Dataset="7769" ID-DOI="10.1515/aemnp-2017-0079" ID-GBIF-Dataset="20436c9c-2b03-46c4-b53c-b14cc674f8e3" ID-ISSN="0374-1036" ID-Zenodo-Dep="4504000" ID-ZooBank="9FB0A4B6-2876-4331-A235-0B5A3781BF07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612462577153" checkinUser="felipe" docAuthor="Bremer, Hans J. &amp; Grimm, Roland" docDate="2017" docId="A67A7341D437FFD3FEA0FBEB899D481D" docLanguage="en" docName="ActaEntMusNatPra.57.2.331-343.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57 (2)" docSource="http://dx.doi.org/10.1515/aemnp-2017-0079" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Paragonocnemis (Borneogonocnemis) ardoini Bremer 1991" docType="treatment" docVersion="8" lastPageNumber="341" masterDocId="5A430B39D43EFFD9FFABFF808B034B0E" masterDocTitle="The species of Gonocnemis and Paragonocnemis from Borneo with notes on synonymy in the Oriental species (Coleoptera: Tenebrionidae)" masterLastPageNumber="343" masterPageNumber="331" pageNumber="340" updateTime="1698927250942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A9E01FC081E287476AFF1B3731836E5F">The species of Gonocnemis and Paragonocnemis from Borneo with notes on synonymy in the Oriental species (Coleoptera: Tenebrionidae)</mods:title>
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<mods:affiliation id="0F7931241AD2BEA85E646DF3B826DE95">) &amp;) Diakonie Wohnstift am Westerberg, Bergstrasse 35 B, 49076 Osnabrück, Germany; e-mail: hjbremer @ live. de</mods:affiliation>
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<mods:affiliation id="9FBFD2B9E3A4CB672D8605B100020F58">) &amp;) Unterer Sägerweg 74, 75305 Neuenbürg, Germany; e-mail: grimm. tenebrio @ t-online. de</mods:affiliation>
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<treatment id="A67A7341D437FFD3FEA0FBEB899D481D" ID-DOI="http://doi.org/10.5281/zenodo.4602457" ID-GBIF-Taxon="180331202" ID-Zenodo-Dep="4602457" LSID="urn:lsid:plazi:treatment:A67A7341D437FFD3FEA0FBEB899D481D" httpUri="http://treatment.plazi.org/id/A67A7341D437FFD3FEA0FBEB899D481D" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340">
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<paragraph id="2E6CC257D437FFD0FEA0FBEB88A54F8D" blockId="9.[267,934,1131,1155]" box="[267,934,1131,1155]" pageId="9" pageNumber="340">
<heading id="7524753BD437FFD0FEA0FBEB88A54F8D" bold="true" box="[267,934,1131,1155]" centered="true" fontSize="8" level="2" pageId="9" pageNumber="340" reason="3">
<taxonomicName id="E9D3B9D4D437FFD0FEA0FBEB88A54F8D" authority="Bremer, 1991" authorityName="Bremer" authorityYear="1991" box="[267,934,1131,1155]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ardoini" subGenus="Borneogonocnemis">
<emphasis id="1CA71E45D437FFD0FEA0FBEB88A54F8D" bold="true" box="[267,934,1131,1155]" pageId="9" pageNumber="340">
<emphasis id="1CA71E45D437FFD0FEA0FBEB8AC74F8D" bold="true" box="[267,452,1131,1155]" italics="true" pageId="9" pageNumber="340">Paragonocnemis</emphasis>
(
<emphasis id="1CA71E45D437FFD0FE7FFBEB89AA4F8D" bold="true" box="[468,681,1131,1155]" italics="true" pageId="9" pageNumber="340">Borneogonocnemis</emphasis>
)
<emphasis id="1CA71E45D437FFD0FD12FBEB880A4F8D" bold="true" box="[697,777,1131,1155]" italics="true" pageId="9" pageNumber="340">ardoini</emphasis>
<bibRefCitation id="4A42BFA6D437FFD0FCBBFBEC88A54F8D" author="BREMER H. J." box="[784,934,1132,1155]" pageId="9" pageNumber="340" pagination="149 - 156" refId="ref7359" refString="BREMER H. J. 1991: Anmerkungen zur Gattung Azarelius Fairmaire, 1892, sowie Beschreibung einer neuen orientalischen Paragonocnemis-Art. (Coleoptera, Tenebrionidae, Amarygmini). Entomofauna 12: 149 - 156." type="journal article" year="1991">Bremer, 1991</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="2E6CC257D437FFD0FD88FB0A898C4F93" blockId="9.[113,987,1162,1242]" box="[547,655,1162,1181]" pageId="9" pageNumber="340">(Figs 6a, 6b)</paragraph>
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<paragraph id="2E6CC257D437FFD0FFDAFB2D888D4FCE" blockId="9.[113,987,1162,1242]" box="[113,910,1196,1216]" pageId="9" pageNumber="340">
<treatmentCitation id="AF72E446D437FFD0FFDAFB2D89CE4FCE" author="BREMER H. J." box="[113,717,1196,1216]" page="153" pageId="9" pageNumber="340" year="1991">
<taxonomicName id="E9D3B9D4D437FFD0FFDAFB2D89CE4FCE" authority="Bremer, 1991: 151 - 153" authorityName="Bremer" authorityPageNumber="151 - 153" authorityYear="1991" box="[113,717,1196,1216]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ardoini" subGenus="Borneogonocnemis">
<emphasis id="1CA71E45D437FFD0FFDAFB2D8A014FB1" box="[113,258,1197,1215]" italics="true" pageId="9" pageNumber="340">Paragonocnemis</emphasis>
(
<emphasis id="1CA71E45D437FFD0FEA5FB2D8AB04FB1" box="[270,435,1197,1215]" italics="true" pageId="9" pageNumber="340">Borneogonocnemis</emphasis>
)
<emphasis id="1CA71E45D437FFD0FE14FB2C8AFE4FB1" box="[447,509,1196,1215]" italics="true" pageId="9" pageNumber="340">ardoini</emphasis>
<bibRefCitation id="4A42BFA6D437FFD0FDA8FB2D89CE4FCE" author="BREMER H. J." box="[515,717,1197,1216]" pageId="9" pageNumber="340" pagination="149 - 156" refId="ref7359" refString="BREMER H. J. 1991: Anmerkungen zur Gattung Azarelius Fairmaire, 1892, sowie Beschreibung einer neuen orientalischen Paragonocnemis-Art. (Coleoptera, Tenebrionidae, Amarygmini). Entomofauna 12: 149 - 156." type="journal article" year="1991">Bremer, 1991: 151153</bibRefCitation>
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(original description).
</paragraph>
<paragraph id="2E6CC257D437FFD0FFDAFB4788D84FD4" blockId="9.[113,987,1162,1242]" box="[113,987,1223,1242]" pageId="9" pageNumber="340">
<treatmentCitation id="AF72E446D437FFD0FFDAFB4789C44FD4" author="BREMER H. J." box="[113,711,1223,1242]" page="56" pageId="9" pageNumber="340" year="1998">
<taxonomicName id="E9D3B9D4D437FFD0FFDAFB4789C44FD4" authority="Bremer, 1998: 54 - 56" authorityName="Bremer" authorityPageNumber="54 - 56" authorityYear="1998" box="[113,711,1223,1242]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="velutinus" subGenus="Borneogonocnemis">
<emphasis id="1CA71E45D437FFD0FFDAFB478A014FD7" box="[113,258,1223,1241]" italics="true" pageId="9" pageNumber="340">Paragonocnemis</emphasis>
(
<emphasis id="1CA71E45D437FFD0FEA5FB478AB04FD7" box="[270,435,1223,1241]" italics="true" pageId="9" pageNumber="340">Borneogonocnemis</emphasis>
)
<emphasis id="1CA71E45D437FFD0FE14FB47890F4FD4" box="[447,524,1223,1242]" italics="true" pageId="9" pageNumber="340">velutinus</emphasis>
<bibRefCitation id="4A42BFA6D437FFD0FDB9FB4789C44FD4" author="BREMER H. J." box="[530,711,1223,1242]" pageId="9" pageNumber="340" pagination="51 - 57" refId="ref7460" refString="BREMER H. J. 1998: Neue Tenebrionidae der Triben Lupropini und Amarygmini aus der orientalischen Region (Coleoptera). Acta Coleopterologica 14 (1): 51 - 57." type="journal article" year="1998">Bremer, 1998: 5456</bibRefCitation>
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(original description),
<emphasis id="1CA71E45D437FFD0FC27FB4788D84FD4" bold="true" box="[908,987,1223,1242]" pageId="9" pageNumber="340">syn. nov.</emphasis>
</paragraph>
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<paragraph id="2E6CC257D437FFD0FFDAFB7189354E11" blockId="9.[113,1090,1265,1444]" pageId="9" pageNumber="340">
<emphasis id="1CA71E45D437FFD0FFDAFB718A574E0A" bold="true" box="[113,340,1265,1284]" pageId="9" pageNumber="340">Type material examined.</emphasis>
<taxonomicName id="E9D3B9D4D437FFD0FEF1FB7189F54E0A" authorityName="Bremer" authorityYear="1998" box="[346,758,1265,1284]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="velutinus" subGenus="Borneogonocnemis">
<emphasis id="1CA71E45D437FFD0FEF1FB718AE84E0D" box="[346,491,1265,1283]" italics="true" pageId="9" pageNumber="340">Paragonocnemis</emphasis>
(
<emphasis id="1CA71E45D437FFD0FE5CFB71899F4E0D" box="[503,668,1265,1283]" italics="true" pageId="9" pageNumber="340">Borneogonocnemis</emphasis>
)
<emphasis id="1CA71E45D437FFD0FD02FB7189F54E0A" box="[681,758,1265,1284]" italics="true" pageId="9" pageNumber="340">velutinus</emphasis>
</taxonomicName>
:
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<typeStatus id="F1687CF5D437FFD0FCA9FB7188564E0C" box="[770,853,1265,1284]" pageId="9" pageNumber="340" type="holotype">HOLOTYPE</typeStatus>
:
<location id="2B0C948CD437FFD0FCCAFB7188A04E0A" LSID="urn:lsid:plazi:treatment:A67A7341D437FFD3FEA0FBEB899D481D:2B0C948CD437FFD0FCCAFB7188A04E0A" box="[865,931,1265,1284]" country="Malaysia" county="Long Linau" municipality="Belaga" name="Borneo" pageId="9" pageNumber="340">Borneo</location>
:
<collectingCountry id="56C482C7D437FFD0FC00FB7188F44E0A" box="[939,1015,1265,1284]" name="Malaysia" pageId="9" pageNumber="340">Sarawak</collectingCountry>
,
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,
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,
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</date>
,
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leg. (
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).
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</paragraph>
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<emphasis id="1CA71E45D437FFD0FFDAFAA6895D4E37" bold="true" box="[113,606,1318,1337]" pageId="9" pageNumber="340">
Additional material examined.
<collectingCountry id="56C482C7D437FFD0FE20FAA689024E37" box="[395,513,1318,1337]" name="Malaysia" pageId="9" pageNumber="340">MALAYSIA</collectingCountry>
:
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:
</emphasis>
Santubong Peninsula, Permai Rainforest Resort,
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,
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,
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(at light),
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,
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,
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(at light),
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,
<specimenCount id="38D509DED437FFD0FC0CFAC188E74E5A" box="[935,996,1345,1364]" pageId="9" pageNumber="340" type="generic">1 spec.</specimenCount>
,
<collectionCode id="48C25A92D437FFD0FC40FAC188FF4E5A" box="[1003,1020,1345,1364]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="340">R</collectionCode>
. Grimm leg.(3
<collectionCode id="48C25A92D437FFD0FF01FADC8BE44E61" box="[170,231,1372,1391]" pageId="9" pageNumber="340">RGCN</collectionCode>
, 1
<collectionCode id="48C25A92D437FFD0FF57FADC8A284E61" box="[252,299,1372,1391]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="340" type="Museum">ZSM</collectionCode>
).
<emphasis id="1CA71E45D437FFD0FE93FADC895E4E61" bold="true" box="[312,605,1372,1391]" pageId="9" pageNumber="340">
<collectingCountry id="56C482C7D437FFD0FE93FADC8AB04E61" box="[312,435,1372,1391]" name="Indonesia" pageId="9" pageNumber="340">INDONESIA</collectingCountry>
:
<collectingRegion id="EC170CB5D437FFD0FE10FADC8AE54E63" box="[443,486,1372,1391]" country="Indonesia" name="Kalimantan Timur" pageId="9" pageNumber="340">EAST</collectingRegion>
KALIMANTAN:
</emphasis>
Ca.
<quantity id="E92B6FB2D437FFD0FD2AFADC89B64E61" box="[641,693,1372,1391]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="340" unit="km" value="55.0">55 km</quantity>
W Balikpapan,PT Fajar Surya Swadaya [area], base camp surrounded with
<taxonomicName id="E9D3B9D4D437FFD0FECAFAF68A9F4E87" box="[353,412,1398,1417]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="340" phylum="Tracheophyta" rank="genus">
<emphasis id="1CA71E45D437FFD0FECAFAF68A9F4E87" box="[353,412,1398,1417]" italics="true" pageId="9" pageNumber="340">Acacia</emphasis>
</taxonomicName>
plantation,
<geoCoordinate id="4BE7A490D437FFD0FDA9FAF689544E87" box="[514,599,1398,1417]" degrees="01" direction="south" minutes="16.4" orientation="latitude" pageId="9" pageNumber="340" precision="92" value="-1.2733333">01°16.4S</geoCoordinate>
,
<geoCoordinate id="4BE7A490D437FFD0FDCAFAF689C14E87" box="[609,706,1398,1417]" degrees="116" direction="east" minutes="21.1" orientation="longitude" pageId="9" pageNumber="340" precision="92" value="116.35167">116°21.1E</geoCoordinate>
,
<quantity id="E92B6FB2D437FFD0FD60FAF689FB4E87" box="[715,760,1398,1417]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="340" unit="m" value="82.0">82 m</quantity>
,
<date id="5A6DE497D437FFD0FCAAFAF688974E87" box="[769,916,1398,1417]" pageId="9" pageNumber="340" value="2011-11-23" valueMax="2011-12-01" valueMin="2011-11-23">23.xi.1.xii.2011</date>
,
<specimenCount id="38D509DED437FFD0FC36FAF688D84E87" box="[925,987,1398,1417]" pageId="9" pageNumber="340" type="generic">1 spec.</specimenCount>
, (individual collecting on vegetation, dead wood and in puddles &amp; light trap), P. Votruba, J. Schneider &amp; J. Hájek leg. (
<collectionCode id="48C25A92D437FFD0FC5CFA118F3B4EAA" box="[1015,1080,1425,1444]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="340">NMPC</collectionCode>
).
</paragraph>
<paragraph id="2E6CC257D437FFD0FFDAFA3D8A4E4D1A" blockId="9.[113,1089,1469,1556]" pageId="9" pageNumber="340">
<emphasis id="1CA71E45D437FFD0FFDAFA3D8A7A4EDA" bold="true" box="[113,377,1469,1492]" pageId="9" pageNumber="340">Diagnostic description.</emphasis>
Body length
<quantity id="E92B6FB2D437FFD0FDBAFA3D89B14EDA" box="[529,690,1469,1492]" metricMagnitude="-3" metricUnit="m" metricValue="3.085" metricValueMax="3.35" metricValueMin="2.82" pageId="9" pageNumber="340" unit="mm" value="3.085" valueMax="3.35" valueMin="2.82">2.823.35 mm</quantity>
. Body width
<quantity id="E92B6FB2D437FFD0FCE3FA3D88EA4EDA" box="[840,1001,1469,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.505" metricValueMax="1.71" metricValueMin="1.3" pageId="9" pageNumber="340" unit="mm" value="1.505" valueMax="1.71" valueMin="1.3">1.301.71 mm</quantity>
. Ratios: Pronotum length / width 0.88-0.96. Elytra length / width 1.481.63; length elytra / length pronotum 2.63-2.89.
</paragraph>
<paragraph id="2E6CC257D434FFD3FF1EFF5C89B14A7D" blockId="10.[151,1130,220,787]" pageId="10" pageNumber="341">
<taxonomicName id="E9D3B9D4D434FFD3FF1EFF5C8AC34BFA" authorityName="Bremer" authorityYear="1991" box="[181,448,220,244]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FF1EFF5C8AC34BFA" box="[181,448,220,244]" italics="true" pageId="10" pageNumber="341">Paragonocnemis ardoini</emphasis>
</taxonomicName>
presents yellow to light brown ground color on upper side and underside; on each elytron there is dark, transverse macula which does not cover median two and lateral three intervals. Inner and outer sides of mesotibiae are mostly markedly bent, in some specimens from
<collectingCountry id="56C482C7D434FFD3FE0CFEBC89054A5D" box="[423,518,316,339]" name="Thailand" pageId="10" pageNumber="341">Thailand</collectingCountry>
only outer side of mesotibiae is bent while inner side is straight. Antennae reach end of basal 1/4 of elytra.
</paragraph>
</subSubSection>
<subSubSection id="66C991DCD434FFD3FF3CFEFC899D481D" pageId="10" pageNumber="341" type="discussion">
<paragraph id="2E6CC257D434FFD3FF3CFEFC899D481D" blockId="10.[151,1130,220,787]" pageId="10" pageNumber="341">
<emphasis id="1CA71E45D434FFD3FF3CFEFC8A054A9D" bold="true" box="[151,262,380,403]" pageId="10" pageNumber="341">Remarks.</emphasis>
This species has a wide distribution, ranging from north and central
<collectingCountry id="56C482C7D434FFD3FC78FEFC8F314A9D" box="[979,1074,380,403]" name="Thailand" pageId="10" pageNumber="341">Thailand</collectingCountry>
over Sumatra to Borneo (the type series of
<taxonomicName id="E9D3B9D4D434FFD3FD89FE1C898A4ABA" authorityName="Bremer" authorityYear="1991" box="[546,649,412,436]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FD89FE1C898A4ABA" box="[546,649,412,436]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
is from central
<collectingCountry id="56C482C7D434FFD3FC80FE1C888E4ABD" box="[811,909,412,435]" name="Thailand" pageId="10" pageNumber="341">Thailand</collectingCountry>
, the description of
<taxonomicName id="E9D3B9D4D434FFD3FBF8FE1C8BF44ADA" authorityName="Bremer" authorityYear="1998" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="1CA71E45D434FFD3FBF8FE1C8BF44ADA" italics="true" pageId="10" pageNumber="341">P. velutinus</emphasis>
</taxonomicName>
is based on a single specimen from Borneo, the
<typeStatus id="F1687CF5D434FFD3FD42FE3C88484ADD" box="[745,843,444,467]" pageId="10" pageNumber="341" type="holotype">holotype</typeStatus>
). Most specimens formerly identified as
<taxonomicName id="E9D3B9D4D434FFD3FEB1FE5C8A7C4AFA" authorityName="Bremer" authorityYear="1991" box="[282,383,476,500]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FEB1FE5C8A7C4AFA" box="[282,383,476,500]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
are smaller (body length
<quantity id="E92B6FB2D434FFD3FD28FE5C88184AFD" box="[643,795,476,499]" metricMagnitude="-3" metricUnit="m" metricValue="2.965" metricValueMax="3.11" metricValueMin="2.82" pageId="10" pageNumber="341" unit="mm" value="2.965" valueMax="3.11" valueMin="2.82">2.823.11 mm</quantity>
) than the specimens determined as
<taxonomicName id="E9D3B9D4D434FFD3FF18FE7C8A2F491A" authorityName="Bremer" authorityYear="1998" box="[179,300,508,532]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="1CA71E45D434FFD3FF18FE7C8A2F491A" box="[179,300,508,532]" italics="true" pageId="10" pageNumber="341">P. velutinus</emphasis>
</taxonomicName>
later on. In the original description of
<taxonomicName id="E9D3B9D4D434FFD3FD6AFE7C8839491A" authorityName="Bremer" authorityYear="1998" box="[705,826,508,532]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="1CA71E45D434FFD3FD6AFE7C8839491A" box="[705,826,508,532]" italics="true" pageId="10" pageNumber="341">P. velutinus</emphasis>
</taxonomicName>
it was stated that in addition to the larger size the mesotibiae are more bent than in
<taxonomicName id="E9D3B9D4D434FFD3FD42FD9C8850493A" authorityName="Bremer" authorityYear="1991" box="[745,851,540,564]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FD42FD9C8850493A" box="[745,851,540,564]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
, and the antennomere XI is allusively longer in
<taxonomicName id="E9D3B9D4D434FFD3FE23FDBC8AF2495A" authorityName="Bremer" authorityYear="1991" box="[392,497,572,596]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FE23FDBC8AF2495A" box="[392,497,572,596]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
. After seeing more specimens from Borneo, but also from
<collectingCountry id="56C482C7D434FFD3FF3CFDDC8BF5497D" box="[151,246,604,627]" name="Thailand" pageId="10" pageNumber="341">Thailand</collectingCountry>
and Sumatra, we are no longer convinced that
<taxonomicName id="E9D3B9D4D434FFD3FD5AFDDC886F497A" authorityName="Bremer" authorityYear="1998" box="[753,876,604,628]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="1CA71E45D434FFD3FD5AFDDC886F497A" box="[753,876,604,628]" italics="true" pageId="10" pageNumber="341">P. velutinus</emphasis>
</taxonomicName>
is really different from
<taxonomicName id="E9D3B9D4D434FFD3FF3CFDFC8A03499A" authorityName="Bremer" authorityYear="1991" box="[151,256,636,660]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FF3CFDFC8A03499A" box="[151,256,636,660]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
. Most specimens of
<taxonomicName id="E9D3B9D4D434FFD3FE4BFDFC894A499A" authorityName="Bremer" authorityYear="1991" box="[480,585,636,660]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FE4BFDFC894A499A" box="[480,585,636,660]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
from
<collectingCountry id="56C482C7D434FFD3FD20FDFC89E9499D" box="[651,746,636,659]" name="Thailand" pageId="10" pageNumber="341">Thailand</collectingCountry>
are indeed smaller than those from Borneo (specimens from Sumatra are mostly in between); however, the other characters are not so obviously different that one may assign specimens to one or the other taxon. Therefore, we can no longer state with certainty that
<taxonomicName id="E9D3B9D4D434FFD3FDE1FD5C89B249FA" authorityName="Bremer" authorityYear="1991" box="[586,689,732,756]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="ardoini">
<emphasis id="1CA71E45D434FFD3FDE1FD5C89B249FA" box="[586,689,732,756]" italics="true" pageId="10" pageNumber="341">P. ardoini</emphasis>
</taxonomicName>
and
<taxonomicName id="E9D3B9D4D434FFD3FD49FD5C885849FA" authorityName="Bremer" authorityYear="1998" box="[738,859,732,756]" class="Insecta" family="Tenebrionidae" genus="Paragonocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="1CA71E45D434FFD3FD49FD5C885849FA" box="[738,859,732,756]" italics="true" pageId="10" pageNumber="341">P. velutinus</emphasis>
</taxonomicName>
are different species, and we therefore decide to synonymize both species.
</paragraph>
</subSubSection>
</treatment>
</document>