The species of Gonocnemis and Paragonocnemis from Borneo with notes on synonymy in the Oriental species (Coleoptera: Tenebrionidae)
Author
Bremer, Hans J.
) &) Diakonie Wohnstift am Westerberg, Bergstrasse 35 B, 49076 Osnabrück, Germany; e-mail: hjbremer @ live. de
Author
Grimm, Roland
) &) Unterer Sägerweg 74, 75305 Neuenbürg, Germany; e-mail: grimm. tenebrio @ t-online. de
text
Acta Entomologica Musei Nationalis Pragae
2017
Acta. Ent. Mus. Natl. Pragae
2017-12-31
57
2
331
343
http://dx.doi.org/10.1515/aemnp-2017-0079
journal article
7769
10.1515/aemnp-2017-0079
20436c9c-2b03-46c4-b53c-b14cc674f8e3
0374-1036
4504000
9FB0A4B6-2876-4331-A235-0B5A3781BF07
Paragonocnemis
(
Borneogonocnemis
)
ardoini
Bremer, 1991
(Figs 6a, 6b)
Paragonocnemis
(
Borneogonocnemis
)
ardoini
Bremer, 1991: 151–153
(original description).
Paragonocnemis
(
Borneogonocnemis
)
velutinus
Bremer, 1998: 54–56
(original description),
syn. nov.
Type material examined.
Paragonocnemis
(
Borneogonocnemis
)
velutinus
:
HOLOTYPE
:
Borneo
:
Sarawak
,
Belaga
,
Long Linau
,
17.–21.iii.1990
,
A. Riedel
leg. (
SMNS
).
Additional material examined.
MALAYSIA
:
SARAWAK
:
Santubong Peninsula, Permai Rainforest Resort,
10–200 m
,
11.–14.ix.2008
,
2 spec.
(at light),
30–150 m
,
10.–14.ii.2012
,
1 spec.
(at light),
15.–21.iv.2016
,
1 spec.
,
R
. Grimm leg.(3
RGCN
, 1
ZSM
).
INDONESIA
:
EAST
KALIMANTAN:
Ca.
55 km
W Balikpapan,PT Fajar Surya Swadaya [area], base camp surrounded with
Acacia
plantation,
01°16.4′S
,
116°21.1′E
,
82 m
,
23.xi.–1.xii.2011
,
1 spec.
, (individual collecting on vegetation, dead wood and in puddles & light trap), P. Votruba, J. Schneider & J. Hájek leg. (
NMPC
).
Diagnostic description.
Body length
2.82–3.35 mm
. Body width
1.30–1.71 mm
. Ratios: Pronotum length / width 0.88-0.96. Elytra length / width 1.48–1.63; length elytra / length pronotum 2.63-2.89.
Paragonocnemis ardoini
presents yellow to light brown ground color on upper side and underside; on each elytron there is dark, transverse macula which does not cover median two and lateral three intervals. Inner and outer sides of mesotibiae are mostly markedly bent, in some specimens from
Thailand
only outer side of mesotibiae is bent while inner side is straight. Antennae reach end of basal 1/4 of elytra.
Remarks.
This species has a wide distribution, ranging from north and central
Thailand
over Sumatra to Borneo (the type series of
P. ardoini
is from central
Thailand
, the description of
P. velutinus
is based on a single specimen from Borneo, the
holotype
). Most specimens formerly identified as
P. ardoini
are smaller (body length
2.82–3.11 mm
) than the specimens determined as
P. velutinus
later on. In the original description of
P. velutinus
it was stated that in addition to the larger size the mesotibiae are more bent than in
P. ardoini
, and the antennomere XI is allusively longer in
P. ardoini
. After seeing more specimens from Borneo, but also from
Thailand
and Sumatra, we are no longer convinced that
P. velutinus
is really different from
P. ardoini
. Most specimens of
P. ardoini
from
Thailand
are indeed smaller than those from Borneo (specimens from Sumatra are mostly in between); however, the other characters are not so obviously different that one may assign specimens to one or the other taxon. Therefore, we can no longer state with certainty that
P. ardoini
and
P. velutinus
are different species, and we therefore decide to synonymize both species.