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<document id="A1999D7A92E4E6EFB7591EA81121A6D6" ID-CLB-Dataset="121554" ID-DOI="10.11646/zootaxa.5174.4.6" ID-GBIF-Dataset="b009d163-c746-470f-a3d7-95dbc8726627" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986469" ID-ZooBank="196884A6-1AF7-4C16-B2C5-D05C7F78E1CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1660327463178" checkinUser="plazi" docAuthor="Kiałka, Agata &amp; Ruta, Rafał" docDate="2022" docId="A0384152FFBC02164DE7FD91FA05FC21" docLanguage="en" docName="zootaxa.5174.4.6.pdf" docOrigin="Zootaxa 5174 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atopida White" docType="key" docVersion="5" lastPageNumber="438" masterDocId="5C01392AFF9802334D70FFB3FF9BFFAA" masterDocTitle="Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae)" masterLastPageNumber="443" masterPageNumber="401" pageNumber="437" updateTime="1699367011641" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="261295F5FE18DE18942719B63747C4DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="39620BB7C6C967E8AE92A824C41B2265">
<mods:title id="8D090E662B8D18B75B4742FC54E5FAE1">Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae)</mods:title>
</mods:titleInfo>
<mods:name id="C8144932234813E9783CCD6E47871FEE" type="personal">
<mods:role id="FB6CBEA89321DE22CCCE4BE441BB1E8B">
<mods:roleTerm id="69DBBB39A8ADA0127FD987A90D1561DD">Author</mods:roleTerm>
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<mods:namePart id="5501C397A070B5A4E7465BE63EEBFFDA">Kiałka, Agata</mods:namePart>
<mods:nameIdentifier id="A6B90D2870E770653E9E85E317995B5A" type="ORCID">0000-0001-8883-9035</mods:nameIdentifier>
<mods:affiliation id="FCB6C04ECAD92758248B57DC40DEEE68">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. &amp; agata. kialka @ gmail. com; https: // orcid. org / 0000 - 0001 - 8883 - 9035</mods:affiliation>
<mods:nameIdentifier id="D252DD3D9F9509A1196C9ACA972F5ECB" type="email">agata.kialka@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1DE82FD66ECEBAFFB97BF3D43A4A796A">Ruta, Rafał</mods:namePart>
<mods:nameIdentifier id="682B85FAE2708C9BFA05F6EA84760567" type="ORCID">0000-0001-8515-2385</mods:nameIdentifier>
<mods:affiliation id="031DDBA9DA1B5993DDE9970873DF3A22">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. &amp; rafal. ruta @ uwr. edu. pl; https: // orcid. org / 0000 - 0001 - 8515 - 2385</mods:affiliation>
<mods:nameIdentifier id="F7A74F0092AD9024B1D8407E827DDE00" type="email">rafal.ruta@uwr.edu.pl</mods:nameIdentifier>
</mods:name>
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<mods:title id="8B1A123FA3E45C69CC8EE7F308ED77B4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7E8A4C139653D3DDF413F7B271F08B68">
<mods:date id="38974C8E60C67AF4C05266E2709EA695">2022</mods:date>
<mods:detail id="02B2F357CF7CE3FC2751BF0FFD80EA0C" type="pubDate">
<mods:number id="DB1C4D8EA8A79F0C79EB28C020DB3897">2022-08-11</mods:number>
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<mods:number id="E45D32ABD28BE9DB72C47BF1174ED978">5174</mods:number>
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<mods:identifier id="A592A29ACA71183A1A64278CEC0361D4" type="DOI">10.11646/zootaxa.5174.4.6</mods:identifier>
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<treatment id="A0384152FFBC02164DE7FD91FA05FC21" ID-DOI="http://doi.org/10.5281/zenodo.6993104" ID-GBIF-Taxon="199094420" ID-Zenodo-Dep="6993104" LSID="urn:lsid:plazi:treatment:A0384152FFBC02164DE7FD91FA05FC21" httpUri="http://treatment.plazi.org/id/A0384152FFBC02164DE7FD91FA05FC21" lastPageId="37" lastPageNumber="438" pageId="36" pageNumber="437">
<subSubSection id="608BA3CFFFBC02174DE7FD91FE79FD97" box="[151,482,546,573]" pageId="36" pageNumber="437" type="nomenclature">
<paragraph id="282EF044FFBC02174DE7FD91FE79FD97" blockId="36.[151,482,546,573]" box="[151,482,546,573]" pageId="36" pageNumber="437">
<heading id="73664728FFBC02174DE7FD91FE79FD97" bold="true" box="[151,482,546,573]" fontSize="11" level="1" pageId="36" pageNumber="437" reason="1">
<emphasis id="1AE52C56FFBC02174DE7FD91FE79FD97" bold="true" box="[151,482,546,573]" pageId="36" pageNumber="437">
A key to species of
<taxonomicName id="EF918BC7FFBC02174CF3FD91FE79FD97" authority="White" authorityName="White" box="[387,482,546,573]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="1AE52C56FFBC02174CF3FD91FE79FD97" bold="true" box="[387,482,546,573]" italics="true" pageId="36" pageNumber="437">Atopida</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="608BA3CFFFBC02174DE7FDD8FBC7FC93" pageId="36" pageNumber="437" type="multiple">
<paragraph id="282EF044FFBC02174DE7FDD8FBC7FC93" blockId="36.[151,1437,619,825]" pageId="36" pageNumber="437">
It is relatively difficult to identify species of
<taxonomicName id="EF918BC7FFBC02174FFEFDD8FD7CFD2F" box="[654,743,619,645]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="1AE52C56FFBC02174FFEFDD8FD7CFD2F" box="[654,743,619,645]" italics="true" pageId="36" pageNumber="437">Atopida</emphasis>
</taxonomicName>
. The main sources of problems with identification of
<taxonomicName id="EF918BC7FFBC02174834FDD8FA06FD2F" box="[1348,1437,619,645]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="1AE52C56FFBC02174834FDD8FA06FD2F" box="[1348,1437,619,645]" italics="true" pageId="36" pageNumber="437">Atopida</emphasis>
</taxonomicName>
are: significant sexual dimorphism, large infraspecific variability, similarity of male genitalia of several species groups, and the fact that bursal sclerites of females of several species are not very distinctive, often rod-like. With the following key, it is possible to identify males of all known species and females of most species (females of undescribed species of the
<taxonomicName id="EF918BC7FFBC02174CB6FD48FDB3FCBE" authorityName=", Broun" authorityYear="1921" box="[454,552,763,788]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="1AE52C56FFBC02174CB6FD48FDB3FCBE" box="[454,552,763,788]" italics="true" pageId="36" pageNumber="437">montana</emphasis>
</taxonomicName>
complex are included). In most cases dissection of genitalia is needed, as well as examination of abdominal setal patches, bursal sclerites and hind wings of females.
</paragraph>
</subSubSection>
<subSubSection id="608BA3CFFFBC02164DE7FCD7FA05FC21" lastPageId="37" lastPageNumber="438" pageId="36" pageNumber="437" type="key">
<key id="5C00A775FFBC02164DE7FCD7FA05FC21" lastPageId="37" lastPageNumber="438" pageId="36" pageNumber="437">
<keyStep id="9365E8E1FFBC02174DE7FCD7FA07FC19" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FCD7FA07FC3D" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FCD7FA07FC3D" pageId="36" pageNumber="437">
1. Head and pronotum covered with adherent setae, in lateral view no semierect setae on head and pronotum can be noticed..............................................................................................
<taxonomicName id="EF918BC7FFBC02174856FC32FA07FC3D" authority="Sharp" authorityName="Sharp" authorityYear="1878" box="[1318,1436,897,919]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="proba">
<emphasis id="1AE52C56FFBC02174856FC32FAC5FC3C" box="[1318,1374,897,918]" italics="true" pageId="36" pageNumber="437">proba</emphasis>
Sharp
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FC2CFA07FC19" blockId="36.[151,1437,868,2034]" box="[151,1436,926,948]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FC2CFA07FC19" box="[151,1436,926,948]" pageId="36" pageNumber="437">-. At least head and pronotum (usually also elytra) covered with erect setae, clearly visible in lateral view................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FC0FFA07FC44" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FC0FFA07FC7B" blockId="36.[151,1437,868,2034]" box="[151,1436,956,978]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FC0FFA07FC7B" box="[151,1436,956,978]" pageId="36" pageNumber="437">2. Male............................................................................................... 3</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FC6AFA07FC44" blockId="36.[151,1437,868,2034]" box="[151,1436,985,1007]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FC6AFA07FC44" box="[151,1436,985,1007]" pageId="36" pageNumber="437">-. Female............................................................................................ 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FC45FA07FB83" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FC45FA07FBA6" blockId="36.[151,1437,868,2034]" box="[151,1436,1014,1036]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FC45FA07FBA6" box="[151,1436,1014,1036]" pageId="36" pageNumber="437">
3. Punctation of pronotum not granulate.........................................................
<taxonomicName id="EF918BC7FFBC02174874FC44FA07FBA6" authority="Broun" authorityName="Broun" authorityYear="1914" box="[1284,1436,1015,1036]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="impressa">
<emphasis id="1AE52C56FFBC02174874FC44FAC1FBA6" box="[1284,1370,1015,1036]" italics="true" pageId="36" pageNumber="437">impressa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FBA7FA07FB83" blockId="36.[151,1437,868,2034]" box="[151,1436,1044,1066]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FBA7FA07FB83" box="[151,1436,1044,1066]" pageId="36" pageNumber="437">-. Punctation of pronotum granulate, granulation easily noticeable at least on lateral portions of pronotum................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FB82FA07FBCE" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FB82FA07FBEC" blockId="36.[151,1437,868,2034]" box="[151,1436,1073,1095]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB82FA07FBEC" box="[151,1436,1073,1095]" pageId="36" pageNumber="437">
4. Trigonium of penis very wide, bulbous (
<figureCitation id="B0AAECC1FFBC02174F15FB82FD20FBED" box="[613,699,1073,1095]" captionStart="FIGURE 15" captionStartId="26.[151,250,1934,1958]" captionTargetBox="[151,1436,777,1910]" captionTargetId="figure-332@26.[151,1436,777,1910]" captionTargetPageId="26" captionText="FIGURE 15. Atopida, male genitalia, penis (left), tegmen (right). A) A. proba, B) A. suffusa, C) A. suturalis, D) A. tuhua sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986533" httpUri="https://zenodo.org/record/6986533/files/figure.png" pageId="36" pageNumber="437">Fig. 15B</figureCitation>
).................................................
<taxonomicName id="EF918BC7FFBC02174866FB82FA07FBEC" authority="Broun" authorityName="Broun" authorityYear="1910" box="[1302,1436,1073,1094]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="1AE52C56FFBC02174866FB82FAC1FBEC" box="[1302,1370,1073,1094]" italics="true" pageId="36" pageNumber="437">suffusa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FBFCFA07FBCE" blockId="36.[151,1437,868,2034]" box="[151,1436,1102,1124]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FBFCFA07FBCE" box="[151,1436,1102,1124]" pageId="36" pageNumber="437">-. Trigonium never bulbous, usually narrow with subparallel sides or triangular...................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FBD8FA07FB34" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FBD8FA07FB28" blockId="36.[151,1437,868,2034]" box="[151,1436,1131,1154]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FBD8FA07FB28" box="[151,1436,1131,1154]" pageId="36" pageNumber="437">5. Pronotum widest in the middle of its length................................................................. 6</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FB3AFA07FB34" blockId="36.[151,1437,868,2034]" box="[151,1436,1161,1183]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB3AFA07FB34" box="[151,1436,1161,1183]" pageId="36" pageNumber="437">-. Pronotum widest before the middle of its length............................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FB15FA07FB73" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FB15FA07FB11" blockId="36.[151,1437,868,2034]" box="[151,1436,1190,1212]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB15FA07FB11" box="[151,1436,1190,1212]" pageId="36" pageNumber="437">
6. Trigonium shorter than parameroids (
<figureCitation id="B0AAECC1FFBC02174F39FB15FD3AFB16" box="[585,673,1190,1212]" captionStart="FIGURE 16" captionStartId="33.[151,250,1982,2006]" captionTargetBox="[165,1421,813,1959]" captionTargetId="figure-287@33.[165,1421,813,1959]" captionTargetPageId="33" captionText="FIGURE 16. Atopida, male genitalia, penis (left), tegmen (right).A) A. villosa, B) A. waipoua sp. nov., C) A. walkerae sp. nov., D) A. westlandensis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986535" httpUri="https://zenodo.org/record/6986535/files/figure.png" pageId="36" pageNumber="437">Fig. 16B</figureCitation>
), occurs in North Island..............................
<taxonomicName id="EF918BC7FFBC02174987FB14FAD1FB16" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1271,1354,1191,1212]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="waipoua" status="sp. nov.">
<emphasis id="1AE52C56FFBC02174987FB14FAD1FB16" box="[1271,1354,1191,1212]" italics="true" pageId="36" pageNumber="437">waipoua</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820FB15FA07FB11" bold="true" box="[1360,1436,1190,1211]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820FB15FA07FB11" box="[1360,1436,1190,1211]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FB77FA07FB73" blockId="36.[151,1437,868,2034]" box="[151,1436,1220,1242]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB77FA07FB73" box="[151,1436,1220,1242]" pageId="36" pageNumber="437">
-. Trigonium as long as parameroids (
<figureCitation id="B0AAECC1FFBC02174F46FB77FD15FB70" box="[566,654,1220,1242]" captionStart="FIGURE 16" captionStartId="33.[151,250,1982,2006]" captionTargetBox="[165,1421,813,1959]" captionTargetId="figure-287@33.[165,1421,813,1959]" captionTargetPageId="33" captionText="FIGURE 16. Atopida, male genitalia, penis (left), tegmen (right).A) A. villosa, B) A. waipoua sp. nov., C) A. walkerae sp. nov., D) A. westlandensis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986535" httpUri="https://zenodo.org/record/6986535/files/figure.png" pageId="36" pageNumber="437">Fig. 16C</figureCitation>
), occurs in South Island...............................
<taxonomicName id="EF918BC7FFBC02174982FB77FAD1FB73" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1266,1354,1220,1241]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="walkerae" status="sp. nov.">
<emphasis id="1AE52C56FFBC02174982FB77FAD1FB73" box="[1266,1354,1220,1241]" italics="true" pageId="36" pageNumber="437">walkerae</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820FB77FA07FB73" bold="true" box="[1360,1436,1220,1241]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820FB77FA07FB73" box="[1360,1436,1220,1241]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FB52FA07FAB9" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FB52FA07FB5C" blockId="36.[151,1437,868,2034]" box="[151,1436,1249,1271]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB52FA07FB5C" box="[151,1436,1249,1271]" pageId="36" pageNumber="437">7. Lateral carinae of elytra crenulate, in most species penis with explanate lateral lobes............................... 8</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FB4CFA07FAB9" blockId="36.[151,1437,868,2034]" box="[151,1436,1278,1300]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FB4CFA07FAB9" box="[151,1436,1278,1300]" pageId="36" pageNumber="437">-. Lateral carinae of elytra not crenulate, penis always without explanate lateral lobes................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FAAFFA07FAE4" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FAAFFA07FA9B" blockId="36.[151,1437,868,2034]" box="[151,1436,1308,1330]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FAAFFA07FA9B" box="[151,1436,1308,1330]" pageId="36" pageNumber="437">
8. Trigonium straight, triangular (e.g.
<figureCitation id="B0AAECC1FFBC02174F46FAAFFD33FA9B" box="[566,680,1308,1330]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13A, C</figureCitation>
)............................................................. 9
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FA8AFA07FAE4" blockId="36.[151,1437,868,2034]" box="[151,1436,1337,1359]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FA8AFA07FAE4" box="[151,1436,1337,1359]" pageId="36" pageNumber="437">
-. Trigonium curved (e.g.
<figureCitation id="B0AAECC1FFBC02174CB6FA8AFD87FAE4" box="[454,540,1337,1359]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13D</figureCitation>
)........................................................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FAE5FA07FA69" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FAE5FA07FA23" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FAE5FA07FA23" pageId="36" pageNumber="437">
9. Sides of pronotum curved, projections of supraantennal ridges long (
<figureCitation id="B0AAECC1FFBC02174E09FAE5FC58FAC1" box="[889,963,1366,1388]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="36" pageNumber="437">Fig. 3E</figureCitation>
), parameroids rounded at apices, outer edges not denticulate (
<figureCitation id="B0AAECC1FFBC02174C28FAC7FE34FA23" box="[344,431,1396,1418]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13A</figureCitation>
).....................................................................
<taxonomicName id="EF918BC7FFBC0217498BFAC7FAD1FA23" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1275,1354,1396,1417]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="1AE52C56FFBC0217498BFAC7FAD1FA23" box="[1275,1354,1396,1417]" italics="true" pageId="36" pageNumber="437">lescheni</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820FAC7FA07FA23" bold="true" box="[1360,1436,1396,1417]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820FAC7FA07FA23" box="[1360,1436,1396,1417]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7FA22FA07FA69" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FA22FA07FA69" pageId="36" pageNumber="437">
-. Sides of pronotum bisinuate, projections of supraantennal ridges elevated, but not forming projections, parameroids pointed at apices, outer edges denticulate (
<figureCitation id="B0AAECC1FFBC02174F64FA1DFDF1FA69" box="[532,618,1454,1476]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13C</figureCitation>
).....................................................
<taxonomicName id="EF918BC7FFBC02174989FA1CFAD2FA6E" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1273,1353,1455,1476]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
<emphasis id="1AE52C56FFBC02174989FA1CFAD2FA6E" box="[1273,1353,1455,1476]" italics="true" pageId="36" pageNumber="437">paparoa</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820FA1DFA07FA69" bold="true" box="[1360,1436,1454,1475]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820FA1DFA07FA69" box="[1360,1436,1454,1475]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7FA7FFA07F9B1" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7FA7FFA07FA54" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7FA7FFA07FA54" pageId="36" pageNumber="437">
10. Apices of parameroids widened, rounded; lateral plates of penis with denticles in apical portion (
<figureCitation id="B0AAECC1FFBC021749F3FA7FFB41FA4B" box="[1155,1242,1484,1506]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13D</figureCitation>
)......................................................................................................
<taxonomicName id="EF918BC7FFBC0217498EFA59FAD1FA55" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1278,1354,1514,1535]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="paringa" status="sp. nov.">
<emphasis id="1AE52C56FFBC0217498EFA59FAD1FA55" box="[1278,1354,1514,1535]" italics="true" pageId="36" pageNumber="437">paringa</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820FA5AFA07FA54" bold="true" box="[1360,1436,1513,1534]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820FA5AFA07FA54" box="[1360,1436,1513,1534]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F9B4FA07F9B1" blockId="36.[151,1437,868,2034]" box="[151,1436,1542,1564]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F9B4FA07F9B1" box="[151,1436,1542,1564]" pageId="36" pageNumber="437">
-. Apices of parameroids triangular, pointed, lateral plates of penis without denticles in apical portion (
<figureCitation id="B0AAECC1FFBC021749D5F9B5FA9AF9B1" box="[1189,1281,1542,1564]" captionStart="FIGURE 12" captionStartId="14.[151,250,1955,1979]" captionTargetBox="[151,1436,832,1932]" captionTargetId="figure-35@14.[151,1436,832,1932]" captionTargetPageId="14" captionText="FIGURE 12. Atopida, male genitalia, penis (left), tegmen (right). A) A. castanea, B) A. hirta, C) A. impressa, D) A. insularis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986513" httpUri="https://zenodo.org/record/6986513/files/figure.png" pageId="36" pageNumber="437">Figs 12D</figureCitation>
,
<figureCitation id="B0AAECC1FFBC0217487BF9B5FAAEF9B6" box="[1291,1333,1542,1564]" captionStart="FIGURE 15" captionStartId="26.[151,250,1934,1958]" captionTargetBox="[151,1436,777,1910]" captionTargetId="figure-332@26.[151,1436,777,1910]" captionTargetPageId="26" captionText="FIGURE 15. Atopida, male genitalia, penis (left), tegmen (right). A) A. proba, B) A. suffusa, C) A. suturalis, D) A. tuhua sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986533" httpUri="https://zenodo.org/record/6986533/files/figure.png" pageId="36" pageNumber="437">15D</figureCitation>
)..... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F997FA07F9FC" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F997FA07F993" blockId="36.[151,1437,868,2034]" box="[151,1436,1572,1594]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F997FA07F993" box="[151,1436,1572,1594]" pageId="36" pageNumber="437">
11. Parameres narrow, triangular (
<figureCitation id="B0AAECC1FFBC02174F75F997FDC0F993" box="[517,603,1572,1594]" captionStart="FIGURE 12" captionStartId="14.[151,250,1955,1979]" captionTargetBox="[151,1436,832,1932]" captionTargetId="figure-35@14.[151,1436,832,1932]" captionTargetPageId="14" captionText="FIGURE 12. Atopida, male genitalia, penis (left), tegmen (right). A) A. castanea, B) A. hirta, C) A. impressa, D) A. insularis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986513" httpUri="https://zenodo.org/record/6986513/files/figure.png" pageId="36" pageNumber="437">Fig. 12D</figureCitation>
)......................................................
<taxonomicName id="EF918BC7FFBC02174985F997FAD2F993" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1269,1353,1572,1593]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
<emphasis id="1AE52C56FFBC02174985F997FAD2F993" box="[1269,1353,1572,1593]" italics="true" pageId="36" pageNumber="437">insularis</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820F997FA07F993" bold="true" box="[1360,1436,1572,1593]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820F997FA07F993" box="[1360,1436,1572,1593]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F9F2FA07F9FC" blockId="36.[151,1437,868,2034]" box="[151,1436,1601,1623]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F9F2FA07F9FC" box="[151,1436,1601,1623]" pageId="36" pageNumber="437">
-. Parameres wide, subrectangular (
<figureCitation id="B0AAECC1FFBC02174F51F9F2FDE3F9FD" box="[545,632,1601,1623]" captionStart="FIGURE 15" captionStartId="26.[151,250,1934,1958]" captionTargetBox="[151,1436,777,1910]" captionTargetId="figure-332@26.[151,1436,777,1910]" captionTargetPageId="26" captionText="FIGURE 15. Atopida, male genitalia, penis (left), tegmen (right). A) A. proba, B) A. suffusa, C) A. suturalis, D) A. tuhua sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986533" httpUri="https://zenodo.org/record/6986533/files/figure.png" pageId="36" pageNumber="437">Fig. 15D</figureCitation>
)......................................................
<taxonomicName id="EF918BC7FFBC02174863F9F2FAD1F9FC" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1299,1354,1601,1622]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="tuhua" status="sp. nov.">
<emphasis id="1AE52C56FFBC02174863F9F2FAD1F9FC" box="[1299,1354,1601,1622]" italics="true" pageId="36" pageNumber="437">tuhua</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820F9F2FA07F9FC" bold="true" box="[1360,1436,1601,1622]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820F9F2FA07F9FC" box="[1360,1436,1601,1622]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F9EDFA07F93B" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F9EDFA07F9D9" blockId="36.[151,1437,868,2034]" box="[151,1436,1630,1652]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F9EDFA07F9D9" box="[151,1436,1630,1652]" pageId="36" pageNumber="437">12. Elytra covered with adherent setae....................................................................... 13</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F9CFFA07F93B" blockId="36.[151,1437,868,2034]" box="[151,1436,1660,1682]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F9CFFA07F93B" box="[151,1436,1660,1682]" pageId="36" pageNumber="437">-. Elytra covered with erect setae.......................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F92AFA06F966" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F92AFA06F905" blockId="36.[151,1437,868,2034]" box="[151,1437,1689,1711]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F92AFA06F905" box="[151,1437,1689,1711]" pageId="36" pageNumber="437">
13. Lateral carinae of pronotum strongly curved, penis long (
<figureCitation id="B0AAECC1FFBC02174E77F92AFCC5F904" box="[775,862,1689,1711]" captionStart="FIGURE 12" captionStartId="14.[151,250,1955,1979]" captionTargetBox="[151,1436,832,1932]" captionTargetId="figure-35@14.[151,1436,832,1932]" captionTargetPageId="14" captionText="FIGURE 12. Atopida, male genitalia, penis (left), tegmen (right). A) A. castanea, B) A. hirta, C) A. impressa, D) A. insularis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986513" httpUri="https://zenodo.org/record/6986513/files/figure.png" pageId="36" pageNumber="437">Fig. 12A</figureCitation>
)...................................
<taxonomicName id="EF918BC7FFBC02174876F929FA06F905" authority="White" authorityName="White" authorityYear="1846" box="[1286,1437,1689,1711]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="1AE52C56FFBC02174876F929FAC0F905" box="[1286,1371,1690,1711]" italics="true" pageId="36" pageNumber="437">castanea</emphasis>
White
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F904FA06F966" blockId="36.[151,1437,868,2034]" box="[151,1437,1718,1740]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F904FA06F966" box="[151,1437,1718,1740]" pageId="36" pageNumber="437">
-. Lateral carinae of pronotum subtly curved, penis less elongated, widest in the middle (
<figureCitation id="B0AAECC1FFBC0217496FF905FBEFF966" box="[1055,1140,1718,1740]" captionStart="FIGURE 15" captionStartId="26.[151,250,1934,1958]" captionTargetBox="[151,1436,777,1910]" captionTargetId="figure-332@26.[151,1436,777,1910]" captionTargetPageId="26" captionText="FIGURE 15. Atopida, male genitalia, penis (left), tegmen (right). A) A. proba, B) A. suffusa, C) A. suturalis, D) A. tuhua sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986533" httpUri="https://zenodo.org/record/6986533/files/figure.png" pageId="36" pageNumber="437">Fig. 15C</figureCitation>
).........
<taxonomicName id="EF918BC7FFBC02174988F905FA06F966" authority="(White)" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[1272,1437,1718,1740]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="1AE52C56FFBC02174988F905FAD7F961" box="[1272,1356,1718,1739]" italics="true" pageId="36" pageNumber="437">suturalis</emphasis>
(White)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F967FA07F8AD" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F967FA07F943" blockId="36.[151,1437,868,2034]" box="[151,1436,1747,1770]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F967FA07F943" box="[151,1436,1747,1770]" pageId="36" pageNumber="437">
14. Body on average smaller (TL
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), narrower........................................................ 15
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F942FA07F8AD" blockId="36.[151,1437,868,2034]" box="[151,1436,1777,1799]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F942FA07F8AD" box="[151,1436,1777,1799]" pageId="36" pageNumber="437">
-. Body on average larger (
<quantity id="EF695DA1FFBC02174CBEF942FDDCF8AC" box="[462,583,1777,1799]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.4" metricValueMin="4.2" pageId="36" pageNumber="437" unit="mm" value="4.8" valueMax="5.4" valueMin="4.2">4.2–5.4 mm</quantity>
), stouter.............................................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F8BDFA07F8EB" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F8BDFA07F88E" blockId="36.[151,1437,868,2034]" box="[151,1436,1806,1828]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F8BDFA07F88E" box="[151,1436,1806,1828]" pageId="36" pageNumber="437">
15. Penis very elongated (
<figureCitation id="B0AAECC1FFBC02174CC7F8BDFD95F889" box="[439,526,1806,1828]" captionStart="FIGURE 12" captionStartId="14.[151,250,1955,1979]" captionTargetBox="[151,1436,832,1932]" captionTargetId="figure-35@14.[151,1436,832,1932]" captionTargetPageId="14" captionText="FIGURE 12. Atopida, male genitalia, penis (left), tegmen (right). A) A. castanea, B) A. hirta, C) A. impressa, D) A. insularis sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986513" httpUri="https://zenodo.org/record/6986513/files/figure.png" pageId="36" pageNumber="437">Fig. 12B</figureCitation>
), species restricted to North Island......................................
<taxonomicName id="EF918BC7FFBC0217485CF8BDFA07F88E" authority="Broun" authorityName="Broun" authorityYear="1880" box="[1324,1436,1806,1828]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="hirta">
<emphasis id="1AE52C56FFBC0217485CF8BDFAC0F889" box="[1324,1371,1806,1827]" italics="true" pageId="36" pageNumber="437">hirta</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F89FFA07F8EB" blockId="36.[151,1437,868,2034]" box="[151,1436,1836,1858]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F89FFA07F8EB" box="[151,1436,1836,1858]" pageId="36" pageNumber="437">
-. Penis less elongated (
<figureCitation id="B0AAECC1FFBC02174CC0F89FFD9CF8EB" box="[432,519,1836,1858]" captionStart="FIGURE 13" captionStartId="20.[151,250,1824,1848]" captionTargetBox="[151,1436,777,1801]" captionTargetId="figure-263@20.[151,1436,777,1801]" captionTargetPageId="20" captionText="FIGURE 13. Atopida, male genitalia, penis (left), tegmen (right). A) A. lescheni sp. nov., B) A. montana, C) A. paparoa sp. nov., D) A. paringa sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6986525" httpUri="https://zenodo.org/record/6986525/files/figure.png" pageId="36" pageNumber="437">Fig. 13B</figureCitation>
), species restricted to South Island....................................
<taxonomicName id="EF918BC7FFBC02174876F89FFA07F8EB" authority="Broun" authorityName=", Broun" authorityYear="1921" box="[1286,1436,1836,1857]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="1AE52C56FFBC02174876F89FFAC1F8EB" box="[1286,1370,1836,1857]" italics="true" pageId="36" pageNumber="437">montana</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F8FAFA07F81C" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F8FAFA07F8D6" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F8FAFA07F8D6" pageId="36" pageNumber="437">
16. Lateral carinae of pronotum curved, anterolateral angles slightly projecting, basal portion of elytra in females deeply excised to fit pronotal base, restricted to North Island.......................................................
<taxonomicName id="EF918BC7FFBC0217486AF8D5FA07F8D6" authority="Broun" authorityName="Broun" authorityYear="1921" box="[1306,1436,1894,1916]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="1AE52C56FFBC0217486AF8D5FAC1F8D1" box="[1306,1370,1894,1915]" italics="true" pageId="36" pageNumber="437">villosa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F837FA07F81C" blockId="36.[151,1437,868,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F837FA07F81C" pageId="36" pageNumber="437">
-. Lateral carinae of pronotum bisinuate, anterolateral angles strongly projecting, basal portion of elytra in females shallowly excised to fit pronotal base, restricted to South Island.......................................
<taxonomicName id="EF918BC7FFBC021749B4F812FAD2F81C" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1220,1353,1953,1974]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="437" phylum="Arthropoda" rank="species" species="westlandensis" status="sp. nov.">
<emphasis id="1AE52C56FFBC021749B4F812FAD2F81C" box="[1220,1353,1953,1974]" italics="true" pageId="36" pageNumber="437">westlandensis</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBC02174820F812FA07F81C" bold="true" box="[1360,1436,1953,1974]" pageId="36" pageNumber="437">
<taxonomicNameLabel id="01D6912DFFBC02174820F812FA07F81C" box="[1360,1436,1953,1974]" pageId="36" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBC02174DE7F80DFA07F85B" pageId="36" pageNumber="437">
<paragraph id="282EF044FFBC02174DE7F80DFA07F879" blockId="36.[151,1437,868,2034]" box="[151,1436,1982,2004]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F80DFA07F879" box="[151,1436,1982,2004]" pageId="36" pageNumber="437">17. Hind wings well developed (macropterous)................................................................ 18</keyLead>
</paragraph>
<paragraph id="282EF044FFBC02174DE7F86FFA07F85B" blockId="36.[151,1437,868,2034]" box="[151,1436,2012,2034]" pageId="36" pageNumber="437">
<keyLead id="93605371FFBC02174DE7F86FFA07F85B" box="[151,1436,2012,2034]" pageId="36" pageNumber="437">-. Hind wings reduced (apterous or brachypterous)............................................................ 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FF2BFA07FF60" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FF2BFA07FF07" blockId="37.[151,1438,151,907]" box="[151,1436,151,173]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FF2BFA07FF07" box="[151,1436,151,173]" pageId="37" pageNumber="438">
18. Small (TL
<quantity id="EF695DA1FFBD02164C3AFF24FE5AFF07" box="[330,449,151,173]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="37" pageNumber="438" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 mm</quantity>
), dorsum covered with erect setae..............................................
<taxonomicName id="EF918BC7FFBD0216485CFF24FA07FF07" authority="Broun" authorityName="Broun" authorityYear="1880" box="[1324,1436,151,173]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="hirta">
<emphasis id="1AE52C56FFBD0216485CFF24FAC0FF06" box="[1324,1371,151,172]" italics="true" pageId="37" pageNumber="438">hirta</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FF06FA07FF60" blockId="37.[151,1438,151,907]" box="[151,1436,181,203]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FF06FA07FF60" box="[151,1436,181,203]" pageId="37" pageNumber="438">
-. Larger (TL&gt;
<quantity id="EF695DA1FFBD02164C15FF06FE29FF60" box="[357,434,181,203]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="37" pageNumber="438" unit="mm" value="4.5">4.5 mm</quantity>
), dorsum covered with adherent or semi-erect setae........................................ 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FF61FA06FEAF" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FF61FA06FF42" blockId="37.[151,1438,151,907]" box="[151,1437,210,232]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FF61FA06FF42" box="[151,1437,210,232]" pageId="37" pageNumber="438">
19. Basal portion of pronotum with sparse, granulate punctation, interspaces shining......................
<taxonomicName id="EF918BC7FFBD02164988FF61FA06FF42" authority="(White)" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[1272,1437,210,232]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="1AE52C56FFBD02164988FF61FAD7FF4D" box="[1272,1356,210,231]" italics="true" pageId="37" pageNumber="438">suturalis</emphasis>
(White)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FF43FA06FEAF" blockId="37.[151,1438,151,907]" box="[151,1437,239,261]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FF43FA06FEAF" box="[151,1437,239,261]" pageId="37" pageNumber="438">
-. Basal portion of pronotum with dense, granulate punctation........................................
<taxonomicName id="EF918BC7FFBD02164877FF43FA06FEAF" authority="White" authorityName="White" authorityYear="1846" box="[1287,1437,239,261]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="1AE52C56FFBD02164877FF43FAC7FEAF" box="[1287,1372,240,261]" italics="true" pageId="37" pageNumber="438">castanea</emphasis>
White
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FEBEFA07FE95" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FEBEFA07FE88" blockId="37.[151,1438,151,907]" box="[151,1436,269,290]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FEBEFA07FE88" box="[151,1436,269,290]" pageId="37" pageNumber="438">20. Apterous........................................................................................... 21</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FE98FA07FE95" blockId="37.[151,1438,151,907]" box="[151,1436,298,320]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE98FA07FE95" box="[151,1436,298,320]" pageId="37" pageNumber="438">-. Brachypterous....................................................................................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FEFBFA07FED0" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FEFBFA05FEF7" blockId="37.[151,1438,151,907]" box="[151,1438,327,349]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FEFBFA05FEF7" box="[151,1438,327,349]" pageId="37" pageNumber="438">
21. Abdomen without setal patches (
<figureCitation id="B0AAECC1FFBD02164F6DFEF4FDEBFEF7" box="[541,624,327,349]" captionStart="FIGURE 14" captionStartId="22.[151,250,1854,1878]" captionTargetBox="[151,1436,633,1830]" captionTargetId="figure-197@22.[151,1436,633,1830]" captionTargetPageId="22" captionText="FIGURE 14. Atopida cf. montana, females: A), D), G)—‘Mt Robinson’, B), E), H)—‘Port Underwood’, C), F), I)—‘Pelorus Bridge’. A–C) dorsal view, D–F) abdominal ventrites 4 &amp; 5, G–I) bursal sclerites." figureDoi="http://doi.org/10.5281/zenodo.6986529" httpUri="https://zenodo.org/record/6986529/files/figure.png" pageId="37" pageNumber="438">Fig. 14F</figureCitation>
)............................................
<taxonomicName id="EF918BC7FFBD021649F6FEF4FB62FEF7" authority="Pelorus" authorityName=", Broun" authorityYear="1921" box="[1158,1273,327,349]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
cf.
<emphasis id="1AE52C56FFBD021649D5FEFBFB62FEF7" box="[1189,1273,328,349]" italics="true" pageId="37" pageNumber="438">montana</emphasis>
</taxonomicName>
‘Pelorus Bridge’
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FED6FA07FED0" blockId="37.[151,1438,151,907]" box="[151,1436,357,379]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FED6FA07FED0" box="[151,1436,357,379]" pageId="37" pageNumber="438">
-. Ventrite 4 with two large, oblique setal patches (
<figureCitation id="B0AAECC1FFBD02164FC1FED6FD67FED0" box="[689,764,357,379]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8C</figureCitation>
), tergite VII with oval depression (
<figureCitation id="B0AAECC1FFBD0216493BFED6FB0DFED0" box="[1099,1174,357,379]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8D</figureCitation>
)........
<taxonomicName id="EF918BC7FFBD02164874FED6FA07FED0" authority="Broun" authorityName="Broun" authorityYear="1914" box="[1284,1436,357,378]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="impressa">
<emphasis id="1AE52C56FFBD02164874FED6FAC1FED0" box="[1284,1370,357,378]" italics="true" pageId="37" pageNumber="438">impressa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FE31FA07FE1F" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FE31FA07FE3D" blockId="37.[151,1438,151,907]" box="[151,1436,386,408]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE31FA07FE3D" box="[151,1436,386,408]" pageId="37" pageNumber="438">22. Setal patches present on ventrites 3 and 4................................................................. 23</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FE13FA07FE1F" blockId="37.[151,1438,151,907]" box="[151,1436,415,437]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE13FA07FE1F" box="[151,1436,415,437]" pageId="37" pageNumber="438">-. Setal patches present only on ventrite 4................................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FE0EFA07FE5A" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FE0EFA07FE78" blockId="37.[151,1438,151,907]" box="[151,1436,445,467]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE0EFA07FE78" box="[151,1436,445,467]" pageId="37" pageNumber="438">
23. Bursal sclerite plate-like, large (
<figureCitation id="B0AAECC1FFBD02164F65FE0EFD39FE78" box="[533,674,445,467]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Figs 10D, F, L</figureCitation>
)............................................................ 24
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FE68FA07FE5A" blockId="37.[151,1438,151,907]" box="[151,1436,474,496]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE68FA07FE5A" box="[151,1436,474,496]" pageId="37" pageNumber="438">
-. Bursal sclerite consisting of paired rods (
<figureCitation id="B0AAECC1FFBD02164F1BFE69FD7EFE45" box="[619,741,474,496]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Figs 10E, M</figureCitation>
)....................................................... 26
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FE4BFA07FDE2" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FE4BFA07FD80" blockId="37.[151,1438,151,907]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FE4BFA07FD80" pageId="37" pageNumber="438">
24. Lateral portions of bursal sclerite with long spines in dorsal view, central portion with pair of tubercles (
<figureCitation id="B0AAECC1FFBD021649B9FE44FA84FDA7" box="[1225,1311,503,525]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Fig. 10D</figureCitation>
)...............................................................................................
<taxonomicName id="EF918BC7FFBD02164985FDA6FAD2FD80" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1269,1353,533,554]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
<emphasis id="1AE52C56FFBD02164985FDA6FAD2FD80" box="[1269,1353,533,554]" italics="true" pageId="37" pageNumber="438">insularis</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBD02164820FDA6FA07FD80" bold="true" box="[1360,1436,533,554]" pageId="37" pageNumber="438">
<taxonomicNameLabel id="01D6912DFFBD02164820FDA6FA07FD80" box="[1360,1436,533,554]" pageId="37" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FD80FA07FDE2" blockId="37.[151,1438,151,907]" box="[151,1436,562,584]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FD80FA07FDE2" box="[151,1436,562,584]" pageId="37" pageNumber="438">
-. Lateral portions of bursal sclerite without long spines in dorsal view, central portion with a single tubercle (
<figureCitation id="B0AAECC1FFBD02164998FD81FAC3FDED" box="[1256,1368,562,584]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Figs 10F, L</figureCitation>
)... 25
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FDFCFA07FD28" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FDFCFA07FDCF" blockId="37.[151,1438,151,907]" box="[151,1436,591,613]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FDFCFA07FDCF" box="[151,1436,591,613]" pageId="37" pageNumber="438">
25. Abdominal setal patches large, transversely oblique (
<figureCitation id="B0AAECC1FFBD02164FAEFDFCFCB2FDCF" box="[734,809,591,613]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8G</figureCitation>
)......................................
<taxonomicName id="EF918BC7FFBD02164989FDE3FAD2FDCF" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1273,1353,592,613]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
<emphasis id="1AE52C56FFBD02164989FDE3FAD2FDCF" box="[1273,1353,592,613]" italics="true" pageId="37" pageNumber="438">paparoa</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBD02164820FDE3FA07FDCF" bold="true" box="[1360,1436,592,613]" pageId="37" pageNumber="438">
<taxonomicNameLabel id="01D6912DFFBD02164820FDE3FA07FDCF" box="[1360,1436,592,613]" pageId="37" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FDDEFA07FD28" blockId="37.[151,1438,151,907]" box="[151,1436,621,643]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FDDEFA07FD28" box="[151,1436,621,643]" pageId="37" pageNumber="438">
-. Abdominal setal patches small, oval (
<figureCitation id="B0AAECC1FFBD02164F3AFDDEFD01FD28" box="[586,666,621,643]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8M</figureCitation>
)....................................................
<taxonomicName id="EF918BC7FFBD02164863FDDEFAD1FD28" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1299,1354,621,642]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="tuhua" status="sp. nov.">
<emphasis id="1AE52C56FFBD02164863FDDEFAD1FD28" box="[1299,1354,621,642]" italics="true" pageId="37" pageNumber="438">tuhua</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBD02164820FDDEFA07FD28" bold="true" box="[1360,1436,621,642]" pageId="37" pageNumber="438">
<taxonomicNameLabel id="01D6912DFFBD02164820FDDEFA07FD28" box="[1360,1436,621,642]" pageId="37" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FD39FA07FD17" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FD39FA07FD0A" blockId="37.[151,1438,151,907]" box="[151,1436,650,672]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FD39FA07FD0A" box="[151,1436,650,672]" pageId="37" pageNumber="438">
26. Abdominal setal patches small, oval, ventrite 5 rounded (
<figureCitation id="B0AAECC1FFBD02164E76FD39FCD6FD35" box="[774,845,650,672]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8F</figureCitation>
)....................................
<taxonomicName id="EF918BC7FFBD02164876FD38FA07FD0A" authority="Broun" authorityName=", Broun" authorityYear="1921" box="[1286,1436,651,672]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="1AE52C56FFBD02164876FD38FAC1FD0A" box="[1286,1370,651,672]" italics="true" pageId="37" pageNumber="438">montana</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FD1BFA07FD17" blockId="37.[151,1438,151,907]" box="[151,1436,679,701]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FD1BFA07FD17" box="[151,1436,679,701]" pageId="37" pageNumber="438">
-. Abdominal setal patches large, transverse, ventrite 5 deeply emarginated (
<figureCitation id="B0AAECC1FFBD02164ED7FD14FC69FD17" box="[935,1010,679,701]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Fig. 8N</figureCitation>
).......................
<taxonomicName id="EF918BC7FFBD0216486AFD14FA07FD17" authority="Broun" authorityName="Broun" authorityYear="1921" box="[1306,1436,679,701]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="villosa">
<emphasis id="1AE52C56FFBD0216486AFD14FAC1FD16" box="[1306,1370,679,700]" italics="true" pageId="37" pageNumber="438">villosa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FD76FA07FD5D" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FD76FA07FD70" blockId="37.[151,1438,151,907]" box="[151,1436,709,731]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FD76FA07FD70" box="[151,1436,709,731]" pageId="37" pageNumber="438">
27. Bursal sclerite consisting of two long rods and a single, v-shaped sclerite (
<figureCitation id="B0AAECC1FFBD02164EDBFD76FB99FD70" box="[939,1026,709,731]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Fig. 10H</figureCitation>
)......................
<taxonomicName id="EF918BC7FFBD02164866FD76FA07FD70" authority="Broun" authorityName="Broun" authorityYear="1910" box="[1302,1436,709,730]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="suffusa">
<emphasis id="1AE52C56FFBD02164866FD76FAC1FD70" box="[1302,1370,709,730]" italics="true" pageId="37" pageNumber="438">suffusa</emphasis>
Broun
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FD50FA07FD5D" blockId="37.[151,1438,151,907]" box="[151,1436,738,760]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FD50FA07FD5D" box="[151,1436,738,760]" pageId="37" pageNumber="438">-. Bursal sclerite consisting of two subtriangular sclerites...................................................... 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FCB3FA07FCE5" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FCB3FA05FC99" blockId="37.[151,1438,151,907]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FCB3FA05FC99" pageId="37" pageNumber="438">
28. Ventrite 4 with very small, puncture-like setal patches, ventrite 5 with two triangular denticles in apical portion (
<figureCitation id="B0AAECC1FFBD02164868FD4CFAF6FCBF" box="[1304,1389,767,789]" captionStart="FIGURE 14" captionStartId="22.[151,250,1854,1878]" captionTargetBox="[151,1436,633,1830]" captionTargetId="figure-197@22.[151,1436,633,1830]" captionTargetPageId="22" captionText="FIGURE 14. Atopida cf. montana, females: A), D), G)—‘Mt Robinson’, B), E), H)—‘Port Underwood’, C), F), I)—‘Pelorus Bridge’. A–C) dorsal view, D–F) abdominal ventrites 4 &amp; 5, G–I) bursal sclerites." figureDoi="http://doi.org/10.5281/zenodo.6986529" httpUri="https://zenodo.org/record/6986529/files/figure.png" pageId="37" pageNumber="438">Fig. 14E</figureCitation>
)...............................................................................
<taxonomicName id="EF918BC7FFBD02164906FCAEFB72FC98" authority="Port" authorityName=", Broun" authorityYear="1921" box="[1142,1257,797,819]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
cf.
<emphasis id="1AE52C56FFBD021649E5FCAEFB72FC98" box="[1173,1257,797,818]" italics="true" pageId="37" pageNumber="438">montana</emphasis>
</taxonomicName>
‘Port Underwood’
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FC88FA07FCE5" blockId="37.[151,1438,151,907]" box="[151,1436,826,848]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FC88FA07FCE5" box="[151,1436,826,848]" pageId="37" pageNumber="438">
-. Ventrite 4 with larger, oval setal patches, ventrite 5 with shallow emargination (
<figureCitation id="B0AAECC1FFBD02164EACFC89FBB7FCE5" box="[988,1068,826,848]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="37" pageNumber="438">Figs 8O</figureCitation>
,
<figureCitation id="B0AAECC1FFBD02164946FC89FBFBFCE5" box="[1078,1120,826,847]" captionStart="FIGURE 14" captionStartId="22.[151,250,1854,1878]" captionTargetBox="[151,1436,633,1830]" captionTargetId="figure-197@22.[151,1436,633,1830]" captionTargetPageId="22" captionText="FIGURE 14. Atopida cf. montana, females: A), D), G)—‘Mt Robinson’, B), E), H)—‘Port Underwood’, C), F), I)—‘Pelorus Bridge’. A–C) dorsal view, D–F) abdominal ventrites 4 &amp; 5, G–I) bursal sclerites." figureDoi="http://doi.org/10.5281/zenodo.6986529" httpUri="https://zenodo.org/record/6986529/files/figure.png" pageId="37" pageNumber="438">14D</figureCitation>
)....................... 29
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9365E8E1FFBD02164DE7FCEBFA05FC21" pageId="37" pageNumber="438">
<paragraph id="282EF044FFBD02164DE7FCEBFA07FCC7" blockId="37.[151,1438,151,907]" box="[151,1436,855,877]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FCEBFA07FCC7" box="[151,1436,855,877]" pageId="37" pageNumber="438">
29. Bursal sclerite with narrowly triangular sclerites (
<figureCitation id="B0AAECC1FFBD02164FCEFCE4FC8EFCC7" box="[702,789,855,877]" captionStart="FIGURE 10" captionStartId="12.[151,250,1441,1465]" captionTargetBox="[151,1436,181,1417]" captionTargetId="figure-19@12.[151,1436,181,1417]" captionTargetPageId="12" captionText="FIGURE 10. Atopida, bursal sclerites, posterior portion on the right side of the photographs. A) A. castanea; B) A. hirta; C) A. impressa; D) A. insularis sp. nov.; E) A. montana; F) A. paparoa sp. nov.; G) A. proba; H) A. suffusa; I) same; J) A. suturalis, lateral view; K) same, dorsal view; L) A. tuhua sp. nov.; M) A. villosa; N) same; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986507" httpUri="https://zenodo.org/record/6986507/files/figure.png" pageId="37" pageNumber="438">Fig. 10O</figureCitation>
)...................................
<taxonomicName id="EF918BC7FFBD021649B4FCE4FAD2FCC6" authority="Kiałka &amp; Ruta, 2022" authorityName="Kiałka &amp; Ruta" authorityYear="2022" box="[1220,1353,855,876]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="westlandensis" status="sp. nov.">
<emphasis id="1AE52C56FFBD021649B4FCE4FAD2FCC6" box="[1220,1353,855,876]" italics="true" pageId="37" pageNumber="438">westlandensis</emphasis>
</taxonomicName>
<emphasis id="1AE52C56FFBD02164820FCEBFA07FCC7" bold="true" box="[1360,1436,856,877]" pageId="37" pageNumber="438">
<taxonomicNameLabel id="01D6912DFFBD02164820FCEBFA07FCC7" box="[1360,1436,856,877]" pageId="37" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="282EF044FFBD02164DE7FCC6FA05FC21" blockId="37.[151,1438,151,907]" box="[151,1438,885,907]" pageId="37" pageNumber="438">
<keyLead id="93605371FFBD02164DE7FCC6FA05FC21" box="[151,1438,885,907]" pageId="37" pageNumber="438">
-. Bursal sclerite with widely triangular sclerites (
<figureCitation id="B0AAECC1FFBD02164FD9FCC6FC9BFC20" box="[681,768,885,907]" captionStart="FIGURE 14" captionStartId="22.[151,250,1854,1878]" captionTargetBox="[151,1436,633,1830]" captionTargetId="figure-197@22.[151,1436,633,1830]" captionTargetPageId="22" captionText="FIGURE 14. Atopida cf. montana, females: A), D), G)—‘Mt Robinson’, B), E), H)—‘Port Underwood’, C), F), I)—‘Pelorus Bridge’. A–C) dorsal view, D–F) abdominal ventrites 4 &amp; 5, G–I) bursal sclerites." figureDoi="http://doi.org/10.5281/zenodo.6986529" httpUri="https://zenodo.org/record/6986529/files/figure.png" pageId="37" pageNumber="438">Fig. 14G</figureCitation>
).................................
<taxonomicName id="EF918BC7FFBD021649E7FCC6FA91FC20" authority="Mt" authorityName=", Broun" authorityYear="1921" box="[1175,1290,885,907]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="438" phylum="Arthropoda" rank="species" species="montana">
cf.
<emphasis id="1AE52C56FFBD021649C6FCC6FA91FC20" box="[1206,1290,885,906]" italics="true" pageId="37" pageNumber="438">montana</emphasis>
</taxonomicName>
‘Mt Robinson’
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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