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<document id="DFDF013B0979ABA58D834B19A69A711E" ID-DOI="10.11646/zootaxa.5512.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849018" ID-ZooBank="0A55482B-7C65-480D-8BA3-FF60F84AB5EB" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1727439976157" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany, Córdoba-Suarez, Eric &amp; Riccardi, Paula Raile" docDate="2024" docId="9C19879A5A334E32FF246967FAE1FAC3" docLanguage="en" docName="zootaxa.5512.3.7.pdf" docOrigin="Zootaxa 5512 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5512.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="440" masterDocId="6020FFE25A304E37FFB36F50FF87FF96" masterDocTitle="Three new species of Apotropina Hendel, 1907 (Diptera: Chloropidae) of the Colombian Andean-Amazon cloud forest" masterLastPageNumber="444" masterPageNumber="435" pageNumber="438" updateTime="1727704630546" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:title id="3244A0B598BBBB66EAA03F3ACFC4D937">Three new species of Apotropina Hendel, 1907 (Diptera: Chloropidae) of the Colombian Andean-Amazon cloud forest</mods:title>
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<mods:roleTerm id="4113D04903F3EE68F444CC82FF06AB2A">Author</mods:roleTerm>
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<mods:namePart id="3662E0692944B8591040B07C867EB9CD">Ramos-Pastrana, Yardany</mods:namePart>
<mods:affiliation id="1E3C8B941093FA0C0E3FF24989A5B333">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:namePart id="BDC55AC0AA2D18F6F4AC95C954FA9C33">Córdoba-Suarez, Eric</mods:namePart>
<mods:affiliation id="B71624D3F105F2F20691166099EB7381">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:namePart id="3CB522D718169DC237E6670C7AB17748">Riccardi, Paula Raile</mods:namePart>
<mods:affiliation id="A79B55FA5085D9D2453F331A3811E3DB">Center for Integrative Biodiversity Discovery, Museum für Naturkunde, Leibniz-Institute for Evolution and Biodiversity Science, Invalidenstr. 43, 10115 Berlin, Germany</mods:affiliation>
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<mods:title id="20C1A8DD4C8FE7E2605433FC1328140E">Zootaxa</mods:title>
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<mods:date id="A3295AC9DE8F142A097D86C6652FA61D">2024</mods:date>
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<mods:number id="31883FF3DC447FB0A9E39E770EBE6EC8">2024-09-20</mods:number>
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<mods:number id="31217FF2E511FA05726436ABAB7F9FD0">5512</mods:number>
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<treatment id="9C19879A5A334E32FF246967FAE1FAC3" ID-DOI="http://doi.org/10.5281/zenodo.13848991" ID-Zenodo-Dep="13848991" LSID="urn:lsid:plazi:treatment:9C19879A5A334E32FF246967FAE1FAC3" httpUri="http://treatment.plazi.org/id/9C19879A5A334E32FF246967FAE1FAC3" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438">
<subSubSection id="5CAA65075A334E34FF246967FE6FF9C7" box="[151,488,1590,1617]" pageId="3" pageNumber="438" type="nomenclature">
<paragraph id="140F368C5A334E34FF246967FE6FF9C7" blockId="3.[151,488,1590,1653]" box="[151,488,1590,1617]" pageId="3" pageNumber="438">
<heading id="4F4781E05A334E34FF246967FE6FF9C7" bold="true" box="[151,488,1590,1617]" fontSize="11" level="1" pageId="3" pageNumber="438" reason="1">
<emphasis id="26C4EA9E5A334E34FF246967FE6FF9C7" bold="true" box="[151,488,1590,1617]" pageId="3" pageNumber="438">
<taxonomicName id="D3B04D0F5A334E34FF246967FE05F9C7" authorityName="Ramos-Pastrana &amp; Córdoba-Suarez &amp; Riccardi" authorityYear="2024" box="[151,386,1590,1617]" class="Insecta" family="Chloropidae" genus="Apotropina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="ruidosa" status="sp. nov.">
<emphasis id="26C4EA9E5A334E34FF246967FE05F9C7" bold="true" box="[151,386,1590,1617]" italics="true" pageId="3" pageNumber="438">Apotropina ruidosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3DF757E55A334E34FE396967FE6FF9C7" box="[394,488,1591,1617]" pageId="3" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CAA65075A334E34FF24690BFEEAF9E3" box="[151,365,1626,1653]" pageId="3" pageNumber="438" type="description">
<paragraph id="140F368C5A334E34FF24690BFEEAF9E3" blockId="3.[151,488,1590,1653]" box="[151,365,1626,1653]" pageId="3" pageNumber="438">
(
<figureCitation id="8C8B2A095A334E34FF2C690BFE95F9E3" box="[159,274,1627,1653]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Figs 9–16</figureCitation>
,
<figureCitation id="8C8B2A095A334E34FEAD690AFEE1F9E3" box="[286,358,1626,1653]" captionStart="FIGURES 25–26. 25" captionStartId="7.[151,264,1855,1879]" captionTargetBox="[154,1432,1284,1828]" captionTargetId="figure-470@7.[151,1435,1281,1831]" captionTargetPageId="7" captionText="FIGURES 25–26. 25, Geographical occurrence for all species of Apotropina of the Colombia; 26, Malaise trap stretched across small stream in habitat of Apotropina longicerci Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov., A. ruidosa Ramos- Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. and A. similis Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13849028" httpUri="https://zenodo.org/record/13849028/files/figure.png" pageId="3" pageNumber="438">25–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5CAA65075A334E34FF2469F2FC0EF8DB" pageId="3" pageNumber="438" type="diagnosis">
<paragraph id="140F368C5A334E34FF2469F2FC0EF8DB" blockId="3.[151,1437,1698,2015]" pageId="3" pageNumber="438">
<emphasis id="26C4EA9E5A334E34FF2469F2FE95F92A" bold="true" box="[151,274,1698,1724]" pageId="3" pageNumber="438">Diagnosis.</emphasis>
Postpedicel rounded, dark brown on upper ¼, yellow on lower ¾ in external surface (
<figureCitation id="8C8B2A095A334E34FB6B69F3FA9AF92B" box="[1240,1309,1699,1725]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Fig. 9</figureCitation>
); fore coxa dark yellow, darker anteriorly (
<figureCitation id="8C8B2A095A334E34FE496997FDC7F977" box="[506,576,1735,1761]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Fig. 9</figureCitation>
); all trochanters yellow; all legs with anterior apex of femur, tibia and first tarsomere yellow (
<figureCitation id="8C8B2A095A334E34FEDA69BBFE29F893" box="[361,430,1771,1797]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Fig. 9</figureCitation>
); tibial organ narrow, dark gray, ~ 3.3× shorter than hind tibial length (
<figureCitation id="8C8B2A095A334E34FB7D69BBFA94F893" box="[1230,1299,1771,1797]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Fig. 9</figureCitation>
); hind tibial spur straight and shorter than tibia width; surstylus narrowed in profile (
<figureCitation id="8C8B2A095A334E34FC7E685FFBA6F8BF" box="[973,1057,1807,1833]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Fig. 13</figureCitation>
); cercus at least 1.5× longer than wide, with straight setae shorter than surstylus length (
<figureCitation id="8C8B2A095A334E34FD496863FCFBF8DB" box="[762,892,1842,1869]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Figs 15–16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5CAA65075A334E32FF746806FE44FB87" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438" type="description">
<paragraph id="140F368C5A334E32FF746806FC59FC7B" blockId="3.[151,1437,1698,2015]" lastBlockId="5.[151,1437,151,1365]" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438">
<emphasis id="26C4EA9E5A334E34FF746806FE30F8E6" bold="true" box="[199,439,1878,1904]" pageId="3" pageNumber="438">Description. MALE</emphasis>
(
<typeStatus id="CB0B882E5A334E34FE7A6807FDB5F8E7" box="[457,562,1879,1905]" pageId="3" pageNumber="438" type="holotype">holotype</typeStatus>
). Body length
<quantity id="D3489B695A334E34FD6C6807FCBBF8E7" box="[735,828,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="438" unit="mm" value="2.6">2.6 mm</quantity>
. Wing length
<quantity id="D3489B695A334E34FC516807FBB9F8E7" box="[994,1086,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="438" unit="mm" value="2.8">2.8 mm</quantity>
. Body dark brown with ochre pruinosity.
<emphasis id="26C4EA9E5A334E34FEA7682BFED3F803" bold="true" box="[276,340,1915,1941]" pageId="3" pageNumber="438">Head</emphasis>
(
<figureCitation id="8C8B2A095A334E34FED1682BFE50F803" box="[354,471,1915,1941]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="3" pageNumber="438">Figs 9–10</figureCitation>
). Wider than long dorsally and longer than high in profile. Ocellar seta as long as inner and outer vertical setae. Postocellar seta cruciate, ~ ⅔ length of ocellar seta. Three strong fronto-orbital setae, the anterior seta ~ ½ length of posterior setae, the two posterior setae ~ each equal to length of ocellar seta; anterior fronto-orbital seta proclinate the two posterior setae lateroclinate. Inner vertical seta proclinate, outer vertical seta lateroclinate. Frons dark brown basally, apical ¼ yellow, slightly wider than long, lateral margins diverging posteriorly to vertex, apical margin sinuous. Ocellar triangle extending to ~ ⅔ length of frons, posterior margin nearly as wide as frons, lateral margins straight, apex acute. Eye oval, long axis slightly oblique. Face pale yellow, about as high as wide; postpedicel rounded, dark brown on upper ¼, yellow on lower ¾ in external surface; arista black, with short, sparse pubescence, as long as ½ frons; genal height equal to height of postpedicel, with whitish pruinosity; one vibrissa; occiput and postgena dark brown; proboscis brown; palpus yellow with yellow setulae; clypeus darkened.
<emphasis id="26C4EA9E5A354E32FED96EC3FE44FE3B" bold="true" box="[362,451,403,429]" pageId="5" pageNumber="440">Thorax</emphasis>
(
<figureCitation id="8C8B2A095A354E32FE616EC3FDC1FE3B" box="[466,582,403,429]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Figs 9, 11</figureCitation>
). Scutum dark brown, ~ 1.2× longer than wide, pruinose; postpronotal lobe concolorous with scutum, with two long setae equal in length to notopleural setae; 1+1 notopleural setae; 5 strong dorsocentral setae, prescutellar dorsocentral seta ~ 2× longer than remaining dorsocentral setae; pleuron with brown pruinosity; katepisternum with pale setulae. Scutellum concolorous with scutum, with black pilosity, apical scutellar seta ~ 3× longer than subapical scutellar seta. Halter stem brown, knob yellowish.
<emphasis id="26C4EA9E5A354E32FB896D72FBFCFDAA" bold="true" box="[1082,1147,546,572]" pageId="5" pageNumber="440">Wing</emphasis>
(
<figureCitation id="8C8B2A095A354E32FB396D73FB59FDAB" box="[1162,1246,547,573]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Fig. 12</figureCitation>
). Hyaline; costal ratios measured from
<emphasis id="26C4EA9E5A354E32FE3C6D17FE1AFDF7" box="[399,413,583,609]" italics="true" pageId="5" pageNumber="440">h</emphasis>
:
<emphasis id="26C4EA9E5A354E32FE1E6D17FE41FDF1" box="[429,454,583,615]" italics="true" pageId="5" pageNumber="440">
R
<subScript id="883434C95A354E32FE0D6D09FE41FDF1" attach="left" box="[446,454,601,615]" fontSize="6" pageId="5" pageNumber="440">1</subScript>
</emphasis>
:
<emphasis id="26C4EA9E5A354E32FE656D17FD84FDF1" box="[470,515,583,615]" italics="true" pageId="5" pageNumber="440">
R
<subScript id="883434C95A354E32FE546D09FD84FDF1" attach="left" box="[487,515,601,615]" fontSize="6" pageId="5" pageNumber="440">2+3</subScript>
</emphasis>
:
<emphasis id="26C4EA9E5A354E32FDA16D17FDB8FDF1" box="[530,575,583,615]" italics="true" pageId="5" pageNumber="440">
R
<subScript id="883434C95A354E32FD906D09FDB8FDF1" attach="left" box="[547,575,601,615]" fontSize="6" pageId="5" pageNumber="440">4+5</subScript>
</emphasis>
is 4: 5: 4: 2; veins
<emphasis id="26C4EA9E5A354E32FCAF6D17FCCFFDF1" box="[796,840,583,615]" italics="true" pageId="5" pageNumber="440">
R
<subScript id="883434C95A354E32FC9E6D09FCCFFDF1" attach="left" box="[813,840,601,615]" fontSize="6" pageId="5" pageNumber="440">4+5</subScript>
</emphasis>
and
<emphasis id="26C4EA9E5A354E32FC326D17FC27FDF1" box="[897,928,583,615]" italics="true" pageId="5" pageNumber="440">
M
<subScript id="883434C95A354E32FC2B6D09FC27FDF1" attach="left" box="[920,928,601,615]" fontSize="6" pageId="5" pageNumber="440">1</subScript>
</emphasis>
subparallel; distance between
<emphasis id="26C4EA9E5A354E32FB4F6D17FAA3FDF6" box="[1276,1316,583,608]" italics="true" pageId="5" pageNumber="440">r-m</emphasis>
and
<emphasis id="26C4EA9E5A354E32FAEF6D17FA1BFDF7" box="[1372,1436,583,609]" italics="true" pageId="5" pageNumber="440">dm-m</emphasis>
4× length of
<emphasis id="26C4EA9E5A354E32FE9E6D3BFED2FD12" box="[301,341,619,644]" italics="true" pageId="5" pageNumber="440">r-m</emphasis>
.
<emphasis id="26C4EA9E5A354E32FED76D3BFE1BFD13" bold="true" box="[356,412,619,645]" pageId="5" pageNumber="440">Legs</emphasis>
(
<figureCitation id="8C8B2A095A354E32FE1E6D3BFE73FD13" box="[429,500,619,645]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Fig. 9</figureCitation>
). Dark brown; fore coxa dark yellow, darker anteriorly; all trochanters yellow; all legs with anterior apex of femur, tibia and first tarsomere yellow; tibial organ narrow, dark gray, ~ 3.3× shorter than hind tibia (
<figureCitation id="8C8B2A095A354E32FEFB6DE3FE0BFD5B" box="[328,396,691,717]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Fig. 9</figureCitation>
); hind tibial spur straight and slightly shorter than width of tibia.
<emphasis id="26C4EA9E5A354E32FBD56DE2FB5CFD5A" bold="true" box="[1126,1243,690,716]" pageId="5" pageNumber="440">Abdomen</emphasis>
(
<figureCitation id="8C8B2A095A354E32FB596DE3FAA9FD5B" box="[1258,1326,691,717]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Fig. 9</figureCitation>
). Tergites brown to dark brown.
<emphasis id="26C4EA9E5A354E32FE3C6D86FD95FD66" bold="true" box="[399,530,726,752]" pageId="5" pageNumber="440">Terminalia</emphasis>
(
<figureCitation id="8C8B2A095A354E32FD936D87FD25FD67" box="[544,674,727,753]" captionStart="FIGURES 9–16" captionStartId="4.[151,264,1865,1889]" captionTargetBox="[155,1431,185,1837]" captionTargetId="figure-26@4.[151,1435,181,1841]" captionTargetPageId="4" captionText="FIGURES 9–16. Apotropina ruidosa Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. (LEUA-67353). Male. 9, Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia; 10, Head, frontal view; 11, Thorax, dorsal view; 12, Wing; 13, Terminalia, lateral view; 14, Terminalia, posterior view; 15, Terminalia, ventral view; 16, Terminalia illustration, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="5" pageNumber="440">Figs 13–16</figureCitation>
). Postabdominal sclerites asymmetric; syntergosternite 7+8 about as long as hypandrium; sternite 6 slightly curved, wider apically; sternite 5 slightly curved, wider medially, tapering to extremes, ~ 10× longer than wide. Epandrium brown, fusiform in profile, with brown pilosity, without laterobasal projection; subepandrial sclerite V-shaped, base narrower than cerci; surstylus with apical half yellow with short, yellow setae, base slightly narrower than epandrial height in profile, apex narrow and rounded; cercus at least 1.5× longer than wide, with straight setae ~ 1× as short as length of surstylus, cerci fused basally; anal lobe indistinct. Hypandrium with arms closed; basiphallus bottle-shaped, slightly longer than distiphallus; distiphallus sclerotized dorsally; pregonite slightly shorter than postgonite; sperm pump indistinct.
</paragraph>
<caption id="40CF66045A344E33FF246819FE9CF85B" ID-DOI="http://doi.org/10.5281/zenodo.13849022" ID-Zenodo-Dep="13849022" httpUri="https://zenodo.org/record/13849022/files/figure.png" pageId="4" pageNumber="439" startId="4.[151,264,1865,1889]" targetBox="[155,1431,185,1837]" targetPageId="4" targetType="figure">
<paragraph id="140F368C5A344E33FF246819FE9CF85B" blockId="4.[151,1437,1865,1997]" pageId="4" pageNumber="439">
<emphasis id="26C4EA9E5A344E33FF246819FEC0F8F7" bold="true" box="[151,327,1865,1889]" pageId="4" pageNumber="439">FIGURES 9–16.</emphasis>
<taxonomicName id="D3B04D0F5A344E33FEFD681AFC60F8F7" authority="Ramos-Pastrana, Cordoba-Suarez &amp; Riccardi" authorityName="Ramos-Pastrana, Cordoba-Suarez &amp; Riccardi" authorityYear="2024" box="[334,999,1865,1890]" class="Insecta" family="Chloropidae" genus="Apotropina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="species" species="ruidosa" status="sp. nov.">
<emphasis id="26C4EA9E5A344E33FEFD681AFD94F8F7" box="[334,531,1865,1889]" italics="true" pageId="4" pageNumber="439">Apotropina ruidosa</emphasis>
Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi
</taxonomicName>
<emphasis id="26C4EA9E5A344E33FC5E681AFBBAF8F4" bold="true" box="[1005,1085,1866,1890]" pageId="4" pageNumber="439">
<taxonomicNameLabel id="3DF757E55A344E33FC5E681AFBBAF8F4" box="[1005,1085,1866,1890]" pageId="4" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(LEUA-67353). Male.
<emphasis id="26C4EA9E5A344E33FA9D6819FAC6F8F7" bold="true" box="[1326,1345,1865,1889]" pageId="4" pageNumber="439">9,</emphasis>
Habitus, left lateral view, black box shows the detail of the tibial organ in the hind tibia;
<emphasis id="26C4EA9E5A344E33FC04683DFC50F813" bold="true" box="[951,983,1901,1925]" pageId="4" pageNumber="439">10,</emphasis>
Head, frontal view;
<emphasis id="26C4EA9E5A344E33FB14683DFB42F813" bold="true" box="[1191,1221,1901,1925]" pageId="4" pageNumber="439">11,</emphasis>
Thorax, dorsal view;
<emphasis id="26C4EA9E5A344E33FF2468C1FF30F83F" bold="true" box="[151,183,1937,1961]" pageId="4" pageNumber="439">12,</emphasis>
Wing;
<emphasis id="26C4EA9E5A344E33FEB268C1FEA6F83F" bold="true" box="[257,289,1937,1961]" pageId="4" pageNumber="439">13,</emphasis>
Terminalia, lateral view;
<emphasis id="26C4EA9E5A344E33FD9668C1FDC3F83F" bold="true" box="[549,580,1937,1961]" pageId="4" pageNumber="439">14,</emphasis>
Terminalia, posterior view;
<emphasis id="26C4EA9E5A344E33FCD768C1FC04F83F" bold="true" box="[868,899,1937,1961]" pageId="4" pageNumber="439">15,</emphasis>
Terminalia, ventral view;
<emphasis id="26C4EA9E5A344E33FB3D68C1FB29F83F" bold="true" box="[1166,1198,1937,1961]" pageId="4" pageNumber="439">16,</emphasis>
Terminalia illustration, ventral view.
</paragraph>
</caption>
<paragraph id="140F368C5A354E32FF746CA6FE44FB87" blockId="5.[151,1437,151,1365]" box="[199,451,1014,1041]" pageId="5" pageNumber="440">
<emphasis id="26C4EA9E5A354E32FF746CA6FEC1FB86" bold="true" box="[199,326,1014,1040]" pageId="5" pageNumber="440">FEMALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5CAA65075A354E32FF746B4BFEC5FB37" pageId="5" pageNumber="440" type="materials_examined">
<paragraph id="140F368C5A354E32FF746B4BFEC5FB37" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="440">
<emphasis id="26C4EA9E5A354E32FF746B4BFEF1FBA2" bold="true" box="[199,374,1050,1077]" pageId="5" pageNumber="440">Type material.</emphasis>
<materialsCitation id="A4D83CD15A354E32FECE6B4AFCCAFBEB" collectingDate="1926-09-28" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana Leg." country="Colombia" elevation="2292" latitude="1.87" location="Florencia" longLatPrecision="21" longitude="-75.66917" pageId="5" pageNumber="440" specimenCode="LEUA-67353" specimenCount="1" specimenCount-male="1" stateProvince="Caqueta" typeStatus="holotype">
<emphasis id="26C4EA9E5A354E32FECE6B4AFD9BFBA2" bold="true" box="[381,540,1050,1076]" pageId="5" pageNumber="440">
<typeStatus id="CB0B882E5A354E32FECE6B4AFD9BFBA2" box="[381,540,1050,1076]" pageId="5" pageNumber="440" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="02B6FD055A354E32FD906B4BFDBDFBA3" box="[547,570,1051,1077]" count="1" pageId="5" pageNumber="440" type="male">♂</specimenCount>
, deposited in
<collectionCode id="72A1AE495A354E32FD6B6B4BFCA0FBA3" box="[728,807,1051,1077]" collectionName="LEUA" pageId="5" pageNumber="440">LEUA</collectionCode>
:
<collectingCountry id="6CA7761C5A354E32FC816B4BFC4DFBA3" box="[818,970,1051,1077]" name="Colombia" pageId="5" pageNumber="440">COLOMBIA</collectingCountry>
,
<collectingRegion id="D674F86E5A354E32FC656B4BFBB2FBA3" box="[982,1077,1051,1077]" country="Colombia" name="Caqueta" pageId="5" pageNumber="440">Caquetá</collectingRegion>
,
<location id="116F60575A354E32FBF36B4BFB2AFBA3" LSID="urn:lsid:plazi:treatment:9C19879A5A334E32FF246967FAE1FAC3:116F60575A354E32FBF36B4BFB2AFBA3" box="[1088,1197,1051,1077]" country="Colombia" latitude="1.87" longLatPrecision="21" longitude="-75.66917" name="Florencia" pageId="5" pageNumber="440" stateProvince="Caqueta">Florencia</location>
, Vda.[Vereda] Sucre,
<geoCoordinate id="7184504B5A354E32FF246B10FE90FBCD" box="[151,279,1088,1115]" degrees="01" direction="north" minutes="52" orientation="latitude" pageId="5" pageNumber="440" precision="15" seconds="12" value="1.87">01º52′12″N</geoCoordinate>
/
<geoCoordinate id="7184504B5A354E32FE936B10FE2CFBCD" box="[288,427,1088,1115]" degrees="75" direction="west" minutes="40" orientation="longitude" pageId="5" pageNumber="440" precision="15" seconds="09" value="-75.66917">75º40′09″W</geoCoordinate>
,
<quantity id="D3489B695A354E32FE076B6FFD8EFBCF" box="[436,521,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.292" pageId="5" pageNumber="440" unit="m" value="2292.0">
<elevation id="9F9DD1BF5A354E32FE076B6FFD8EFBCF" box="[436,521,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.292" pageId="5" pageNumber="440" unit="m" value="2292.0">2292 m</elevation>
</quantity>
[eters],
<date id="600E104C5A354E32FDE96B6FFD52FBCF" box="[602,725,1087,1113]" pageId="5" pageNumber="440" value="1926-09-28">
<collectingDate id="704AE9A45A354E32FDE96B6FFD52FBCF" box="[602,725,1087,1113]" pageId="5" pageNumber="440" value="1926-09-28">26–28.Sep</collectingDate>
</date>
[IX].2023,
<collectorName id="B945535A5A354E32FCF86B6FFBD2FBCF" box="[843,1109,1087,1113]" pageId="5" pageNumber="440">Y. Ramos-Pastrana Leg.</collectorName>
/ Captura con trampa Malaise colocada a nivel de un sistema lótico (al aire) / (
<specimenCode id="44169EF75A354E32FD176B32FCC2FBEB" box="[676,837,1122,1149]" pageId="5" pageNumber="440">LEUA-67353</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="A4D83CD15A354E32FCE56B32FEB8FB37" collectingDate="1926-09-28" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" elevation="2292" latitude="1.87" location="Florencia" longLatPrecision="21" longitude="-75.66917" pageId="5" pageNumber="440" specimenCode="LEUA-67354-67422" specimenCount="69" specimenCount-male="69" stateProvince="Caqueta" typeStatus="paratype">
<emphasis id="26C4EA9E5A354E32FCE56B32FB83FBEA" bold="true" box="[854,1028,1122,1148]" pageId="5" pageNumber="440">
<typeStatus id="CB0B882E5A354E32FCE56B32FC79FBEA" box="[854,1022,1122,1148]" pageId="5" pageNumber="440" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
same data as holotype (
<specimenCount id="02B6FD055A354E32FAB96B33FAC5FBEB" box="[1290,1346,1123,1149]" count="69" pageId="5" pageNumber="440" type="male">69 ♂</specimenCount>
<specimenCode id="44169EF75A354E32FAFB6B33FEB1FB37" pageId="5" pageNumber="440">LEUA-67354–67422</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CAA65075A354E32FF746BFBFAE1FAC3" pageId="5" pageNumber="440" type="etymology">
<paragraph id="140F368C5A354E32FF746BFBFB01FB7F" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="440">
<emphasis id="26C4EA9E5A354E32FF746BFBFEC9FB53" bold="true" box="[199,334,1195,1221]" pageId="5" pageNumber="440">Etymology.</emphasis>
The specific epithet
<taxonomicName id="D3B04D0F5A354E32FD876BFBFD11FB53" authorityName="Ramos-Pastrana, Cordoba-Suarez &amp; Riccardi" authorityYear="2024" box="[564,662,1195,1221]" class="Insecta" family="Chloropidae" genus="Apotropina" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="ruidosa">
“
<emphasis id="26C4EA9E5A354E32FDF36BFBFD11FB53" box="[576,662,1195,1221]" italics="true" pageId="5" pageNumber="440">ruidosa</emphasis>
</taxonomicName>
”, refers to the name “Reserva Natural y Ecoturística La Ruidosa”, the locality where the new species was collected. The name is treated as a noun in apposition.
</paragraph>
<paragraph id="140F368C5A354E32FF746BA2FC4AFA9B" blockId="5.[151,1437,151,1365]" box="[199,973,1266,1293]" pageId="5" pageNumber="440">
<emphasis id="26C4EA9E5A354E32FF746BA2FE7CFA9B" bold="true" box="[199,507,1266,1293]" pageId="5" pageNumber="440">Geographical occurrence.</emphasis>
<collectingCountry id="6CA7761C5A354E32FDB16BA3FDF7FA9B" box="[514,624,1267,1293]" name="Colombia" pageId="5" pageNumber="440">Colombia</collectingCountry>
(
<collectingRegion id="D674F86E5A354E32FDCC6BA3FD58FA9B" box="[639,735,1267,1293]" country="Colombia" name="Caqueta" pageId="5" pageNumber="440">Caquetá</collectingRegion>
, Florencia) (
<figureCitation id="8C8B2A095A354E32FCDE6BA3FC46FA9B" box="[877,961,1266,1293]" captionStart="FIGURES 25–26. 25" captionStartId="7.[151,264,1855,1879]" captionTargetBox="[154,1432,1284,1828]" captionTargetId="figure-470@7.[151,1435,1281,1831]" captionTargetPageId="7" captionText="FIGURES 25–26. 25, Geographical occurrence for all species of Apotropina of the Colombia; 26, Malaise trap stretched across small stream in habitat of Apotropina longicerci Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov., A. ruidosa Ramos- Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. and A. similis Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13849028" httpUri="https://zenodo.org/record/13849028/files/figure.png" pageId="5" pageNumber="440">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="140F368C5A354E32FF746A46FAE1FAC3" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="440">
<emphasis id="26C4EA9E5A354E32FF746A46FEADFAA6" bold="true" box="[199,298,1302,1328]" pageId="5" pageNumber="440">Habitat.</emphasis>
The specimens were collected in a Malaise trap placed over a creek in the low montane rainforest with dense secondary vegetation in the Andean-Amazonian transition corridor of
<collectingCountry id="6CA7761C5A354E32FC5C6A6BFBDAFAC3" box="[1007,1117,1339,1365]" name="Colombia" pageId="5" pageNumber="440">Colombia</collectingCountry>
(
<bibRefCitation id="70214B7D5A354E32FBDF6A6BFB68FAC3" author="CAM &amp; Corporacion Autonoma Regional &amp; Alto Magdalena" box="[1132,1263,1339,1365]" pageId="5" pageNumber="440" pagination="1 - 183" refId="ref5249" refString="CAM [Corporacion Autonoma Regional del Alto Magdalena] (2018) Actualizacion Plan de manejo Ambiental - PmA- Parque Natural regional &quot; Cerro Paramo Miraflores Rigoberto Urriago &quot;. ONF Andina SAS, 2018, 1 - 183." type="journal article" year="2018">CAM 2018</bibRefCitation>
) (
<figureCitation id="8C8B2A095A354E32FAB66A6BFADDFAC3" box="[1285,1370,1339,1365]" captionStart="FIGURES 25–26. 25" captionStartId="7.[151,264,1855,1879]" captionTargetBox="[154,1432,1284,1828]" captionTargetId="figure-470@7.[151,1435,1281,1831]" captionTargetPageId="7" captionText="FIGURES 25–26. 25, Geographical occurrence for all species of Apotropina of the Colombia; 26, Malaise trap stretched across small stream in habitat of Apotropina longicerci Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov., A. ruidosa Ramos- Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov. and A. similis Ramos-Pastrana, Córdoba-Suarez &amp; Riccardi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13849028" httpUri="https://zenodo.org/record/13849028/files/figure.png" pageId="5" pageNumber="440">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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