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<document id="1B64FE416E323BD98D07AA35A8CA601D" ID-DOI="10.5281/zenodo.5372196" ID-ISSN="1638-9395" ID-Zenodo-Dep="5372196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630271865429" checkinUser="marcus" docAuthor="Dumitrica, Paulian &amp; Zügel, Peter" docDate="2003" docId="943E87C0FFC9FF8FFCF46F13FD23F0B6" docLanguage="en" docName="Geodiversitas.2003.25.1.5-72.pdf" docOrigin="Geodiversitas 25 (1)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Anaticapitula Dumitrica &amp; Zügel 2003, n. gen." docType="treatment" docUuid="0D15192D-9BF8-474E-B0E1-FFC0B5D060B3" docUuidSource="ZooBank" docVersion="5" lastPageNumber="54" masterDocId="6807FFB8FFE6FFBEFFCA6929FFB2F469" masterDocTitle="Lower Tithonian mono- and dicyrtid Nassellaria (Radiolaria) from the Solnhofen area (southern Germany)" masterLastPageNumber="72" masterPageNumber="5" pageNumber="52" updateTime="1699207405888" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="64B0982DE1B54F5010274C4BE738CF69">Lower Tithonian mono- and dicyrtid Nassellaria (Radiolaria) from the Solnhofen area (southern Germany)</mods:title>
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<mods:namePart id="5B3099116545CCFE50EC56D7392088B7">Dumitrica, Paulian</mods:namePart>
<mods:affiliation id="B13C555F61C73BA9FB93715DB3842158">Institut de Géologie et Paléontologie, BFSH 2, UNIL, CH- 1015 Lausanne (Switzerland) Paulian. Dumitrica @ igp. unil. ch</mods:affiliation>
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<mods:namePart id="41856D6DA5FA97DAC57D67C8B5F97896">Zügel, Peter</mods:namePart>
<mods:affiliation id="71AB4E1C51D4DE1477F66B5F6A0789CC">Geologisch-Paläontologisches Institut, Johann Wolfgang Goethe-Universität, Senckenberganlage 32 - 34, D- 60054 Frankfurt am Main (Germany) zuegel @ em. uni-frankfurt. de.</mods:affiliation>
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<subSubSection id="548D655DFFC9FF91FCF46F13FBD9F23D" box="[830,1131,1594,1620]" pageId="47" pageNumber="52" type="nomenclature">
<paragraph id="1C2836D6FFC9FF91FCF46F13FBD9F23D" blockId="47.[830,1131,1594,1620]" box="[830,1131,1594,1620]" pageId="47" pageNumber="52">
<heading id="476081BAFFC9FF91FCF46F13FBD9F23D" box="[830,1131,1594,1620]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="52" reason="2">
Genus 
<taxonomicName id="DB974D55FFC9FF91FC416F13FBABF23D" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[907,1049,1594,1620]" class="Polycystina" family="Ultranaporidae" genus="Anaticapitula" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="47" pageNumber="52" phylum="Radiozoa" rank="genus" status="gen. nov.">
<emphasis id="2EE3EAC4FFC9FF91FC416F13FBABF23D" box="[907,1049,1594,1620]" italics="true" pageId="47" pageNumber="52">Anaticapitula</emphasis>
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<taxonomicNameLabel id="35D057BFFFC9FF91FBEA6F13FBD9F23D" box="[1056,1131,1594,1620]" pageId="47" pageNumber="52" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="548D655DFFC9FF91FD006F44FB77F2ED" box="[714,1221,1645,1670]" pageId="47" pageNumber="52" type="materials_examined">
<paragraph id="1C2836D6FFC9FF91FD006F44FB77F2ED" blockId="47.[714,1221,1645,1670]" box="[714,1221,1645,1670]" pageId="47" pageNumber="52">
<collectionCode id="7A86AE13FFC9FF91FD006F44FCFAF2EA" box="[714,840,1645,1668]" pageId="47" pageNumber="52">
<typeStatus id="C32C8874FFC9FF91FD006F44FD49F2EB" box="[714,763,1645,1668]" pageId="47" pageNumber="52">TYPE</typeStatus>
SPECIES
</collectionCode>
. — 
<taxonomicName id="DB974D55FFC9FF91FCBF6F47FB88F2EF" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[885,1082,1646,1670]" class="Polycystina" family="Ultranaporidae" genus="Anaticapitula" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="47" pageNumber="52" phylum="Radiozoa" rank="species" species="clauda" status="n. gen., n. sp.">
<emphasis id="2EE3EAC4FFC9FF91FCBF6F47FB88F2EF" box="[885,1082,1646,1670]" italics="true" pageId="47" pageNumber="52">Anaticapitula clauda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFC9FF91FB8A6F44FB77F2ED" box="[1088,1221,1645,1669]" pageId="47" pageNumber="52" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="48E8665EFFD6FF8EFF5D6F6EFD04F2EA" ID-DOI="http://doi.org/10.5281/zenodo.5372248" ID-Zenodo-Dep="5372248" httpUri="https://zenodo.org/record/5372248/files/figure.png" pageId="48" pageNumber="53" startId="48.[151,162,1607,1624]" targetBox="[178,1218,202,1575]" targetPageId="48">
<paragraph id="1C2836D6FFD6FF8EFF5D6F6EFD04F2EA" blockId="48.[151,1248,1607,1667]" pageId="48" pageNumber="53">
FIG. 26. — 
<emphasis id="2EE3EAC4FFD6FF8EFF3A6F6EFEA0F231" bold="true" box="[240,274,1607,1624]" pageId="48" pageNumber="53">A -C</emphasis>
, 
<emphasis id="2EE3EAC4FFD6FF8EFED66F6EFE98F231" bold="true" box="[284,298,1607,1624]" pageId="48" pageNumber="53">H</emphasis>
, 
<taxonomicName id="DB974D55FFD6FF8EFEFE6F6EFE64F231" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[308,470,1607,1624]" class="Polycystina" family="Tertoniidae" genus="Toritenum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="48" pageNumber="53" phylum="Radiozoa" rank="species" species="hirsutum" status="n. gen., n. sp.">
<emphasis id="2EE3EAC4FFD6FF8EFEFE6F6EFE64F231" box="[308,470,1607,1624]" italics="true" pageId="48" pageNumber="53">Toritenum hirsutum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFD6FF8EFE116F6EFDFDF231" box="[475,591,1607,1624]" pageId="48" pageNumber="53" rank="species">n. gen., n. sp.</taxonomicNameLabel>
; 
<emphasis id="2EE3EAC4FFD6FF8EFD9C6F6EFDD1F231" bold="true" box="[598,611,1607,1624]" pageId="48" pageNumber="53">A</emphasis>
, holotype; 
<emphasis id="2EE3EAC4FFD6FF8EFD756F6EFD7EF231" bold="true" box="[703,716,1607,1624]" pageId="48" pageNumber="53">B</emphasis>
, 
<emphasis id="2EE3EAC4FFD6FF8EFD1C6F6EFD56F231" bold="true" box="[726,740,1607,1624]" pageId="48" pageNumber="53">C</emphasis>
, 
<emphasis id="2EE3EAC4FFD6FF8EFD246F6EFD4EF231" bold="true" box="[750,764,1607,1624]" pageId="48" pageNumber="53">H</emphasis>
, paratypes; 
<emphasis id="2EE3EAC4FFD6FF8EFCA96F6EFCC3F231" bold="true" box="[867,881,1607,1624]" pageId="48" pageNumber="53">C</emphasis>
, broken specimen showing non-segmented internal surface; 
<emphasis id="2EE3EAC4FFD6FF8EFEED6F74FE87F207" bold="true" box="[295,309,1629,1646]" pageId="48" pageNumber="53">H</emphasis>
, broken specimen, basal view showing details of initial spicule; 
<emphasis id="2EE3EAC4FFD6FF8EFCA96F74FC34F207" bold="true" box="[867,902,1629,1646]" pageId="48" pageNumber="53">D -G</emphasis>
, 
<taxonomicName id="DB974D55FFD6FF8EFC586F74FBE8F207" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[914,1114,1629,1646]" class="Polycystina" family="Tertoniidae" genus="Toritenum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="48" pageNumber="53" phylum="Radiozoa" rank="species" species="corpulentum" status="n. gen., n. sp.">
<emphasis id="2EE3EAC4FFD6FF8EFC586F74FBE8F207" box="[914,1114,1629,1646]" italics="true" pageId="48" pageNumber="53">Toritenum corpulentum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFD6FF8EFBAB6F74FB6FF207" box="[1121,1245,1629,1646]" pageId="48" pageNumber="53" rank="species">n. gen., n. sp.</taxonomicNameLabel>
; 
<emphasis id="2EE3EAC4FFD6FF8EFF5D6F5BFF17F2EA" bold="true" box="[151,165,1650,1667]" pageId="48" pageNumber="53">D</emphasis>
, holotype; 
<emphasis id="2EE3EAC4FFD6FF8EFEC86F5BFE91F2EA" bold="true" box="[258,291,1650,1667]" pageId="48" pageNumber="53">E -G</emphasis>
, paratypes. Scale bars: A-G, 100 µm; H, 30 µm.
</paragraph>
</caption>
<subSubSection id="548D655DFFD7FF8FFF5D6980FEAEF586" pageId="49" pageNumber="54" type="etymology">
<paragraph id="1C2836D6FFD7FF8FFF5D6980FE40F4B2" blockId="49.[151,684,169,495]" pageId="49" pageNumber="54">
ETYMOLOGY. — From the Latin 
<emphasis id="2EE3EAC4FFD7FF8FFE176983FDB5F4A8" box="[477,519,170,193]" italics="true" pageId="49" pageNumber="54">anas</emphasis>
: duck; and 
<emphasis id="2EE3EAC4FFD7FF8FFDB76980FF76F4B2" italics="true" pageId="49" pageNumber="54">capitulus</emphasis>
: small head. Feminine gender.
</paragraph>
<paragraph id="1C2836D6FFD7FF8FFF5D69CEFEAEF586" blockId="49.[151,684,169,495]" pageId="49" pageNumber="54">KNOWN RANGE. — Lower Jurassic to Lower Cretaceous. DIAGNOSIS. — Highly ovate dicyrtid test with bladed apical horn and thorax prolonged into a thin-walled terminal tube. Cephalis and thorax continuous externally, without collar stricture. Initial spicule with A, V, D, two L, two l, and a long Ax. Cephalis and thorax with a superimposed network of strong ridges. With or without feet representing external extensions of L and D. Feet, when present, with an outer blade and two lateral blades.</paragraph>
</subSubSection>
<subSubSection id="548D655DFFD7FF8FFF5D6B23FD23F0B6" pageId="49" pageNumber="54" type="discussion">
<paragraph id="1C2836D6FFD7FF8FFF5D6B23FF49F64B" blockId="49.[151,685,522,1247]" box="[151,251,522,548]" pageId="49" pageNumber="54">REMARKS</paragraph>
<paragraph id="1C2836D6FFD7FF8FFF5D6B03FDD0F00D" blockId="49.[151,685,522,1247]" pageId="49" pageNumber="54">
By its general shape and the tubular prolongation of the thorax this genus shows characters in common with the genus 
<taxonomicName id="DB974D55FFD7FF8FFEB46B42FE8CF6CC" authority="Campbell &amp; Clark, 1944" authorityName="Campbell &amp; Clark" authorityYear="1944" class="Polycystina" family="Cannobotryidae" genus="Rhopalosyringium" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus">
<emphasis id="2EE3EAC4FFD7FF8FFEB46B42FD8AF6EC" box="[382,568,619,645]" italics="true" pageId="49" pageNumber="54">Rhopalosyringium</emphasis>
Campbell &amp; Clark, 1944
</taxonomicName>
. A comparable axobate was illustrated by 
<bibRefCitation id="78064B27FFD7FF8FFF356B83FE72F6AC" author="DE WEVER P." box="[255,448,682,709]" pageId="49" pageNumber="54" pagination="189 - 232" refId="ref34292" refString="DE WEVER P. 1982. - Nassellaria (Radiolaires polycystines) du Lias de Turquie. Revue de Micropaleontologie 24: 189 - 232." type="journal article" year="1982">De Wever (1982)</bibRefCitation>
in the Lower Jurassic species 
<taxonomicName id="DB974D55FFD7FF8FFF2D6BE3FDE3F68C" authority="De Wever, 1982" authorityName="De Wever" authorityYear="1982" box="[231,593,714,741]" class="Polycystina" family="Lophophaenidae" genus="Ovum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="species" species="pertusum">
<emphasis id="2EE3EAC4FFD7FF8FFF2D6BE3FE22F68A" box="[231,400,714,740]" italics="true" pageId="49" pageNumber="54">Ovum pertusum</emphasis>
<bibRefCitation id="78064B27FFD7FF8FFE526BE3FDE3F68C" author="DE WEVER P." box="[408,593,714,741]" pageId="49" pageNumber="54" pagination="189 - 232" refId="ref34292" refString="DE WEVER P. 1982. - Nassellaria (Radiolaires polycystines) du Lias de Turquie. Revue de Micropaleontologie 24: 189 - 232." type="journal article" year="1982">De Wever, 1982</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="DB974D55FFD7FF8FFD966BE3FD23F68A" authorityName="De Wever" authorityYear="1982" box="[604,657,714,739]" class="Polycystina" family="Ultranaporidae" genus="Jacus" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus">
<emphasis id="2EE3EAC4FFD7FF8FFD966BE3FD23F68A" box="[604,657,714,739]" italics="true" pageId="49" pageNumber="54">Jacus</emphasis>
</taxonomicName>
(?) 
<emphasis id="2EE3EAC4FFD7FF8FFF5D6BC0FEBFF76A" box="[151,269,745,771]" italics="true" pageId="49" pageNumber="54">anatiformis</emphasis>
<bibRefCitation id="78064B27FFD7FF8FFEDC6BC3FE63F76C" author="DE WEVER P." box="[278,465,746,773]" pageId="49" pageNumber="54" pagination="189 - 232" refId="ref34292" refString="DE WEVER P. 1982. - Nassellaria (Radiolaires polycystines) du Lias de Turquie. Revue de Micropaleontologie 24: 189 - 232." type="journal article" year="1982">De Wever, 1982</bibRefCitation>
, described from the lower Pliensbachian of 
<collectingCountry id="64807646FFD7FF8FFE606A22FDB3F74C" box="[426,513,779,805]" name="Turkey" pageId="49" pageNumber="54">Turkey</collectingCountry>
, is also almost identical to 
<taxonomicName id="DB974D55FFD7FF8FFEFA6A03FE01F72A" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[304,435,810,836]" class="Polycystina" family="Ultranaporidae" genus="Anaticapitula" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="species" species="pennata" status="n. gen., n. sp.">
<emphasis id="2EE3EAC4FFD7FF8FFEFA6A03FEF3F72D" box="[304,321,810,836]" italics="true" pageId="49" pageNumber="54">A</emphasis>
. 
<emphasis id="2EE3EAC4FFD7FF8FFE9F6A03FE01F72A" box="[341,435,810,835]" italics="true" pageId="49" pageNumber="54">pennata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFD7FF8FFE746A03FDDEF72D" box="[446,620,810,836]" pageId="49" pageNumber="54" rank="species">n. gen., n. sp.</taxonomicNameLabel>
and, although 
<bibRefCitation id="78064B27FFD7FF8FFECA6A63FE70F70C" author="DE WEVER P." box="[256,450,842,869]" pageId="49" pageNumber="54" pagination="189 - 232" refId="ref34292" refString="DE WEVER P. 1982. - Nassellaria (Radiolaires polycystines) du Lias de Turquie. Revue de Micropaleontologie 24: 189 - 232." type="journal article" year="1982">De Wever (1982)</bibRefCitation>
did not mention the presence of Ax in his species, a small light-grey spot opposite to the ventral spine, representing probably a small Ax, is visible in a broken specimen figured by him (
<bibRefCitation id="78064B27FFD7FF8FFEB46AE2FD82F78C" author="DE WEVER P." box="[382,560,970,997]" pageId="49" pageNumber="54" pagination="189 - 232" refId="ref34292" refString="DE WEVER P. 1982. - Nassellaria (Radiolaires polycystines) du Lias de Turquie. Revue de Micropaleontologie 24: 189 - 232." type="journal article" year="1982">De Wever 1982</bibRefCitation>
: pl. 11, fig. 13). 
<taxonomicName id="DB974D55FFD7FF8FFF036AC3FF4CF06A" authorityName="De Wever" authorityYear="1982" box="[201,254,1002,1027]" class="Polycystina" family="Ultranaporidae" genus="Jacus" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus">
<emphasis id="2EE3EAC4FFD7FF8FFF036AC3FF4CF06A" box="[201,254,1002,1027]" italics="true" pageId="49" pageNumber="54">Jacus</emphasis>
</taxonomicName>
(?) 
<emphasis id="2EE3EAC4FFD7FF8FFED56AC2FEDBF06C" box="[287,361,1003,1029]" italics="true" pageId="49" pageNumber="54">italicus</emphasis>
<bibRefCitation id="78064B27FFD7FF8FFEA56AC2FE69F06C" author="JUD R." box="[367,475,1003,1029]" pageId="49" pageNumber="54" pagination="1 - 147" refId="ref34810" refString="JUD R. 1994. - Biochronology and systematics of Early Cretaceous Radiolaria of the Western Tethys. Memoires de Geologie 19: 1 - 147." type="journal article" year="1994">Jud, 1994</bibRefCitation>
described from the Lower Cretaceous (
<bibRefCitation id="78064B27FFD7FF8FFEA66D22FE64F04C" author="JUD R." box="[364,470,1035,1061]" pageId="49" pageNumber="54" pagination="1 - 147" refId="ref34810" refString="JUD R. 1994. - Biochronology and systematics of Early Cretaceous Radiolaria of the Western Tethys. Memoires de Geologie 19: 1 - 147." type="journal article" year="1994">Jud 1994</bibRefCitation>
) has a morphology similar to 
<emphasis id="2EE3EAC4FFD7FF8FFEDA6D03FEA8F02A" box="[272,282,1066,1091]" italics="true" pageId="49" pageNumber="54">J</emphasis>
.(?) 
<emphasis id="2EE3EAC4FFD7FF8FFE836D00FE7AF02A" box="[329,456,1065,1091]" italics="true" pageId="49" pageNumber="54">anatiformis</emphasis>
and should also be assigned to the genus 
<taxonomicName id="DB974D55FFD7FF8FFE486D62FDA2F00C" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[386,528,1099,1125]" class="Polycystina" family="Ultranaporidae" genus="Anaticapitula" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus" status="gen. nov.">
<emphasis id="2EE3EAC4FFD7FF8FFE486D62FDA2F00C" box="[386,528,1099,1125]" italics="true" pageId="49" pageNumber="54">Anaticapitula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFD7FF8FFDDD6D63FDD0F00D" box="[535,610,1098,1124]" pageId="49" pageNumber="54" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
<paragraph id="1C2836D6FFD7FF8FFF5D6D40FD23F0B6" blockId="49.[151,685,522,1247]" pageId="49" pageNumber="54">
From 
<taxonomicName id="DB974D55FFD7FF8FFF1C6D40FE93F0EB" authorityName="Pessagno" authorityYear="1977" box="[214,289,1129,1154]" class="Polycystina" family="Ultranaporidae" genus="Napora" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus">
<emphasis id="2EE3EAC4FFD7FF8FFF1C6D40FE93F0EB" box="[214,289,1129,1154]" italics="true" pageId="49" pageNumber="54">Napora</emphasis>
</taxonomicName>
, 
<taxonomicName id="DB974D55FFD7FF8FFEE76D40FE01F0EA" authority="Dumitrica &amp; Zügel, 2003" authorityName="Dumitrica &amp; Zügel" authorityYear="2003" box="[301,435,1129,1155]" class="Polycystina" family="Ultranaporidae" genus="Anaticapitula" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="49" pageNumber="54" phylum="Radiozoa" rank="genus" status="gen. nov.">
<emphasis id="2EE3EAC4FFD7FF8FFEE76D40FE01F0EA" box="[301,435,1129,1155]" italics="true" pageId="49" pageNumber="54">Anaticapitula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35D057BFFFD7FF8FFE736D40FE4DF0EA" box="[441,511,1129,1155]" pageId="49" pageNumber="54" rank="genus">n. gen.</taxonomicNameLabel>
differs by having a well developed Ax in the initial spicule, thorax continuous with the velum, no crown of spines on the apical horn, and a much larger cephalis.
</paragraph>
</subSubSection>
</treatment>
</document>