1027 lines
148 KiB
XML
1027 lines
148 KiB
XML
<document id="86AAF8C0DF910585EAA52B6459293CB9" ID-DOI="10.4202/app.00661.2019" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626646" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707250217550" checkinUser="felipe" docAuthor="Rotatori, Filippo Maria, Moreno-Azanza, Miguel & Mateus, Octávio" docDate="2020" docId="926187EDFFCC9347BC76FD9BFE0EFC55" docLanguage="en" docName="ActaPalaeontolPol.65.1.35-57.pdf" docOrigin="Acta Palaeontologica Polonica 65 (1)" docSource="http://dx.doi.org/10.4202/app.00661.2019" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Ankylopollexia" docType="treatment" docVersion="5" lastPageNumber="48" masterDocId="6E58FF95FFC59349BC14FF90FFA9FFC8" masterDocTitle="New information on ornithopod dinosaurs from the Late Jurassic of Portugal" masterLastPageNumber="57" masterPageNumber="35" pageNumber="44" updateTime="1715215747394" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FC04A78A422D32EBA5C13589E1D62CA4">New information on ornithopod dinosaurs from the Late Jurassic of Portugal</mods:title>
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<mods:namePart id="0DE2FEFE07DC67AEF59E5378A57964F1">Rotatori, Filippo Maria</mods:namePart>
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<mods:namePart id="CBA266EA51622D95F5834268A33777CA">Moreno-Azanza, Miguel</mods:namePart>
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<mods:namePart id="A170CD6F500756A202D2878483332A50">Mateus, Octávio</mods:namePart>
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<mods:title id="CB9499C4148996986A15412C0DF7BCC2">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="D261947CA6392E6021931FE9434DE4C8">2020</mods:date>
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<mods:number id="F108DBC3FC4CB9DCD1357DAC02426D80">2020-03-04</mods:number>
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<treatment id="926187EDFFCC9347BC76FD9BFE0EFC55" ID-DOI="http://doi.org/10.5281/zenodo.11088645" ID-Zenodo-Dep="11088645" LSID="urn:lsid:plazi:treatment:926187EDFFCC9347BC76FD9BFE0EFC55" httpUri="http://treatment.plazi.org/id/926187EDFFCC9347BC76FD9BFE0EFC55" lastPageId="14" lastPageNumber="48" pageId="9" pageNumber="44">
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<subSubSection id="52D26570FFCC9340BC76FD9BFED0FDEF" box="[98,377,523,551]" pageId="9" pageNumber="44" type="nomenclature">
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<paragraph id="1A7736FBFFCC9340BC76FD9BFED0FDEF" blockId="9.[98,377,523,551]" box="[98,377,523,551]" pageId="9" pageNumber="44">
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<heading id="413F8197FFCC9340BC76FD9BFED0FDEF" box="[98,377,523,551]" fontSize="12" level="2" pageId="9" pageNumber="44" reason="2">
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<taxonomicName id="DDC84D78FFCC9340BC76FD9BFE80FDEF" box="[98,297,523,551]" family="Dryosauridae" genus="Ankylopollexia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="44" phylum="Chordata" rank="genus">Ankylopollexia</taxonomicName>
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indet.
|
||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="52D26570FFCC9340BC76FDA8FF6AFD85" box="[98,195,568,589]" pageId="9" pageNumber="44" type="description">
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<paragraph id="1A7736FBFFCC9340BC76FDA8FF6AFD85" blockId="9.[98,195,568,589]" box="[98,195,568,589]" pageId="9" pageNumber="44">
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<figureCitation id="82F32A7EFFCC9340BC76FDA8FF16FD85" box="[98,191,568,589]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="9.[98,131,1907,1927]" captionStartId-1="10.[151,184,1353,1373]" captionStartId-2="11.[98,131,860,880]" captionStartId-3="13.[98,131,1438,1458]" captionTargetBox-0="[238,1341,1075,1881]" captionTargetBox-1="[153,1522,174,1319]" captionTargetBox-2="[542,931,494,833]" captionTargetBox-3="[128,1434,165,1401]" captionTargetId-0="figure-503@9.[238,1294,1075,1881]" captionTargetId-1="figure-286@10.[153,1525,164,1323]" captionTargetId-3="figure-254@13.[176,1438,189,1401]" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="Fig. 5. Cranial material of Ankylopollexia indet. from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Dentary ML 818, in medial (A1, A5), lateral (A2, A6), and dorsal (A3, A7) views, detail of the dentary/surangular contact (A4, A8)." captionText-1="Fig. 6. Dorsal vertebrae Ankylopollexia indet. from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. A. Partial neural arch, ML 864 in anterior (A1), posterior (A2), lateral (A3), and dorsal (A4) views. B, C. Dorsal vertebrae, ML 452a, complete (B) and ML 452b, incomplete (C) specimens, in anterior (B1, C1), posterior (B2, C2), lateral (B3, C3), dorsal (B4, C4), and ventral (digitally modified) (B5, C5) views." captionText-2="Fig. 7. Comparative dorsal vertebrae table of selected Ankylopollexians from the Late Jurassic and Early Cretaceous. A, B. Ankylopollexia indet. Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. A. ML 452 in lateral view (A1, A2). B. ML 864 in right lateral view. C. “Uteodon” SHN.LPP 015 in left lateral view; Praia da corva, Torres Vedras Municipality, Portugal, Lourinhã Formation (Kimmeridgian–Tithonian). D. “Uteodon” aphanoecetes CM 11337 in left lateral view; East end of Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah (USA), Morrison Formation (Kimmeridgian–Tithonian). E. Camptosaurus dispar (unnumbered specimen) in left lateral view; Bone Cabin Quarry, Wyoming USA), Morrison Formation (Kimmeridgian–Tithonian). F. “Cumnoria” prestwichii OUM. J.3303 in lateral view; Oxford, UK, Kimmeridge Clay Formation (Kimmeridgian–Tithonian). G. Hippodraco scutodens UMNH VP 20208 in left lateral view; Andrew’s Site, Grand County, Utah; Upper Yellow Cat Memberof the Cedar Mountain Formation (upper Barremian–lowermost Aptian). H. Iguanacolossus fortis UMNH VP 20205 in right lateral view; Don’s Ridge, Grand County, Utah, Lower Yellow Cat Member, Cedar Mountain Formation (?lower Barremian). I. Mantellisaurus atherfieldensis IRSNB 1551 in left lateral view; Isle of Wight, Wessex Formation (Barremian). J. Barilium dawsoni NHMUK R798 in left lateral view; Shornden, East Sussex, UK, Wadhurst Clay Formation (Valanginian). K. Hypselospinus fittoni NHMUK R604 in lateral view; Shornden Quarry, Hastings, UK, Wadhurst Clay Formation (Valanginian). L. Iguanodon bernissartensis IRSNB “Individu S” in left lateral view; Bernissart, Belgium, Sainte Barbe Clays Formation Barremian). Abbreviations: dia, diapophysis; par, parapophysis. Scale bars 100 mm. Re-drawn from: C, Escaso 2014: fig. 6.5; D, Carpenter and Wilson 2008: fig. 11; E, Carpenter and Galton 2018: fig. 22D; F, Galton and Powell 1980: fig. 4; G, McDonald 2010b: fig. 27; H, McDonald 2010b: fig. 10;, Norman 1980: fig. 37; J, Norman 2011: fig. 4; K, Norman 2015: fig. 22; L, Norman 1980: fig. 31)." captionText-3="Fig. 8. Ankylopollexian appendicular skeleton from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Coracoid ML 2206 (A), scapula ML 2042 (B), in lateral (A1, B1) and medial (A2, B2) views." figureDoi-0="http://doi.org/10.5281/zenodo.11088519" figureDoi-1="http://doi.org/10.5281/zenodo.10626658" figureDoi-2="http://doi.org/10.5281/zenodo.10626660" figureDoi-3="http://doi.org/10.5281/zenodo.10626662" httpUri-0="https://zenodo.org/record/11088519/files/figure.png" httpUri-1="https://zenodo.org/record/10626658/files/figure.png" httpUri-2="https://zenodo.org/record/10626660/files/figure.png" httpUri-3="https://zenodo.org/record/10626662/files/figure.png" pageId="9" pageNumber="44">Figs. 5–8</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52D26570FFCC9340BC76FDF0FE7DFCF5" pageId="9" pageNumber="44" type="materials_examined">
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<paragraph id="1A7736FBFFCC9340BC76FDF0FE7DFCF5" blockId="9.[98,773,608,829]" pageId="9" pageNumber="44">
|
||
<emphasis id="28BCEAE9FFCC9340BC76FDF0FF6FFDB1" box="[98,198,608,633]" italics="true" pageId="9" pageNumber="44">Material</emphasis>
|
||
.—
|
||
<collectionCode id="7CD9AE3EFFCC9340BCF3FDF0FEBDFDB1" box="[231,276,608,633]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
2042, isolated, almost complete scapula fromPraiadaPeralta;
|
||
<collectionCode id="7CD9AE3EFFCC9340BD43FD11FE2DFD52" box="[343,388,641,666]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
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||
452,twoassociatedvertebrae,
|
||
<collectionCode id="7CD9AE3EFFCC9340BECCFD11FCACFD52" box="[728,773,641,666]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
864, an isolated neural arch (not found in association with
|
||
<collectionCode id="7CD9AE3EFFCC9340BC76FD52FF24FD13" box="[98,141,706,731]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
452) from Praia de
|
||
<collectingRegion id="D80CF819FFCC9340BD66FD52FE06FD13" box="[370,431,706,731]" country="Portugal" name="Porto" pageId="9" pageNumber="44">Porto</collectingRegion>
|
||
das Barcas;
|
||
<collectionCode id="7CD9AE3EFFCC9340BE56FD52FDC4FD13" box="[578,621,706,731]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
818, isolated dentary from Praia Vale Frades;
|
||
<collectionCode id="7CD9AE3EFFCC9340BDF2FD73FDB8FD34" box="[486,529,739,764]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
2206, isolated coracoids from Lourinhã coastline. All from Kimmeridgian– Tithonian of Lourinhã,
|
||
<collectingCountry id="62DF766BFFCC9340BD78FCB4FE79FCF5" box="[364,464,804,829]" name="Portugal" pageId="9" pageNumber="44">Portugal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52D26570FFCC9347BC76FCDFFE0EFC55" lastPageId="14" lastPageNumber="49" pageId="9" pageNumber="44" type="description">
|
||
<paragraph id="1A7736FBFFCC9340BC76FCDFFBCAFDF4" blockId="9.[98,771,847,1035]" lastBlockId="9.[808,1481,160,927]" pageId="9" pageNumber="44">
|
||
<emphasis id="28BCEAE9FFCC9340BC76FCDFFF43FCA0" box="[98,234,847,872]" italics="true" pageId="9" pageNumber="44">Description</emphasis>
|
||
.—
|
||
<emphasis id="28BCEAE9FFCC9340BD1AFCDFFE7BFCA0" box="[270,466,847,872]" italics="true" pageId="9" pageNumber="44">Cranial skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFCC9340BDF3FCDFFDEFFCA0" box="[487,582,847,872]" italics="true" pageId="9" pageNumber="44">Dentary</emphasis>
|
||
(
|
||
<figureCitation id="82F32A7EFFCC9340BE48FCDFFD15FCA0" box="[604,700,847,872]" captionStart="Fig" captionStartId="9.[98,131,1907,1927]" captionTargetBox="[238,1341,1075,1881]" captionTargetId="figure-503@9.[238,1294,1075,1881]" captionText="Fig. 5. Cranial material of Ankylopollexia indet. from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Dentary ML 818, in medial (A1, A5), lateral (A2, A6), and dorsal (A3, A7) views, detail of the dentary/surangular contact (A4, A8)." figureDoi="http://doi.org/10.5281/zenodo.11088519" httpUri="https://zenodo.org/record/11088519/files/figure.png" pageId="9" pageNumber="44">Fig. 5A</figureCitation>
|
||
): The partial right dentary
|
||
<collectionCode id="7CD9AE3EFFCC9340BD44FCE0FED2FC41" box="[336,379,880,905]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
818 is broken at both anterior and posterior extremities, measuring
|
||
<quantity id="DD309B1EFFCC9340BDF3FC00FDEEFC61" box="[487,583,912,937]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="44" unit="mm" value="150.0">150 mm</quantity>
|
||
in total length. The maximum depth is reached in the posterior extremity being
|
||
<quantity id="DD309B1EFFCC9340BCBFFC42FF54FC23" box="[171,253,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="44" unit="mm" value="62.0">62 mm</quantity>
|
||
tall, while the anterior extremity is
|
||
<quantity id="DD309B1EFFCC9340BEA4FC42FCABFC23" box="[688,770,978,1003]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="9" pageNumber="44" unit="mm" value="49.0">49 mm</quantity>
|
||
tall. In general, the bone appears stout and compact, being heavily eroded towards the anterior-most part. The lateral surface is heavily eroded, although preservation of the coronoid process is fairly complete, missing just the dorsal-most end. The dorsal and ventral margins appear to be parallel to one another. The ventral surface is smooth, lacking a ventral flange (character 121: 0). The tooth row preserves ten, possibly eleven, distinguishable close-packed alveoli (character 136: 1). The tooth row, which ends medial to the coronoid process, is encased in a parapet-like shelf that is smoothly arched medially (character 122: 0). Two nutrient foramina are located on the marginal shelf of the tooth row (character 145: 0). Teeth are not preserved and the tooth-sockets do not show any interdental plates.
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFCC9340BF5AFDD3FA07FD15" blockId="9.[808,1481,160,927]" pageId="9" pageNumber="44">On the medial surface, dorsally to the Meckelian sulcus, a longitudinally striated surface extends almost to the anterior end of the specimen. Two other bones may have articulated against this surface: the splenial and the pre-articular. There is no difference in rugosity to distinguish these two areas.</paragraph>
|
||
<paragraph id="1A7736FBFFCC9340BF5AFD75FB5BFC57" blockId="9.[808,1481,160,927]" pageId="9" pageNumber="44">On the lateral surface the coronoid process projects postero-laterally (characters 124: 1, 125: 1) with respect to the tooth-row and is inclined at about 30° to the horizontal. The contact with the surangular corresponds to the 4th alveolus counted from the coronoid process cranially, so it reaches further anteriorly than coronoid process.</paragraph>
|
||
<paragraph id="1A7736FBFFCC9342BF3CFC21FC4AF86F" blockId="9.[808,1481,945,1035]" lastBlockId="11.[808,1481,1294,1959]" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44">
|
||
<emphasis id="28BCEAE9FFCC9340BF3CFC21FC60FC02" box="[808,969,945,970]" italics="true" pageId="9" pageNumber="44">Axial skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFCC9340BFCEFC21FACBFC02" box="[986,1378,945,970]" italics="true" pageId="9" pageNumber="44">Dorsal vertebrae and neural arch</emphasis>
|
||
(
|
||
<figureCitation id="82F32A7EFFCC9340B967FC21FA6AFC02" box="[1395,1475,945,970]" captionStart="Fig" captionStartId="10.[151,184,1353,1373]" captionTargetBox="[153,1522,174,1319]" captionTargetId="figure-286@10.[153,1525,164,1323]" captionTargetPageId="10" captionText="Fig. 6. Dorsal vertebrae Ankylopollexia indet. from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. A. Partial neural arch, ML 864 in anterior (A1), posterior (A2), lateral (A3), and dorsal (A4) views. B, C. Dorsal vertebrae, ML 452a, complete (B) and ML 452b, incomplete (C) specimens, in anterior (B1, C1), posterior (B2, C2), lateral (B3, C3), dorsal (B4, C4), and ventral (digitally modified) (B5, C5) views." figureDoi="http://doi.org/10.5281/zenodo.10626658" httpUri="https://zenodo.org/record/10626658/files/figure.png" pageId="9" pageNumber="44">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="82F32A7EFFCC9340BF3CFC41FC91FC22" box="[808,824,977,1002]" captionStart="Fig" captionStartId="11.[98,131,860,880]" captionTargetBox="[542,931,494,833]" captionTargetPageId="11" captionText="Fig. 7. Comparative dorsal vertebrae table of selected Ankylopollexians from the Late Jurassic and Early Cretaceous. A, B. Ankylopollexia indet. Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. A. ML 452 in lateral view (A1, A2). B. ML 864 in right lateral view. C. “Uteodon” SHN.LPP 015 in left lateral view; Praia da corva, Torres Vedras Municipality, Portugal, Lourinhã Formation (Kimmeridgian–Tithonian). D. “Uteodon” aphanoecetes CM 11337 in left lateral view; East end of Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah (USA), Morrison Formation (Kimmeridgian–Tithonian). E. Camptosaurus dispar (unnumbered specimen) in left lateral view; Bone Cabin Quarry, Wyoming USA), Morrison Formation (Kimmeridgian–Tithonian). F. “Cumnoria” prestwichii OUM. J.3303 in lateral view; Oxford, UK, Kimmeridge Clay Formation (Kimmeridgian–Tithonian). G. Hippodraco scutodens UMNH VP 20208 in left lateral view; Andrew’s Site, Grand County, Utah; Upper Yellow Cat Memberof the Cedar Mountain Formation (upper Barremian–lowermost Aptian). H. Iguanacolossus fortis UMNH VP 20205 in right lateral view; Don’s Ridge, Grand County, Utah, Lower Yellow Cat Member, Cedar Mountain Formation (?lower Barremian). I. Mantellisaurus atherfieldensis IRSNB 1551 in left lateral view; Isle of Wight, Wessex Formation (Barremian). J. Barilium dawsoni NHMUK R798 in left lateral view; Shornden, East Sussex, UK, Wadhurst Clay Formation (Valanginian). K. Hypselospinus fittoni NHMUK R604 in lateral view; Shornden Quarry, Hastings, UK, Wadhurst Clay Formation (Valanginian). L. Iguanodon bernissartensis IRSNB “Individu S” in left lateral view; Bernissart, Belgium, Sainte Barbe Clays Formation Barremian). Abbreviations: dia, diapophysis; par, parapophysis. Scale bars 100 mm. Re-drawn from: C, Escaso 2014: fig. 6.5; D, Carpenter and Wilson 2008: fig. 11; E, Carpenter and Galton 2018: fig. 22D; F, Galton and Powell 1980: fig. 4; G, McDonald 2010b: fig. 27; H, McDonald 2010b: fig. 10;, Norman 1980: fig. 37; J, Norman 2011: fig. 4; K, Norman 2015: fig. 22; L, Norman 1980: fig. 31)." figureDoi="http://doi.org/10.5281/zenodo.10626660" httpUri="https://zenodo.org/record/10626660/files/figure.png" pageId="9" pageNumber="44">7</figureCitation>
|
||
):
|
||
<collectionCode id="7CD9AE3EFFCC9340BF5AFC41FCD0FC22" box="[846,889,977,1002]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
452 are two partially distorted and fractured associated vertebrae, while
|
||
<collectionCode id="7CD9AE3EFFCC9340B80AFC62FBE0FBC3" box="[1054,1097,1010,1035]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
864 is a broken and isolated neural arch, which preserves both postzygapophyses and one prezygapophysis. The two vertebrae
|
||
<collectionCode id="7CD9AE3EFFCF9343BE5EFA47FDDCFA38" box="[586,629,1495,1520]" country="France" name="Musee de Lectoure" pageId="10" pageNumber="45">ML</collectionCode>
|
||
452 were found in association and therefore are interpreted to represent the same individual, while
|
||
<collectionCode id="7CD9AE3EFFCF9343BDBCF98BFE7AF9FC" box="[424,467,1563,1588]" country="France" name="Musee de Lectoure" pageId="10" pageNumber="45">ML</collectionCode>
|
||
864 is a second individual recovered from a different locality but comparable in size. Compared to the axial series of
|
||
<taxonomicName id="DDC84D78FFCF9343BE1AF9CEFCACF9BF" box="[526,773,1630,1655]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="45" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFCF9343BE1AF9CEFCACF9BF" box="[526,773,1630,1655]" italics="true" pageId="10" pageNumber="45">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC84D78FFCF9343BC83F910FD98F951" authority=", ML" authorityName=", ML" authorityYear="2042" box="[151,561,1664,1689]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="45" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCF9343BC83F910FE5CF951" box="[151,501,1664,1689]" italics="true" pageId="10" pageNumber="45">Mantellisaurus atherfieldensis</emphasis>
|
||
, ML
|
||
</taxonomicName>
|
||
864 is most likely located at the 6–8
|
||
<superScript id="EDBD9BB3FFCF9343BD53F90EFEFEF967" attach="left" box="[327,343,1694,1711]" fontSize="7" pageId="10" pageNumber="45">th</superScript>
|
||
position of the dorsal series, while
|
||
<collectionCode id="7CD9AE3EFFCF9343BECBF932FCA3F973" box="[735,778,1698,1723]" country="France" name="Musee de Lectoure" pageId="10" pageNumber="45">ML</collectionCode>
|
||
452 are located after the 10th but before the 16th position of the dorsal series (
|
||
<bibRefCitation id="7E594B0AFFCF9343BD21F976FE7FF937" author="Gilmore, C. W." box="[309,470,1766,1791]" pageId="10" pageNumber="45" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCF9343BDF6F976FD28F937" author="Norman, D. B." box="[482,641,1766,1791]" pageId="10" pageNumber="45" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
|
||
). The preserved centra are stout ranging
|
||
<quantity id="DD309B1EFFCF9343BDAAF897FD97F8E8" box="[446,574,1799,1824]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="10" pageNumber="45" unit="mm" value="62.5" valueMax="65.0" valueMin="60.0">60–65 mm</quantity>
|
||
and consistently sub cylindrical in shape as in Ankylopollexia (Norman 2004), slightly amphicoelous with the anterior facet slightly bigger than the posterior (anterior facet height
|
||
<quantity id="DD309B1EFFCF9343BE61F8FDFD65F84E" box="[629,716,1901,1926]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="45" unit="mm" value="60.0">60 mm</quantity>
|
||
,
|
||
<quantity id="DD309B1EFFCF9343BEC8F8FDFC9DF84E" box="[732,820,1901,1926]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="45" unit="mm" value="65.0">65 mm</quantity>
|
||
; anterior width
|
||
<quantity id="DD309B1EFFCF9343BD5DF81FFE36F860" box="[329,415,1935,1960]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="45" unit="mm" value="65.0">65 mm</quantity>
|
||
,
|
||
<quantity id="DD309B1EFFCF9343BDA4F81FFDA1F860" box="[432,520,1935,1960]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="45" unit="mm" value="73.0">73 mm</quantity>
|
||
; posterior height
|
||
<quantity id="DD309B1EFFCF9343BEC8F81FFC9AF860" box="[732,819,1935,1960]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="45" unit="mm" value="60.0">60 mm</quantity>
|
||
,
|
||
<quantity id="DD309B1EFFCF9343BF4AFA25FC1CFA06" box="[862,949,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="45" unit="mm" value="73.0">73 mm</quantity>
|
||
; posterior width
|
||
<quantity id="DD309B1EFFCF9343B894FA25FB7CFA06" box="[1152,1237,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="45" unit="mm" value="75.0">75 mm</quantity>
|
||
,
|
||
<quantity id="DD309B1EFFCF9343B8F1FA25FA95FA06" box="[1253,1340,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="45" unit="mm" value="70.0">70 mm</quantity>
|
||
). The slight amphicoelous condition indicates a close proximity to the sacrum. On the ventral surface, a slight constriction forms a smooth keel. The lateral surfaces of the centrum are longitudinally convex, showing the presence of small vascular foramina. The neural canal is fully open in both articulated vertebrae, being sub-circular in shape. Dorsally, two clearly distinguishable prezygapophysis are arranged in an angle of 45° with respect to one another, as in other iguanodontians (Norman 2004). Posterolateral to the prezygapophyses, the stout transverse processes originate, strongly inflecting dorsally in the partial neural arch. The associated vertebrae
|
||
<collectionCode id="7CD9AE3EFFCF9343B9C0F8B9FA56F88A" box="[1492,1535,1833,1858]" country="France" name="Musee de Lectoure" pageId="10" pageNumber="45">ML</collectionCode>
|
||
452 show a strong lateroventral inflection on the left lateral side and a weaker dorsolateral inflection on the right side. The similar orientation of the transverse processes caused by the same diagenetic alteration further supports the association of these
|
||
<specimenCount id="0CCEFD72FFCE9342BD0FFABEFE61FA8F" box="[283,456,1326,1351]" pageId="11" pageNumber="46" type="generic">two specimens</specimenCount>
|
||
. Similar cases are reported by
|
||
<bibRefCitation id="7E594B0AFFCE9342BC91FADEFE91FAAF" author="Janensch, W." box="[133,312,1358,1383]" pageId="11" pageNumber="46" pagination="105 - 176" refId="ref17818" refString="Janensch, W. 1955. Der Ornithopode Dysalotosaurus der Tendaguruschichten. Palaeontographica-Supplementbande 3: 105 - 176." type="journal article" year="1955">Janensch (1955)</bibRefCitation>
|
||
describing bone beds of
|
||
<emphasis id="28BCEAE9FFCE9342BE4DFADEFD52FAAF" box="[601,763,1358,1383]" italics="true" pageId="11" pageNumber="46">Dysalotosaurs</emphasis>
|
||
. Nevertheless, it is here noted that in Ornithopoda, the transverse processes generally are not directed ventrally, therefore it is suggested the lateral left side has been affected by taphonomical processes more than the right one. The right transverse processes show an inclination which goes from weakly anterodorsal to nearly sub-horizontal, in the case of the most incomplete vertebrae of the pair. Comparing with other taxa (
|
||
<bibRefCitation id="7E594B0AFFCE9342BCFCF9DEFE25F9AF" author="Gilmore, C. W." box="[232,396,1614,1639]" pageId="11" pageNumber="46" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342BD8EF9DEFD92F9AF" author="Norman, D. B." box="[410,571,1614,1639]" pageId="11" pageNumber="46" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFCE9342BE5EF9DEFD2AF9AF" author="Norman, D. B." box="[586,643,1614,1639]" pageId="11" pageNumber="46" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342BE85F9DEFE8FF94F" author="Carpenter, K. & Wilson, Y." pageId="11" pageNumber="46" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342BD26F9FEFD2FF94F" author="Carpenter, K. & Lamanna, M. C." box="[306,646,1646,1671]" pageId="11" pageNumber="46" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342BE85F9FEFE89F96F" author="Carpenter, K. & Galton, P. M." pageId="11" pageNumber="46" pagination="167 - 207" refId="ref16462" refString="Carpenter, K. and Galton, P. M. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter and Galton 2018</bibRefCitation>
|
||
) we note that this degree of inclination is consistent with undistorted specimens. Therefore, it is suggested that the right side has not been strongly affected by taphonomy, the inclination being different on the right lateral side a genuine character to discriminate the position of the vertebrae along the vertebral column. The different inclination of the transverse processes indicates that
|
||
<collectionCode id="7CD9AE3EFFCE9342BECCF8DEFCAAF8AF" box="[728,771,1870,1895]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="46">ML</collectionCode>
|
||
864 is more cranially positioned than
|
||
<collectionCode id="7CD9AE3EFFCE9342BE3CF8FEFDFAF84F" box="[552,595,1902,1927]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="46">ML</collectionCode>
|
||
452 vertebrae. This interpretation is corroborated also by the position of the parapophyses with respect to the diapophyses, being on two well separated planes in
|
||
<collectionCode id="7CD9AE3EFFCE9342B88DFABEFB6DFA8F" box="[1177,1220,1326,1351]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="46">ML</collectionCode>
|
||
864 and almost on the same one in
|
||
<collectionCode id="7CD9AE3EFFCE9342BFACFADEFC4AFAAF" box="[952,995,1358,1383]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="46">ML</collectionCode>
|
||
452 vertebrae, reflecting the general trend within Iguanodontia (Norman 2004). The neural spines rise immediately posterior to the prezygapophyses, with an anterior lamina encased by two lateral grooves, as in
|
||
<taxonomicName id="DDC84D78FFCE9342BF3CFA5EFC64FA2F" authorityName="Marsh" authorityYear="1885" box="[808,973,1486,1511]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCE9342BF3CFA5EFC64FA2F" box="[808,973,1486,1511]" italics="true" pageId="11" pageNumber="46">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCE9342BFCBFA5EFA94FA2F" authorityName="IRSNB" authorityYear="1551" box="[991,1341,1486,1511]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCE9342BFCBFA5EFA94FA2F" box="[991,1341,1486,1511]" italics="true" pageId="11" pageNumber="46">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCE9342B95AFA5EFC7DF9CF" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFCE9342B95AFA5EFC7DF9CF" italics="true" pageId="11" pageNumber="46">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and other ankylopollexians (
|
||
<bibRefCitation id="7E594B0AFFCE9342B930FA7EFA6CF9CF" author="Gilmore, C. W." box="[1316,1477,1518,1543]" pageId="11" pageNumber="46" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342BF3CF99EFC61F9EF" author="Norman, D. B." box="[808,968,1550,1575]" pageId="11" pageNumber="46" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFCE9342BFC2F99EFBA6F9EF" author="Norman, D. B." box="[982,1039,1550,1575]" pageId="11" pageNumber="46" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
, 2004). In the most-complete vertebra of the sample, the neural spine is highly elevated with respect to the centrum as in other ankylopollexians, while dryosaurids have comparatively lower spines (
|
||
<bibRefCitation id="7E594B0AFFCE9342B926F9FEFA14F94F" author="Galton, P. M." box="[1330,1469,1646,1671]" pageId="11" pageNumber="46" pagination="271 - 312" refId="ref17007" refString="Galton, P. M. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton. Palaontologische Zeitschrift 55: 271 - 312." type="journal article" year="1981">Galton 1981</bibRefCitation>
|
||
). The overall shape of the neural spine is rectangular, a characteristic shared both by
|
||
<collectionCode id="7CD9AE3EFFCE9342B859F93EFBD1F90F" box="[1101,1144,1710,1735]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="46">ML</collectionCode>
|
||
452 and 864 individuals and different from the condition shown by
|
||
<taxonomicName id="DDC84D78FFCE9342B8C1F95EFA61F92F" box="[1237,1480,1742,1767]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFCE9342B8C1F95EFA61F92F" box="[1237,1480,1742,1767]" italics="true" pageId="11" pageNumber="46">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC84D78FFCE9342BF4CF97EFBBDF8CF" box="[856,1044,1774,1799]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFCE9342BF4CF97EFBBDF8CF" box="[856,1044,1774,1799]" italics="true" pageId="11" pageNumber="46">C. aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFCE9342B837F97EFB6DF8CF" author="Gilmore, C. W." box="[1059,1220,1774,1799]" pageId="11" pageNumber="46" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCE9342B8DBF97EFCC8F8EF" author="Carpenter, K. & Wilson, Y." pageId="11" pageNumber="46" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
) but similar to that of
|
||
<taxonomicName id="DDC84D78FFCE9342B872F89EFA6BF8EF" authorityName="IRSNB" authorityYear="1551" box="[1126,1474,1806,1831]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCE9342B872F89EFA6BF8EF" box="[1126,1474,1806,1831]" italics="true" pageId="11" pageNumber="46">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
. Moderately deep grooves for the attachment of ossified tendons are visible on the apical-most part of the neural spine. Posteriorly, the postzygapophyses originate from the base of the neural spine.
|
||
</paragraph>
|
||
<caption id="4EB76673FFCC9340BC76F8E3FB0DF86B" ID-DOI="http://doi.org/10.5281/zenodo.11088519" ID-Zenodo-Dep="11088519" httpUri="https://zenodo.org/record/11088519/files/figure.png" pageId="9" pageNumber="44" startId="9.[98,131,1907,1927]" targetBox="[238,1341,1075,1881]" targetPageId="9">
|
||
<paragraph id="1A7736FBFFCC9340BC76F8E3FB0DF86B" pageId="9" pageNumber="44">
|
||
Fig. 5. Cranial material of Ankylopollexia indet. from the Lourinhã municipality,
|
||
<collectingCountry id="62DF766BFFCC9340BF73F8E3FC1FF84F" box="[871,950,1907,1927]" name="Portugal" pageId="9" pageNumber="44">Portugal</collectingCountry>
|
||
, Lourinhã Formation, Kimmeridgian–Tithonian. Dentary
|
||
<collectionCode id="7CD9AE3EFFCC9340BC76F81FFF2AF86B" box="[98,131,1935,1955]" country="France" name="Musee de Lectoure" pageId="9" pageNumber="44">ML</collectionCode>
|
||
818, in medial (A
|
||
<subScript id="864C34BEFFCC9340BD3CF808FE99F86E" attach="left" box="[296,304,1944,1958]" fontSize="6" pageId="9" pageNumber="44">1</subScript>
|
||
, A
|
||
<subScript id="864C34BEFFCC9340BD5FF808FEFAF86E" attach="right" box="[331,339,1944,1958]" fontSize="6" pageId="9" pageNumber="44">5</subScript>
|
||
), lateral (A
|
||
<subScript id="864C34BEFFCC9340BDA8F808FE6DF86E" attach="right" box="[444,452,1944,1958]" fontSize="6" pageId="9" pageNumber="44">2</subScript>
|
||
, A
|
||
<subScript id="864C34BEFFCC9340BDCAF808FE4FF86E" attach="left" box="[478,486,1944,1958]" fontSize="6" pageId="9" pageNumber="44">6</subScript>
|
||
), and dorsal (A
|
||
<subScript id="864C34BEFFCC9340BE60F808FDD5F86E" attach="left" box="[628,636,1944,1958]" fontSize="6" pageId="9" pageNumber="44">3</subScript>
|
||
, A
|
||
<subScript id="864C34BEFFCC9340BE83F808FD36F86E" attach="right" box="[663,671,1944,1958]" fontSize="6" pageId="9" pageNumber="44">7</subScript>
|
||
) views, detail of the dentary/surangular contact (A
|
||
<subScript id="864C34BEFFCC9340B879F808FBDCF86E" attach="right" box="[1133,1141,1944,1958]" fontSize="6" pageId="9" pageNumber="44">4</subScript>
|
||
, A
|
||
<subScript id="864C34BEFFCC9340B89BF808FB3EF86E" attach="left" box="[1167,1175,1944,1958]" fontSize="6" pageId="9" pageNumber="44">8</subScript>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4EB76673FFCF9343BC83FAD9FAFEFA5D" ID-DOI="http://doi.org/10.5281/zenodo.10626658" ID-Zenodo-Dep="10626658" httpUri="https://zenodo.org/record/10626658/files/figure.png" pageId="10" pageNumber="45" startId="10.[151,184,1353,1373]" targetBox="[153,1522,174,1319]" targetPageId="10" targetType="figure">
|
||
<paragraph id="1A7736FBFFCF9343BC83FAD9FAFEFA5D" blockId="10.[151,1535,1351,1432]" pageId="10" pageNumber="45">
|
||
Fig. 6. Dorsal vertebrae Ankylopollexia indet. from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian.
|
||
<emphasis id="28BCEAE9FFCF9343B97EFAD7FAD3FA94" bold="true" box="[1386,1402,1351,1372]" pageId="10" pageNumber="45">A</emphasis>
|
||
. Partial neural arch, ML 864 in anterior (A
|
||
<subScript id="864C34BEFFCF9343BD9BFAFEFE3EFAB4" attach="right" box="[399,407,1390,1404]" fontSize="6" pageId="10" pageNumber="45">1</subScript>
|
||
), posterior (A
|
||
<subScript id="864C34BEFFCF9343BE02FAFEFDB7FAB4" attach="right" box="[534,542,1390,1404]" fontSize="6" pageId="10" pageNumber="45">2</subScript>
|
||
), lateral (A
|
||
<subScript id="864C34BEFFCF9343BE91FAFEFD24FAB4" attach="right" box="[645,653,1390,1404]" fontSize="6" pageId="10" pageNumber="45">3</subScript>
|
||
), and dorsal (A
|
||
<subScript id="864C34BEFFCF9343BF0CFAFEFC89FAB4" attach="left" box="[792,800,1390,1404]" fontSize="6" pageId="10" pageNumber="45">4</subScript>
|
||
) views.
|
||
<emphasis id="28BCEAE9FFCF9343BF78FAF4FCD2FAB1" bold="true" box="[876,891,1380,1401]" pageId="10" pageNumber="45">B</emphasis>
|
||
,
|
||
<emphasis id="28BCEAE9FFCF9343BF91FAF3FC3CFAB1" bold="true" box="[901,917,1379,1401]" pageId="10" pageNumber="45">C</emphasis>
|
||
. Dorsal vertebrae, ML 452a, complete (
|
||
<emphasis id="28BCEAE9FFCF9343B8EFFAF4FAA3FAB1" bold="true" box="[1275,1290,1380,1401]" pageId="10" pageNumber="45">B</emphasis>
|
||
) and ML 452b, incomplete (
|
||
<emphasis id="28BCEAE9FFCF9343BC8BFAEFFF06FA5D" bold="true" box="[159,175,1407,1429]" pageId="10" pageNumber="45">C</emphasis>
|
||
) specimens, in anterior (B
|
||
<subScript id="864C34BEFFCF9343BDB4FA1AFE01FA50" attach="left" box="[416,424,1418,1432]" fontSize="6" pageId="10" pageNumber="45">1</subScript>
|
||
, C
|
||
<subScript id="864C34BEFFCF9343BDD7FA1AFE62FA50" attach="right" box="[451,459,1418,1432]" fontSize="6" pageId="10" pageNumber="45">1</subScript>
|
||
), posterior (B
|
||
<subScript id="864C34BEFFCF9343BE5EFA1AFDFBFA50" attach="left" box="[586,594,1418,1432]" fontSize="6" pageId="10" pageNumber="45">2</subScript>
|
||
, C
|
||
<subScript id="864C34BEFFCF9343BE79FA1AFDDCFA50" attach="left" box="[621,629,1418,1432]" fontSize="6" pageId="10" pageNumber="45">2</subScript>
|
||
), lateral (B
|
||
<subScript id="864C34BEFFCF9343BEC8FA1AFD4DFA50" attach="left" box="[732,740,1418,1432]" fontSize="6" pageId="10" pageNumber="45">3</subScript>
|
||
, C
|
||
<subScript id="864C34BEFFCF9343BEEBFA1AFCAEFA50" attach="left" box="[767,775,1418,1432]" fontSize="6" pageId="10" pageNumber="45">3</subScript>
|
||
), dorsal (B
|
||
<subScript id="864C34BEFFCF9343BF79FA1AFCDCFA50" attach="left" box="[877,885,1418,1432]" fontSize="6" pageId="10" pageNumber="45">4</subScript>
|
||
, C
|
||
<subScript id="864C34BEFFCF9343BF84FA1AFC31FA50" attach="left" box="[912,920,1418,1432]" fontSize="6" pageId="10" pageNumber="45">4</subScript>
|
||
), and ventral (digitally modified) (B
|
||
<subScript id="864C34BEFFCF9343B8F0FA1AFB45FA50" attach="left" box="[1252,1260,1418,1432]" fontSize="6" pageId="10" pageNumber="45">5</subScript>
|
||
, C
|
||
<subScript id="864C34BEFFCF9343B912FA1AFAA7FA50" attach="left" box="[1286,1294,1418,1432]" fontSize="6" pageId="10" pageNumber="45">5</subScript>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1A7736FBFFCE9342BC70FF33FC7FFD67" pageId="11" pageNumber="46">
|
||
A
|
||
<subScript id="864C34BEFFCE9342BC6AFF22FF27FF00" attach="left" box="[126,142,178,200]" fontSize="9" pageId="11" pageNumber="46">1</subScript>
|
||
B C D E F A dia par 2 dia par par dia par par dia par par dia dia I J K L G H dia dia par dia dia dia par par par par
|
||
</paragraph>
|
||
<caption id="4EB76673FFCE9342BC76FCCCFC43FB30" ID-DOI="http://doi.org/10.5281/zenodo.10626660" ID-Zenodo-Dep="10626660" httpUri="https://zenodo.org/record/10626660/files/figure.png" pageId="11" pageNumber="46" startId="11.[98,131,860,880]" targetBox="[542,931,494,833]" targetPageId="11" targetType="figure">
|
||
<paragraph id="1A7736FBFFCE9342BC76FCCCFC43FB30" blockId="11.[98,1482,858,1272]" pageId="11" pageNumber="46">
|
||
Fig. 7. Comparative dorsal vertebrae table of selected Ankylopollexians from the Late Jurassic and Early Cretaceous.
|
||
<emphasis id="28BCEAE9FFCE9342B8D0FCCAFB7DFCA7" bold="true" box="[1220,1236,858,879]" pageId="11" pageNumber="46">A</emphasis>
|
||
,
|
||
<emphasis id="28BCEAE9FFCE9342B8F0FCCAFB5AFCA7" bold="true" box="[1252,1267,858,879]" pageId="11" pageNumber="46">B</emphasis>
|
||
. Ankylopollexia indet. Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian.
|
||
<emphasis id="28BCEAE9FFCE9342BF5DFCE6FCF0FC43" bold="true" box="[841,857,886,907]" pageId="11" pageNumber="46">A</emphasis>
|
||
. ML 452 in lateral view (A
|
||
<subScript id="864C34BEFFCE9342B84AFC11FBCFFC47" attach="right" box="[1118,1126,897,911]" fontSize="6" pageId="11" pageNumber="46">1</subScript>
|
||
, A
|
||
<subScript id="864C34BEFFCE9342B896FC11FB23FC47" attach="left" box="[1154,1162,897,911]" fontSize="6" pageId="11" pageNumber="46">2</subScript>
|
||
).
|
||
<emphasis id="28BCEAE9FFCE9342B88BFCE6FB07FC43" bold="true" box="[1183,1198,886,907]" pageId="11" pageNumber="46">B</emphasis>
|
||
. ML 864 in right lateral view.
|
||
<emphasis id="28BCEAE9FFCE9342BC76FC02FFDBFC60" bold="true" box="[98,114,914,936]" pageId="11" pageNumber="46">C</emphasis>
|
||
. “
|
||
<taxonomicName id="DDC84D78FFCE9342BC9CFC04FF7CFC60" authority="SHN.LPP" authorityName="McDonald" authorityYear="2011" box="[136,213,916,936]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCE9342BC9CFC04FF7CFC60" box="[136,213,916,936]" italics="true" pageId="11" pageNumber="46">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
” SHN.LPP 015 in left lateral view; Praia da corva, Torres Vedras Municipality, Portugal, Lourinhã Formation (Kimmeridgian–Tithonian).
|
||
<emphasis id="28BCEAE9FFCE9342BC76FC3EFFDBFC0B" bold="true" box="[98,114,942,963]" pageId="11" pageNumber="46">D</emphasis>
|
||
. “
|
||
<taxonomicName id="DDC84D78FFCE9342BC9CFC20FF7CFC0C" authorityName="McDonald" authorityYear="2011" box="[136,213,944,964]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCE9342BC9CFC20FF7CFC0C" box="[136,213,944,964]" italics="true" pageId="11" pageNumber="46">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="DDC84D78FFCE9342BCF2FC20FE23FC0C" authority="CM" authorityName="CM" box="[230,394,944,964]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFCE9342BCF2FC20FEC9FC0C" box="[230,352,944,964]" italics="true" pageId="11" pageNumber="46">aphanoecetes</emphasis>
|
||
CM
|
||
</taxonomicName>
|
||
11337 in left lateral view; East end of Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah (USA), Morrison Formation (Kimmeridgian–Tithonian).
|
||
<emphasis id="28BCEAE9FFCE9342BE31FC5AFD9DFC17" bold="true" box="[549,564,970,991]" pageId="11" pageNumber="46">E</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BE56FC5CFCAFFC28" box="[578,774,972,992]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFCE9342BE56FC5CFCAFFC28" box="[578,774,972,992]" italics="true" pageId="11" pageNumber="46">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
(unnumbered specimen) in left lateral view; Bone Cabin Quarry, Wyoming USA), Morrison Formation (Kimmeridgian–Tithonian).
|
||
<emphasis id="28BCEAE9FFCE9342BE63FC76FD2CFC33" bold="true" box="[631,645,998,1019]" pageId="11" pageNumber="46">F</emphasis>
|
||
. “
|
||
<taxonomicName id="DDC84D78FFCE9342BE8BFC78FD53FC34" authorityName="Seeley" authorityYear="1888" box="[671,762,1000,1020]" class="Reptilia" genus="Cumnoria" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCE9342BE8BFC78FD53FC34" box="[671,762,1000,1020]" italics="true" pageId="11" pageNumber="46">Cumnoria</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="DDC84D78FFCE9342BF1AFC78FC64FC34" authority="OUM. J." authorityName="OUM. J." box="[782,973,1000,1020]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFCE9342BF1AFC78FCD8FC34" box="[782,881,1000,1020]" italics="true" pageId="11" pageNumber="46">prestwichii</emphasis>
|
||
OUM. J.
|
||
</taxonomicName>
|
||
3303 in lateral view; Oxford, UK, Kimmeridge Clay Formation (Kimmeridgian–Tithonian).
|
||
<emphasis id="28BCEAE9FFCE9342BDDDFB92FE72FBD0" bold="true" box="[457,475,1026,1048]" pageId="11" pageNumber="46">G</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BDFDFB94FC81FBD0" authority="UMNH VP" authorityName="UMNH VP" box="[489,808,1028,1048]" class="Reptilia" genus="Hippodraco" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="scutodens">
|
||
<emphasis id="28BCEAE9FFCE9342BDFDFB94FD1FFBD0" box="[489,694,1028,1048]" italics="true" pageId="11" pageNumber="46">Hippodraco scutodens</emphasis>
|
||
UMNH VP
|
||
</taxonomicName>
|
||
20208 in left lateral view; Andrew’s Site, Grand County, Utah; Upper Yellow Cat Memberof the Cedar Mountain Formation (upper Barremian–lowermost Aptian).
|
||
<emphasis id="28BCEAE9FFCE9342BFA1FB8EFC6EFBFB" bold="true" box="[949,967,1054,1075]" pageId="11" pageNumber="46">H</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BFC7FBB0FAABFBFC" authority="UMNH VP" authorityName="UMNH VP" box="[979,1282,1056,1076]" class="Reptilia" genus="Iguanacolossus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="fortis">
|
||
<emphasis id="28BCEAE9FFCE9342BFC7FBB0FB3DFBFC" box="[979,1172,1056,1076]" italics="true" pageId="11" pageNumber="46">Iguanacolossus fortis</emphasis>
|
||
UMNH VP
|
||
</taxonomicName>
|
||
20205 in right lateral view; Don’s Ridge, Grand County, Utah, Lower Yellow Cat Member, Cedar Mountain Formation (?lower Barremian).
|
||
<emphasis id="28BCEAE9FFCE9342B8B6FBAAFB02FB87" bold="true" box="[1186,1195,1082,1103]" pageId="11" pageNumber="46">I</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342B8A3FBACFF7EFBA4" authority="IRSNB 1551" authorityName="IRSNB" authorityYear="1551" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCE9342B8A3FBACFA60FB98" box="[1207,1481,1084,1104]" italics="true" pageId="11" pageNumber="46">Mantellisaurus atherfieldensis</emphasis>
|
||
IRSNB 1551
|
||
</taxonomicName>
|
||
in left lateral view; Isle of Wight, Wessex Formation (Barremian).
|
||
<emphasis id="28BCEAE9FFCE9342BF2FFBC6FCEFFBA3" bold="true" box="[827,838,1110,1131]" pageId="11" pageNumber="46">J</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BF47FBC8FBCCFBA4" authority="NHMUK R" authorityName="NHMUK R" box="[851,1125,1112,1132]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="dawsoni">
|
||
<emphasis id="28BCEAE9FFCE9342BF47FBC8FC5DFBA4" box="[851,1012,1112,1132]" italics="true" pageId="11" pageNumber="46">Barilium dawsoni</emphasis>
|
||
NHMUK R
|
||
</taxonomicName>
|
||
798 in left lateral view; Shornden, East Sussex, UK, Wadhurst Clay Formation (Valanginian).
|
||
<emphasis id="28BCEAE9FFCE9342BE5DFBE2FDF2FB4F" bold="true" box="[585,603,1138,1159]" pageId="11" pageNumber="46">K</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BE71FBE4FC3BFB40" authority="NHMUK R" authorityName="NHMUK R" box="[613,914,1140,1160]" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFCE9342BE71FBE4FC8BFB40" box="[613,802,1140,1160]" italics="true" pageId="11" pageNumber="46">Hypselospinus fittoni</emphasis>
|
||
NHMUK R
|
||
</taxonomicName>
|
||
604 in lateral view; Shornden Quarry, Hastings, UK, Wadhurst Clay Formation (Valanginian).
|
||
<emphasis id="28BCEAE9FFCE9342BD6EFB1EFE20FB6B" bold="true" box="[378,393,1166,1187]" pageId="11" pageNumber="46">L</emphasis>
|
||
.
|
||
<taxonomicName id="DDC84D78FFCE9342BD81FB00FD62FB6C" authority="IRSNB" authorityName="IRSNB" box="[405,715,1168,1188]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFCE9342BD81FB00FD2DFB6C" box="[405,644,1168,1188]" italics="true" pageId="11" pageNumber="46">Iguanodon bernissartensis</emphasis>
|
||
IRSNB
|
||
</taxonomicName>
|
||
“Individu S” in left lateral view; Bernissart, Belgium, Sainte Barbe Clays Formation Barremian). Abbreviations: dia, diapophysis; par, parapophysis. Scale bars 100 mm. Re-drawn from: C, Escaso 2014: fig. 6.5; D,
|
||
<bibRefCitation id="7E594B0AFFCE9342B917FB3CFF3BFB14" author="Carpenter, K. & Wilson, Y." pageId="11" pageNumber="46" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
: fig. 11; E,
|
||
<bibRefCitation id="7E594B0AFFCE9342BD14FB58FE54FB14" author="Carpenter, K. & Galton, P. M." box="[256,509,1224,1244]" pageId="11" pageNumber="46" pagination="167 - 207" refId="ref16462" refString="Carpenter, K. and Galton, P. M. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter and Galton 2018</bibRefCitation>
|
||
: fig. 22D; F,
|
||
<bibRefCitation id="7E594B0AFFCE9342BE6EFB58FCF5FB14" author="Galton, P. M. & Powell, H. P." box="[634,860,1224,1244]" pageId="11" pageNumber="46" pagination="411 - 443" refId="ref17197" refString="Galton, P. M. and Powell, H. P. 1980. The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology 23: 411 - 443." type="journal article" year="1980">Galton and Powell 1980</bibRefCitation>
|
||
: fig. 4; G,
|
||
<bibRefCitation id="7E594B0AFFCE9342BFD0FB58FBCEFB14" author="McDonald, A. T. & Kirkland, J. I. & DeBlieux, D. D. & Madsen, S. K. & Cavin, J. & Milner, A. R. & Panzarin, L." box="[964,1127,1224,1244]" pageId="11" pageNumber="46" pagination="14075" refId="ref18478" refString="McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R., and Panzarin, L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5 (11): e 14075." type="journal article" year="2010">McDonald 2010b</bibRefCitation>
|
||
: fig. 27; H,
|
||
<bibRefCitation id="7E594B0AFFCE9342B8CDFB58FAD5FB14" author="McDonald, A. T. & Kirkland, J. I. & DeBlieux, D. D. & Madsen, S. K. & Cavin, J. & Milner, A. R. & Panzarin, L." box="[1241,1404,1224,1244]" pageId="11" pageNumber="46" pagination="14075" refId="ref18478" refString="McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R., and Panzarin, L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5 (11): e 14075." type="journal article" year="2010">McDonald 2010b</bibRefCitation>
|
||
: fig. 10;,
|
||
<bibRefCitation id="7E594B0AFFCE9342BC60FB74FF5AFB30" author="Norman, D. B." box="[116,243,1252,1272]" pageId="11" pageNumber="46" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
: fig. 37; J,
|
||
<bibRefCitation id="7E594B0AFFCE9342BD42FB74FE7CFB30" author="Norman, D. B." box="[342,469,1252,1272]" pageId="11" pageNumber="46" pagination="165 - 194" refId="ref18777" refString="Norman, D. B. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">Norman 2011</bibRefCitation>
|
||
: fig. 4;
|
||
<bibRefCitation id="7E594B0AFFCE9342BE0CFB74FD1BFB30" author="Carpenter, K. & Lamanna, M. C." box="[536,690,1252,1272]" pageId="11" pageNumber="46" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">K, Norman 2015</bibRefCitation>
|
||
: fig. 22; L,
|
||
<bibRefCitation id="7E594B0AFFCE9342BF0EFB74FC30FB30" author="Norman, D. B." box="[794,921,1252,1272]" pageId="11" pageNumber="46" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
: fig. 31).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1A7736FBFFC99345BC83FF30FD36FCC9" blockId="12.[151,825,160,1581]" pageId="12" pageNumber="47">
|
||
<emphasis id="28BCEAE9FFC99345BC83FF30FE3EFF71" box="[151,407,160,185]" italics="true" pageId="12" pageNumber="47">Appendicular skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFC99345BDBDFF30FDBAFF71" box="[425,531,160,185]" italics="true" pageId="12" pageNumber="47">Coracoid</emphasis>
|
||
(
|
||
<figureCitation id="82F32A7EFFC99345BE32FF30FD28FF71" box="[550,641,160,185]" captionStart="Fig" captionStartId="13.[98,131,1438,1458]" captionTargetBox="[128,1434,165,1401]" captionTargetId="figure-254@13.[176,1438,189,1401]" captionTargetPageId="13" captionText="Fig. 8. Ankylopollexian appendicular skeleton from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Coracoid ML 2206 (A), scapula ML 2042 (B), in lateral (A1, B1) and medial (A2, B2) views." figureDoi="http://doi.org/10.5281/zenodo.10626662" httpUri="https://zenodo.org/record/10626662/files/figure.png" pageId="12" pageNumber="47">Fig. 8A</figureCitation>
|
||
): The left coracoid
|
||
<collectionCode id="7CD9AE3EFFC99345BCC4FF50FF50FF11" box="[208,249,192,217]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
2206 is complete with the marginal sides slightly eroded. It is sub-rectangular in shape, bowed medio-laterally, and measures
|
||
<quantity id="DD309B1EFFC99345BD8DFE91FE55FED2" box="[409,508,257,282]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="47" unit="mm" value="130.0">130 mm</quantity>
|
||
in antero-posterior length and
|
||
<quantity id="DD309B1EFFC99345BCDFFEB2FE82FEF3" box="[203,299,290,315]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="12" pageNumber="47" unit="mm" value="108.0">108 mm</quantity>
|
||
in dorsoventral width. The dorsal margin is straight, while the ventral margin is deeply arched. The scapular surface is slightly concave and relegated to the dorsal-most part of the coracoid, while the glenoid surface is wider and slightly convex. Ventrally, the glenoid deflects abruptly, forming with the bowed ventral margin a conspicuous labrum. The coracoid foramen is dorsally located and totally enclosed on the lateral surface. On the medial surface, instead, it is open along the scapula-coracoid suture as in
|
||
<taxonomicName id="DDC84D78FFC99345BCC8FDD6FE28FD97" authorityName="Marsh" authorityYear="1885" box="[220,385,582,607]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFC99345BCC8FDD6FE28FD97" box="[220,385,582,607]" italics="true" pageId="12" pageNumber="47">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BD80FDD6FDA6FD97" authorityName="Mantell" authorityYear="1825" box="[404,527,582,607]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFC99345BD80FDD6FDA6FD97" box="[404,527,582,607]" italics="true" pageId="12" pageNumber="47">Iguanodon</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BE36FDD6FD64FD97" authorityName="Paul" authorityYear="2007" box="[546,717,582,607]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFC99345BE36FDD6FD64FD97" box="[546,717,582,607]" italics="true" pageId="12" pageNumber="47">Mantellisaurus</emphasis>
|
||
</taxonomicName>
|
||
, and derived iguanodontians (
|
||
<bibRefCitation id="7E594B0AFFC99345BD8CFDF6FD90FDB7" author="Gilmore, C. W." box="[408,569,614,639]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BE52FDF6FD4CFDB7" author="Norman, D. B." box="[582,741,614,639]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BEE6FDF6FC85FDB7" author="Norman, D. B." box="[754,812,614,639]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
). With derived iguanodontians it also shares the width/length ratio falling between 70–100% (
|
||
<bibRefCitation id="7E594B0AFFC99345BE14FD37FD52FD08" author="Dieudonne, P. - E. & Tortosa, T. & Fernandez-Baldor, F. T. & Canudo, J. I. & Diaz-Martinez, I." box="[512,763,679,704]" pageId="12" pageNumber="47" pagination="0156251" refId="ref16604" refString="Dieudonne, P. - E., Tortosa, T., Fernandez-Baldor, F. T., Canudo, J. I., and Diaz-Martinez, I. 2016. An unexpected early rhabdodontid from Europe (Lower Cretaceous of Salas de los Infantes, Burgos Province, Spain) and a re-examination of basal iguanodontian relationships. PloS ONE 11 (6): e 0156251." type="journal article" year="2016">Dieudonné et al. 2016</bibRefCitation>
|
||
). The sternal process is wide and broad, a plesiomorphic condition within Ornithopoda (
|
||
<bibRefCitation id="7E594B0AFFC99345BD92FD78FD3AFCC9" author="Weishampel, D. B. & Jianu, C. - M. & Csiki, Z. & Norman, D. B." box="[390,659,744,769]" pageId="12" pageNumber="47" pagination="65 - 123" refId="ref19649" refString="Weishampel, D. B., Jianu, C. - M., Csiki, Z., and Norman, D. B. 2003. Osteology and phylogeny of Zalmoxes, an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology 1: 65 - 123." type="journal article" year="2003">Weishampel et al. 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFC99345BCA9FC99FD9AF9E5" blockId="12.[151,825,160,1581]" pageId="12" pageNumber="47">
|
||
<emphasis id="28BCEAE9FFC99345BCA9FC99FEB1FCEA" box="[189,280,777,802]" italics="true" pageId="12" pageNumber="47">Scapula</emphasis>
|
||
(
|
||
<figureCitation id="82F32A7EFFC99345BD33FC99FED2FCEA" box="[295,379,777,802]" captionStart="Fig" captionStartId="13.[98,131,1438,1458]" captionTargetBox="[128,1434,165,1401]" captionTargetId="figure-254@13.[176,1438,189,1401]" captionTargetPageId="13" captionText="Fig. 8. Ankylopollexian appendicular skeleton from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Coracoid ML 2206 (A), scapula ML 2042 (B), in lateral (A1, B1) and medial (A2, B2) views." figureDoi="http://doi.org/10.5281/zenodo.10626662" httpUri="https://zenodo.org/record/10626662/files/figure.png" pageId="12" pageNumber="47">Fig. 8B</figureCitation>
|
||
):
|
||
<collectionCode id="7CD9AE3EFFC99345BD9BFC99FE10FCEA" box="[399,441,777,802]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
2042 is an incomplete and eroded scapula measuring
|
||
<quantity id="DD309B1EFFC99345BD65FCB9FE7BFC8A" box="[369,466,809,834]" metricMagnitude="-1" metricUnit="m" metricValue="4.4" pageId="12" pageNumber="47" unit="mm" value="440.0">440 mm</quantity>
|
||
, missing the distal-most part of the blade and being partially eroded in the proximal end. The missing part is estimated to be not more than 10% of the total length of the whole scapular blade. The preserved dorsal and ventral margins appear to be parallel and progressively converging towards the distal part of the blade as seen in
|
||
<taxonomicName id="DDC84D78FFC99345BD09FC7CFDD5FBCD" authorityName="IRSNB" authorityYear="1551" box="[285,636,1004,1029]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BD09FC7CFDD5FBCD" box="[285,636,1004,1029]" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC84D78FFC99345BEAAFC7CFEEBFBEE" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345BEAAFC7CFEEBFBEE" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BD40FB9DFE5CFBEE" author="Norman, D. B." box="[340,501,1037,1062]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BE10FB9DFD94FBEE" author="Norman, D. B." box="[516,573,1037,1062]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
), differently from the condition exhibited by
|
||
<taxonomicName id="DDC84D78FFC99345BD83FBBDFD2EFB8E" box="[407,647,1069,1094]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFC99345BD83FBBDFD2EFB8E" box="[407,647,1069,1094]" italics="true" pageId="12" pageNumber="47">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BE80FBBDFF1CFBAF" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFC99345BE80FBBDFF1CFBAF" italics="true" pageId="12" pageNumber="47">C. aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345BCEDFBDEFE30FBAF" box="[249,409,1102,1127]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFC99345BCEDFBDEFE30FBAF" box="[249,409,1102,1127]" italics="true" pageId="12" pageNumber="47">C. prestwichii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BDBEFBDEFDE2FBAF" author="Gilmore, C. W." box="[426,587,1102,1127]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BE4DFBDEFEAAFB4F" author="Carpenter, K. & Wilson, Y." pageId="12" pageNumber="47" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BD04FBFEFE6FFB4F" author="McDonald, A. T." box="[272,454,1134,1159]" pageId="12" pageNumber="47" pagination="52 - 68" refId="ref18369" refString="McDonald, A. T. 2011. The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa 2783: 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="62DF766BFFC99345BD04FBFEFE2FFB4F" box="[272,390,1134,1159]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BDC0FBFEFC84FB4F" author="Carpenter, K. & Lamanna, M. C." box="[468,813,1134,1159]" pageId="12" pageNumber="47" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna 2015</bibRefCitation>
|
||
), which display a strong distal expansion towards the middle of the blade. The scapular blade is dorsoventrally bowed as in
|
||
<taxonomicName id="DDC84D78FFC99345BCA1FB5FFDA5FB20" authorityName="IRSNB" authorityYear="1551" box="[181,524,1231,1256]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BCA1FB5FFDA5FB20" box="[181,524,1231,1256]" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC84D78FFC99345BE57FB5FFF7EFAC1" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345BE57FB5FFF7EFAC1" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BCF1FB60FE2BFAC1" author="Norman, D. B." box="[229,386,1264,1289]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BD99FB60FE6EFAC1" author="Norman, D. B." box="[397,455,1264,1289]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
). The proximal part flares gently, being concave on the lateral surface. The acromion process is slightly rounded and forwardly directed, the underlying coracoid suture is straight and ventrally deflects into the glenoid. Contrarily to
|
||
<taxonomicName id="DDC84D78FFC99345BD67FAE2FDCFFA43" box="[371,614,1394,1419]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFC99345BD67FAE2FDCFFA43" box="[371,614,1394,1419]" italics="true" pageId="12" pageNumber="47">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BE62FAE2FC98FA43" box="[630,817,1394,1419]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFC99345BE62FAE2FC98FA43" box="[630,817,1394,1419]" italics="true" pageId="12" pageNumber="47">C. aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345BCDCFA02FECEFA63" box="[200,359,1426,1451]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFC99345BCDCFA02FECEFA63" box="[200,359,1426,1451]" italics="true" pageId="12" pageNumber="47">C. prestwichii</emphasis>
|
||
</taxonomicName>
|
||
but similar to
|
||
<taxonomicName id="DDC84D78FFC99345BE1BFA02FEECFA04" authority=", ML 2042" authorityName=", ML" authorityYear="2042" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BE1BFA02FF66FA04" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
, ML 2042
|
||
</taxonomicName>
|
||
does not possess high protuberances corresponding to the acromion process and glenoid (
|
||
<bibRefCitation id="7E594B0AFFC99345BE82FA43FC9AFA24" author="Gilmore, C. W." box="[662,819,1491,1516]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BC83FA64FE9BF9C5" author="Norman, D. B." box="[151,306,1524,1549]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BD29FA64FEDCF9C5" author="Norman, D. B." box="[317,373,1524,1549]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BD94FA64FD1EF9C5" author="Carpenter, K. & Wilson, Y." box="[384,695,1524,1549]" pageId="12" pageNumber="47" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BED6FA64FF64F9E5" author="McDonald, A. T." pageId="12" pageNumber="47" pagination="52 - 68" refId="ref18369" refString="McDonald, A. T. 2011. The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa 2783: 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="62DF766BFFC99345BED6FA64FC91F9C5" box="[706,824,1524,1549]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BCCCF984FD8EF9E5" author="Carpenter, K. & Lamanna, M. C." box="[216,551,1556,1581]" pageId="12" pageNumber="47" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFC99345BC83F9DAFA2FFDD6" blockId="12.[151,827,1610,1960]" lastBlockId="12.[862,1535,160,542]" pageId="12" pageNumber="47">
|
||
<emphasis id="28BCEAE9FFC99345BC83F9DAFF5EF9AB" box="[151,247,1610,1635]" italics="true" pageId="12" pageNumber="47">Remarks</emphasis>
|
||
.—
|
||
<emphasis id="28BCEAE9FFC99345BD0FF9DAFE7CF9AB" box="[283,469,1610,1635]" italics="true" pageId="12" pageNumber="47">Cranial skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFC99345BDF2F9DAFDEBF9AB" box="[486,578,1610,1635]" italics="true" pageId="12" pageNumber="47">Dentary</emphasis>
|
||
:
|
||
<collectionCode id="7CD9AE3EFFC99345BE47F9DAFDD4F9AB" box="[595,637,1610,1635]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
818 shares with other ankylopollexians, such as
|
||
<taxonomicName id="DDC84D78FFC99345BE1AF9FAFD40F94B" box="[526,745,1642,1667]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC99345BE1AF9FAFD18F94B" box="[526,689,1642,1667]" italics="true" pageId="12" pageNumber="47">Camptosaurus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BEE8F9FAFE14F96C" authorityName=", ML" authorityYear="2042" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BEE8F9FAFE14F96C" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BDC4F91BFD05F96C" authorityName="Brill & Carpenter" authorityYear="2007" box="[464,684,1675,1700]" genus="Theiophytalia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="kerri">
|
||
<emphasis id="28BCEAE9FFC99345BDC4F91BFD05F96C" box="[464,684,1675,1700]" italics="true" pageId="12" pageNumber="47">Theiophytalia kerri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BEABF91BFE96F90C" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345BEABF91BFE96F90C" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345BD94F93BFDDBF90C" authorityName="McDonald, Barrett & Chapman" authorityYear="2010" box="[384,626,1707,1732]" class="Reptilia" genus="Kukufeldia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="tilgatensis">
|
||
<emphasis id="28BCEAE9FFC99345BD94F93BFDDBF90C" box="[384,626,1707,1732]" italics="true" pageId="12" pageNumber="47">Kukufeldia tilgatensis</emphasis>
|
||
</taxonomicName>
|
||
, parallel margins of the dentary, closed packed alveoli, and a highly emarginated parapet-like structure which constrains the tooth row (
|
||
<bibRefCitation id="7E594B0AFFC99345BC8BF89DFEEAF8EE" author="Gilmore, C. W." box="[159,323,1805,1830]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BD41F89DFE5FF8EE" author="Norman, D. B." box="[341,502,1805,1830]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BE1CF89DFDE9F8EE" author="Norman, D. B." box="[520,576,1805,1830]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BE46F89DFF78F88E" author="Brill, K. & Carpenter, K." pageId="12" pageNumber="47" pagination="49 - 67" refId="ref16254" refString="Brill, K. and Carpenter, K. 2006. A description of a new ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of Camptosaurus. In: K. Carpenter (ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, 49 - 67. Indiana University Press, Bloomington." type="book chapter" year="2006">Brill and Carpenter 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BCF4F8BDFE4CF88E" author="McDonald, A. T. & Barrett, P. M. & Chapman, S. D." box="[224,485,1837,1862]" pageId="12" pageNumber="47" pagination="1 - 43" refId="ref18428" refString="McDonald, A. T., Barrett, P. M., and Chapman, S. D. 2010 a. A new basal iguanodont (Dinosauria: Ornithischia) from the Wealden (Lower Cretaceous) of England. Zootaxa 2569: 1 - 43." type="journal article" year="2010">
|
||
<collectingCountry id="62DF766BFFC99345BCF4F8BDFEFCF88E" box="[224,341,1837,1862]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
et al. 2010a
|
||
</bibRefCitation>
|
||
). These characters suggest an ankylopollexian affinity for
|
||
<collectionCode id="7CD9AE3EFFC99345BDDAF8DEFE51F8AF" box="[462,504,1870,1895]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
818. However, it differs from thesetaxainhavingamorestronglyinclinedcoronoidprocess, a condition shared with dryosaurids and
|
||
<taxonomicName id="DDC84D78FFC99345BE4BF81FFC90F860" box="[607,825,1935,1960]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC99345BE4BF81FFCAAF860" box="[607,771,1935,1960]" italics="true" pageId="12" pageNumber="47">Tenontosaurus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BF71FF30FC44FF71" author="Galton, P. M." box="[869,1005,160,185]" pageId="12" pageNumber="47" pagination="207 - 243" refId="ref17040" refString="Galton, P. M. 1983. The cranial anatomy of Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and East Africa, with a review of hypsilophodontids from the Upper Jurassic of North America. Geologica et Palaeontologica 17: 207 - 243." type="journal article" year="1983">Galton 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345BFE2FF30FB27FF71" author="Thomas, D. A." box="[1014,1166,160,185]" pageId="12" pageNumber="47" pagination="1 - 99" refId="ref19483" refString="Thomas, D. A. 2015. The cranial anatomy of Tenontosaurus tilletti Ostrom, 1970 (Dinosauria, Ornithopoda). Palaeontologia Electronica 18 (2): 1 - 99." type="journal article" year="2015">Thomas 2015</bibRefCitation>
|
||
). Furthermore,
|
||
<collectionCode id="7CD9AE3EFFC99345B922FF30FAC9FF71" box="[1334,1376,160,185]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
818 has a dentary/surangular contact placed further anterior to the coronoid process as in basal taxa:
|
||
<taxonomicName id="DDC84D78FFC99345B890FF71FAC1FF32" baseAuthorityName="Huxley" baseAuthorityYear="1870" box="[1156,1384,225,250]" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="foxii">
|
||
<emphasis id="28BCEAE9FFC99345B890FF71FAC1FF32" box="[1156,1384,225,250]" italics="true" pageId="12" pageNumber="47">Hypsilophodon foxii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345B96DFF71FC3AFED2" baseAuthorityName="Marsh" baseAuthorityYear="1878" family="Dryosauridae" genus="Dryosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="altus">
|
||
<emphasis id="28BCEAE9FFC99345B96DFF71FC3AFED2" italics="true" pageId="12" pageNumber="47">Dryosaurus altus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BFB4FE91FB50FED2" authorityName=", ML" authorityYear="1851" box="[928,1273,257,282]" class="Reptilia" family="Dryosauridae" genus="Dysalotosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="lettowvorbecki">
|
||
<emphasis id="28BCEAE9FFC99345BFB4FE91FB50FED2" box="[928,1273,257,282]" italics="true" pageId="12" pageNumber="47">Dysalotosaurus lettowvorbecki</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345B921FE91FA56FED2" box="[1333,1535,257,282]" class="Reptilia" family="Hypsilophodontidae" genus="Tenontosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC99345B921FE91FA70FED2" box="[1333,1497,257,282]" italics="true" pageId="12" pageNumber="47">Tenontosaurus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BF71FEB2FC5DFEF3" author="Galton, P. M." box="[869,1012,290,315]" pageId="12" pageNumber="47" pagination="1 - 152" refId="ref16936" refString="Galton, P. M. 1974. The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology 25: 1 - 152." type="journal article" year="1974">Galton 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345B811FEB2FB95FEF3" author="Galton, P. M." box="[1029,1084,290,315]" pageId="12" pageNumber="47" pagination="207 - 243" refId="ref17040" refString="Galton, P. M. 1983. The cranial anatomy of Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and East Africa, with a review of hypsilophodontids from the Upper Jurassic of North America. Geologica et Palaeontologica 17: 207 - 243." type="journal article" year="1983">1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345B859FEB2FB44FEF3" author="Thomas, D. A." box="[1101,1261,290,315]" pageId="12" pageNumber="47" pagination="1 - 99" refId="ref19483" refString="Thomas, D. A. 2015. The cranial anatomy of Tenontosaurus tilletti Ostrom, 1970 (Dinosauria, Ornithopoda). Palaeontologia Electronica 18 (2): 1 - 99." type="journal article" year="2015">Thomas 2015</bibRefCitation>
|
||
). This condition is different from other akylopollexians such as
|
||
<taxonomicName id="DDC84D78FFC99345B94EFED2FC23FEB4" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC99345B94EFED2FA56FE93" box="[1370,1535,322,347]" italics="true" pageId="12" pageNumber="47">Camptosaurus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BFB4FEF3FAA9FEB4" authorityName=", ML" authorityYear="2042" box="[928,1280,355,380]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BFB4FEF3FAA9FEB4" box="[928,1280,355,380]" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345B901FEF3FA51FEB4" authorityName="Brill & Carpenter" authorityYear="2007" box="[1301,1528,355,380]" genus="Theiophytalia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="kerri">
|
||
<emphasis id="28BCEAE9FFC99345B901FEF3FA51FEB4" box="[1301,1528,355,380]" italics="true" pageId="12" pageNumber="47">Theiophytalia kerri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BF4AFE13FB38FE54" box="[862,1169,387,412]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345BF4AFE13FB38FE54" box="[862,1169,387,412]" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345B8C9FE13FA73FE54" authorityName="McDonald, Barrett & Chapman" authorityYear="2010" box="[1245,1498,387,412]" class="Reptilia" genus="Kukufeldia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="tilgatensis">
|
||
<emphasis id="28BCEAE9FFC99345B8C9FE13FA73FE54" box="[1245,1498,387,412]" italics="true" pageId="12" pageNumber="47">Kukufeldia tilgatensis</emphasis>
|
||
</taxonomicName>
|
||
in which the dentary/surangular contact is placed on the same axis or immediately posterior to the coronoid process in a nearly perpendicular fashion (
|
||
<bibRefCitation id="7E594B0AFFC99345B8BBFE74FAE7FE35" author="Gilmore, C. W." box="[1199,1358,484,509]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
; Norman 2004;
|
||
<bibRefCitation id="7E594B0AFFC99345BF4AFD95FBDCFDD6" author="Brill, K. & Carpenter, K." box="[862,1141,517,542]" pageId="12" pageNumber="47" pagination="49 - 67" refId="ref16254" refString="Brill, K. and Carpenter, K. 2006. A description of a new ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of Camptosaurus. In: K. Carpenter (ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, 49 - 67. Indiana University Press, Bloomington." type="book chapter" year="2006">Brill and Carpenter 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345B894FD95FAD2FDD6" author="McDonald, A. T. & Barrett, P. M. & Chapman, S. D." box="[1152,1403,517,542]" pageId="12" pageNumber="47" pagination="1 - 43" refId="ref18428" refString="McDonald, A. T., Barrett, P. M., and Chapman, S. D. 2010 a. A new basal iguanodont (Dinosauria: Ornithischia) from the Wealden (Lower Cretaceous) of England. Zootaxa 2569: 1 - 43." type="journal article" year="2010">
|
||
<collectingCountry id="62DF766BFFC99345B894FD95FB5CFDD6" box="[1152,1269,517,542]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
et al. 2010a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFC99345BF4AFDA0FA7AF933" blockId="12.[862,1535,560,1787]" pageId="12" pageNumber="47">
|
||
<emphasis id="28BCEAE9FFC99345BF4AFDA0FC57FD81" box="[862,1022,560,585]" italics="true" pageId="12" pageNumber="47">Axial skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFC99345B819FDA0FA26FD81" box="[1037,1423,560,585]" italics="true" pageId="12" pageNumber="47">Dorsal vertebrae and neural arch</emphasis>
|
||
: Ankylopollexians (and ornithopods in general) display a conservative axial skeleton during their evolutionary history, thus a taxonomic attribution of isolated material is problematic. The dorsal vertebrae
|
||
<collectionCode id="7CD9AE3EFFC99345B8ADFD22FB4DFD03" box="[1209,1252,690,715]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
452 and the partial dorsal neural arch
|
||
<collectionCode id="7CD9AE3EFFC99345B803FD42FBEBFD23" box="[1047,1090,722,747]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
864 are noteworthy in terms of size, they are bigger than most of the material recovered from the Lourinhã Formation, although it is comparable in size with
|
||
<taxonomicName id="DDC84D78FFC99345BF8CFCA4FBCCFC85" box="[920,1125,820,845]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC99345BF8CFCA4FB97FC85" box="[920,1086,820,845]" italics="true" pageId="12" pageNumber="47">Camptosaurus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
dorsal vertebrae from the Morrison Formation, in particular USNM 4282 (
|
||
<bibRefCitation id="7E594B0AFFC99345B936FCC4FA69FCA5" author="Gilmore, C. W." box="[1314,1472,852,877]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
) and the specimen discussed in
|
||
<bibRefCitation id="7E594B0AFFC99345B8BCFCE5FA52FC46" author="Carpenter, K. & Galton, P. M." box="[1192,1531,885,910]" pageId="12" pageNumber="47" pagination="167 - 207" refId="ref16462" refString="Carpenter, K. and Galton, P. M. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter and Galton (2018</bibRefCitation>
|
||
: fig. 19G). The
|
||
<typeStatus id="C5738859FFC99345B802FC05FBD3FC66" box="[1046,1146,917,942]" pageId="12" pageNumber="47">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="DDC84D78FFC99345B8BEFC05FA25FC66" baseAuthorityName="Mateus and Antunes" baseAuthorityYear="2001" box="[1194,1420,917,942]" class="Reptilia" genus="Draconyx" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="loureiroi">
|
||
<emphasis id="28BCEAE9FFC99345B8BEFC05FA25FC66" box="[1194,1420,917,942]" italics="true" pageId="12" pageNumber="47">Draconyx loureiroi</emphasis>
|
||
</taxonomicName>
|
||
is a mature individual (
|
||
<bibRefCitation id="7E594B0AFFC99345B800FC26FAE8FC07" author="Waskow, K. & Mateus, O." box="[1044,1345,950,975]" pageId="12" pageNumber="47" pagination="425 - 439" refId="ref19576" refString="Waskow, K. and Mateus, O. 2017. Dorsal rib histology of dinosaurs and a crocodylomorph from western Portugal: skeletochronological implications on age determination and life history traits. Comptes Rendus Palevol 16: 425 - 439." type="journal article" year="2017">Waskow and Mateus 2017</bibRefCitation>
|
||
), the caudal centra preserved ranges
|
||
<quantity id="DD309B1EFFC99345B842FC46FB7AFC27" box="[1110,1235,982,1007]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="5.8" metricValueMin="4.6" pageId="12" pageNumber="47" unit="mm" value="52.0" valueMax="58.0" valueMin="46.0">46–58 mm</quantity>
|
||
in length, being sensibly smaller than the ones presented here (
|
||
<bibRefCitation id="7E594B0AFFC99345B91BFC67FC3EFBF8" author="Mateus, O. & Antunes, M." pageId="12" pageNumber="47" pagination="61 - 73" refId="ref18214" refString="Mateus, O. and Antunes, M. 2001. Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales de Paleontologie 87: 61 - 73." type="journal article" year="2001">Mateus and Antunes 2001</bibRefCitation>
|
||
). The general size proportions also closely resemble the ones of
|
||
<taxonomicName id="DDC84D78FFC99345BFF6FBA8FA94FB99" authorityName=", ML" authorityYear="2042" box="[994,1341,1080,1105]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BFF6FBA8FA94FB99" box="[994,1341,1080,1105]" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC84D78FFC99345B962FBA8FC66FBB9" class="Reptilia" genus="Hippodraco" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="scutodens">
|
||
<emphasis id="28BCEAE9FFC99345B962FBA8FC66FBB9" italics="true" pageId="12" pageNumber="47">Hippodraco scutodens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BFF7FBC8FB20FBB9" author="Norman, D. B." box="[995,1161,1112,1137]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">Norman 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345B88FFBC8FA19FBB9" author="McDonald, A. T. & Kirkland, J. I. & DeBlieux, D. D. & Madsen, S. K. & Cavin, J. & Milner, A. R. & Panzarin, L." box="[1179,1456,1112,1137]" pageId="12" pageNumber="47" pagination="14075" refId="ref18478" refString="McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R., and Panzarin, L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5 (11): e 14075." type="journal article" year="2010">
|
||
<collectingCountry id="62DF766BFFC99345B88FFBC8FABAFBB9" box="[1179,1299,1112,1137]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
et al. 2010b
|
||
</bibRefCitation>
|
||
). Both
|
||
<collectionCode id="7CD9AE3EFFC99345BF4AFBE9FC20FB5A" box="[862,905,1145,1170]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
452 and
|
||
<collectionCode id="7CD9AE3EFFC99345BFE4FBE9FBB2FB5A" box="[1008,1051,1145,1170]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
864 share with
|
||
<taxonomicName id="DDC84D78FFC99345B8C6FBE9FC3EFB7A" authorityName=", ML" authorityYear="2042" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345B8C6FBE9FC3EFB7A" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BFBEFB09FB06FB7A" box="[938,1199,1177,1202]" class="Reptilia" genus="Hippodraco" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="scutodens">
|
||
<emphasis id="28BCEAE9FFC99345BFBEFB09FB06FB7A" box="[938,1199,1177,1202]" italics="true" pageId="12" pageNumber="47">Hippodraco scutodens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345B8D7FB09FA51FB7A" box="[1219,1528,1177,1202]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345B8D7FB09FA51FB7A" box="[1219,1528,1177,1202]" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BF4AFB2AFB9DFB1B" box="[862,1076,1210,1235]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dawsoni">
|
||
<emphasis id="28BCEAE9FFC99345BF4AFB2AFB9DFB1B" box="[862,1076,1210,1235]" italics="true" pageId="12" pageNumber="47">Barilium dawsoni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345B892FB2AFA2CFB1B" box="[1158,1413,1210,1235]" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFC99345B892FB2AFA2CFB1B" box="[1158,1413,1210,1235]" italics="true" pageId="12" pageNumber="47">Hypselospinus fittoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345B989FB2AFC3EFB3B" author="Norman, D. B." pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BFB7FB4AFC75FB3B" author="Norman, D. B." box="[931,988,1242,1267]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345BFFDFB4AFBB6FB3B" author="Norman, D. B." box="[1001,1055,1242,1267]" pageId="12" pageNumber="47" pagination="165 - 194" refId="ref18777" refString="Norman, D. B. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345B83FFB4AFBCAFB3B" author="Norman, D. B." box="[1067,1123,1242,1267]" pageId="12" pageNumber="47" pagination="92 - 189" refId="ref18814" refString="Norman, D. B. 2015. On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna). Zoological Journal of the Linnean Society 173: 92 - 189." type="journal article" year="2015">2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345B864FB4AFADAFB3B" author="McDonald, A. T. & Kirkland, J. I. & DeBlieux, D. D. & Madsen, S. K. & Cavin, J. & Milner, A. R. & Panzarin, L." box="[1136,1395,1242,1267]" pageId="12" pageNumber="47" pagination="14075" refId="ref18478" refString="McDonald, A. T., Kirkland, J. I., DeBlieux, D. D., Madsen, S. K., Cavin, J., Milner, A. R., and Panzarin, L. 2010 b. New basal iguanodonts from the Cedar Mountain Formation of Utah and the evolution of thumb-spiked dinosaurs. PLoS One 5 (11): e 14075." type="journal article" year="2010">
|
||
<collectingCountry id="62DF766BFFC99345B864FB4AFB4EFB3B" box="[1136,1255,1242,1267]" name="Heard Island and McDonald Islands" pageId="12" pageNumber="47">McDonald</collectingCountry>
|
||
et al. 2010b
|
||
</bibRefCitation>
|
||
) the antero-posterior stout proportions, the rectangular shape of the neural spine, and its ratio with respect to the centrum; while
|
||
<taxonomicName id="DDC84D78FFC99345BF4AFAACFBFBFA9D" box="[862,1106,1340,1365]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFC99345BF4AFAACFBFBFA9D" box="[862,1106,1340,1365]" italics="true" pageId="12" pageNumber="47">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
shows a paddle like structure towards the dorsal-most part of the spine and generally slender proportions (
|
||
<figureCitation id="82F32A7EFFC99345BFDFFAECFBA6FA5D" box="[971,1039,1404,1429]" captionStart="Fig" captionStartId="11.[98,131,860,880]" captionTargetBox="[542,931,494,833]" captionTargetPageId="11" captionText="Fig. 7. Comparative dorsal vertebrae table of selected Ankylopollexians from the Late Jurassic and Early Cretaceous. A, B. Ankylopollexia indet. Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. A. ML 452 in lateral view (A1, A2). B. ML 864 in right lateral view. C. “Uteodon” SHN.LPP 015 in left lateral view; Praia da corva, Torres Vedras Municipality, Portugal, Lourinhã Formation (Kimmeridgian–Tithonian). D. “Uteodon” aphanoecetes CM 11337 in left lateral view; East end of Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah (USA), Morrison Formation (Kimmeridgian–Tithonian). E. Camptosaurus dispar (unnumbered specimen) in left lateral view; Bone Cabin Quarry, Wyoming USA), Morrison Formation (Kimmeridgian–Tithonian). F. “Cumnoria” prestwichii OUM. J.3303 in lateral view; Oxford, UK, Kimmeridge Clay Formation (Kimmeridgian–Tithonian). G. Hippodraco scutodens UMNH VP 20208 in left lateral view; Andrew’s Site, Grand County, Utah; Upper Yellow Cat Memberof the Cedar Mountain Formation (upper Barremian–lowermost Aptian). H. Iguanacolossus fortis UMNH VP 20205 in right lateral view; Don’s Ridge, Grand County, Utah, Lower Yellow Cat Member, Cedar Mountain Formation (?lower Barremian). I. Mantellisaurus atherfieldensis IRSNB 1551 in left lateral view; Isle of Wight, Wessex Formation (Barremian). J. Barilium dawsoni NHMUK R798 in left lateral view; Shornden, East Sussex, UK, Wadhurst Clay Formation (Valanginian). K. Hypselospinus fittoni NHMUK R604 in lateral view; Shornden Quarry, Hastings, UK, Wadhurst Clay Formation (Valanginian). L. Iguanodon bernissartensis IRSNB “Individu S” in left lateral view; Bernissart, Belgium, Sainte Barbe Clays Formation Barremian). Abbreviations: dia, diapophysis; par, parapophysis. Scale bars 100 mm. Re-drawn from: C, Escaso 2014: fig. 6.5; D, Carpenter and Wilson 2008: fig. 11; E, Carpenter and Galton 2018: fig. 22D; F, Galton and Powell 1980: fig. 4; G, McDonald 2010b: fig. 27; H, McDonald 2010b: fig. 10;, Norman 1980: fig. 37; J, Norman 2011: fig. 4; K, Norman 2015: fig. 22; L, Norman 1980: fig. 31)." figureDoi="http://doi.org/10.5281/zenodo.10626660" httpUri="https://zenodo.org/record/10626660/files/figure.png" pageId="12" pageNumber="47">Fig. 7</figureCitation>
|
||
). Both
|
||
<collectionCode id="7CD9AE3EFFC99345B877FAECFB27FA5D" box="[1123,1166,1404,1429]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
452 and
|
||
<collectionCode id="7CD9AE3EFFC99345B8E1FAECFA89FA5D" box="[1269,1312,1404,1429]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
864 display a more developed neural spine with respect to the transverse processes as in
|
||
<taxonomicName id="DDC84D78FFC99345BFF2FA2DFAE9FA1E" authorityName=", ML" authorityYear="2042" box="[998,1344,1469,1494]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC99345BFF2FA2DFAE9FA1E" box="[998,1344,1469,1494]" italics="true" pageId="12" pageNumber="47">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345B95AFA2DFC4DFA3F" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC99345B95AFA2DFC4DFA3F" italics="true" pageId="12" pageNumber="47">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC99345BFE2FA4EFB6AFA3F" box="[1014,1219,1502,1527]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dawsoni">
|
||
<emphasis id="28BCEAE9FFC99345BFE2FA4EFB6AFA3F" box="[1014,1219,1502,1527]" italics="true" pageId="12" pageNumber="47">Barilium dawsoni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC99345B91EFA4EFA56FA3F" box="[1290,1535,1502,1527]" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFC99345B91EFA4EFA56FA3F" box="[1290,1535,1502,1527]" italics="true" pageId="12" pageNumber="47">Hypselospinus fittoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC99345BF71FA6EFBA1F9DF" author="Norman, D. B." box="[869,1032,1534,1559]" pageId="12" pageNumber="47" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345B802FA6EFBE6F9DF" author="Norman, D. B." box="[1046,1103,1534,1559]" pageId="12" pageNumber="47" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345B849FA6EFB3AF9DF" author="Norman, D. B." box="[1117,1171,1534,1559]" pageId="12" pageNumber="47" pagination="165 - 194" refId="ref18777" refString="Norman, D. B. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC99345B8B5FA6EFB72F9DF" author="Norman, D. B." box="[1185,1243,1534,1559]" pageId="12" pageNumber="47" pagination="92 - 189" refId="ref18814" refString="Norman, D. B. 2015. On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna). Zoological Journal of the Linnean Society 173: 92 - 189." type="journal article" year="2015">2015</bibRefCitation>
|
||
); while in
|
||
<taxonomicName id="DDC84D78FFC99345B94DFA6EFC0EF9F0" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFC99345B94DFA6EFC0EF9F0" italics="true" pageId="12" pageNumber="47">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
specimens it is less developed (
|
||
<bibRefCitation id="7E594B0AFFC99345B95AF98FFA50F9F0" author="Gilmore, C. W." box="[1358,1529,1567,1592]" pageId="12" pageNumber="47" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
; Norman 2004;
|
||
<bibRefCitation id="7E594B0AFFC99345B819F9AFFAE7F990" author="Carpenter, K. & Wilson, Y." box="[1037,1358,1599,1624]" pageId="12" pageNumber="47" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC99345B948F9AFFC58F9B1" author="Carpenter, K. & Galton, P. M." pageId="12" pageNumber="47" pagination="167 - 207" refId="ref16462" refString="Carpenter, K. and Galton, P. M. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter and Galton 2018</bibRefCitation>
|
||
). Herein,
|
||
<collectionCode id="7CD9AE3EFFC99345B87EF9F0FB3CF9B1" box="[1130,1173,1632,1657]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
452 and
|
||
<collectionCode id="7CD9AE3EFFC99345B919F9F0FA91F9B1" box="[1293,1336,1632,1657]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
864 are conservatively assigned to Ankylopollexia, although the antero-posterior extension of the neural spine and its rectangular shape, its development respect to centrum and transverse processes, suggests a possible affinity to Styracosterna.
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFC99344BF4AF89DFD56F8AF" blockId="12.[862,1535,1805,1960]" lastBlockId="13.[98,771,1518,1959]" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47">
|
||
<emphasis id="28BCEAE9FFC99345BF4AF89DFBF1F8EE" box="[862,1112,1805,1830]" italics="true" pageId="12" pageNumber="47">Appendicular skeleton</emphasis>
|
||
:
|
||
<emphasis id="28BCEAE9FFC99345B872F89DFB66F8EE" box="[1126,1231,1805,1830]" italics="true" pageId="12" pageNumber="47">Coracoid</emphasis>
|
||
: The isolated coracoid
|
||
<collectionCode id="7CD9AE3EFFC99345B9C0F89DFA56F8EE" box="[1492,1535,1805,1830]" country="France" name="Musee de Lectoure" pageId="12" pageNumber="47">ML</collectionCode>
|
||
2206 possesses a short and broad sternal process, a typical basal condition within
|
||
<taxonomicName id="DDC84D78FFC99345B87BF8DEFB56F8AF" box="[1135,1279,1870,1895]" order="Ornithischia" pageId="12" pageNumber="57" rank="order">Ornithischia</taxonomicName>
|
||
. It shares with ankylopollexians, basal ornithopods (non-dryomorphans) and neornithischians, a width/length ratio between 70–100% It shares with dryomorphans,
|
||
<taxonomicName id="DDC84D78FFC89344BDA1FA7EFD24F9CF" authorityName="Coria & Calvo" authorityYear="2002" box="[437,653,1518,1543]" class="Reptilia" genus="Anabisetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="saldivai">
|
||
<emphasis id="28BCEAE9FFC89344BDA1FA7EFD24F9CF" box="[437,653,1518,1543]" italics="true" pageId="13" pageNumber="48">Anabisetia saldivai</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC89344BED8FA7EFEC8F9EF" authorityName="Bartholomai & Molnar" authorityYear="1981" family="Dryosauridae" genus="Muttaburrasurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="langdoni">
|
||
<emphasis id="28BCEAE9FFC89344BED8FA7EFEC8F9EF" italics="true" pageId="13" pageNumber="48">Muttaburrasurus langdoni</emphasis>
|
||
</taxonomicName>
|
||
an open coracoid foramen along the scapula-coracoid suture. The size of
|
||
<collectionCode id="7CD9AE3EFFC89344BE0DF9BEFD97F98F" box="[537,574,1582,1607]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="48">ML</collectionCode>
|
||
2206 is greatly bigger than dryosaurids and other neornithischians (
|
||
<bibRefCitation id="7E594B0AFFC89344BEA2F9DEFF30F94F" author="Galton, P. M." pageId="13" pageNumber="48" pagination="1 - 152" refId="ref16936" refString="Galton, P. M. 1974. The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History), Geology 25: 1 - 152." type="journal article" year="1974">Galton 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BCB1F9FEFE2DF94F" author="Norman, D. B. & Sues, H. - D. & Witmer, L. M. & Coria, R. A." box="[165,388,1646,1671]" pageId="13" pageNumber="48" pagination="393 - 412" refId="ref18859" refString="Norman, D. B., Sues, H. - D., Witmer, L. M., and Coria, R. A. 2004. Basal Ornithopoda. In: D. B. Weishampel, P. Dodson, and H. Osmolska (eds.), The Dinosauria, 2 nd edition, 393 - 412. University of California Press, Berkeley." type="book chapter" year="2004">Norman et al. 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BD85F9FEFD8AF94F" author="Hubner, T." box="[401,547,1646,1671]" pageId="13" pageNumber="48" pagination="43 - 120" refId="ref17733" refString="Hubner, T. 2018. The postcranial ontogeny of Dysalotosaurus lettowvorbecki (Ornithischia: Iguanodontia) and implications for the evolution of ornithopod dinosaurs. Palaeontographica A 43 - 120." type="book chapter" year="2018">Hübner 2018</bibRefCitation>
|
||
), while it is comparable with ankylopollexians (
|
||
<bibRefCitation id="7E594B0AFFC89344BDA4F91EFDF8F96F" author="Gilmore, C. W." box="[432,593,1678,1703]" pageId="13" pageNumber="48" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BE4BF91EFD57F96F" author="Norman, D. B." box="[607,766,1678,1703]" pageId="13" pageNumber="48" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344BC76F93EFF33F90F" author="Norman, D. B." box="[98,154,1710,1735]" pageId="13" pageNumber="48" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BCB3F93EFE4AF90F" author="Carpenter, K. & Wilson, Y." box="[167,483,1710,1735]" pageId="13" pageNumber="48" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BDE4F93EFF30F92F" author="Carpenter, K. & Lamanna, M. C." pageId="13" pageNumber="48" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BCB3F95EFE4BF92F" author="Carpenter, K. & Galton, P. M." box="[167,482,1742,1767]" pageId="13" pageNumber="48" pagination="167 - 207" refId="ref16462" refString="Carpenter, K. and Galton, P. M. 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207." type="journal article" year="2018">Carpenter and Galton 2018</bibRefCitation>
|
||
). The coracoid
|
||
<collectionCode id="7CD9AE3EFFC89344BE82F95EFD69F92F" box="[662,704,1742,1767]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="48">ML</collectionCode>
|
||
2206 does not present any distinguishable characters with respect to the variability shown by other ankylopollexians present in the Late Jurassic, such as
|
||
<taxonomicName id="DDC84D78FFC89344BD9CF8BEFDFDF88F" box="[392,596,1838,1863]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="28BCEAE9FFC89344BD9CF8BEFD85F88F" box="[392,556,1838,1863]" italics="true" pageId="13" pageNumber="48">Camptosaurus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFC89344BE70F8BEFD57F88F" author="Dodson, P." box="[612,766,1838,1863]" pageId="13" pageNumber="48" pagination="81 - 85" refId="ref16676" refString="Dodson, P. 1980. Comparative osteology of the American ornithopods Camptosaurus and Tenontosaurus. Memoirs of the Society of Geology, France 139: 81 - 85." type="journal article" year="1980">Dodson 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BC76F8DEFE30F8AF" author="Carpenter, K. & Wilson, Y." box="[98,409,1870,1895]" pageId="13" pageNumber="48" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BDB0F8DEFD5AF8AF" author="Carpenter, K. & Lamanna, M. C." box="[420,755,1870,1895]" pageId="13" pageNumber="48" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna 2015</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1A7736FBFFC99345B9EFF81FFA56F860" blockId="12.[862,1535,1805,1960]" box="[1531,1535,1935,1960]" pageId="12" pageNumber="47">.</paragraph>
|
||
<paragraph id="1A7736FBFFC89344BCFBFAE1FE9BFA4C" box="[239,306,1393,1412]" pageId="13" pageNumber="48">glenoid</paragraph>
|
||
<caption id="4EB76673FFC89344BC76FA0EFCE4FA06" ID-DOI="http://doi.org/10.5281/zenodo.10626662" ID-Zenodo-Dep="10626662" httpUri="https://zenodo.org/record/10626662/files/figure.png" pageId="13" pageNumber="48" startId="13.[98,131,1438,1458]" targetBox="[128,1434,165,1401]" targetPageId="13" targetType="figure">
|
||
<paragraph id="1A7736FBFFC89344BC76FA0EFCE4FA06" pageId="13" pageNumber="48">
|
||
Fig. 8. Ankylopollexian appendicular skeleton from the Lourinhã municipality, Portugal, Lourinhã Formation, Kimmeridgian–Tithonian. Coracoid ML 2206 (
|
||
<emphasis id="28BCEAE9FFC89344BCD7FA29FF7AFA06" bold="true" box="[195,211,1465,1486]" pageId="13" pageNumber="48">A</emphasis>
|
||
), scapula ML 2042 (
|
||
<emphasis id="28BCEAE9FFC89344BD85FA29FE09FA06" bold="true" box="[401,416,1465,1486]" pageId="13" pageNumber="48">B</emphasis>
|
||
), in lateral (A
|
||
<subScript id="864C34BEFFC89344BE35FA53FD80FA19" attach="right" box="[545,553,1475,1489]" fontSize="6" pageId="13" pageNumber="48">1</subScript>
|
||
, B
|
||
<subScript id="864C34BEFFC89344BE57FA53FDE2FA19" attach="left" box="[579,587,1475,1489]" fontSize="6" pageId="13" pageNumber="48">1</subScript>
|
||
) and medial (A
|
||
<subScript id="864C34BEFFC89344BECEFA53FD4BFA19" attach="right" box="[730,738,1475,1489]" fontSize="6" pageId="13" pageNumber="48">2</subScript>
|
||
, B
|
||
<subScript id="864C34BEFFC89344BEE8FA53FCADFA19" attach="left" box="[764,772,1475,1489]" fontSize="6" pageId="13" pageNumber="48">2</subScript>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1A7736FBFFC89347BC93F8FEFE0EFC55" blockId="13.[98,771,1518,1959]" lastBlockId="14.[151,824,160,925]" lastPageId="14" lastPageNumber="49" pageId="13" pageNumber="48">
|
||
<emphasis id="28BCEAE9FFC89344BC93F8FEFF48F84F" box="[135,225,1902,1927]" italics="true" pageId="13" pageNumber="48">Scapula</emphasis>
|
||
:
|
||
<collectionCode id="7CD9AE3EFFC89344BCE1F8FEFE89F84F" box="[245,288,1902,1927]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="48">ML</collectionCode>
|
||
2042 shows characters which are widespread among Ankylopollexia, although they hint to more derived affinities. It is differentiated from
|
||
<taxonomicName id="DDC84D78FFC89344B937FA7EFCC6F9EF" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFC89344B937FA7EFCC6F9EF" italics="true" pageId="13" pageNumber="48">Camptosaurus dispar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344BF94F99EFB8BF9EF" box="[896,1058,1550,1575]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFC89344BF94F99EFB8BF9EF" box="[896,1058,1550,1575]" italics="true" pageId="13" pageNumber="48">C. prestwichii</emphasis>
|
||
</taxonomicName>
|
||
, and dryosaurids in having the dorsal margin of the scapular blade convex and slender proportions of the proximal end with respect to the scapular blade.
|
||
<collectionCode id="7CD9AE3EFFC89344BF66F9FEFC34F94F" box="[882,925,1646,1671]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="48">ML</collectionCode>
|
||
2042 shares with
|
||
<taxonomicName id="DDC84D78FFC89344B87EF9FEFA9AF94F" box="[1130,1331,1646,1671]" class="Reptilia" family="Iguanodontidae" genus="Barilium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="dawsoni">
|
||
<emphasis id="28BCEAE9FFC89344B87EF9FEFA9AF94F" box="[1130,1331,1646,1671]" italics="true" pageId="13" pageNumber="48">Barilium dawsoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344B955F9FEFC30F96F" class="Reptilia" genus="Hippodraco" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="scutodens">
|
||
<emphasis id="28BCEAE9FFC89344B955F9FEFC30F96F" italics="true" pageId="13" pageNumber="48">Hippodraco scutodens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344BFA4F91EFB05F96F" box="[944,1196,1678,1703]" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFC89344BFA4F91EFB05F96F" box="[944,1196,1678,1703]" italics="true" pageId="13" pageNumber="48">Hypselospinus fittoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344B8D7F91EFA6BF96F" box="[1219,1474,1678,1703]" class="Reptilia" genus="Iguanacolossus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="fortis">
|
||
<emphasis id="28BCEAE9FFC89344B8D7F91EFA6BF96F" box="[1219,1474,1678,1703]" italics="true" pageId="13" pageNumber="48">Iguanacolossus fortis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344BF3CF93EFB2DF90F" authorityName=", ML" authorityYear="2042" box="[808,1156,1710,1735]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC89344BF3CF93EFB2DF90F" box="[808,1156,1710,1735]" italics="true" pageId="13" pageNumber="48">Mantellisaurus atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344B887F93EFA6BF90F" box="[1171,1474,1710,1735]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC89344B887F93EFA6BF90F" box="[1171,1474,1710,1735]" italics="true" pageId="13" pageNumber="48">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and more derived Hadrosauriformes the general bowed outline of the scapular blade (
|
||
<bibRefCitation id="7E594B0AFFC89344B84CF97EFB50F8CF" author="Norman, D. B." box="[1112,1273,1774,1799]" pageId="13" pageNumber="48" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344B911F97EFA97F8CF" author="Norman, D. B." box="[1285,1342,1774,1799]" pageId="13" pageNumber="48" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344B95FF97EFA28F8CF" author="Norman, D. B." box="[1355,1409,1774,1799]" pageId="13" pageNumber="48" pagination="165 - 194" refId="ref18777" refString="Norman, D. B. 2011. On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna). Special Papers in Palaeontology 86: 165 - 194." type="journal article" year="2011">2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344B999F97EFA6CF8CF" author="Norman, D. B." box="[1421,1477,1774,1799]" pageId="13" pageNumber="48" pagination="92 - 189" refId="ref18814" refString="Norman, D. B. 2015. On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna). Zoological Journal of the Linnean Society 173: 92 - 189." type="journal article" year="2015">2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BF3CF89EFC53F8EF" author="Horner, J. R. & Weishampel, D. B. & Forster, C. A." box="[808,1018,1806,1831]" pageId="13" pageNumber="48" pagination="438 - 463" refId="ref17635" refString="Horner, J. R., Weishampel, D. B., and Forster, C. A. 2004. Hadrosauridae. In: D. B. Weishampel, P. Dodson, and H. Osmolska (eds.), The Dinosauria, 2 nd Edition, 438 - 463. University of California Press, Berkeley." type="book chapter" year="2004">Horner et al. 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344B812F89EFAA1F8EF" author="McDonald, A. T. & Barrett, P. M. & Chapman, S. D." box="[1030,1288,1806,1831]" pageId="13" pageNumber="48" pagination="1 - 43" refId="ref18428" refString="McDonald, A. T., Barrett, P. M., and Chapman, S. D. 2010 a. A new basal iguanodont (Dinosauria: Ornithischia) from the Wealden (Lower Cretaceous) of England. Zootaxa 2569: 1 - 43." type="journal article" year="2010">
|
||
<collectingCountry id="62DF766BFFC89344B812F89EFBD4F8EF" box="[1030,1149,1806,1831]" name="Heard Island and McDonald Islands" pageId="13" pageNumber="48">McDonald</collectingCountry>
|
||
et al. 2010a
|
||
</bibRefCitation>
|
||
, b). The scapular blade in
|
||
<taxonomicName id="DDC84D78FFC89344BF9BF8BEFB98F88F" box="[911,1073,1838,1863]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFC89344BF9BF8BEFB98F88F" box="[911,1073,1838,1863]" italics="true" pageId="13" pageNumber="48">C. prestwichii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344B857F8BEFABAF88F" authorityName=", ML" authorityYear="2042" box="[1091,1299,1838,1863]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFC89344B857F8BEFABAF88F" box="[1091,1299,1838,1863]" italics="true" pageId="13" pageNumber="48">M. atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFC89344B930F8BEFCC4F8AF" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFC89344B930F8BEFCC4F8AF" italics="true" pageId="13" pageNumber="48">Hypselospinus fittoni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFC89344BFA5F8DEFBD1F8AF" box="[945,1144,1870,1895]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFC89344BFA5F8DEFBD1F8AF" box="[945,1144,1870,1895]" italics="true" pageId="13" pageNumber="48">I. bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
displays a concave proximal end (
|
||
<bibRefCitation id="7E594B0AFFC89344BF4AF8FEFC54F84F" author="Norman, D. B." box="[862,1021,1902,1927]" pageId="13" pageNumber="48" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344B813F8FEFBE9F84F" author="Norman, D. B." box="[1031,1088,1902,1927]" pageId="13" pageNumber="48" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFC89344B85FF8FEFB2AF84F" author="Norman, D. B." box="[1099,1155,1902,1927]" pageId="13" pageNumber="48" pagination="92 - 189" refId="ref18814" refString="Norman, D. B. 2015. On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna). Zoological Journal of the Linnean Society 173: 92 - 189." type="journal article" year="2015">2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344B899F8FEFA6DF84F" author="Carpenter, K. & Wilson, Y." box="[1165,1476,1902,1927]" pageId="13" pageNumber="48" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFC89344BF3CF81EFC74F86F" author="McDonald, A. T." box="[808,989,1934,1959]" pageId="13" pageNumber="48" pagination="52 - 68" refId="ref18369" refString="McDonald, A. T. 2011. The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa 2783: 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="62DF766BFFC89344BF3CF81EFC34F86F" box="[808,925,1934,1959]" name="Heard Island and McDonald Islands" pageId="13" pageNumber="48">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
). The acromion process is rounded, different from
|
||
<taxonomicName id="DDC84D78FFCB9347BD15FF30FEC3FF71" box="[257,362,160,185]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFCB9347BD15FF30FEC3FF71" box="[257,362,160,185]" italics="true" pageId="14" pageNumber="49">C. dispar</emphasis>
|
||
</taxonomicName>
|
||
, but similar to
|
||
<taxonomicName id="DDC84D78FFCB9347BE0DFF30FD7CFF71" box="[537,725,160,185]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFCB9347BE0DFF30FD7CFF71" box="[537,725,160,185]" italics="true" pageId="14" pageNumber="49">C. aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCB9347BEF0FF30FEB2FF11" authorityName=", ML" authorityYear="2042" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCB9347BEF0FF30FEB2FF11" italics="true" pageId="14" pageNumber="49">M. atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCB9347BD3DFF50FDFFFF11" box="[297,598,192,217]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFCB9347BD3DFF50FDFFFF11" box="[297,598,192,217]" italics="true" pageId="14" pageNumber="49">Iguanodon bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFCB9347BE87FF50FF73FF31" class="Reptilia" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="fittoni">
|
||
<emphasis id="28BCEAE9FFCB9347BE87FF50FF73FF31" italics="true" pageId="14" pageNumber="49">Hypselospinus fittoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7E594B0AFFCB9347BCFFFF70FE22FF31" author="Gilmore, C. W." box="[235,395,224,249]" pageId="14" pageNumber="49" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCB9347BD83FF70FD9DFF31" author="Norman, D. B." box="[407,564,224,249]" pageId="14" pageNumber="49" pagination="1 - 105" refId="ref18617" refString="Norman, D. B. 1980. On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower Cretaceous of Bernissart (Belgium). Memoir de l'Institut Royal des Sciences Naturelles de Belgique 178: 1 - 105." type="journal article" year="1980">Norman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFCB9347BE2BFF70FDD1FF31" author="Norman, D. B." box="[575,632,224,249]" pageId="14" pageNumber="49" pagination="281 - 372" refId="ref18692" refString="Norman, D. B. 1986. On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda). Bulletin de l'Institut royal des Sciences naturelles de Belgique 56: 281 - 372." type="journal article" year="1986">1986</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7E594B0AFFCB9347BE90FF70FD15FF31" author="Norman, D. B." box="[644,700,224,249]" pageId="14" pageNumber="49" pagination="92 - 189" refId="ref18814" refString="Norman, D. B. 2015. On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna). Zoological Journal of the Linnean Society 173: 92 - 189." type="journal article" year="2015">2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCB9347BED3FF70FEF4FED1" author="Carpenter, K. & Wilson, Y." pageId="14" pageNumber="49" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCB9347BD7FFE90FD88FED1" author="McDonald, A. T." box="[363,545,256,281]" pageId="14" pageNumber="49" pagination="52 - 68" refId="ref18369" refString="McDonald, A. T. 2011. The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa 2783: 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="62DF766BFFCB9347BD7FFE90FE4BFED1" box="[363,482,256,281]" name="Heard Island and McDonald Islands" pageId="14" pageNumber="49">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
). The ventral and dorsal margins, being sub-parallel towards the middle of the blade, resemble the condition seen in
|
||
<taxonomicName id="DDC84D78FFCB9347BE1EFED1FD72FE92" authorityName=", ML" authorityYear="2042" box="[522,731,321,346]" class="Reptilia" family="Iguanodontidae" genus="Mantellisaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="atherfieldensis">
|
||
<emphasis id="28BCEAE9FFCB9347BE1EFED1FD72FE92" box="[522,731,321,346]" italics="true" pageId="14" pageNumber="49">M. atherfieldensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCB9347BEF9FED1FEB7FEB2" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="bernissartensis">
|
||
<emphasis id="28BCEAE9FFCB9347BEF9FED1FEB7FEB2" italics="true" pageId="14" pageNumber="49">I. bernissartensis</emphasis>
|
||
</taxonomicName>
|
||
, and other derived styracosternans (Norman 2004) but differ from
|
||
<taxonomicName id="DDC84D78FFCB9347BD89FE11FDA0FE52" box="[413,521,385,410]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="dispar">
|
||
<emphasis id="28BCEAE9FFCB9347BD89FE11FDA0FE52" box="[413,521,385,410]" italics="true" pageId="14" pageNumber="49">C. dispar</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="DDC84D78FFCB9347BE0EFE11FD70FE52" box="[538,729,385,410]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFCB9347BE0EFE11FD70FE52" box="[538,729,385,410]" italics="true" pageId="14" pageNumber="49">C. aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DDC84D78FFCB9347BF0AFE11FEBCFE72" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFCB9347BF0AFE11FEBCFE72" italics="true" pageId="14" pageNumber="49">C. prestwichii</emphasis>
|
||
</taxonomicName>
|
||
which have the ventral margin slightly diverging from the mid-blade or immediately distal to the mid-blade (
|
||
<bibRefCitation id="7E594B0AFFCB9347BC8BFE72FEEEFE33" author="Gilmore, C. W." box="[159,327,482,507]" pageId="14" pageNumber="49" pagination="197 - 332" refId="ref17231" refString="Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus with a revision of the species of the genus, and description of two new species. Proceedings of the United States National Musuem (36): 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCB9347BD4EFE72FD04FE33" author="Carpenter, K. & Wilson, Y." box="[346,685,482,507]" pageId="14" pageNumber="49" pagination="227 - 264" refId="ref16550" refString="Carpenter, K. and Wilson, Y. 2008. A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of Carnegie Museum 76: 227 - 264." type="journal article" year="2008">Carpenter and Wilson 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7E594B0AFFCB9347BED4FE72FF67FDD3" author="McDonald, A. T." pageId="14" pageNumber="49" pagination="52 - 68" refId="ref18369" refString="McDonald, A. T. 2011. The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda). Zootaxa 2783: 52 - 68." type="journal article" year="2011">
|
||
<collectingCountry id="62DF766BFFCB9347BED4FE72FC91FE33" box="[704,824,482,507]" name="Heard Island and McDonald Islands" pageId="14" pageNumber="49">McDonald</collectingCountry>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="7E594B0AFFCB9347BCF4FD92FD96FDD3" author="Carpenter, K. & Lamanna, M. C." box="[224,575,514,539]" pageId="14" pageNumber="49" pagination="149 - 166" refId="ref16502" refString="Carpenter, K. and Lamanna, M. C. 2015. The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy. Annals of Carnegie Museum 83 149 - 166." type="journal article" year="2015">Carpenter and Lamanna (2015)</bibRefCitation>
|
||
lumped the once considered oldest styracosternan “
|
||
<taxonomicName id="DDC84D78FFCB9347BDE2FDB2FDFEFDF3" authorityName="McDonald" authorityYear="2011" box="[502,599,546,571]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCB9347BDE2FDB2FDFEFDF3" box="[502,599,546,571]" italics="true" pageId="14" pageNumber="49">Uteodon</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="DDC84D78FFCB9347BE78FDB2FCAEFDF3" box="[620,775,546,571]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="aphanoecetes">
|
||
<emphasis id="28BCEAE9FFCB9347BE78FDB2FCAEFDF3" box="[620,775,546,571]" italics="true" pageId="14" pageNumber="49">aphanoecetes</emphasis>
|
||
</taxonomicName>
|
||
and “
|
||
<taxonomicName id="DDC84D78FFCB9347BCB7FDD2FEBFFD93" authorityName="Seeley" authorityYear="1888" box="[163,278,578,603]" class="Reptilia" genus="Cumnoria" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCB9347BCB7FDD2FEBFFD93" box="[163,278,578,603]" italics="true" pageId="14" pageNumber="49">Cumnoria</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="DDC84D78FFCB9347BD3DFDD2FE0EFD93" box="[297,423,578,603]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="species" species="prestwichii">
|
||
<emphasis id="28BCEAE9FFCB9347BD3DFDD2FE0EFD93" box="[297,423,578,603]" italics="true" pageId="14" pageNumber="49">prestwichii</emphasis>
|
||
</taxonomicName>
|
||
into the genus
|
||
<taxonomicName id="DDC84D78FFCB9347BE46FDD2FD5EFD93" authorityName="Marsh" authorityYear="1885" box="[594,759,578,603]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="28BCEAE9FFCB9347BE46FDD2FD5EFD93" box="[594,759,578,603]" italics="true" pageId="14" pageNumber="49">Camptosaurus</emphasis>
|
||
</taxonomicName>
|
||
. This lumping has great implications for the attribution of
|
||
<collectionCode id="7CD9AE3EFFCB9347BF19FDF2FC91FDB3" box="[781,824,610,635]" country="France" name="Musee de Lectoure" pageId="14" pageNumber="49">ML</collectionCode>
|
||
2042 to Styracosterna, since some of the characters considered diagnostic present in the scapular blade (i.e., dorsal scapular margin convex, rounded acromion process) are now ambiguously widespread among Ankylopollexia.
|
||
<collectionCode id="7CD9AE3EFFCB9347BF19FD73FC91FD34" box="[781,824,739,764]" country="France" name="Musee de Lectoure" pageId="14" pageNumber="49">ML</collectionCode>
|
||
2042 is here conservatively considered Ankylopollexia, although the presence of a dorsoventrally bow of the scapular blade, sub-parallel ventral and dorsal margins at mid-blade, and a generally rounded acromion indicates a possible attribution to Styracosterna.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |