treatments-xml/data/92/48/87/924887EDFFFDFFFDFB97FAB1E1AEF840.xml

<document id="8C742D62EB8582E28C283B5248B2379C" ID-CLB-Dataset="231396" ID-DOI="10.1655/HERPETOLOGICA-D-16-00069" ID-GBIF-Dataset="39128a81-7c6c-4fb9-8e48-a23259b36f23" ID-ISSN="1938-5099" ID-Zenodo-Dep="7712269" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1678362835023" checkinUser="felipe" docAuthor="Malagoli, Leo R., de Sá, Fábio P., Canedo, Clarissa &amp; Haddad, Célio F. B." docDate="2017" docId="924887EDFFFDFFFDFB97FAB1E1AEF840" docLanguage="en" docName="Herpetologica.73.2.136-147.pdf" docOrigin="Herpetologica 73 (2)" docSource="http://dx.doi.org/10.1655/herpetologica-d-16-00069" docStyle="DocumentStyle:7157CB4A8C4DC19627D49B44D928F812.2:Herpetologica.2015-.journal_article.1cover" docStyleId="7157CB4A8C4DC19627D49B44D928F812" docStyleName="Herpetologica.2015-.journal_article.1cover" docStyleVersion="2" docTitle="Hnlodes caete Malagoli &amp; de Sá &amp; Canedo &amp; Haddad 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="143" masterDocId="6E71FF95FFFEFFF5FF94FFB6E50CFF91" masterDocTitle="A New Species of Hŋlodes (Anura, Hylodidae) from Serra do Mar, Southeastern Brazil: The Fourth with Nuptial Thumb Tubercles" masterLastPageNumber="147" masterPageNumber="136" pageNumber="138" updateTime="1699476429410" updateUser="ExternalLinkService">
<mods:mods id="C1BF032EB20130EA720CA39D8F4B0A0C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="787D239537E967183234ACE8859CA54C">
<mods:title id="DD6B2FAC24F31EC59C7ED10D43685D1C">A New Species of Hŋlodes (Anura, Hylodidae) from Serra do Mar, Southeastern Brazil: The Fourth with Nuptial Thumb Tubercles</mods:title>
</mods:titleInfo>
<mods:name id="90B7EE272F347F85A3F5CEA4F589A1C9" type="personal">
<mods:role id="D78C64CA11553C952CA37E241BB33D15">
<mods:roleTerm id="3D477ADCF7651C2D4F3B3EACB70D2C95">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="591E673219C4FDCC11578CA6C8525764">Malagoli, Leo R.</mods:namePart>
</mods:name>
<mods:name id="832C25AFDE146FF2CB89A1CBE06A8C35" type="personal">
<mods:role id="E236114F3E2CC8F3B893B32E32B40BFA">
<mods:roleTerm id="619E891D6572C1C373667F5C8C6A64C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD4D37A16D15C4F17BD663BDD0D4879E">de Sá, Fábio P.</mods:namePart>
</mods:name>
<mods:name id="87922DBAE0187F3834141DBB1E45D344" type="personal">
<mods:role id="0B3982DC3828C2432B67B15B8BFE4562">
<mods:roleTerm id="B2619626D49575A8F4A8AC5585892276">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08C85CE405072527398221CE24192C61">Canedo, Clarissa</mods:namePart>
</mods:name>
<mods:name id="4699644CA1BCB6B0358548478B04EEA7" type="personal">
<mods:role id="6EFB4D9D535EF5C2466998633872BE99">
<mods:roleTerm id="F3E494F81CC3ED2BC941A8926FB6CD00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEEB6485F13A7B310E53C6CB5F120E0F">Haddad, Célio F. B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="75260EC5221527A9C7E77EE066C1CBA1">text</mods:typeOfResource>
<mods:relatedItem id="8A4E8C2D36A90696FA45D10FF3906D19" type="host">
<mods:titleInfo id="BDAE015A51F65D471DA2D9E4B3645F62">
<mods:title id="09A1E483A19AD310D4CE1192866E0EDF">Herpetologica</mods:title>
</mods:titleInfo>
<mods:part id="2497D1FD71E2321155C9164773F49456">
<mods:date id="90F580EC6AFE1FB363A52A3115F00D16">2017</mods:date>
<mods:detail id="5F42927A2BB8BFC7834F2B2F62DE95F8" type="pubDate">
<mods:number id="04652734EE36E86AE72CCF6A798787E6">2017-06-01</mods:number>
</mods:detail>
<mods:detail id="5C013F5D23A16820679C5F5016A8E410" type="volume">
<mods:number id="2D25EC58FBBBE2CEC0D8FAA2A8AED508">73</mods:number>
</mods:detail>
<mods:detail id="4DD942829DBEF57938C0B3A089FAC94D" type="issue">
<mods:number id="D044CC187010A3F7C17B707AC659F31E">2</mods:number>
</mods:detail>
<mods:extent id="7A74F6F82C9F8F00A8159092F4AAE88E" unit="page">
<mods:start id="28059869A7F9143DE073C83919DFB632">136</mods:start>
<mods:end id="9B2B2B362173454F0A4182EC3ECAF5E5">147</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B0E2DDB6AAEF95EA5CBF61485E6E6276">
<mods:url id="E23FDEA7B70A3C79C194152B96BEF049">http://dx.doi.org/10.1655/herpetologica-d-16-00069</mods:url>
</mods:location>
<mods:classification id="7B3298EE24E083E888F3F3C51201AA2F">journal article</mods:classification>
<mods:identifier id="F52CB78B308EB8904BD81285F3507102" type="CLB-Dataset">231396</mods:identifier>
<mods:identifier id="7275DC75882FED5AC316A92CACD743D8" type="DOI">10.1655/HERPETOLOGICA-D-16-00069</mods:identifier>
<mods:identifier id="6D2CEE2365069199947DC0738AD0E5C2" type="GBIF-Dataset">39128a81-7c6c-4fb9-8e48-a23259b36f23</mods:identifier>
<mods:identifier id="4B3FC0DEF0F8B331595EEA09EA718D7A" type="ISSN">1938-5099</mods:identifier>
<mods:identifier id="AED0AA4170A0D3FD6C26DBD8882FCA76" type="Zenodo-Dep">7712269</mods:identifier>
</mods:mods>
<treatment id="924887EDFFFDFFFDFB97FAB1E1AEF840" ID-DOI="http://doi.org/10.5281/zenodo.7716565" ID-GBIF-Taxon="209435021" ID-Zenodo-Dep="7716565" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840" httpUri="http://treatment.plazi.org/id/924887EDFFFDFFFDFB97FAB1E1AEF840" lastPageId="8" lastPageNumber="143" pageId="3" pageNumber="138">
<subSubSection id="52FB6570FFFDFFF6FB97FAB1E00FFA8F" box="[1027,1283,1287,1311]" pageId="3" pageNumber="138" type="nomenclature">
<paragraph id="1A5E36FBFFFDFFF6FB97FAB1E00FFA8F" blockId="3.[1027,1283,1287,1311]" box="[1027,1283,1287,1311]" pageId="3" pageNumber="138">
<heading id="41168197FFFDFFF6FB97FAB1E00FFA8F" box="[1027,1283,1287,1311]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="138" reason="2">
<taxonomicName id="DDE14D78FFFDFFF6FB97FAB1E1AFFA8F" authority="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad, 2017" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1027,1187,1287,1311]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="147" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
<emphasis id="2895EAE9FFFDFFF6FB97FAB1E1AFFA8F" box="[1027,1187,1287,1311]" italics="true" pageId="3" pageNumber="138">
Hŋlodes 
<emphasis id="2895EAE9FFFDFFF6FBF1FAB1E1AFFA8F" bold="true" box="[1125,1187,1287,1310]" italics="true" pageId="3" pageNumber="138">caete</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A65792FFFDFFF6FB38FAB1E00FFA8F" box="[1196,1283,1287,1310]" pageId="3" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFFDFFF6FB81FA92E1FDFAAC" box="[1045,1265,1316,1341]" pageId="3" pageNumber="138" type="description">
<paragraph id="1A5E36FBFFFDFFF6FB81FA92E1FDFAAC" blockId="3.[1045,1265,1316,1341]" box="[1045,1265,1316,1341]" pageId="3" pageNumber="138">
(
<figureCitation id="82DA2A7EFFFDFFF6FB89FA92E189FAAC" box="[1053,1157,1316,1341]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="Fig" captionStartId-0="3.[142,156,1909,1928]" captionStartId-1="4.[166,180,898,917]" captionStartId-2="5.[142,156,1866,1885]" captionStartId-3="6.[868,900,815,834]" captionTargetBox-0="[118,788,1398,1883]" captionTargetBox-1="[142,1515,191,872]" captionTargetBox-2="[119,1488,193,1839]" captionTargetBox-3="[844,1515,193,790]" captionTargetId-0="figure-805@3.[118,788,1398,1883]" captionTargetId-1="figure-795@4.[142,1516,191,872]" captionTargetId-2="figure-0@5.[118,1491,191,1840]" captionTargetId-3="figure-488@6.[844,1516,191,790]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIG. 1.—Live adult male Hŋlodes caete sp. nov. (holotype CFBH 40524; snout–vent length ¼ 31.6 mm). Photograph taken in Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online." captionText-1="FIG. 2.—Hŋlodes caete, CFBH 40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm." captionText-2="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." captionText-3="Fig. 4.—Distribution of nuptial tubercles on the right thumb of an adult male Hŋlodes caete, CFBH 40524 (holotype). (A) Drawing in lateral view. (B) Picture in dorsal view, with dashed line indicating the elliptical distribution of tubercles. Scale bars ¼ 1 mm. A color version of this figure is available online." figureDoi-0="http://doi.org/10.5281/zenodo.7712273" figureDoi-1="http://doi.org/10.5281/zenodo.7712275" figureDoi-2="http://doi.org/10.5281/zenodo.7712277" figureDoi-3="http://doi.org/10.5281/zenodo.7712279" httpUri-0="https://zenodo.org/record/7712273/files/figure.png" httpUri-1="https://zenodo.org/record/7712275/files/figure.png" httpUri-2="https://zenodo.org/record/7712277/files/figure.png" httpUri-3="https://zenodo.org/record/7712279/files/figure.png" pageId="3" pageNumber="138">Figs. 1–4</figureCitation>
; 
<tableCitation id="57630340FFFDFFF6FB07FA92E1E4FAAC" box="[1171,1256,1316,1341]" captionStart="TABLE" captionStartId="6.[169,182,188,207]" captionTargetPageId="6" captionText="TABLE 3.—Measurements (mm, reported as ¯X ± 1 SD) of males (n ¼ 16) and females (n ¼ 5) of the type series of Hŋlodes caete sp. nov." httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" pageId="3" pageNumber="138" tableUuid="4E9E6673FFF8FFF3FF3DFF0AE7C7FF76">Table 3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFFDFFF6FCA0FAE4E084FAFB" box="[820,1416,1362,1386]" pageId="3" pageNumber="138" type="reference_group">
<paragraph id="1A5E36FBFFFDFFF6FCA0FAE4E084FAFB" blockId="3.[820,1416,1362,1386]" box="[820,1416,1362,1386]" pageId="3" pageNumber="138">
<treatmentCitationGroup id="3AF111D5FFFDFFF6FCA0FAE4E084FAFB" box="[820,1416,1362,1386]" pageId="3" pageNumber="138">
<taxonomicName id="DDE14D78FFFDFFF6FCA0FAE4E6B9FAFB" box="[820,949,1362,1386]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="138" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="2895EAE9FFFDFFF6FCA0FAE4E681FAFB" box="[820,909,1362,1386]" italics="true" pageId="3" pageNumber="138">Hŋlodes</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="DDE14D78FFFDFFF6FC51FAE4E156FAFB" box="[965,1114,1362,1386]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" isUncertain="true" kingdom="Animalia" order="Anura" pageId="3" pageNumber="138" phylum="Chordata" rank="species" species="phnllodes">
aff. 
<emphasis id="2895EAE9FFFDFFF6FC66FAE4E156FAFB" box="[1010,1114,1362,1386]" italics="true" pageId="3" pageNumber="138">phŋllodes</emphasis>
</taxonomicName>
); 
<treatmentCitation id="9B4010EAFFFDFFF6FBE7FAE4E084FAFB" author="Trevine, V. &amp; M. C. Forlani &amp; C. F. B. Haddad &amp; H. Zaher" box="[1139,1416,1362,1386]" page="130" pageId="3" pageNumber="138" year="2014">
<bibRefCitation id="7E704B0AFFFDFFF6FBE7FAE4E050FAFB" author="Trevine, V. &amp; M. C. Forlani &amp; C. F. B. Haddad &amp; H. Zaher" box="[1139,1372,1362,1386]" pageId="3" pageNumber="138" pagination="126 - 146" refId="ref11738" refString="Trevine, V., M. C. Forlani, C. F. B. Haddad, and H. Zaher. 2014. Herpetofauna of Paranapiacaba: Expanding our knowledge on a historical region in the Atlantic forest of southeastern Brazil. Zoologia 31: 126 - 146." type="journal article" year="2014">Trevine et al. (2014)</bibRefCitation>
:130
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFFDFFF1FCDAFA30E60CF973" lastPageId="4" lastPageNumber="139" pageId="3" pageNumber="138" type="materials_examined">
<paragraph id="1A5E36FBFFFDFFF6FCDAFA30E670F9C1" blockId="3.[820,1490,1414,2001]" pageId="3" pageNumber="138">
<materialsCitation id="AA893CA6FFFDFFF6FCDAFA30E670F9C1" ID-GBIF-Occurrence="4052833307" collectingDate="2015-03-11" collectionCode="CFBH" collectorName="Leo R. Malagoli &amp; Paulo D. P. Pinheiro" country="Brazil" county="Southeastern" elevation="775" latitude="23.59" location="Rio Camburi" longitude="46.44" municipality="Itanhaem" pageId="3" pageNumber="138" specimenCode="40524" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="2895EAE9FFFDFFF6FCDAFA30E6CAFA0F" bold="true" box="[846,966,1414,1438]" pageId="3" pageNumber="138">
<typeStatus id="C55A8859FFFDFFF6FCDAFA30E6CDFA0F" box="[846,961,1414,1438]" pageId="3" pageNumber="138" type="holotype">Holotype</typeStatus>
.
</emphasis>
— 
<collectionCode id="7CF0AE3EFFFDFFF6FC77FA30E120FA0F" box="[995,1068,1414,1438]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FBA8FA30E172FA0F" box="[1084,1150,1414,1438]" pageId="3" pageNumber="138">40524</specimenCode>
(
<figureCitation id="82DA2A7EFFFDFFF6FB02FA30E00BFA0F" box="[1174,1287,1414,1438]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="Fig" captionStartId-0="3.[142,156,1909,1928]" captionStartId-1="4.[166,180,898,917]" captionStartId-2="5.[142,156,1866,1885]" captionStartId-3="6.[868,900,815,834]" captionTargetBox-0="[118,788,1398,1883]" captionTargetBox-1="[142,1515,191,872]" captionTargetBox-2="[119,1488,193,1839]" captionTargetBox-3="[844,1515,193,790]" captionTargetId-0="figure-805@3.[118,788,1398,1883]" captionTargetId-1="figure-795@4.[142,1516,191,872]" captionTargetId-2="figure-0@5.[118,1491,191,1840]" captionTargetId-3="figure-488@6.[844,1516,191,790]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIG. 1.—Live adult male Hŋlodes caete sp. nov. (holotype CFBH 40524; snout–vent length ¼ 31.6 mm). Photograph taken in Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online." captionText-1="FIG. 2.—Hŋlodes caete, CFBH 40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm." captionText-2="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." captionText-3="Fig. 4.—Distribution of nuptial tubercles on the right thumb of an adult male Hŋlodes caete, CFBH 40524 (holotype). (A) Drawing in lateral view. (B) Picture in dorsal view, with dashed line indicating the elliptical distribution of tubercles. Scale bars ¼ 1 mm. A color version of this figure is available online." figureDoi-0="http://doi.org/10.5281/zenodo.7712273" figureDoi-1="http://doi.org/10.5281/zenodo.7712275" figureDoi-2="http://doi.org/10.5281/zenodo.7712277" figureDoi-3="http://doi.org/10.5281/zenodo.7712279" httpUri-0="https://zenodo.org/record/7712273/files/figure.png" httpUri-1="https://zenodo.org/record/7712275/files/figure.png" httpUri-2="https://zenodo.org/record/7712277/files/figure.png" httpUri-3="https://zenodo.org/record/7712279/files/figure.png" pageId="3" pageNumber="138">Figs. 1–4</figureCitation>
), an 
<specimenCount id="0CE7FD72FFFDFFF6FADAFA30E0C2FA0F" box="[1358,1486,1414,1438]" pageId="3" pageNumber="138" type="adult">adult male</specimenCount>
, collected by 
<collectorName id="B714532DFFFDFFF6FC54FA12E17DFA2A" box="[960,1137,1443,1467]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFDFFF6FB3CFA15E081FA2A" box="[1192,1421,1443,1467]" pageId="3" pageNumber="138">Paulo D. P. Pinheiro</collectorName>
on 
<date id="6E5F103BFFFDFFF6FA23FA12E6CDFA48" pageId="3" pageNumber="138" value="2015-03-11">
<collectingDate id="7E1BE9D3FFFDFFF6FA23FA12E6CDFA48" pageId="3" pageNumber="138" value="2015-03-11">11 March 2015</collectingDate>
</date>
at 
<location id="1F3E6020FFFDFFF6FC60FA77E183FA48" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFDFFF6FC60FA77E183FA48" box="[1012,1167,1473,1497]" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Rio Camburi" pageId="3" pageNumber="138" stateProvince="Sao Paulo">Rio Camburi</location>
, 
<location id="1F3E6020FFFDFFF6FB36FA77E063FA48" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFDFFF6FB36FA77E063FA48" box="[1186,1391,1473,1497]" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Nucleo Curucutu" pageId="3" pageNumber="138" stateProvince="Sao Paulo">Núcleo Curucutu</location>
, 
<location id="1F3E6020FFFDFFF6FA16FA74E151FA67" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFDFFF6FA16FA74E151FA67" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Parque Estadual da Serra do Mar" pageId="3" pageNumber="138" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFDFFF6FBFBFA69E01DFA66" box="[1135,1297,1499,1527]" degrees="23" direction="south" minutes="59" orientation="latitude" pageId="3" pageNumber="138" seconds="56.76" value="23.59">
23°59 
<emphasis id="2895EAE9FFFDFFF6FB24FA6DE1B9FA7D" box="[1200,1205,1499,1516]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FB24FA6DE1B9FA7D" attach="none" box="[1200,1205,1499,1516]" fontSize="7" pageId="3" pageNumber="138">′</superScript>
</emphasis>
56.76 
<emphasis id="2895EAE9FFFDFFF6FB67FA6DE00CFA7D" box="[1267,1280,1499,1516]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FB67FA6DE00CFA7D" attach="right" box="[1267,1280,1499,1516]" fontSize="7" pageId="3" pageNumber="138">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFDFFF6FAB5FA69E0C2FA67" box="[1313,1486,1499,1527]" degrees="46" direction="west" minutes="44" orientation="longitude" pageId="3" pageNumber="138" seconds="17.69" value="46.44">
46°44 
<emphasis id="2895EAE9FFFDFFF6FAF4FA6DE069FA7D" box="[1376,1381,1499,1516]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FAF4FA6DE069FA7D" attach="none" box="[1376,1381,1499,1516]" fontSize="7" pageId="3" pageNumber="138">′</superScript>
</emphasis>
17.69 
<emphasis id="2895EAE9FFFDFFF6FA30FA6DE0BCFA7D" box="[1444,1456,1499,1516]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FA30FA6DE0BCFA7D" attach="right" box="[1444,1456,1499,1516]" fontSize="7" pageId="3" pageNumber="138">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<quantity id="DD199B1EFFFDFFF6FCA0FA4AE675F985" box="[820,889,1532,1556]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="3" pageNumber="138" unit="m" value="775.0">
<elevation id="91CCD1C8FFFDFFF6FCA0FA4AE675F985" box="[820,889,1532,1556]" metricMagnitude="2" metricUnit="m" metricValue="7.75" pageId="3" pageNumber="138" unit="m" value="775.0">775 m</elevation>
</quantity>
above sea level [a.s.l.]; in all cases, datum ¼ WGS84), municipality of 
<collectingMunicipality id="FA3AAC81FFFDFFF6FC72F9ACE15EF9A3" box="[998,1106,1562,1586]" pageId="3" pageNumber="138">Itanhaém</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFDFFF6FB54F9ACE03FF9A3" box="[1216,1331,1562,1586]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="138">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFDFFF6FCA0F98EE674F9C1" box="[820,888,1592,1616]" name="Brazil" pageId="3" pageNumber="138">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="1A5E36FBFFFDFFF1FCDAF9E0E60CF973" blockId="3.[820,1490,1414,2001]" lastBlockId="4.[142,813,962,2001]" lastPageId="4" lastPageNumber="139" pageId="3" pageNumber="138">
<materialsCitation id="AA893CA6FFFDFFF6FCDAF9E0E1ECF938" ID-GBIF-Occurrence="4052833302" collectingDate="2005-03-17" collectingDateMax="2016-02-13" collectingDateMin="2005-03-17" collectionCode="CFBH" collectorName="Ricardo J. Sawaya; Fausto Barbo; Fernando Couto &amp; Leo R. Malagoli" country="Brazil" county="Southeastern" elevation="818" location="Parque Estadual da Serra do Mar" municipality="State" pageId="3" pageNumber="138" specimenCode="12220" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis id="2895EAE9FFFDFFF6FCDAF9E0E6C2F9FF" bold="true" box="[846,974,1622,1646]" pageId="3" pageNumber="138">
<typeStatus id="C55A8859FFFDFFF6FCDAF9E0E6C5F9FF" box="[846,969,1622,1646]" pageId="3" pageNumber="138" type="paratype">Paratypes</typeStatus>
.
</emphasis>
— 
<specimenCount id="0CE7FD72FFFDFFF6FC7EF9E3E1CDF9FC" box="[1002,1217,1621,1645]" pageId="3" pageNumber="138">Fifteen adult males</specimenCount>
: 
<collectionCode id="7CF0AE3EFFFDFFF6FB58F9E0E01BF9FF" box="[1228,1303,1622,1646]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FA8AF9E0E06FF9FF" box="[1310,1379,1622,1646]" pageId="3" pageNumber="138">12220</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FCC9F9C5E11DF91A" box="[861,1041,1651,1675]" pageId="3" pageNumber="138" value="2005-03-17">
<collectingDate id="7E1BE9D3FFFDFFF6FCC9F9C5E11DF91A" box="[861,1041,1651,1675]" pageId="3" pageNumber="138" value="2005-03-17">17 March 2005</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FBD3F9C5E011F91A" box="[1095,1309,1651,1675]" pageId="3" pageNumber="138">Ricardo J. Sawaya</collectorName>
, 
<collectorName id="B714532DFFFDFFF6FABBF9C2E0C2F91A" box="[1327,1486,1651,1675]" pageId="3" pageNumber="138">Fausto Barbo</collectorName>
, 
<collectorName id="B714532DFFFDFFF6FCA0F927E6E2F938" box="[820,1006,1681,1705]" pageId="3" pageNumber="138">Fernando Couto</collectorName>
, and 
<collectorName id="B714532DFFFDFFF6FBBCF927E1D0F938" box="[1064,1244,1681,1705]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
;
</materialsCitation>
<materialsCitation id="AA893CA6FFFDFFF6FB72F927E075F957" ID-GBIF-Occurrence="4052833301" collectingDate="2008-01-06" collectingDateMax="2016-02-13" collectingDateMin="2005-03-17" collectionCode="CFBH" collectorName="Leo R. Malagoli" country="Brazil" county="Southeastern" elevation="818" location="Parque Estadual da Serra do Mar" municipality="State" pageId="3" pageNumber="138" specimenCode="19370-19371" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFDFFF6FB72F927E03DF938" box="[1254,1329,1681,1705]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FAADF927E0C1F938" box="[1337,1485,1681,1705]" pageId="3" pageNumber="138">19370–19371</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FC59F918E173F956" box="[973,1151,1710,1735]" pageId="3" pageNumber="138" value="2008-01-06">
<collectingDate id="7E1BE9D3FFFDFFF6FC59F918E173F956" box="[973,1151,1710,1735]" pageId="3" pageNumber="138" value="2008-01-06">6 January 2008</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FB20F919E079F957" box="[1204,1397,1710,1734]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
;
</materialsCitation>
<materialsCitation id="AA893CA6FFFDFFF6FA12F919E12CF893" ID-GBIF-Occurrence="4052833304" collectingDate="2015-03-11" collectingDateMax="2016-02-13" collectingDateMin="2005-03-17" collectionCode="CFBH" collectorName="Leo R. Malagoli &amp; Paulo D. P. Pinheiro" country="Brazil" county="Southeastern" elevation="818" location="Parque Estadual da Serra do Mar" municipality="State" pageId="3" pageNumber="138" specimenCode="40525" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFDFFF6FA12F919E0DDF956" box="[1414,1489,1711,1735]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FCA0F97AE675F975" box="[820,889,1740,1764]" pageId="3" pageNumber="138">40525</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FB82F97AE1CCF975" box="[1046,1216,1740,1764]" pageId="3" pageNumber="138" value="2015-03-11">
<collectingDate id="7E1BE9D3FFFDFFF6FB82F97AE1CCF975" box="[1046,1216,1740,1764]" pageId="3" pageNumber="138" value="2015-03-11">11 March 2015</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FB7FF97BE0ACF975" box="[1259,1440,1740,1764]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFDFFF6FCA0F95CE110F893" box="[820,1052,1770,1794]" pageId="3" pageNumber="138">Paulo D. P. Pinheiro</collectorName>
;
</materialsCitation>
<materialsCitation id="AA893CA6FFFDFFF6FBBCF95CE16CF8AC" ID-GBIF-Occurrence="4052833303" collectingDate="2016-02-13" collectingDateMax="2016-02-13" collectingDateMin="2005-03-17" collectionCode="CFBH" collectorName="Leo R. Malagoli &amp; Alba N. Lozano" country="Brazil" county="Southeastern" elevation="818" location="Parque Estadual da Serra do Mar" municipality="State" pageId="3" pageNumber="138" specimenCode="40528-40529" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
and 
<collectionCode id="7CF0AE3EFFFDFFF6FBCCF95CE1AFF893" box="[1112,1187,1770,1794]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FB3FF95CE04CF893" box="[1195,1344,1770,1794]" pageId="3" pageNumber="138">40528–40529</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FCA0F8B1E6F0F88E" box="[820,1020,1799,1823]" pageId="3" pageNumber="138" value="2016-02-13">
<collectingDate id="7E1BE9D3FFFDFFF6FCA0F8B1E6F0F88E" box="[820,1020,1799,1823]" pageId="3" pageNumber="138" value="2016-02-13">13 February 2016</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FBBDF8BEE1D2F88E" box="[1065,1246,1799,1823]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFDFFF6FA8DF8B1E0C1F88E" box="[1305,1485,1799,1823]" pageId="3" pageNumber="138">Alba N. Lozano</collectorName>
; same locality as holotype.
</materialsCitation>
<materialsCitation id="AA893CA6FFFDFFF6FBF9F893E077F825" ID-GBIF-Occurrence="4052833306" collectingDate="2007-10-12" collectingDateMax="2016-02-13" collectingDateMin="2005-03-17" collectionCode="CFBH" collectorName="Leo R. Malagoli; Januaria M. Onca &amp; Nucleo Curucutu" country="Brazil" county="Southeastern" elevation="818" latitude="23.59" location="Parque Estadual da Serra do Mar" longitude="46.44" municipality="Itanhaem" pageId="3" pageNumber="138" specimenCode="17667" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFDFFF6FBF9F893E1B4F8AC" box="[1133,1208,1829,1853]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FB52F893E007F8AC" box="[1222,1291,1829,1853]" pageId="3" pageNumber="138">17667</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FA23F893E6C1F8CA" pageId="3" pageNumber="138" value="2007-10-12">
<collectingDate id="7E1BE9D3FFFDFFF6FA23F893E6C1F8CA" pageId="3" pageNumber="138" value="2007-10-12">12 October 2007</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FC6EF8F5E1BCF8CA" box="[1018,1200,1859,1883]" pageId="3" pageNumber="138">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFDFFF6FB78F8F5E0B8F8CA" box="[1260,1460,1859,1883]" pageId="3" pageNumber="138">Januária M. Onça</collectorName>
at 
<collectorName id="B714532DFFFDFFF6FCA0F8D6E104F8E8" box="[820,1032,1888,1913]" pageId="3" pageNumber="138">Núcleo Curucutu</collectorName>
, 
<location id="1F3E6020FFFDFFF6FB8AF8D7E0DDF8E9" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFDFFF6FB8AF8D7E0DDF8E9" box="[1054,1489,1888,1913]" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Parque Estadual da Serra do Mar" pageId="3" pageNumber="138" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFDFFF6FCA8F8C8E6D2F807" box="[828,990,1914,1942]" degrees="23" direction="south" minutes="59" orientation="latitude" pageId="3" pageNumber="138" seconds="56.16" value="23.59">
23°59 
<emphasis id="2895EAE9FFFDFFF6FCE8F8CCE68DF81A" box="[892,897,1914,1931]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FCE8F8CCE68DF81A" attach="none" box="[892,897,1914,1931]" fontSize="7" pageId="3" pageNumber="138">′</superScript>
</emphasis>
50.16 
<emphasis id="2895EAE9FFFDFFF6FC54F8CCE6C0F81A" box="[960,972,1914,1931]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FC54F8CCE6C0F81A" attach="right" box="[960,972,1914,1931]" fontSize="7" pageId="3" pageNumber="138">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFDFFF6FC7AF8C8E197F807" box="[1006,1179,1914,1942]" degrees="46" direction="west" minutes="44" orientation="longitude" pageId="3" pageNumber="138" seconds="41.46" value="46.44">
46°44 
<emphasis id="2895EAE9FFFDFFF6FBB9F8CCE13EF81A" box="[1069,1074,1914,1931]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FBB9F8CCE13EF81A" attach="none" box="[1069,1074,1914,1931]" fontSize="7" pageId="3" pageNumber="138">′</superScript>
</emphasis>
41.46 
<emphasis id="2895EAE9FFFDFFF6FBE5F8CCE172F81A" box="[1137,1150,1914,1931]" italics="true" pageId="3" pageNumber="138">
<superScript id="ED949BB3FFFDFFF6FBE5F8CCE172F81A" attach="right" box="[1137,1150,1914,1931]" fontSize="7" pageId="3" pageNumber="138">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<elevation id="91CCD1C8FFFDFFF6FB3EF8C8E022F807" box="[1194,1326,1918,1942]" metricMagnitude="2" metricUnit="m" metricValue="8.18" pageId="3" pageNumber="138" unit="m" value="818.0">
<quantity id="DD199B1EFFFDFFF6FB3EF8C8E1FFF807" box="[1194,1267,1918,1942]" metricMagnitude="2" metricUnit="m" metricValue="8.18" pageId="3" pageNumber="138" unit="m" value="818.0">818 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="FA3AAC81FFFDFFF6FCC5F82AE6B1F825" box="[849,957,1948,1972]" pageId="3" pageNumber="138">Itanhaém</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFDFFF6FBB0F82AE198F825" box="[1060,1172,1948,1972]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="138">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFDFFF6FAA3F82AE077F825" box="[1335,1403,1948,1972]" name="Brazil" pageId="3" pageNumber="138">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="AA893CA6FFFDFFF1FA12F82AE47DFBFE" ID-GBIF-Occurrence="4052833311" collectingDate="2007-11-16" collectingDateMax="2009-01-12" collectingDateMin="2007-11-16" collectionCode="CFBH" collectorName="Fabio Schunck" country="Brazil" county="Southeastern" elevation="719" lastPageId="4" lastPageNumber="139" latitude="24.0" location="Rio Mambu" longitude="46.46" municipality="Itanhaem" pageId="3" pageNumber="138" specimenCode="19351, 25808, 25810-25811" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFDFFF6FA12F82AE0DDF825" box="[1414,1489,1948,1972]" collectionName="CFBH" pageId="3" pageNumber="138">CFBH</collectionCode>
<specimenCode id="4A479E80FFFDFFF6FCA0F80FE675F840" box="[820,889,1977,2001]" pageId="3" pageNumber="138">19351</specimenCode>
, collected on 
<date id="6E5F103BFFFDFFF6FB8EF80FE1F8F840" box="[1050,1268,1977,2001]" pageId="3" pageNumber="138" value="2007-11-16">
<collectingDate id="7E1BE9D3FFFDFFF6FB8EF80FE1F8F840" box="[1050,1268,1977,2001]" pageId="3" pageNumber="138" value="2007-11-16">16 November 2007</collectingDate>
</date>
by 
<collectorName id="B714532DFFFDFFF6FAB7F80FE0C2F840" box="[1315,1486,1977,2001]" pageId="3" pageNumber="138">Fabio Schunck</collectorName>
; 
<collectionCode id="7CF0AE3EFFFAFFF1FF1AFC74E5D5FC4B" box="[142,217,962,986]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FF7FFC74E43CFC4B" box="[235,304,962,986]" pageId="4" pageNumber="139">25808</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FE7CFC74E7A3FC4B" box="[488,687,962,986]" pageId="4" pageNumber="139" value="2009-01-11">
<collectingDate id="7E1BE9D3FFFAFFF1FE7CFC74E7A3FC4B" box="[488,687,962,986]" pageId="4" pageNumber="139" value="2009-01-11">11 January 2009</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FD78FC74E5E3FC69" pageId="4" pageNumber="139">Fabio Schunck</collectorName>
; 
<collectionCode id="7CF0AE3EFFFAFFF1FF6EFC56E449FC69" box="[250,325,992,1016]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FED9FC56E4EDFC69" box="[333,481,992,1016]" pageId="4" pageNumber="139">25810–25811</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FDEEFC56E620FC69" box="[634,812,992,1016]" pageId="4" pageNumber="139" value="2009-01-12">
<collectingDate id="7E1BE9D3FFFAFFF1FDEEFC56E620FC69" box="[634,812,992,1016]" pageId="4" pageNumber="139" value="2009-01-12">12 January 2009</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FF24FC48E459FB87" box="[176,341,1022,1046]" pageId="4" pageNumber="139">Fabio Schunck</collectorName>
at 
<location id="1F3E6020FFFAFFF1FEEDFC48E70DFB87" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FEEDFC48E70DFB87" box="[377,513,1022,1046]" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Rio Mambu" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Rio Mambu</location>
, Núcleo Curucutu, 
<location id="1F3E6020FFFAFFF1FD49FC48E4CCFBA2" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FD49FC48E4CCFBA2" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Parque Estadual da Serra do Mar" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFAFFF1FE41FBAAE766FBA5" box="[469,618,1047,1076]" degrees="24" direction="south" minutes="0" orientation="latitude" pageId="4" pageNumber="139" seconds="36.15" value="24.0">
24°0 
<emphasis id="2895EAE9FFFAFFF1FD9CFBA1E701FBB9" box="[520,525,1047,1064]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FD9CFBA1E701FBB9" attach="right" box="[520,525,1047,1064]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
36.15 
<emphasis id="2895EAE9FFFAFFF1FDD8FBA1E754FBB9" box="[588,600,1047,1064]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FDD8FBA1E754FBB9" attach="right" box="[588,600,1047,1064]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFAFFF1FDEFFBADE624FBA2" box="[635,808,1047,1075]" degrees="46" direction="west" minutes="46" orientation="longitude" pageId="4" pageNumber="139" seconds="57.83" value="46.46">
46°46 
<emphasis id="2895EAE9FFFAFFF1FD2FFBA1E7CCFBB9" box="[699,704,1047,1064]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FD2FFBA1E7CCFBB9" attach="none" box="[699,704,1047,1064]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
57.83 
<emphasis id="2895EAE9FFFAFFF1FD6AFBA1E607FBB9" box="[766,779,1047,1064]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FD6AFBA1E607FBB9" attach="right" box="[766,779,1047,1064]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<elevation id="91CCD1C8FFFAFFF1FF1AFB8FE41CFBC0" box="[142,272,1081,1105]" metricMagnitude="2" metricUnit="m" metricValue="7.19" pageId="4" pageNumber="139" unit="m" value="719.0">
<quantity id="DD199B1EFFFAFFF1FF1AFB8FE5DAFBC0" box="[142,214,1081,1105]" metricMagnitude="2" metricUnit="m" metricValue="7.19" pageId="4" pageNumber="139" unit="m" value="719.0">719 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="FA3AAC81FFFAFFF1FE4FFB8FE74BFBC0" box="[475,583,1081,1105]" pageId="4" pageNumber="139">Itanhaém</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFAFFF1FD22FB8FE624FBC0" box="[694,808,1081,1105]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="139">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFAFFF1FEB9FBE1E47DFBFE" box="[301,369,1111,1135]" name="Brazil" pageId="4" pageNumber="139">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="AA893CA6FFFAFFF1FE17FBE1E625FB76" ID-GBIF-Occurrence="4052833305" collectingDate="2015-12-02" collectionCode="CFBH" collectorName="Leo R. Malagoli &amp; Delio B. Baeta" country="Brazil" elevation="805" latitude="23.58" location="Parque Estadual da Serra do Mar" longitude="46.35" municipality="Sao Vicente" pageId="4" pageNumber="139" specimenCode="40532-40533" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFAFFF1FE17FBE1E4C2FBFE" box="[387,462,1111,1135]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FE49FBE1E77EFBFE" box="[477,626,1111,1135]" pageId="4" pageNumber="139">40532–40533</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FC8BFBE1E44FFB1C" pageId="4" pageNumber="139" value="2015-12-02">
<collectingDate id="7E1BE9D3FFFAFFF1FC8BFBE1E44FFB1C" pageId="4" pageNumber="139" value="2015-12-02">2 December 2015</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FEE6FBC3E726FB1C" box="[370,554,1141,1165]" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFAFFF1FDF3FBC3E7A8FB1C" box="[615,676,1141,1165]" pageId="4" pageNumber="139">Délio</collectorName>
P. 
<collectorName id="B714532DFFFAFFF1FD5BFBC3E602FB1C" box="[719,782,1141,1165]" pageId="4" pageNumber="139">Baêta</collectorName>
at 
<collectorName id="B714532DFFFAFFF1FF1AFB25E49CFB3A" box="[142,400,1171,1195]" pageId="4" pageNumber="139">Núcleo Itutinga-Pilões</collectorName>
, 
<location id="1F3E6020FFFAFFF1FE0BFB25E620FB3B" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FE0BFB25E620FB3B" box="[415,812,1171,1195]" country="Brazil" municipality="Sao Vicente" name="Parque Estadual da Serra do Mar" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFAFFF1FF02FB07E4FAFB59" box="[150,502,1197,1225]" degrees="46" direction="west" minutes="35" orientation="longitude" pageId="4" pageNumber="139" seconds="31.59" value="46.35">
23°58 
<emphasis id="2895EAE9FFFAFFF1FF43FB1BE5D0FB2F" box="[215,220,1197,1214]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FF43FB1BE5D0FB2F" attach="none" box="[215,220,1197,1214]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
59.15 
<emphasis id="2895EAE9FFFAFFF1FE8EFB1BE42BFB2F" box="[282,295,1197,1214]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE8EFB1BE42BFB2F" attach="right" box="[282,295,1197,1214]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
S, 
<collectorName id="B714532DFFFAFFF1FEDDFB07E4FAFB59" box="[329,502,1197,1225]" pageId="4" pageNumber="139">
46°35 
<emphasis id="2895EAE9FFFAFFF1FE1CFB1BE481FB2F" box="[392,397,1197,1214]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE1CFB1BE481FB2F" attach="none" box="[392,397,1197,1214]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
31.59 
<emphasis id="2895EAE9FFFAFFF1FE58FB1BE4D4FB2F" box="[460,472,1197,1214]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE58FB1BE4D4FB2F" attach="right" box="[460,472,1197,1214]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
W
</collectorName>
</geoCoordinate>
; 
<elevation id="91CCD1C8FFFAFFF1FD91FB06E785FB58" box="[517,649,1200,1225]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="4" pageNumber="139" unit="m" value="805.0">
<quantity id="DD199B1EFFFAFFF1FD91FB06E742FB59" box="[517,590,1200,1224]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="4" pageNumber="139" unit="m" value="805.0">805 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="FA3AAC81FFFAFFF1FF27FB79E433FB76" box="[179,319,1231,1255]" pageId="4" pageNumber="139">São Vicente</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFAFFF1FE28FB79E73FFB76" box="[444,563,1231,1255]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="139">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFAFFF1FD71FB79E625FB76" box="[741,809,1231,1255]" name="Brazil" pageId="4" pageNumber="139">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="AA893CA6FFFAFFF1FF1AFB5AE7FEFAED" ID-GBIF-Occurrence="4052833312" collectingDate="2016-01-26" collectingDateMax="2008-11-29" collectingDateMin="2005-12-16" collectionCode="CFBH" collectorName="Leo R. Malagoli; Wesley P. Soares &amp; Kleber E. Rodrigues" country="Brazil" county="Southeastern" elevation="718" latitude="23.57" location="Nucleo Curucutu" longitude="46.38" municipality="Sao Paulo" pageId="4" pageNumber="139" specimenCode="40526-40527" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFAFFF1FF1AFB5AE5D5FA95" box="[142,217,1260,1284]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FF70FB5AE475FA95" box="[228,377,1260,1284]" pageId="4" pageNumber="139">40526–40527</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FD8EFB5AE7DEFA95" box="[538,722,1260,1285]" pageId="4" pageNumber="139" value="2016-01-26">
<collectingDate id="7E1BE9D3FFFAFFF1FD8EFB5AE7DEFA95" box="[538,722,1260,1285]" pageId="4" pageNumber="139" value="2016-01-26">26 January 2016</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FC95FB5BE41FFAB3" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
, 
<collectorName id="B714532DFFFAFFF1FEB5FABCE4E9FAB3" box="[289,485,1290,1314]" pageId="4" pageNumber="139">Wesley P. Soares</collectorName>
, and 
<collectorName id="B714532DFFFAFFF1FDB1FABCE602FAB3" box="[549,782,1290,1314]" pageId="4" pageNumber="139">Kleber E. Rodrigues</collectorName>
at 
<location id="1F3E6020FFFAFFF1FF1AFA9EE46EFAD1" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FF1AFA9EE46EFAD1" box="[142,354,1320,1344]" country="Brazil" county="Southeastern" municipality="Sao Paulo" name="Nucleo Curucutu" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Núcleo Curucutu</location>
, 
<location id="1F3E6020FFFAFFF1FEEDFA9FE620FAD1" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FEEDFA9FE620FAD1" box="[377,812,1320,1344]" country="Brazil" county="Southeastern" municipality="Sao Paulo" name="Parque Estadual da Serra do Mar" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFAFFF1FF02FAF0E434FACF" box="[150,312,1346,1374]" degrees="23" direction="south" minutes="57" orientation="latitude" pageId="4" pageNumber="139" seconds="54.62" value="23.57">
23°57 
<emphasis id="2895EAE9FFFAFFF1FF43FAF4E5D0FAC2" box="[215,220,1346,1363]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FF43FAF4E5D0FAC2" attach="none" box="[215,220,1346,1363]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
54.62 
<emphasis id="2895EAE9FFFAFFF1FE8EFAF4E42BFAC2" box="[282,295,1346,1363]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE8EFAF4E42BFAC2" attach="right" box="[282,295,1346,1363]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFAFFF1FEDDFAF0E4FAFACF" box="[329,502,1346,1374]" degrees="46" direction="west" minutes="38" orientation="longitude" pageId="4" pageNumber="139" seconds="58.24" value="46.38">
46°38 
<emphasis id="2895EAE9FFFAFFF1FE1CFAF4E481FAC2" box="[392,397,1346,1363]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE1CFAF4E481FAC2" attach="none" box="[392,397,1346,1363]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
58.24 
<emphasis id="2895EAE9FFFAFFF1FE58FAF4E4D4FAC2" box="[460,472,1346,1363]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE58FAF4E4D4FAC2" attach="right" box="[460,472,1346,1363]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<elevation id="91CCD1C8FFFAFFF1FD91FAF0E785FACF" box="[517,649,1350,1374]" metricMagnitude="2" metricUnit="m" metricValue="7.18" pageId="4" pageNumber="139" unit="m" value="718.0">
<quantity id="DD199B1EFFFAFFF1FD91FAF0E742FACF" box="[517,590,1350,1374]" metricMagnitude="2" metricUnit="m" metricValue="7.18" pageId="4" pageNumber="139" unit="m" value="718.0">718 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="FA3AAC81FFFAFFF1FF3AFAD2E42CFAED" box="[174,288,1380,1404]" pageId="4" pageNumber="139">São Paulo</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFAFFF1FE1BFAD2E70DFAED" box="[399,513,1380,1404]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="139">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFAFFF1FD3EFAD2E7E2FAED" box="[682,750,1380,1404]" name="Brazil" pageId="4" pageNumber="139">Brazil</collectingCountry>
.
</materialsCitation>
<materialsCitation id="AA893CA6FFFAFFF1FD6FFAD2E4B1FA26" ID-GBIF-Occurrence="4052833310" collectingDate="2005-12-16" collectingDateMax="2008-11-29" collectingDateMin="2005-12-16" collectionCode="CFBH" collectorName="Leo R. Malagoli" country="Brazil" county="Southeastern" elevation="718" location="Nucleo Curucutu" municipality="State" pageId="4" pageNumber="139" specimenCode="11172" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCount id="0CE7FD72FFFAFFF1FD6FFAD2E421FA0B" pageId="4" pageNumber="139">Five adult females</specimenCount>
: 
<collectionCode id="7CF0AE3EFFFAFFF1FEABFA34E486FA0B" box="[319,394,1410,1434]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FE0DFA34E4D2FA0B" box="[409,478,1410,1434]" pageId="4" pageNumber="139">11172</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FD19FA34E5CFFA26" pageId="4" pageNumber="139" value="2005-12-16">
<collectingDate id="7E1BE9D3FFFAFFF1FD19FA34E5CFFA26" pageId="4" pageNumber="139" value="2005-12-16">16 December 2005</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FF6CFA16E4B5FA26" box="[248,441,1439,1463]" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
;
</materialsCitation>
<materialsCitation id="AA893CA6FFFAFFF1FE5EFA16E5F9FA62" ID-GBIF-Occurrence="4052833309" collectingDate="2008-11-29" collectingDateMax="2008-11-29" collectingDateMin="2005-12-16" collectionCode="CFBH" collectorName="Leo R. Malagoli" country="Brazil" county="Southeastern" elevation="718" location="Nucleo Curucutu" municipality="State" pageId="4" pageNumber="139" specimenCode="22131" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFAFFF1FE5EFA16E719FA29" box="[458,533,1440,1464]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FDB6FA16E76BFA29" box="[546,615,1440,1464]" pageId="4" pageNumber="139">22131</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FC86FA16E447FA47" pageId="4" pageNumber="139" value="2008-11-29">
<collectingDate id="7E1BE9D3FFFAFFF1FC86FA16E447FA47" pageId="4" pageNumber="139" value="2008-11-29">29 November 2008</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FE1EFA08E759FA44" box="[394,597,1469,1493]" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
; same locality as holotype.
</materialsCitation>
<materialsCitation id="AA893CA6FFFAFFF1FE95FA6DE47CF9FA" ID-GBIF-Occurrence="4052833308" collectingDate="2007-09-12" collectingDateMax="2008-11-29" collectingDateMin="2005-12-16" collectionCode="CFBH" collectorName="Leo R. Malagoli" country="Brazil" county="Southeastern" elevation="818" latitude="23.59" location="Parque Estadual da Serra do Mar" longitude="46.44" municipality="Itanhaem" pageId="4" pageNumber="139" specimenCode="17655" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFAFFF1FE95FA6DE440FA62" box="[257,332,1499,1523]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FECEFA6DE493FA62" box="[346,415,1499,1523]" pageId="4" pageNumber="139">17655</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FDDDFA6DE620FA62" box="[585,812,1499,1523]" pageId="4" pageNumber="139" value="2007-09-12">
<collectingDate id="7E1BE9D3FFFAFFF1FDDDFA6DE620FA62" box="[585,812,1499,1523]" pageId="4" pageNumber="139" value="2007-09-12">12 September 2007</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FF24FA4CE46DF980" box="[176,353,1529,1553]" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
at 
<collectorName id="B714532DFFFAFFF1FE10FA4FE744F980" box="[388,584,1529,1553]" pageId="4" pageNumber="139">Núcleo Curucutu</collectorName>
, 
<location id="1F3E6020FFFAFFF1FDC0FA4CE428F9BF" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FDC0FA4CE428F9BF" country="Brazil" county="Southeastern" municipality="Itanhaem" name="Parque Estadual da Serra do Mar" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFAFFF1FEA1F9A1E4DBF9BE" box="[309,471,1555,1583]" degrees="23" direction="south" minutes="59" orientation="latitude" pageId="4" pageNumber="139" seconds="50.16" value="23.59">
23°59 
<emphasis id="2895EAE9FFFAFFF1FEE1F9A5E476F9B5" box="[373,378,1555,1572]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FEE1F9A5E476F9B5" attach="none" box="[373,378,1555,1572]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
50.16 
<emphasis id="2895EAE9FFFAFFF1FE2DF9A5E4C9F9B5" box="[441,453,1555,1572]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE2DF9A5E4C9F9B5" attach="right" box="[441,453,1555,1572]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFAFFF1FE70F9A1E79DF9BF" box="[484,657,1555,1583]" degrees="46" direction="west" minutes="44" orientation="longitude" pageId="4" pageNumber="139" seconds="41.46" value="46.44">
46°44 
<emphasis id="2895EAE9FFFAFFF1FDB0F9A5E725F9B5" box="[548,553,1555,1572]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FDB0F9A5E725F9B5" attach="none" box="[548,553,1555,1572]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
41.46 
<emphasis id="2895EAE9FFFAFFF1FDF3F9A5E778F9B5" box="[615,628,1555,1572]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FDF3F9A5E778F9B5" attach="right" box="[615,628,1555,1572]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<quantity id="DD199B1EFFFAFFF1FD0AF9A1E7E8F9BF" box="[670,740,1559,1583]" metricMagnitude="2" metricUnit="m" metricValue="8.18" pageId="4" pageNumber="139" unit="m" value="818.0">
<elevation id="91CCD1C8FFFAFFF1FD0AF9A1E7E8F9BF" box="[670,740,1559,1583]" metricMagnitude="2" metricUnit="m" metricValue="8.18" pageId="4" pageNumber="139" unit="m" value="818.0">818 m</elevation>
</quantity>
above sea level), municipality of 
<collectingMunicipality id="FA3AAC81FFFAFFF1FE5EF983E73AF9DC" box="[458,566,1589,1613]" pageId="4" pageNumber="139">Itanhaém</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFAFFF1FD26F983E624F9DC" box="[690,808,1589,1613]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="139">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFAFFF1FEB8F9E5E47CF9FA" box="[300,368,1619,1643]" name="Brazil" pageId="4" pageNumber="139">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="AA893CA6FFFAFFF1FE15F9E5E60CF973" ID-GBIF-Occurrence="4052833313" collectingDate="2015-12-02" collectionCode="CFBH" collectorName="Leo R. Malagoli &amp; Delio P. Baeta" country="Brazil" elevation="805" latitude="23.58" location="Parque Estadual da Serra do Mar" longitude="46.35" municipality="Sao Vicente" pageId="4" pageNumber="139" specimenCode="40531, 40534" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectionCode id="7CF0AE3EFFFAFFF1FE15F9E5E4C0F9FA" box="[385,460,1619,1643]" collectionName="CFBH" pageId="4" pageNumber="139">CFBH</collectionCode>
<specimenCode id="4A479E80FFFAFFF1FE4DF9E5E712F9FA" box="[473,542,1619,1643]" pageId="4" pageNumber="139">40531</specimenCode>
, 
<specimenCode id="4A479E80FFFAFFF1FDA4F9E5E779F9FA" box="[560,629,1619,1643]" collectionCode="CFBH" country="Brazil" httpUri="http://grbio.org/cool/3q7f-dnij" name="Universidade Estadual Paulista" pageId="4" pageNumber="139" type="University or college">40534</specimenCode>
, collected on 
<date id="6E5F103BFFFAFFF1FC8BF9E5E44FF919" pageId="4" pageNumber="139" value="2015-12-02">
<collectingDate id="7E1BE9D3FFFAFFF1FC8BF9E5E44FF919" pageId="4" pageNumber="139" value="2015-12-02">2 December 2015</collectingDate>
</date>
by 
<collectorName id="B714532DFFFAFFF1FEE6F9C7E726F918" box="[370,554,1649,1673]" pageId="4" pageNumber="139">Leo R. Malagoli</collectorName>
and 
<collectorName id="B714532DFFFAFFF1FDF3F9C7E602F918" box="[615,782,1649,1673]" pageId="4" pageNumber="139">Délio P. Baêta</collectorName>
at 
<collectorName id="B714532DFFFAFFF1FF1AF938E49CF937" box="[142,400,1678,1702]" pageId="4" pageNumber="139">Núcleo Itutinga-Pilões</collectorName>
, 
<location id="1F3E6020FFFAFFF1FE0BF939E620F937" LSID="urn:lsid:plazi:treatment:924887EDFFFDFFFDFB97FAB1E1AEF840:1F3E6020FFFAFFF1FE0BF939E620F937" box="[415,812,1678,1703]" country="Brazil" municipality="Sao Vicente" name="Parque Estadual da Serra do Mar" pageId="4" pageNumber="139" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar</location>
(
<geoCoordinate id="7FD5503CFFFAFFF1FF02F91AE434F955" box="[150,312,1704,1732]" degrees="23" direction="south" minutes="58" orientation="latitude" pageId="4" pageNumber="139" seconds="59.15" value="23.58">
23°58 
<emphasis id="2895EAE9FFFAFFF1FF43F91EE5D0F928" box="[215,220,1704,1721]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FF43F91EE5D0F928" attach="none" box="[215,220,1704,1721]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
59.15 
<emphasis id="2895EAE9FFFAFFF1FE8EF91EE42BF928" box="[282,295,1704,1721]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE8EF91EE42BF928" attach="right" box="[282,295,1704,1721]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="7FD5503CFFFAFFF1FEDDF91AE4FAF955" box="[329,502,1704,1732]" degrees="46" direction="west" minutes="35" orientation="longitude" pageId="4" pageNumber="139" seconds="31.59" value="46.35">
46°35 
<emphasis id="2895EAE9FFFAFFF1FE1CF91EE481F928" box="[392,397,1704,1721]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE1CF91EE481F928" attach="none" box="[392,397,1704,1721]" fontSize="7" pageId="4" pageNumber="139">′</superScript>
</emphasis>
31.59 
<emphasis id="2895EAE9FFFAFFF1FE58F91EE4D4F928" box="[460,472,1704,1721]" italics="true" pageId="4" pageNumber="139">
<superScript id="ED949BB3FFFAFFF1FE58F91EE4D4F928" attach="right" box="[460,472,1704,1721]" fontSize="7" pageId="4" pageNumber="139">′ ′</superScript>
</emphasis>
W
</geoCoordinate>
; 
<elevation id="91CCD1C8FFFAFFF1FD91F91AE785F955" box="[517,649,1708,1732]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="4" pageNumber="139" unit="m" value="805.0">
<quantity id="DD199B1EFFFAFFF1FD91F91AE742F955" box="[517,590,1708,1732]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="4" pageNumber="139" unit="m" value="805.0">805 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="FA3AAC81FFFAFFF1FF39F97CE43FF973" box="[173,307,1738,1762]" pageId="4" pageNumber="139">São Vicente</collectingMunicipality>
, State of 
<collectingRegion id="D825F819FFFAFFF1FE34F97CE71DF973" box="[416,529,1738,1762]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="139">São Paulo</collectingRegion>
, Southeastern 
<collectingCountry id="62F6766BFFFAFFF1FD2CF97CE7F0F973" box="[696,764,1738,1762]" name="Brazil" pageId="4" pageNumber="139">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="4E9E6673FFFDFFF6FF1AF8C3E76FF85E" ID-DOI="http://doi.org/10.5281/zenodo.7712273" ID-Zenodo-Dep="7712273" httpUri="https://zenodo.org/record/7712273/files/figure.png" pageId="3" pageNumber="138" startId="3.[142,156,1909,1928]" targetBox="[118,788,1398,1883]" targetPageId="3">
<paragraph id="1A5E36FBFFFDFFF6FF1AF8C3E76FF85E" blockId="3.[118,789,1908,1999]" pageId="3" pageNumber="138">
FIG. 1.—Live adult male 
<taxonomicName id="DDE14D78FFFDFFF6FEE5F8C2E4E6F816" authority="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad, 2017" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[369,490,1908,1927]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="138" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
<emphasis id="2895EAE9FFFDFFF6FEE5F8C2E4E6F816" box="[369,490,1908,1927]" italics="true" pageId="3" pageNumber="138">Hŋlodes caete</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A65792FFFDFFF6FE65F8C3E73BF819" box="[497,567,1909,1928]" pageId="3" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
(holotype CFBH 40524; snout–vent length ¼ 31.6 mm). Photograph taken in Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online.
</paragraph>
</caption>
<caption id="4E9E6673FFFAFFF1FF32FC34E1B3FC04" ID-DOI="http://doi.org/10.5281/zenodo.7712275" ID-Zenodo-Dep="7712275" box="[166,1215,897,918]" httpUri="https://zenodo.org/record/7712275/files/figure.png" pageId="4" pageNumber="139" startId="4.[166,180,898,917]" targetBox="[142,1515,191,872]" targetPageId="4">
<paragraph id="1A5E36FBFFFAFFF1FF32FC34E1B3FC04" blockId="4.[166,1215,897,918]" box="[166,1215,897,918]" pageId="4" pageNumber="139">
FIG. 2.— 
<taxonomicName id="DDE14D78FFFAFFF1FF63FC34E4B5FC04" authority=", CFBH" authorityName="CFBH" box="[247,441,898,917]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FF63FC34E47CFC04" box="[247,368,898,917]" italics="true" pageId="4" pageNumber="139">Hŋlodes caete</emphasis>
, CFBH
</taxonomicName>
40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm.
</paragraph>
</caption>
<subSubSection id="52FB6570FFFAFFF3FF3DF95EE010FB4D" lastPageId="6" lastPageNumber="141" pageId="4" pageNumber="139" type="diagnosis">
<paragraph id="1A5E36FBFFFAFFF1FF3DF95EE1F8FB3A" blockId="4.[142,813,962,2001]" lastBlockId="4.[844,1514,962,2001]" pageId="4" pageNumber="139">
<emphasis id="2895EAE9FFFAFFF1FF3DF95EE42BF891" bold="true" box="[169,295,1768,1792]" pageId="4" pageNumber="139">Diagnosis.</emphasis>
— 
<taxonomicName id="DDE14D78FFFAFFF1FED6F95EE4D6F891" box="[322,474,1768,1792]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FED6F95EE4D6F891" box="[322,474,1768,1792]" italics="true" pageId="4" pageNumber="139">Hŋlodes caete</emphasis>
</taxonomicName>
is a slender species with light, oblique lateral stripes and relatively smooth dorsal surfaces (
<figureCitation id="82DA2A7EFFFAFFF1FF03F892E5FDF8AD" box="[151,241,1828,1852]" captionStart="FIG" captionStartId="3.[142,156,1909,1928]" captionTargetBox="[118,788,1398,1883]" captionTargetId="figure-805@3.[118,788,1398,1883]" captionTargetPageId="3" captionText="FIG. 1.—Live adult male Hŋlodes caete sp. nov. (holotype CFBH 40524; snout–vent length ¼ 31.6 mm). Photograph taken in Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712273" httpUri="https://zenodo.org/record/7712273/files/figure.png" pageId="4" pageNumber="139">Figs. 1</figureCitation>
and 
<figureCitation id="82DA2A7EFFFAFFF1FED6F892E46AF8AD" box="[322,358,1828,1852]" captionStart="FIG" captionStartId="4.[166,180,898,917]" captionTargetBox="[142,1515,191,872]" captionTargetId="figure-795@4.[142,1516,191,872]" captionTargetPageId="4" captionText="FIG. 2.—Hŋlodes caete, CFBH 40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7712275" httpUri="https://zenodo.org/record/7712275/files/figure.png" pageId="4" pageNumber="139">2A</figureCitation>
). It is diagnosed by the following combination of characters: (1) pointed nuptial tubercles distributed in an elliptical area at the base of the dorsal surface of the thumb of male individuals (
<figureCitation id="82DA2A7EFFFAFFF1FD36F8C8E7E2F807" box="[674,750,1918,1942]" captionStart="Fig" captionStartId="6.[868,900,815,834]" captionTargetBox="[844,1515,193,790]" captionTargetId="figure-488@6.[844,1516,191,790]" captionTargetPageId="6" captionText="Fig. 4.—Distribution of nuptial tubercles on the right thumb of an adult male Hŋlodes caete, CFBH 40524 (holotype). (A) Drawing in lateral view. (B) Picture in dorsal view, with dashed line indicating the elliptical distribution of tubercles. Scale bars ¼ 1 mm. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712279" httpUri="https://zenodo.org/record/7712279/files/figure.png" pageId="4" pageNumber="139">Fig. 4</figureCitation>
); (2) medium-sized (SVL about 31.1–34.0 mm in adult males and 
<quantity id="DD199B1EFFFAFFF1FF1AF80FE421F840" box="[142,301,1977,2001]" metricMagnitude="-2" metricUnit="m" metricValue="3.575" metricValueMax="3.8299999999999996" metricValueMin="3.32" pageId="4" pageNumber="139" unit="mm" value="35.75" valueMax="38.3" valueMin="33.2">33.2–38.3 mm</quantity>
in adult females; variation reported in 
<tableCitation id="57630340FFFAFFF1FD7AF80FE656FC4B" box="[750,858,962,2001]" captionStart="TABLE" captionStartId="6.[169,182,188,207]" captionTargetPageId="6" captionText="TABLE 3.—Measurements (mm, reported as ¯X ± 1 SD) of males (n ¼ 16) and females (n ¼ 5) of the type series of Hŋlodes caete sp. nov." httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" pageId="4" pageNumber="139" tableUuid="4E9E6673FFF8FFF3FF3DFF0AE7C7FF76">Table 3</tableCitation>
); (3) advertisement call with a call duration of 1.22– 
<geoCoordinate id="7FD5503CFFFAFFF1FA36FC74E0E8FC48" box="[1442,1508,962,986]" degrees="2.75" direction="south" orientation="latitude" pageId="4" pageNumber="139" precision="555" value="-2.75">2.75 s</geoCoordinate>
, 20–45 notes per call, note repetition rate of 14.8–18.3/s, note duration of 
<geoCoordinate id="7FD5503CFFFAFFF1FC46FC48E102FB87" box="[978,1038,1022,1046]" degrees="0.009" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="55" value="0.009">0.009</geoCoordinate>
<geoCoordinate id="7FD5503CFFFAFFF1FB9AFC48E15BFB87" box="[1038,1111,1022,1046]" degrees="0.023" direction="west" orientation="longitude" pageId="4" pageNumber="139" precision="55" value="-0.023">
– 
<geoCoordinate id="7FD5503CFFFAFFF1FB88FC48E15BFB87" box="[1052,1111,1022,1046]" degrees="0.023" direction="south" orientation="latitude" pageId="4" pageNumber="139" precision="55" value="-0.023">0.023</geoCoordinate>
</geoCoordinate>
s, and internote duration of 
<geoCoordinate id="7FD5503CFFFAFFF1FA35FC4BE0D1FB84" box="[1441,1501,1021,1045]" degrees="0.036" direction="north" orientation="latitude" pageId="4" pageNumber="139" precision="55" value="0.036">0.036</geoCoordinate>
<geoCoordinate id="7FD5503CFFFAFFF1FA48FC4BE684FBA2" degrees="0.058" direction="west" orientation="longitude" pageId="4" pageNumber="139" precision="55" value="-0.058">
– 
<geoCoordinate id="7FD5503CFFFAFFF1FCD8FBADE684FBA2" box="[844,904,1051,1075]" degrees="0.058" direction="south" orientation="latitude" pageId="4" pageNumber="139" precision="55" value="-0.058">0.058</geoCoordinate>
</geoCoordinate>
s (
<tableCitation id="57630340FFFAFFF1FC33FBADE6F1FBA5" box="[935,1021,1051,1076]" captionStart="TABLE" captionStartId="6.[169,182,1654,1673]" captionTargetPageId="6" captionText="TABLE 4.—Advertisement calls of Hŋlodes caete sp. nov. and of the other three Hŋlodes species with males having nuptial tubercles on the thumb: Hŋlodes fredi (Canedo and Pombal 2007), H. phŋllodes (Heyer and Cocroft 1986), and H. pipilans (Canedo and Pombal 2007). Values are reported as ¯X ± 1 SD. (—) in cells indicates unavailable data in cited references." httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DF9C0E761F929" pageId="4" pageNumber="139" tableUuid="4E9E6673FFF8FFF3FF3DF9C0E761F929">Table 4</tableCitation>
); and (4) uncorrected 
<emphasis id="2895EAE9FFFAFFF1FB61FBAAE00FFBA2" box="[1269,1283,1052,1075]" italics="true" pageId="4" pageNumber="139">p</emphasis>
distance of 10.5% ± 0.0003 from 
<taxonomicName id="DDE14D78FFFAFFF1FC4EFB8CE13CFBC0" box="[986,1072,1081,1105]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FC4EFB8CE13CFBC0" box="[986,1072,1081,1105]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
, 10.2% ± 0.001 from 
<taxonomicName id="DDE14D78FFFAFFF1FABCFB8CE0BEFBC0" box="[1320,1458,1081,1105]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FABCFB8CE0BEFBC0" box="[1320,1458,1081,1105]" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
, and 9.5% ± 0.001 from 
<taxonomicName id="DDE14D78FFFAFFF1FBBBFBEEE1A6FBFE" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1071,1194,1111,1135]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FBBBFBEEE1A6FBFE" box="[1071,1194,1111,1135]" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
for complete 16S gene, and of 6.5% from 
<taxonomicName id="DDE14D78FFFAFFF1FC64FBC0E146FB1C" box="[1008,1098,1141,1165]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FC64FBC0E146FB1C" box="[1008,1098,1141,1165]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
, 6.2% from 
<taxonomicName id="DDE14D78FFFAFFF1FB4AFBC0E060FB1C" box="[1246,1388,1141,1165]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FB4AFBC0E060FB1C" box="[1246,1388,1141,1165]" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
, and 6.2% from 
<taxonomicName id="DDE14D78FFFAFFF1FC1EFB22E10AFB3A" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[906,1030,1171,1195]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FC1EFB22E10AFB3A" box="[906,1030,1171,1195]" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
for partial 16S gene.
</paragraph>
<paragraph id="1A5E36FBFFFAFFF1FCF3FB07E69DF8E9" blockId="4.[844,1514,962,2001]" pageId="4" pageNumber="139">
<emphasis id="2895EAE9FFFAFFF1FCF3FB07E001FB58" bold="true" box="[871,1293,1201,1225]" pageId="4" pageNumber="139">Comparisons with other species.</emphasis>
— 
<taxonomicName id="DDE14D78FFFAFFF1FABCFB07E0C4FB58" box="[1320,1480,1201,1225]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FABCFB07E0C4FB58" box="[1320,1480,1201,1225]" italics="true" pageId="4" pageNumber="139">Hŋlodes caete</emphasis>
</taxonomicName>
is easily differentiated from all other species of 
<taxonomicName id="DDE14D78FFFAFFF1FAD7FB79E090FB76" authorityName="Fitzinger" authorityYear="1826" box="[1347,1436,1231,1255]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="2895EAE9FFFAFFF1FAD7FB79E090FB76" box="[1347,1436,1231,1255]" italics="true" pageId="4" pageNumber="139">Hŋlodes</emphasis>
</taxonomicName>
by the presence of nuptial tubercles on the thumb of males, except for 
<taxonomicName id="DDE14D78FFFAFFF1FCE0FABDE6C6FAB3" box="[884,970,1290,1314]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FCE0FABDE6C6FAB3" box="[884,970,1290,1314]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFFAFFF1FC4DFABDE168FAB3" box="[985,1124,1290,1314]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FC4DFABDE168FAB3" box="[985,1124,1290,1314]" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFFAFFF1FB30FABDE012FAB3" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1188,1310,1290,1314]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FB30FABDE012FAB3" box="[1188,1310,1290,1314]" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
(nuptial tubercles present on thumb of males in these species; 
<bibRefCitation id="7E704B0AFFFAFFF1FAFAFA9FE6EDFACF" author="Heyer, W. R. &amp; R. B. Cocroft" pageId="4" pageNumber="139" pagination="100 - 109" refId="ref11139" refString="Heyer, W. R., and R. B. Cocroft. 1986. Descriptions of two new species of Hnlodes from the Atlantic Forests of Brazil (Amphibia: Leptodactylidae). Proceedings of Biological Society of Washington 99: 100 - 109." type="journal article" year="1986">Heyer and Cocroft 1986</bibRefCitation>
; 
<bibRefCitation id="7E704B0AFFFAFFF1FC7AFAF0E01AFACF" author="Canedo, C. &amp; J. P. Pombal, Jr." box="[1006,1302,1350,1374]" pageId="4" pageNumber="139" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
). The males of the four species with nuptial tubercles differ in SVL (
<emphasis id="2895EAE9FFFAFFF1FAE0FAD3E088FAED" box="[1396,1412,1381,1404]" italics="true" pageId="4" pageNumber="139">F</emphasis>
¼ 369.6; 
<emphasis id="2895EAE9FFFAFFF1FCD8FA35E657FA0B" box="[844,859,1411,1434]" italics="true" pageId="4" pageNumber="139">P</emphasis>
&lt;0.0001; 
<emphasis id="2895EAE9FFFAFFF1FC48FA35E6E6FA0B" box="[988,1002,1411,1434]" italics="true" pageId="4" pageNumber="139">n</emphasis>
¼ 120). Males of 
<taxonomicName id="DDE14D78FFFAFFF1FB28FA35E01BFA0B" box="[1212,1303,1411,1434]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FB28FA35E01BFA0B" box="[1212,1303,1411,1434]" italics="true" pageId="4" pageNumber="139">H. caete</emphasis>
</taxonomicName>
(32.3 ± 
<quantity id="DD199B1EFFFAFFF1FA14FA34E0EAFA08" box="[1408,1510,1410,1434]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="139" unit="mm" value="0.88">0.88 mm</quantity>
, 31.1–34.0 mm; 
<emphasis id="2895EAE9FFFAFFF1FC6BFA16E101FA26" box="[1023,1037,1440,1463]" italics="true" pageId="4" pageNumber="139">n</emphasis>
¼ 16) are distinguishable by their larger size than males of 
<taxonomicName id="DDE14D78FFFAFFF1FB88FA08E1ABFA47" box="[1052,1191,1470,1494]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FB88FA08E1ABFA47" box="[1052,1191,1470,1494]" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
(27.8 ± 
<quantity id="DD199B1EFFFAFFF1FA9AFA08E07FFA44" box="[1294,1395,1470,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="4" pageNumber="139" unit="mm" value="1.01">1.01 mm</quantity>
, 
<quantity id="DD199B1EFFFAFFF1FAEBFA0BE676FA62" metricMagnitude="-2" metricUnit="m" metricValue="2.755" metricValueMax="2.9699999999999998" metricValueMin="2.54" pageId="4" pageNumber="139" unit="mm" value="27.549999999999997" valueMax="29.7" valueMin="25.4">25.4–29.7 mm</quantity>
; 
<emphasis id="2895EAE9FFFAFFF1FC13FA6AE699FA62" box="[903,917,1500,1523]" italics="true" pageId="4" pageNumber="139">n</emphasis>
¼ 76; personal measurements of 
<typeStatus id="C55A8859FFFAFFF1FA81FA6AE048FA65" box="[1301,1348,1500,1524]" pageId="4" pageNumber="139">type</typeStatus>
series) and 
<taxonomicName id="DDE14D78FFFAFFF1FA5BFA6AE6A8F980" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FA5BFA6AE6A8F980" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
(24.1 ± 
<quantity id="DD199B1EFFFAFFF1FB87FA4FE176F980" box="[1043,1146,1529,1553]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="4" pageNumber="139" unit="mm" value="0.71">0.71 mm</quantity>
, 23.0– 
<quantity id="DD199B1EFFFAFFF1FB53FA4FE021F980" box="[1223,1325,1529,1553]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="139" unit="mm" value="25.1">25.1 mm</quantity>
; 
<emphasis id="2895EAE9FFFAFFF1FAA8FA4CE046F980" box="[1340,1354,1530,1553]" italics="true" pageId="4" pageNumber="139">n</emphasis>
¼ 9; 
<bibRefCitation id="7E704B0AFFFAFFF1FA00FA4FE11EF9BE" author="Canedo, C. &amp; J. P. Pombal, Jr." pageId="4" pageNumber="139" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
). The mean SVL of male 
<taxonomicName id="DDE14D78FFFAFFF1FABBF9AEE085F9BE" box="[1327,1417,1560,1583]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FABBF9AEE085F9BE" box="[1327,1417,1560,1583]" italics="true" pageId="4" pageNumber="139">H. caete</emphasis>
</taxonomicName>
is larger than the SVL of male 
<taxonomicName id="DDE14D78FFFAFFF1FBABF980E1C5F9DC" box="[1087,1225,1589,1613]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FBABF980E1C5F9DC" box="[1087,1225,1589,1613]" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
(Tukey test; 
<emphasis id="2895EAE9FFFAFFF1FACFF983E063F9DD" box="[1371,1391,1589,1612]" italics="true" pageId="4" pageNumber="139">Q</emphasis>
¼ 23.85; 
<emphasis id="2895EAE9FFFAFFF1FA4FF980E0E6F9DC" box="[1499,1514,1590,1613]" italics="true" pageId="4" pageNumber="139">P</emphasis>
&lt;0.01) and 
<taxonomicName id="DDE14D78FFFAFFF1FC74F9E2E150F9FA" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[992,1116,1619,1643]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FC74F9E2E150F9FA" box="[992,1116,1619,1643]" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
(
<emphasis id="2895EAE9FFFAFFF1FBFBF9E5E18FF9FB" box="[1135,1155,1619,1642]" italics="true" pageId="4" pageNumber="139">Q</emphasis>
¼ 28.73; 
<emphasis id="2895EAE9FFFAFFF1FB63F9E2E00AF9FA" box="[1271,1286,1620,1643]" italics="true" pageId="4" pageNumber="139">P</emphasis>
&lt;0.01). The mean SVL of male 
<taxonomicName id="DDE14D78FFFAFFF1FC49F9C7E13AF919" box="[989,1078,1649,1672]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FC49F9C7E13AF919" box="[989,1078,1649,1672]" italics="true" pageId="4" pageNumber="139">H. caete</emphasis>
</taxonomicName>
is smaller than that of male 
<taxonomicName id="DDE14D78FFFAFFF1FAE4F9C7E0CAF918" box="[1392,1478,1649,1673]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FAE4F9C7E0CAF918" box="[1392,1478,1649,1673]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
(
<emphasis id="2895EAE9FFFAFFF1FA42F9C7E0E6F919" box="[1494,1514,1649,1672]" italics="true" pageId="4" pageNumber="139">Q</emphasis>
¼ 8.84, 
<emphasis id="2895EAE9FFFAFFF1FC36F939E6BDF937" box="[930,945,1679,1702]" italics="true" pageId="4" pageNumber="139">P</emphasis>
&lt;0.01), although there is overlap in their body size (male SVL 34.4 ± 
<quantity id="DD199B1EFFFAFFF1FBB7F91AE177F955" box="[1059,1147,1708,1732]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="139" unit="mm" value="0.9">0.9 mm</quantity>
, 
<quantity id="DD199B1EFFFAFFF1FB13F91AE025F955" box="[1159,1321,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="3.475" metricValueMax="3.67" metricValueMin="3.28" pageId="4" pageNumber="139" unit="mm" value="34.75" valueMax="36.7" valueMin="32.8">32.8–36.7 mm</quantity>
; 
<emphasis id="2895EAE9FFFAFFF1FAA1F91BE04FF955" box="[1333,1347,1709,1732]" italics="true" pageId="4" pageNumber="139">n</emphasis>
¼ 19; 
<bibRefCitation id="7E704B0AFFFAFFF1FA00F91AE112F973" author="Canedo, C. &amp; J. P. Pombal, Jr." pageId="4" pageNumber="139" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
). Furthermore, 
<taxonomicName id="DDE14D78FFFAFFF1FB4FF97DE036F973" box="[1243,1338,1739,1762]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FB4FF97DE036F973" box="[1243,1338,1739,1762]" italics="true" pageId="4" pageNumber="139">H. caete</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="DDE14D78FFFAFFF1FC62F95FE158F891" box="[1014,1108,1768,1792]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FC62F95FE158F891" box="[1014,1108,1768,1792]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
by the distribution of its nuptial tubercles, which are arranged in an elliptical area at the base of the thumb compared to them being distributed all over the dorsum of the thumb in 
<taxonomicName id="DDE14D78FFFAFFF1FB03F8F4E682F8E9" authority="(Canedo and Pombal 2007)" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FB03F8F4E1FCF8C8" box="[1175,1264,1857,1881]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
(
<bibRefCitation id="7E704B0AFFFAFFF1FA96F8F7E68AF8E9" author="Canedo, C. &amp; J. P. Pombal, Jr." pageId="4" pageNumber="139" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="1A5E36FBFFFAFFF3FCF8F8C8E495FBA6" blockId="4.[844,1514,962,2001]" lastBlockId="6.[142,813,730,1611]" lastPageId="6" lastPageNumber="141" pageId="4" pageNumber="139">
In addition to differences in size and nuptial tubercle distribution, 
<taxonomicName id="DDE14D78FFFAFFF1FC48F82AE13BF822" box="[988,1079,1948,1971]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFAFFF1FC48F82AE13BF822" box="[988,1079,1948,1971]" italics="true" pageId="4" pageNumber="139">H. caete</emphasis>
</taxonomicName>
is also distinguished from 
<taxonomicName id="DDE14D78FFFAFFF1FAF3F82AE0B3F822" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1383,1471,1947,1971]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFFAFFF1FAF3F82AE0B3F822" box="[1383,1471,1947,1971]" italics="true" pageId="4" pageNumber="139">H. fredi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFFAFFF1FA5BF82AE6B8F840" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFFAFFF1FA5BF82AE6B8F840" italics="true" pageId="4" pageNumber="139">H. phŋllodes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFFAFFF1FC61F80CE17CF840" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1013,1136,1977,2001]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="139" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFFAFFF1FC61F80CE17CF840" box="[1013,1136,1977,2001]" italics="true" pageId="4" pageNumber="139">H. pipilans</emphasis>
</taxonomicName>
by its advertisement call (see call comparisons in 
<tableCitation id="57630340FFF8FFF3FEDEFD6CE4ABFD62" box="[330,423,730,755]" captionStart="TABLE" captionStartId="6.[169,182,1654,1673]" captionTargetPageId="6" captionText="TABLE 4.—Advertisement calls of Hŋlodes caete sp. nov. and of the other three Hŋlodes species with males having nuptial tubercles on the thumb: Hŋlodes fredi (Canedo and Pombal 2007), H. phŋllodes (Heyer and Cocroft 1986), and H. pipilans (Canedo and Pombal 2007). Values are reported as ¯X ± 1 SD. (—) in cells indicates unavailable data in cited references." httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DF9C0E761F929" pageId="6" pageNumber="141" tableUuid="4E9E6673FFF8FFF3FF3DF9C0E761F929">Table 4</tableCitation>
). The call notes of 
<taxonomicName id="DDE14D78FFF8FFF3FD0AFD6DE7F7FD63" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[670,763,730,754]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FD0AFD6DE7F7FD63" box="[670,763,730,754]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
are emitted at a lower rate (6.06–9.5 notes/s) with a longer interval (0.06– 
<geoCoordinate id="7FD5503CFFF8FFF3FEBAFCA0E47CFCBC" box="[302,368,790,814]" degrees="0.16" direction="south" orientation="latitude" pageId="6" pageNumber="141" precision="555" value="-0.16">0.16 s</geoCoordinate>
; 
<bibRefCitation id="7E704B0AFFF8FFF3FEEAFCA0E7AEFCBF" author="Canedo, C. &amp; J. P. Pombal, Jr." box="[382,674,790,814]" pageId="6" pageNumber="141" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
). The call of 
<taxonomicName id="DDE14D78FFF8FFF3FF1AFC82E415FCDA" box="[142,281,819,843]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FF1AFC82E415FCDA" box="[142,281,819,843]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
emits a lower number of notes per call (12–20) at a lower rate (8–11 notes/s) and with a longer duration (0.05– 
<geoCoordinate id="7FD5503CFFF8FFF3FF46FCD8E410FC17" box="[210,284,878,902]" degrees="0.06" direction="south" orientation="latitude" pageId="6" pageNumber="141" precision="555" value="-0.06">0.06 s</geoCoordinate>
; 
<bibRefCitation id="7E704B0AFFF8FFF3FEA4FCD9E754FC17" author="Heyer, W. R. &amp; R. B. Cocroft" box="[304,600,878,903]" pageId="6" pageNumber="141" pagination="100 - 109" refId="ref11139" refString="Heyer, W. R., and R. B. Cocroft. 1986. Descriptions of two new species of Hnlodes from the Atlantic Forests of Brazil (Amphibia: Leptodactylidae). Proceedings of Biological Society of Washington 99: 100 - 109." type="journal article" year="1986">Heyer and Cocroft 1986</bibRefCitation>
). The calls of 
<taxonomicName id="DDE14D78FFF8FFF3FC85FCD9E5EAFC35" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FC85FCD9E5EAFC35" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
are shorter (0.06– 
<geoCoordinate id="7FD5503CFFF8FFF3FE53FC3AE70CFC35" box="[455,512,908,932]" degrees="0.1" direction="south" orientation="latitude" pageId="6" pageNumber="141" precision="5555" value="-0.1">0.1 s</geoCoordinate>
), with fewer notes being emitted per call (2) and at a higher rate (20.58–32.79 notes/s; 
<bibRefCitation id="7E704B0AFFF8FFF3FF1AFC71E4CEFC4E" author="Canedo, C. &amp; J. P. Pombal, Jr." box="[142,450,967,991]" pageId="6" pageNumber="141" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
). Finally, 
<taxonomicName id="DDE14D78FFF8FFF3FDAEFC71E795FC4F" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[570,665,967,990]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FDAEFC71E795FC4F" box="[570,665,967,990]" italics="true" pageId="6" pageNumber="141">H. caete</emphasis>
</taxonomicName>
can also be differentiated from 
<taxonomicName id="DDE14D78FFF8FFF3FEFCFC53E4B1FC6D" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[360,445,996,1020]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FEFCFC53E4B1FC6D" box="[360,445,996,1020]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FE5FFC53E758FC6D" box="[459,596,996,1020]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FE5FFC53E758FC6D" box="[459,596,996,1020]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFF8FFF3FD06FC53E607FC6D" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[658,779,996,1020]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FD06FC53E607FC6D" box="[658,779,996,1020]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
by molecular evidence which corroborates our phenotypical analyses (see 
<tableCitation id="57630340FFF8FFF3FEB2FBA9E487FBA6" box="[294,395,1055,1079]" captionStart="TABLE" captionStartId="2.[169,182,188,207]" captionTargetPageId="2" captionText="TABLE 1.—Specimens included in genetic analyses to ascertain species identity within Hŋlodes and the phylogenetic relationships of clades within" httpUri="http://table.plazi.org/id/4E9E6673FFFCFFF7FF3DFF0AE18AFF76" pageId="6" pageNumber="141" tableUuid="4E9E6673FFFCFFF7FF3DFF0AE18AFF76">Table S1</tableCitation>
).
</paragraph>
<caption id="4E9E6673FFFBFFF0FF1AF8FCE42EF8E4" ID-DOI="http://doi.org/10.5281/zenodo.7712277" ID-Zenodo-Dep="7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="5" pageNumber="140" startId="5.[142,156,1866,1885]" targetBox="[119,1488,193,1839]" targetPageId="5">
<paragraph id="1A5E36FBFFFBFFF0FF1AF8FCE42EF8E4" blockId="5.[118,1491,1865,1909]" pageId="5" pageNumber="140">
FIG. 3.— 
<taxonomicName id="DDE14D78FFFBFFF0FF49F8FFE491F8CD" authority=", CFBH" authorityName="CFBH" box="[221,413,1865,1885]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="140" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFFBFFF0FF49F8FFE45AF8CD" box="[221,342,1865,1884]" italics="true" pageId="5" pageNumber="140">Hŋlodes caete</emphasis>
, CFBH
</taxonomicName>
40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 
<quantity id="DD199B1EFFFBFFF0FF73F8D7E412F8E4" box="[231,286,1889,1909]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="140" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
<caption id="4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" ID-Table-UUID="4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" pageId="6" pageNumber="141" startId="6.[169,182,188,207]" targetBox="[142,813,266,617]" targetIsTable="true" targetPageId="6">
<paragraph id="1A5E36FBFFF8FFF3FF3DFF0AE7C7FF76" blockId="6.[142,813,184,232]" pageId="6" pageNumber="141">
TABLE 3.—Measurements (mm, reported as 
<emphasis id="2895EAE9FFF8FFF3FDBBFF0EE730FF5B" box="[559,572,184,202]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± 1 SD) of males (
<emphasis id="2895EAE9FFF8FFF3FD7DFF0BE7F8FF5E" box="[745,756,189,207]" italics="true" pageId="6" pageNumber="141">n</emphasis>
¼ 16) and females (
<emphasis id="2895EAE9FFF8FFF3FE93FF63E41EFF76" box="[263,274,213,231]" italics="true" pageId="6" pageNumber="141">n</emphasis>
¼ 5) of the type series of 
<taxonomicName id="DDE14D78FFF8FFF3FD90FF62E771FF76" authority="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad, 2017" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[516,637,212,231]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
<emphasis id="2895EAE9FFF8FFF3FD90FF62E771FF76" box="[516,637,212,231]" italics="true" pageId="6" pageNumber="141">Hŋlodes caete</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A65792FFF8FFF3FD10FF62E7C7FF76" box="[644,715,212,231]" pageId="6" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1A5E36FBFFF8FFF3FEC9FEBCE621FDF9" pageId="6" pageNumber="141">
<table id="68E1C45BFFF8000AFF1AFEBCE621FDF8" box="[142,813,266,617]" gridcols="5" gridrows="14" pageId="6" pageNumber="141">
<tr id="A4D134B9FFF8000AFF1AFEBCE621FE89" box="[142,813,266,280]" gridrow="0" pageId="6" pageNumber="141" rowspan-0="1">
<th id="E7005DC5FFF8000AFF7FFEBCE4FAFE89" box="[235,502,266,280]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="141">Males</th>
<th id="E7005DC5FFF8000AFDB6FEBCE621FE89" box="[546,813,266,280]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="6" pageNumber="141">Females</th>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE9EE621FEAB" box="[142,813,296,314]" gridrow="1" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE9EE5B3FEAB" box="[142,191,296,314]" gridcol="0" gridrow="1" pageId="6" pageNumber="141">Trait A</th>
<td id="E7005DC5FFF8000AFF7FFE9EE46CFEAB" box="[235,352,296,314]" gridcol="1" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FE9EFE9EE418FEA7" box="[266,276,296,310]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFE18FE9EE4FAFEAB" box="[396,502,296,314]" gridcol="2" gridrow="1" pageId="6" pageNumber="141">Range</td>
<td id="E7005DC5FFF8000AFDB6FE9EE79AFEAB" box="[546,662,296,314]" gridcol="3" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FDD4FE9EE746FEA7" box="[576,586,296,310]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFD56FE9EE621FEAB" box="[706,813,296,314]" gridcol="4" gridrow="1" pageId="6" pageNumber="141">Range</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFEF8E621FEF0" box="[142,813,334,353]" gridrow="2" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFEF8E5B3FEF0" box="[142,191,334,353]" gridcol="0" gridrow="2" pageId="6" pageNumber="141">SVL</th>
<td id="E7005DC5FFF8000AFF7FFEF8E46CFEF0" box="[235,352,334,353]" gridcol="1" gridrow="2" pageId="6" pageNumber="141">32.3 ± 0.88</td>
<td id="E7005DC5FFF8000AFE18FEF8E4FAFEF0" box="[396,502,334,353]" gridcol="2" gridrow="2" pageId="6" pageNumber="141">31.1–34.0</td>
<td id="E7005DC5FFF8000AFDB6FEF8E79AFEF0" box="[546,662,334,353]" gridcol="3" gridrow="2" pageId="6" pageNumber="141">35.24 ± 1.75</td>
<td id="E7005DC5FFF8000AFD56FEF8E621FEF0" box="[706,813,334,353]" gridcol="4" gridrow="2" pageId="6" pageNumber="141">33.2–38.3</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFED0E621FEE8" box="[142,813,358,377]" gridrow="3" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFED0E5B3FEE8" box="[142,191,358,377]" gridcol="0" gridrow="3" pageId="6" pageNumber="141">HL</th>
<td id="E7005DC5FFF8000AFF7FFED0E46CFEE8" box="[235,352,358,377]" gridcol="1" gridrow="3" pageId="6" pageNumber="141">12.48 ± 0.31</td>
<td id="E7005DC5FFF8000AFE18FED0E4FAFEE8" box="[396,502,358,377]" gridcol="2" gridrow="3" pageId="6" pageNumber="141">12.06–13.02</td>
<td id="E7005DC5FFF8000AFDB6FED0E79AFEE8" box="[546,662,358,377]" gridcol="3" gridrow="3" pageId="6" pageNumber="141">12.82 ± 0.72</td>
<td id="E7005DC5FFF8000AFD56FED0E621FEE8" box="[706,813,358,377]" gridcol="4" gridrow="3" pageId="6" pageNumber="141">11.82–13.8</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFEC8E621FE00" box="[142,813,382,401]" gridrow="4" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFEC8E5B3FE00" box="[142,191,382,401]" gridcol="0" gridrow="4" pageId="6" pageNumber="141">HW</th>
<td id="E7005DC5FFF8000AFF7FFEC8E46CFE00" box="[235,352,382,401]" gridcol="1" gridrow="4" pageId="6" pageNumber="141">10.63 ± 0.26</td>
<td id="E7005DC5FFF8000AFE18FEC8E4FAFE00" box="[396,502,382,401]" gridcol="2" gridrow="4" pageId="6" pageNumber="141">10.14–11.01</td>
<td id="E7005DC5FFF8000AFDB6FEC8E79AFE00" box="[546,662,382,401]" gridcol="3" gridrow="4" pageId="6" pageNumber="141">11.13 ± 0.34</td>
<td id="E7005DC5FFF8000AFD56FEC8E621FE00" box="[706,813,382,401]" gridcol="4" gridrow="4" pageId="6" pageNumber="141">10.74–11.64</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE23E621FE38" box="[142,813,405,425]" gridrow="5" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE23E5B3FE38" box="[142,191,405,425]" gridcol="0" gridrow="5" pageId="6" pageNumber="141">ED</th>
<td id="E7005DC5FFF8000AFF7FFE23E46CFE38" box="[235,352,405,425]" gridcol="1" gridrow="5" pageId="6" pageNumber="141">5.11 ± 0.25</td>
<td id="E7005DC5FFF8000AFE18FE23E4FAFE38" box="[396,502,405,425]" gridcol="2" gridrow="5" pageId="6" pageNumber="141">4.62–5.52</td>
<td id="E7005DC5FFF8000AFDB6FE23E79AFE38" box="[546,662,405,425]" gridcol="3" gridrow="5" pageId="6" pageNumber="141">5.23 ± 0.23</td>
<td id="E7005DC5FFF8000AFD56FE23E621FE38" box="[706,813,405,425]" gridcol="4" gridrow="5" pageId="6" pageNumber="141">4.92–5.53</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE18E621FE50" box="[142,813,430,449]" gridrow="6" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE18E5B3FE50" box="[142,191,430,449]" gridcol="0" gridrow="6" pageId="6" pageNumber="141">TD</th>
<td id="E7005DC5FFF8000AFF7FFE18E46CFE50" box="[235,352,430,449]" gridcol="1" gridrow="6" pageId="6" pageNumber="141">2.47 ± 0.19</td>
<td id="E7005DC5FFF8000AFE18FE18E4FAFE50" box="[396,502,430,449]" gridcol="2" gridrow="6" pageId="6" pageNumber="141">2.04–2.78</td>
<td id="E7005DC5FFF8000AFDB6FE18E79AFE50" box="[546,662,430,449]" gridcol="3" gridrow="6" pageId="6" pageNumber="141">2.63 ± 0.18</td>
<td id="E7005DC5FFF8000AFD56FE18E621FE50" box="[706,813,430,449]" gridcol="4" gridrow="6" pageId="6" pageNumber="141">2.33–2.91</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE73E621FE48" box="[142,813,453,473]" gridrow="7" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE73E5B3FE48" box="[142,191,453,473]" gridcol="0" gridrow="7" pageId="6" pageNumber="141">END</th>
<td id="E7005DC5FFF8000AFF7FFE73E46CFE48" box="[235,352,453,473]" gridcol="1" gridrow="7" pageId="6" pageNumber="141">2.42 ± 0.10</td>
<td id="E7005DC5FFF8000AFE18FE73E4FAFE48" box="[396,502,453,473]" gridcol="2" gridrow="7" pageId="6" pageNumber="141">2.28–2.63</td>
<td id="E7005DC5FFF8000AFDB6FE73E79AFE48" box="[546,662,453,473]" gridcol="3" gridrow="7" pageId="6" pageNumber="141">2.48 ± 0.15</td>
<td id="E7005DC5FFF8000AFD56FE73E621FE48" box="[706,813,453,473]" gridcol="4" gridrow="7" pageId="6" pageNumber="141">2.31–2.78</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE6BE621FE60" box="[142,813,477,497]" gridrow="8" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE6BE5B3FE60" box="[142,191,477,497]" gridcol="0" gridrow="8" pageId="6" pageNumber="141">IOD</th>
<td id="E7005DC5FFF8000AFF7FFE6BE46CFE60" box="[235,352,477,497]" gridcol="1" gridrow="8" pageId="6" pageNumber="141">3.34 ± 0.14</td>
<td id="E7005DC5FFF8000AFE18FE6BE4FAFE60" box="[396,502,477,497]" gridcol="2" gridrow="8" pageId="6" pageNumber="141">3.12–3.58</td>
<td id="E7005DC5FFF8000AFDB6FE6BE79AFE60" box="[546,662,477,497]" gridcol="3" gridrow="8" pageId="6" pageNumber="141">3.32 ± 0.12</td>
<td id="E7005DC5FFF8000AFD56FE6BE621FE60" box="[706,813,477,497]" gridcol="4" gridrow="8" pageId="6" pageNumber="141">3.1–3.46</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFE43E621FD98" box="[142,813,501,521]" gridrow="9" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFE43E5B3FD98" box="[142,191,501,521]" gridcol="0" gridrow="9" pageId="6" pageNumber="141">IND</th>
<td id="E7005DC5FFF8000AFF7FFE43E46CFD98" box="[235,352,501,521]" gridcol="1" gridrow="9" pageId="6" pageNumber="141">4.38 ± 0.18</td>
<td id="E7005DC5FFF8000AFE18FE43E4FAFD98" box="[396,502,501,521]" gridcol="2" gridrow="9" pageId="6" pageNumber="141">4.1–4.75</td>
<td id="E7005DC5FFF8000AFDB6FE43E79AFD98" box="[546,662,501,521]" gridcol="3" gridrow="9" pageId="6" pageNumber="141">4.51 ± 0.16</td>
<td id="E7005DC5FFF8000AFD56FE43E621FD98" box="[706,813,501,521]" gridcol="4" gridrow="9" pageId="6" pageNumber="141">4.28–4.71</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFDBBE621FDB0" box="[142,813,525,545]" gridrow="10" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFDBBE5B3FDB0" box="[142,191,525,545]" gridcol="0" gridrow="10" pageId="6" pageNumber="141">THL</th>
<td id="E7005DC5FFF8000AFF7FFDBBE46CFDB0" box="[235,352,525,545]" gridcol="1" gridrow="10" pageId="6" pageNumber="141">16.02 ± 0.63</td>
<td id="E7005DC5FFF8000AFE18FDBBE4FAFDB0" box="[396,502,525,545]" gridcol="2" gridrow="10" pageId="6" pageNumber="141">15.03–17.53</td>
<td id="E7005DC5FFF8000AFDB6FDBBE79AFDB0" box="[546,662,525,545]" gridcol="3" gridrow="10" pageId="6" pageNumber="141">16.62 ± 0.23</td>
<td id="E7005DC5FFF8000AFD56FDBBE621FDB0" box="[706,813,525,545]" gridcol="4" gridrow="10" pageId="6" pageNumber="141">16.45–17.8</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFD93E621FDA8" box="[142,813,549,569]" gridrow="11" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFD93E5B3FDA8" box="[142,191,549,569]" gridcol="0" gridrow="11" pageId="6" pageNumber="141">TBL</th>
<td id="E7005DC5FFF8000AFF7FFD93E46CFDA8" box="[235,352,549,569]" gridcol="1" gridrow="11" pageId="6" pageNumber="141">17.83 ± 0.53</td>
<td id="E7005DC5FFF8000AFE18FD93E4FAFDA8" box="[396,502,549,569]" gridcol="2" gridrow="11" pageId="6" pageNumber="141">16.52–18.73</td>
<td id="E7005DC5FFF8000AFDB6FD93E79AFDA8" box="[546,662,549,569]" gridcol="3" gridrow="11" pageId="6" pageNumber="141">19.18 ± 0.44</td>
<td id="E7005DC5FFF8000AFD56FD93E621FDA8" box="[706,813,549,569]" gridcol="4" gridrow="11" pageId="6" pageNumber="141">18.5–19.84</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFD8BE621FDC1" box="[142,813,573,592]" gridrow="12" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFD8BE5B3FDC1" box="[142,191,573,592]" gridcol="0" gridrow="12" pageId="6" pageNumber="141">TAL</th>
<td id="E7005DC5FFF8000AFF7FFD8BE46CFDC1" box="[235,352,573,592]" gridcol="1" gridrow="12" pageId="6" pageNumber="141">9.14 ± 0.26</td>
<td id="E7005DC5FFF8000AFE18FD8BE4FAFDC1" box="[396,502,573,592]" gridcol="2" gridrow="12" pageId="6" pageNumber="141">8.75–9.65</td>
<td id="E7005DC5FFF8000AFDB6FD8BE79AFDC1" box="[546,662,573,592]" gridcol="3" gridrow="12" pageId="6" pageNumber="141">9.61 ± 0.21</td>
<td id="E7005DC5FFF8000AFD56FD8BE621FDC1" box="[706,813,573,592]" gridcol="4" gridrow="12" pageId="6" pageNumber="141">9.29–9.82</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AFDE3E621FDF8" box="[142,813,597,617]" gridrow="13" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AFDE3E5B3FDF8" box="[142,191,597,617]" gridcol="0" gridrow="13" pageId="6" pageNumber="141">FL</th>
<td id="E7005DC5FFF8000AFF7FFDE3E46CFDF8" box="[235,352,597,617]" gridcol="1" gridrow="13" pageId="6" pageNumber="141">17.01 ± 0.70</td>
<td id="E7005DC5FFF8000AFE18FDE3E4FAFDF8" box="[396,502,597,617]" gridcol="2" gridrow="13" pageId="6" pageNumber="141">15.72–18.22</td>
<td id="E7005DC5FFF8000AFDB6FDE3E79AFDF8" box="[546,662,597,617]" gridcol="3" gridrow="13" pageId="6" pageNumber="141">18.34 ± 0.84</td>
<td id="E7005DC5FFF8000AFD56FDE3E621FDF8" box="[706,813,597,617]" gridcol="4" gridrow="13" pageId="6" pageNumber="141">17.51–19.53</td>
</tr>
</table>
</paragraph>
<paragraph id="1A5E36FBFFF8FFF3FF3DFDC3E7F5FD3D" blockId="6.[142,812,629,684]" pageId="6" pageNumber="141">
<tableNote id="E7073775FFF8FFF3FF3DFDC3E7F5FD3D" pageId="6" pageNumber="141" targetBox="[142,813,266,617]" targetPageId="6">
<superScript id="ED949BB3FFF8FFF3FF3DFDC3E5BDFDEE" attach="right" box="[169,177,629,639]" fontSize="4" pageId="6" pageNumber="141">A</superScript>
SVL ¼ snout–vent length; HL ¼ head length; HW ¼ head width; ED ¼ eye diameter; TD ¼ tympanum diameter; END ¼ eye–nostril distance; IOD ¼ interorbital distance; IND ¼ internostril distance; THL ¼ thigh length; TBL ¼ tibia length; TAL ¼ tarsus length; FL ¼ foot length.
</tableNote>
</paragraph>
<caption id="4E9E6673FFF8FFF3FCF0FC99E6D1FC33" ID-DOI="http://doi.org/10.5281/zenodo.7712279" ID-Zenodo-Dep="7712279" httpUri="https://zenodo.org/record/7712279/files/figure.png" pageId="6" pageNumber="141" startId="6.[868,900,815,834]" targetBox="[844,1515,193,790]" targetPageId="6">
<paragraph id="1A5E36FBFFF8FFF3FCF0FC99E6D1FC33" blockId="6.[844,1515,815,930]" pageId="6" pageNumber="141">
Fig. 4.—Distribution of nuptial tubercles on the right thumb of an adult male 
<taxonomicName id="DDE14D78FFF8FFF3FCEAFCF1E14EFCCB" authority=", CFBH" authorityName="CFBH" box="[894,1090,839,859]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FCEAFCF1E6F5FCCB" box="[894,1017,839,858]" italics="true" pageId="6" pageNumber="141">Hŋlodes caete</emphasis>
, CFBH
</taxonomicName>
40524 (holotype). (A) Drawing in lateral view. (B) Picture in dorsal view, with dashed line indicating the elliptical distribution of tubercles. Scale bars ¼ 1 mm. A color version of this figure is available online.
</paragraph>
</caption>
<paragraph id="1A5E36FBFFF8FFF3FF3DFB8BE402FAED" blockId="6.[142,813,730,1611]" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FF3DFB8BE479FBC4" bold="true" box="[169,373,1085,1109]" pageId="6" pageNumber="141">Genetic analysis.</emphasis>
—Our uncorrected 
<emphasis id="2895EAE9FFF8FFF3FDDBFB88E751FBC4" box="[591,605,1086,1109]" italics="true" pageId="6" pageNumber="141">p</emphasis>
distance estimates support 
<taxonomicName id="DDE14D78FFF8FFF3FF7EFBEDE44EFBE3" box="[234,322,1115,1138]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FF7EFBEDE44EFBE3" box="[234,322,1115,1138]" italics="true" pageId="6" pageNumber="141">H. caete</emphasis>
</taxonomicName>
being a new species (
<tableCitation id="57630340FFF8FFF3FDBAFBECE79DFBE3" box="[558,657,1114,1138]" captionStart="TABLE" captionStartId="2.[169,182,188,207]" captionTargetPageId="2" captionText="TABLE 1.—Specimens included in genetic analyses to ascertain species identity within Hŋlodes and the phylogenetic relationships of clades within" httpUri="http://table.plazi.org/id/4E9E6673FFFCFFF7FF3DFF0AE18AFF76" pageId="6" pageNumber="141" tableUuid="4E9E6673FFFCFFF7FF3DFF0AE18AFF76">Table S1</tableCitation>
). For the new species, and particularly for the others having males with nuptial tubercles, intraspecific distances range from 0 (
<taxonomicName id="DDE14D78FFF8FFF3FC85FB20E5EAFB5A" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FC85FB20E5EAFB5A" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
) to 0.3% ± 0.005 (
<taxonomicName id="DDE14D78FFF8FFF3FE5FFB02E72BFB5A" box="[459,551,1204,1227]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FE5FFB02E72BFB5A" box="[459,551,1204,1227]" italics="true" pageId="6" pageNumber="141">H. caete</emphasis>
</taxonomicName>
) for the complete 16S gene and from 0 (
<taxonomicName id="DDE14D78FFF8FFF3FEF3FB67E4EAFB79" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[359,486,1232,1256]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FEF3FB67E4EAFB79" box="[359,486,1232,1256]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
) to 0.5% (
<taxonomicName id="DDE14D78FFF8FFF3FDFCFB67E7FBFB79" box="[616,759,1232,1256]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FDFCFB67E7FBFB79" box="[616,759,1232,1256]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
) for partial 16S sequences; interspecific distances range from 9.5% (
<taxonomicName id="DDE14D78FFF8FFF3FF48FABAE43BFAB2" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[220,311,1291,1315]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FF48FABAE43BFAB2" box="[220,311,1291,1315]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="DDE14D78FFF8FFF3FEF9FABAE4E1FAB2" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[365,493,1291,1315]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FEF9FABAE4E1FAB2" box="[365,493,1291,1315]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
) to 11.3% (
<taxonomicName id="DDE14D78FFF8FFF3FDEBFABAE7D7FAB2" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[639,731,1291,1315]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FDEBFABAE7D7FAB2" box="[639,731,1291,1315]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="DDE14D78FFF8FFF3FC85FABAE5FAFAD0" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FC85FABAE5FAFAD0" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
) for complete 16S sequences and from 6% (
<taxonomicName id="DDE14D78FFF8FFF3FC85FA9CE5FAFACF" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FC85FA9CE5FAFACF" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="DDE14D78FFF8FFF3FEB7FAF1E492FACF" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[291,414,1350,1374]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FEB7FAF1E492FACF" box="[291,414,1350,1374]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
) to 7.2 (
<taxonomicName id="DDE14D78FFF8FFF3FE6AFAF1E759FACF" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[510,597,1350,1374]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FE6AFAF1E759FACF" box="[510,597,1350,1374]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="DDE14D78FFF8FFF3FD15FAF1E7F0FACF" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[641,764,1350,1374]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FD15FAF1E7F0FACF" box="[641,764,1350,1374]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
) for partial 16S.
</paragraph>
<paragraph id="1A5E36FBFFF8FFF3FF3AFA34E010FB4D" blockId="6.[142,813,730,1611]" lastBlockId="6.[844,1514,975,1611]" pageId="6" pageNumber="141">
Our reconstructed tree from a BI analysis (of the complete mitochondrial gene 16S) distinguishes all included species as independent evolutionary lineages, including 
<taxonomicName id="DDE14D78FFF8FFF3FF1AFA6CE42BFA63" box="[142,295,1498,1522]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FF1AFA6CE42BFA63" box="[142,295,1498,1522]" italics="true" pageId="6" pageNumber="141">Hŋlodes caete</emphasis>
</taxonomicName>
(
<figureCitation id="82DA2A7EFFF8FFF3FEACFA6CE471FA63" box="[312,381,1498,1522]" captionStart="FIG" captionStartId="7.[142,156,786,805]" captionTargetBox="[118,1490,192,760]" captionTargetId="figure-702@7.[118,1490,191,760]" captionTargetPageId="7" captionText="FIG. 5.—Phylogenetic tree reconstructed for representative hylodid anurans from a Bayesian inference analysis of the complete 16S mitochondrial gene. Values adjacent to each node are posterior probabilities." figureDoi="http://doi.org/10.5281/zenodo.7712283" httpUri="https://zenodo.org/record/7712283/files/figure.png" pageId="6" pageNumber="141">Fig. 5</figureCitation>
). Monophyly of the genus 
<taxonomicName id="DDE14D78FFF8FFF3FD3AFA6CE60BFA63" authorityName="Fitzinger" authorityYear="1826" box="[686,775,1498,1522]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="genus">
<emphasis id="2895EAE9FFF8FFF3FD3AFA6CE60BFA63" box="[686,775,1498,1522]" italics="true" pageId="6" pageNumber="141">Hŋlodes</emphasis>
</taxonomicName>
, as well as monophyly of the 
<taxonomicName id="DDE14D78FFF8FFF3FE27FA4FE77CF981" baseAuthorityName="Baumann" baseAuthorityYear="1912" box="[435,624,1528,1552]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="lateristrigatus">
<emphasis id="2895EAE9FFF8FFF3FE27FA4FE77CF981" box="[435,624,1528,1552]" italics="true" pageId="6" pageNumber="141">H. lateristrigatus</emphasis>
</taxonomicName>
group, are wellsupported. Within the 
<taxonomicName id="DDE14D78FFF8FFF3FE04F9A0E740F9BC" baseAuthorityName="Baumann" baseAuthorityYear="1912" box="[400,588,1557,1581]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="lateristrigatus">
<emphasis id="2895EAE9FFF8FFF3FE04F9A0E740F9BC" box="[400,588,1557,1581]" italics="true" pageId="6" pageNumber="141">H. lateristrigatus</emphasis>
</taxonomicName>
group (represented in our phylogenetic study by 
<taxonomicName id="DDE14D78FFF8FFF3FE47F982E754F9DA" box="[467,600,1587,1611]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="2895EAE9FFF8FFF3FE47F982E754F9DA" box="[467,600,1587,1611]" italics="true" pageId="6" pageNumber="141">H. amnicola</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FDF3F982E7CCF9DA" box="[615,704,1588,1611]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FDF3F982E7CCF9DA" box="[615,704,1588,1611]" italics="true" pageId="6" pageNumber="141">H. caete</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FD5BF982E62AF9DA" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[719,806,1587,1611]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FD5BF982E62AF9DA" box="[719,806,1587,1611]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FCD8FC66E695FC76" box="[844,921,976,999]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="japi">
<emphasis id="2895EAE9FFF8FFF3FCD8FC66E695FC76" box="[844,921,976,999]" italics="true" pageId="6" pageNumber="141">H. japi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FC3DFC66E138FC76" box="[937,1076,975,999]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FC3DFC66E138FC76" box="[937,1076,975,999]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FBD0FC66E1CCFC76" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1092,1216,975,999]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FBD0FC66E1CCFC76" box="[1092,1216,975,999]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFF8FFF3FA95FC66E07BFC76" box="[1281,1399,976,999]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="2895EAE9FFF8FFF3FA95FC66E07BFC76" box="[1281,1399,976,999]" italics="true" pageId="6" pageNumber="141">H. ornatus</emphasis>
</taxonomicName>
), the new species and all other 
<taxonomicName id="DDE14D78FFF8FFF3FBCDFC58E1FCFB94" box="[1113,1264,1006,1030]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="2895EAE9FFF8FFF3FBCDFC58E1BEFB97" box="[1113,1202,1006,1030]" italics="true" pageId="6" pageNumber="141">Hŋlodes</emphasis>
spp.
</taxonomicName>
that possess nuptial tubercles form a well-supported clade. 
<taxonomicName id="DDE14D78FFF8FFF3FA92FBBAE0C3FBB5" box="[1286,1487,1036,1060]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FA92FBBAE0C3FBB5" box="[1286,1487,1036,1060]" italics="true" pageId="6" pageNumber="141">Hŋlodes phŋllodes</emphasis>
</taxonomicName>
is recovered as the sister taxon of a clade that includes 
<taxonomicName id="DDE14D78FFF8FFF3FA05FB9DE68CFBF0" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FA05FB9DE68CFBF0" italics="true" pageId="6" pageNumber="141">Hŋlodes fredi</emphasis>
</taxonomicName>
and 
<taxonomicName id="DDE14D78FFF8FFF3FC59FBFCE15DFBF0" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[973,1105,1097,1121]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FC59FBFCE15DFBF0" box="[973,1105,1097,1121]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
(which were recovered as sister species). The new species is found to be the sister taxon of a clade composed of 
<taxonomicName id="DDE14D78FFF8FFF3FBACFB31E183FB0F" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1080,1167,1158,1182]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FBACFB31E183FB0F" box="[1080,1167,1158,1182]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FB09FB31E024FB0F" box="[1181,1320,1158,1182]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FB09FB31E024FB0F" box="[1181,1320,1158,1182]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFF8FFF3FAFCFB31E0EFFB0F" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1384,1507,1158,1182]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FAFCFB31E0EFFB0F" box="[1384,1507,1158,1182]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
. Posterior probabilities, particularly within the clade of 
<taxonomicName id="DDE14D78FFF8FFF3FCD8FB72E6A9FB4D" authorityName="Fitzinger" authorityYear="1826" box="[844,933,1220,1244]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="genus">
<emphasis id="2895EAE9FFF8FFF3FCD8FB72E6A9FB4D" box="[844,933,1220,1244]" italics="true" pageId="6" pageNumber="141">Hŋlodes</emphasis>
</taxonomicName>
with nuptial tubercles, are&gt;0.9.
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFF8FFFDFCF3FB54E7B6FB7B" lastPageId="8" lastPageNumber="143" pageId="6" pageNumber="141" type="description">
<paragraph id="1A5E36FBFFF8FFF2FCF3FB54E7ECFB87" blockId="6.[844,1514,975,1611]" lastBlockId="7.[118,788,873,2001]" lastPageId="7" lastPageNumber="142" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FCF3FB54E198FB6A" bold="true" box="[871,1172,1250,1275]" pageId="6" pageNumber="141">
Description of 
<typeStatus id="C55A8859FFF8FFF3FBB7FB55E183FB6A" box="[1059,1167,1251,1275]" pageId="6" pageNumber="141" type="holotype">holotype</typeStatus>
.
</emphasis>
—Body slender (
<figureCitation id="82DA2A7EFFF8FFF3FAC3FB55E090FB6A" box="[1367,1436,1251,1275]" captionStart="FIG" captionStartId="4.[166,180,898,917]" captionTargetBox="[142,1515,191,872]" captionTargetId="figure-795@4.[142,1516,191,872]" captionTargetPageId="4" captionText="FIG. 2.—Hŋlodes caete, CFBH 40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7712275" httpUri="https://zenodo.org/record/7712275/files/figure.png" pageId="6" pageNumber="141">Fig. 2</figureCitation>
); head longer than wide; snout nearly rounded in dorsal view and protruding in lateral view (
<figureCitation id="82DA2A7EFFF8FFF3FB01FA96E005FAA9" box="[1173,1289,1312,1336]" captionStart="FIG" captionStartId="5.[142,156,1866,1885]" captionTargetBox="[119,1488,193,1839]" captionTargetId="figure-0@5.[118,1491,191,1840]" captionTargetPageId="5" captionText="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="6" pageNumber="141">Fig. 3A,B</figureCitation>
); nostrils elliptical, slightly protruding, laterally directed; canthus rostralis distinct, slightly curved; loreal region concave; row of small, light-colored tubercles uniformly distributed on the edge and along the entire upper lip; tympanum visible, nearly rounded, diameter larger than half of eye diameter; supratympanic fold well developed, extending from posterior corner of eye to posterior edge of the shoulder, reaching the dorsal tympanic annulus; oblique lateral fold weak and continuous from eye, above supratympanic fold, to inguinal region; tubercles absent lateral to the oblique lateral fold; paired lateral vocal sacs, widely expanded externally; vocal slits present; tongue large, nearly ovoid, free for its distal 1/3; vomerine teeth in two small series between choanae, bearing three teeth each; choanae small and nearly round, reaching the palatine process of maxilla; maxillary teeth present.
</paragraph>
<caption id="4E9E6673FFF8FFF3FF3DF9C0E761F929" ID-Table-UUID="4E9E6673FFF8FFF3FF3DF9C0E761F929" httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DF9C0E761F929" pageId="6" pageNumber="141" startId="6.[169,182,1654,1673]" targetBox="[142,1513,1755,1986]" targetIsTable="true" targetPageId="6">
<paragraph id="1A5E36FBFFF8FFF3FF3DF9C0E761F929" blockId="6.[142,1514,1653,1720]" pageId="6" pageNumber="141">
TABLE 4.—Advertisement calls of 
<taxonomicName id="DDE14D78FFF8FFF3FE5BF9C3E74BF919" authority="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad, 2017" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[463,583,1653,1672]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete" status="sp. nov.">
<emphasis id="2895EAE9FFF8FFF3FE5BF9C3E74BF919" box="[463,583,1653,1672]" italics="true" pageId="6" pageNumber="141">Hŋlodes caete</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A65792FFF8FFF3FDD8F9C0E79CF918" box="[588,656,1654,1673]" pageId="6" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
and of the other three 
<taxonomicName id="DDE14D78FFF8FFF3FCCCF9C3E6EFF919" box="[856,995,1653,1672]" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FCCCF9C3E693F919" box="[856,927,1653,1672]" italics="true" pageId="6" pageNumber="141">Hŋlodes</emphasis>
species
</taxonomicName>
with males having nuptial tubercles on the thumb: 
<taxonomicName id="DDE14D78FFF8FFF3FA37F9C3E4A3F931" authority="(Canedo and Pombal 2007)" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FA37F9C3E5BBF931" italics="true" pageId="6" pageNumber="141">Hŋlodes fredi</emphasis>
(
<bibRefCitation id="7E704B0AFFF8FFF3FF57F93BE4A5F931" author="Canedo, C. &amp; J. P. Pombal, Jr." box="[195,425,1677,1696]" pageId="6" pageNumber="141" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="DDE14D78FFF8FFF3FE23F938E603F931" authority="(Heyer and Cocroft 1986)" baseAuthorityName="Heyer and Cocroft" baseAuthorityYear="1986" box="[439,783,1677,1696]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FE23F938E729F931" box="[439,549,1677,1696]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
(
<bibRefCitation id="7E704B0AFFF8FFF3FDA5F93BE605F931" author="Heyer, W. R. &amp; R. B. Cocroft" box="[561,777,1677,1696]" pageId="6" pageNumber="141" pagination="100 - 109" refId="ref11139" refString="Heyer, W. R., and R. B. Cocroft. 1986. Descriptions of two new species of Hnlodes from the Atlantic Forests of Brazil (Amphibia: Leptodactylidae). Proceedings of Biological Society of Washington 99: 100 - 109." type="journal article" year="1986">Heyer and Cocroft 1986</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="DDE14D78FFF8FFF3FCAAF938E19AF931" authority="(Canedo and Pombal 2007)" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[830,1174,1677,1696]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FCAAF938E692F931" box="[830,926,1677,1696]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
(
<bibRefCitation id="7E704B0AFFF8FFF3FC3DF93BE19CF931" author="Canedo, C. &amp; J. P. Pombal, Jr." box="[937,1168,1677,1696]" pageId="6" pageNumber="141" pagination="224 - 235" refId="ref10423" refString="Canedo, C., and J. P. Pombal, Jr. 2007. Two new species of torrent frog of the genus Hnlodes (Anura, Hylodidae) with nuptial thumb tubercles. Herpetologica 63: 224 - 235." type="journal article" year="2007">Canedo and Pombal 2007</bibRefCitation>
)
</taxonomicName>
. Values are reported as 
<emphasis id="2895EAE9FFF8FFF3FAF2F93FE07FF90A" box="[1382,1395,1673,1691]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± 1 SD. (—) in cells indicates unavailable data in cited references.
</paragraph>
</caption>
<paragraph id="1A5E36FBFFF8FFF3FE49F96AE0E9F853" pageId="6" pageNumber="141">
<table id="68E1C45BFFF8000AFF1AF96DE0E5F853" box="[142,1513,1755,1986]" gridcols="9" gridrows="9" pageId="6" pageNumber="141">
<tr id="A4D134B9FFF8000AFF1AF96DE0E5F97B" box="[142,1513,1755,1770]" gridrow="0" pageId="6" pageNumber="141" rowspan-0="1">
<th id="E7005DC5FFF8000AFE03F96DE7C6F97B" box="[407,714,1755,1770]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="141">
<taxonomicName id="DDE14D78FFF8FFF3FE49F96AE71FF97B" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[477,531,1756,1770]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF8FFF3FE49F96AE71FF97B" box="[477,531,1756,1770]" italics="true" pageId="6" pageNumber="141">H. caete</emphasis>
</taxonomicName>
(at 20.8–22.8°C)
</th>
<th id="E7005DC5FFF8000AFD7FF96DE6D3F97B" box="[747,991,1755,1770]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="6" pageNumber="141">
<taxonomicName id="DDE14D78FFF8FFF3FCBEF96AE652F978" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[810,862,1755,1770]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="fredi">
<emphasis id="2895EAE9FFF8FFF3FCBEF96AE652F978" box="[810,862,1755,1770]" italics="true" pageId="6" pageNumber="141">H. fredi</emphasis>
</taxonomicName>
(at 25°C)
</th>
<th id="E7005DC5FFF8000AFB96F96DE1B2F97B" box="[1026,1214,1755,1770]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="6" pageNumber="141">
<taxonomicName id="DDE14D78FFF8FFF3FB9BF96AE16FF978" baseAuthorityName="Heyer and Cocroft" baseAuthorityYear="1986" box="[1039,1123,1755,1770]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF8FFF3FB9BF96AE16FF978" box="[1039,1123,1755,1770]" italics="true" pageId="6" pageNumber="141">H. phŋllodes</emphasis>
</taxonomicName>
(at 20.4°C)
</th>
<th id="E7005DC5FFF8000AFB75F96DE0E5F97B" box="[1249,1513,1755,1770]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="6" pageNumber="141">
<taxonomicName id="DDE14D78FFF8FFF3FA8BF96AE066F978" baseAuthorityName="Canedo and Pombal" baseAuthorityYear="2007" box="[1311,1386,1755,1770]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="141" phylum="Chordata" rank="species" species="pipilans">
<emphasis id="2895EAE9FFF8FFF3FA8BF96AE066F978" box="[1311,1386,1755,1770]" italics="true" pageId="6" pageNumber="141">H. pipilans</emphasis>
</taxonomicName>
(at 22°C)
</th>
</tr>
<tr id="A4D134B9FFF8000AFF1AF94CE0E5F89A" box="[142,1513,1786,1803]" gridrow="1" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF94CE47BF89A" box="[142,375,1786,1803]" gridcol="0" gridrow="1" pageId="6" pageNumber="141">Call parameters</th>
<td id="E7005DC5FFF8000AFE03F94CE720F89A" box="[407,556,1786,1803]" gridcol="1" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FE52F94CE4DCF899" box="[454,464,1786,1800]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFDD8F94CE7C6F89A" box="[588,714,1786,1803]" gridcol="2" gridrow="1" pageId="6" pageNumber="141">Range</td>
<td id="E7005DC5FFF8000AFD7FF94CE653F89A" box="[747,863,1786,1803]" gridcol="3" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FC9DF94CE61FF899" box="[777,787,1786,1800]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFCEBF94CE6D3F89A" box="[895,991,1786,1803]" gridcol="4" gridrow="1" pageId="6" pageNumber="141">Range</td>
<td id="E7005DC5FFF8000AFB96F94CE135F89A" box="[1026,1081,1786,1803]" gridcol="5" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FB96F94CE100F899" box="[1026,1036,1786,1800]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFBCAF94CE1B2F89A" box="[1118,1214,1786,1803]" gridcol="6" gridrow="1" pageId="6" pageNumber="141">Range</td>
<td id="E7005DC5FFF8000AFB75F94CE06CF89A" box="[1249,1376,1786,1803]" gridcol="7" gridrow="1" pageId="6" pageNumber="141">
<emphasis id="2895EAE9FFF8FFF3FA91F94CE003F899" box="[1285,1295,1786,1800]" italics="true" pageId="6" pageNumber="141">¯X</emphasis>
± SD
</td>
<td id="E7005DC5FFF8000AFA14F94CE0E5F89A" box="[1408,1513,1786,1803]" gridcol="8" gridrow="1" pageId="6" pageNumber="141">Range</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF8A9E0E5F8A2" box="[142,1513,1823,1843]" gridrow="2" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF8A9E47BF8A2" box="[142,375,1823,1843]" gridcol="0" gridrow="2" pageId="6" pageNumber="141">Call duration (s)</th>
<td id="E7005DC5FFF8000AFE03F8A9E720F8A2" box="[407,556,1823,1843]" gridcol="1" gridrow="2" pageId="6" pageNumber="141">1.79 ± 0.38</td>
<td id="E7005DC5FFF8000AFDD8F8A9E7C6F8A2" box="[588,714,1823,1843]" gridcol="2" gridrow="2" pageId="6" pageNumber="141">1.22–2.72</td>
<td id="E7005DC5FFF8000AFD7FF8A9E653F8A2" box="[747,863,1823,1843]" gridcol="3" gridrow="2" pageId="6" pageNumber="141">1.35 ± 0.25</td>
<td id="E7005DC5FFF8000AFCEBF8A9E6D3F8A2" box="[895,991,1823,1843]" gridcol="4" gridrow="2" pageId="6" pageNumber="141">0.59–3.36</td>
<td id="E7005DC5FFF8000AFB96F8A9E135F8A2" box="[1026,1081,1823,1843]" gridcol="5" gridrow="2" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF8A9E1B2F8A2" box="[1118,1214,1823,1843]" gridcol="6" gridrow="2" pageId="6" pageNumber="141">1.05–2.1</td>
<td id="E7005DC5FFF8000AFB75F8A9E06CF8A2" box="[1249,1376,1823,1843]" gridcol="7" gridrow="2" pageId="6" pageNumber="141">0.08 ± 0.43</td>
<td id="E7005DC5FFF8000AFA14F8A9E0E5F8A2" box="[1408,1513,1823,1843]" gridcol="8" gridrow="2" pageId="6" pageNumber="141">0.06–0.10</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF881E0E5F8DA" box="[142,1513,1847,1867]" gridrow="3" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF881E47BF8DA" box="[142,375,1847,1867]" gridcol="0" gridrow="3" pageId="6" pageNumber="141">Intercall duration (s)</th>
<td id="E7005DC5FFF8000AFE03F881E720F8DA" box="[407,556,1847,1867]" gridcol="1" gridrow="3" pageId="6" pageNumber="141">3.54 ± 1.19</td>
<td id="E7005DC5FFF8000AFDD8F881E7C6F8DA" box="[588,714,1847,1867]" gridcol="2" gridrow="3" pageId="6" pageNumber="141">1.85–7.72</td>
<td id="E7005DC5FFF8000AFD7FF881E653F8DA" box="[747,863,1847,1867]" gridcol="3" gridrow="3" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFCEBF881E6D3F8DA" box="[895,991,1847,1867]" gridcol="4" gridrow="3" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFB96F881E135F8DA" box="[1026,1081,1847,1867]" gridcol="5" gridrow="3" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF881E1B2F8DA" box="[1118,1214,1847,1867]" gridcol="6" gridrow="3" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFB75F881E06CF8DA" box="[1249,1376,1847,1867]" gridcol="7" gridrow="3" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFA14F881E0E5F8DA" box="[1408,1513,1847,1867]" gridcol="8" gridrow="3" pageId="6" pageNumber="141">—</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF8F9E0E5F8F2" box="[142,1513,1871,1891]" gridrow="4" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF8F9E47BF8F2" box="[142,375,1871,1891]" gridcol="0" gridrow="4" pageId="6" pageNumber="141">Notes per call</th>
<td id="E7005DC5FFF8000AFE03F8F9E720F8F2" box="[407,556,1871,1891]" gridcol="1" gridrow="4" pageId="6" pageNumber="141">30 ± 6.67</td>
<td id="E7005DC5FFF8000AFDD8F8F9E7C6F8F2" box="[588,714,1871,1891]" gridcol="2" gridrow="4" pageId="6" pageNumber="141">20–45</td>
<td id="E7005DC5FFF8000AFD7FF8F9E653F8F2" box="[747,863,1871,1891]" gridcol="3" gridrow="4" pageId="6" pageNumber="141">10.49 ± 1.16</td>
<td id="E7005DC5FFF8000AFCEBF8F9E6D3F8F2" box="[895,991,1871,1891]" gridcol="4" gridrow="4" pageId="6" pageNumber="141">5–23</td>
<td id="E7005DC5FFF8000AFB96F8F9E135F8F2" box="[1026,1081,1871,1891]" gridcol="5" gridrow="4" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF8F9E1B2F8F2" box="[1118,1214,1871,1891]" gridcol="6" gridrow="4" pageId="6" pageNumber="141">12–20</td>
<td id="E7005DC5FFF8000AFB75F8F9E06CF8F2" box="[1249,1376,1871,1891]" gridcol="7" gridrow="4" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFA14F8F9E0E5F8F2" box="[1408,1513,1871,1891]" gridcol="8" gridrow="4" pageId="6" pageNumber="141">2</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF8D1E0E5F8EA" box="[142,1513,1895,1915]" gridrow="5" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF8D1E47BF8EA" box="[142,375,1895,1915]" gridcol="0" gridrow="5" pageId="6" pageNumber="141">Rate notes/s</th>
<td id="E7005DC5FFF8000AFE03F8D1E720F8EA" box="[407,556,1895,1915]" gridcol="1" gridrow="5" pageId="6" pageNumber="141">16.6 ± 1.02</td>
<td id="E7005DC5FFF8000AFDD8F8D1E7C6F8EA" box="[588,714,1895,1915]" gridcol="2" gridrow="5" pageId="6" pageNumber="141">14.8–18.3</td>
<td id="E7005DC5FFF8000AFD7FF8D1E653F8EA" box="[747,863,1895,1915]" gridcol="3" gridrow="5" pageId="6" pageNumber="141">7.85 ± 0.6</td>
<td id="E7005DC5FFF8000AFCEBF8D1E6D3F8EA" box="[895,991,1895,1915]" gridcol="4" gridrow="5" pageId="6" pageNumber="141">6.06–9.50</td>
<td id="E7005DC5FFF8000AFB96F8D1E135F8EA" box="[1026,1081,1895,1915]" gridcol="5" gridrow="5" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF8D1E1B2F8EA" box="[1118,1214,1895,1915]" gridcol="6" gridrow="5" pageId="6" pageNumber="141">8–11</td>
<td id="E7005DC5FFF8000AFB75F8D1E06CF8EA" box="[1249,1376,1895,1915]" gridcol="7" gridrow="5" pageId="6" pageNumber="141">26.64 ± 3.67</td>
<td id="E7005DC5FFF8000AFA14F8D1E0E5F8EA" box="[1408,1513,1895,1915]" gridcol="8" gridrow="5" pageId="6" pageNumber="141">20.58–32.79</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF8C9E0E5F803" box="[142,1513,1919,1938]" gridrow="6" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF8C9E47BF803" box="[142,375,1919,1938]" gridcol="0" gridrow="6" pageId="6" pageNumber="141">Note duration (s)</th>
<td id="E7005DC5FFF8000AFE03F8C9E720F803" box="[407,556,1919,1938]" gridcol="1" gridrow="6" pageId="6" pageNumber="141">0.016 ± 0.002</td>
<td id="E7005DC5FFF8000AFDD8F8C9E7C6F803" box="[588,714,1919,1938]" gridcol="2" gridrow="6" pageId="6" pageNumber="141">0.009–0.023</td>
<td id="E7005DC5FFF8000AFD7FF8C9E653F803" box="[747,863,1919,1938]" gridcol="3" gridrow="6" pageId="6" pageNumber="141">0.03 ± 0.01</td>
<td id="E7005DC5FFF8000AFCEBF8C9E6D3F803" box="[895,991,1919,1938]" gridcol="4" gridrow="6" pageId="6" pageNumber="141">0.02–0.04</td>
<td id="E7005DC5FFF8000AFB96F8C9E135F803" box="[1026,1081,1919,1938]" gridcol="5" gridrow="6" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF8C9E1B2F803" box="[1118,1214,1919,1938]" gridcol="6" gridrow="6" pageId="6" pageNumber="141">0.05–0.06</td>
<td id="E7005DC5FFF8000AFB75F8C9E06CF803" box="[1249,1376,1919,1938]" gridcol="7" gridrow="6" pageId="6" pageNumber="141">0.02 ± 0.01</td>
<td id="E7005DC5FFF8000AFA14F8C9E0E5F803" box="[1408,1513,1919,1938]" gridcol="8" gridrow="6" pageId="6" pageNumber="141">0.02–0.03</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF821E0E5F83A" box="[142,1513,1943,1963]" gridrow="7" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF821E47BF83A" box="[142,375,1943,1963]" gridcol="0" gridrow="7" pageId="6" pageNumber="141">Internote duration (s)</th>
<td id="E7005DC5FFF8000AFE03F821E720F83A" box="[407,556,1943,1963]" gridcol="1" gridrow="7" pageId="6" pageNumber="141">0.045 ± 0.005</td>
<td id="E7005DC5FFF8000AFDD8F821E7C6F83A" box="[588,714,1943,1963]" gridcol="2" gridrow="7" pageId="6" pageNumber="141">0.036–0.058</td>
<td id="E7005DC5FFF8000AFD7FF821E653F83A" box="[747,863,1943,1963]" gridcol="3" gridrow="7" pageId="6" pageNumber="141">0.1 ± 0.01</td>
<td id="E7005DC5FFF8000AFCEBF821E6D3F83A" box="[895,991,1943,1963]" gridcol="4" gridrow="7" pageId="6" pageNumber="141">0.06–0.16</td>
<td id="E7005DC5FFF8000AFB96F821E135F83A" box="[1026,1081,1943,1963]" gridcol="5" gridrow="7" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF821E1B2F83A" box="[1118,1214,1943,1963]" gridcol="6" gridrow="7" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFB75F821E06CF83A" box="[1249,1376,1943,1963]" gridcol="7" gridrow="7" pageId="6" pageNumber="141">0.03 ± 0.005</td>
<td id="E7005DC5FFF8000AFA14F821E0E5F83A" box="[1408,1513,1943,1963]" gridcol="8" gridrow="7" pageId="6" pageNumber="141">0.02–0.04</td>
</tr>
<tr id="A4D134B9FFF8000AFF1AF819E0E5F853" box="[142,1513,1967,1986]" gridrow="8" pageId="6" pageNumber="141">
<th id="E7005DC5FFF8000AFF1AF819E47BF853" box="[142,375,1967,1986]" gridcol="0" gridrow="8" pageId="6" pageNumber="141">Dominant frequency (Hz)</th>
<td id="E7005DC5FFF8000AFE03F819E720F853" box="[407,556,1967,1986]" gridcol="1" gridrow="8" pageId="6" pageNumber="141">4640.63 ± 28.47</td>
<td id="E7005DC5FFF8000AFDD8F819E7C6F853" box="[588,714,1967,1986]" gridcol="2" gridrow="8" pageId="6" pageNumber="141">3937.5–5062.5</td>
<td id="E7005DC5FFF8000AFD7FF819E653F853" box="[747,863,1967,1986]" gridcol="3" gridrow="8" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFCEBF819E6D3F853" box="[895,991,1967,1986]" gridcol="4" gridrow="8" pageId="6" pageNumber="141">3600–4500</td>
<td id="E7005DC5FFF8000AFB96F819E135F853" box="[1026,1081,1967,1986]" gridcol="5" gridrow="8" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFBCAF819E1B2F853" box="[1118,1214,1967,1986]" gridcol="6" gridrow="8" pageId="6" pageNumber="141">4100–5700</td>
<td id="E7005DC5FFF8000AFB75F819E06CF853" box="[1249,1376,1967,1986]" gridcol="7" gridrow="8" pageId="6" pageNumber="141">—</td>
<td id="E7005DC5FFF8000AFA14F819E0E5F853" box="[1408,1513,1967,1986]" gridcol="8" gridrow="8" pageId="6" pageNumber="141">4600–5400</td>
</tr>
</table>
</paragraph>
<caption id="4E9E6673FFF9FFF2FF1AFCA4E763FCAD" ID-DOI="http://doi.org/10.5281/zenodo.7712283" ID-Zenodo-Dep="7712283" httpUri="https://zenodo.org/record/7712283/files/figure.png" pageId="7" pageNumber="142" startId="7.[142,156,786,805]" targetBox="[118,1490,192,760]" targetPageId="7">
<paragraph id="1A5E36FBFFF9FFF2FF1AFCA4E763FCAD" blockId="7.[118,1489,785,829]" pageId="7" pageNumber="142">FIG. 5.—Phylogenetic tree reconstructed for representative hylodid anurans from a Bayesian inference analysis of the complete 16S mitochondrial gene. Values adjacent to each node are posterior probabilities.</paragraph>
</caption>
<paragraph id="1A5E36FBFFF9FFF2FF02FBABE773F8E6" blockId="7.[118,788,873,2001]" pageId="7" pageNumber="142">
Arms moderately slender and forearms moderately robust; subarticular tubercles single, round (
<figureCitation id="82DA2A7EFFF9FFF2FDFFFB8CE7CFFBC3" box="[619,707,1082,1106]" captionStart="FIG" captionStartId="5.[142,156,1866,1885]" captionTargetBox="[119,1488,193,1839]" captionTargetId="figure-0@5.[118,1491,191,1840]" captionTargetPageId="5" captionText="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="7" pageNumber="142">Fig. 3C</figureCitation>
); outer metacarpal tubercle large, round; inner metacarpal tubercle elliptical, smaller than outer metacarpal tubercle; supernumerary tubercles absent from hands; relative lengths of Fingers II &lt;I = IV &lt;III; corneous, pointed whitish nuptial tubercles in an elliptical area at the base of the dorsal surface of thumbs (
<figureCitation id="82DA2A7EFFF9FFF2FE94FB5BE444FA94" box="[256,328,1261,1285]" captionStart="Fig" captionStartId="6.[868,900,815,834]" captionTargetBox="[844,1515,193,790]" captionTargetId="figure-488@6.[844,1516,191,790]" captionTargetPageId="6" captionText="Fig. 4.—Distribution of nuptial tubercles on the right thumb of an adult male Hŋlodes caete, CFBH 40524 (holotype). (A) Drawing in lateral view. (B) Picture in dorsal view, with dashed line indicating the elliptical distribution of tubercles. Scale bars ¼ 1 mm. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712279" httpUri="https://zenodo.org/record/7712279/files/figure.png" pageId="7" pageNumber="142">Fig. 4</figureCitation>
); thumb slightly fringed on both sides; Finger II fringed on both sides; Finger III fringed laterally from proximal level of proximal subarticular tubercle to disc on both sides; Finger IV fringed along inner margin from distal edge of proximal subarticular tubercle to disc and along outer margin from medial level of proximal subarticular tubercle to disc; finger discs dilated; discs on fingertips nearly oval from ventral view; disc of Finger I small (
<figureCitation id="82DA2A7EFFF9FFF2FD7CFA08E594FA65" captionStart="FIG" captionStartId="5.[142,156,1866,1885]" captionTargetBox="[119,1488,193,1839]" captionTargetId="figure-0@5.[118,1491,191,1840]" captionTargetPageId="5" captionText="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="7" pageNumber="142">Fig. 3C</figureCitation>
); paired scutes on dorsal surfaces of finger discs well developed. Legs slender, tibia slightly longer than thigh; foot with elongated oval inner metatarsal tubercle and a smaller, protruding, round outer metatarsal tubercle (
<figureCitation id="82DA2A7EFFF9FFF2FD37F983E60AF9DC" box="[675,774,1589,1613]" captionStart="FIG" captionStartId="5.[142,156,1866,1885]" captionTargetBox="[119,1488,193,1839]" captionTargetId="figure-0@5.[118,1491,191,1840]" captionTargetPageId="5" captionText="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="7" pageNumber="142">Fig. 3D</figureCitation>
); subarticular tubercles single, protruding, slightly elliptical, and slightly larger on Toe I; supernumerary tubercles absent from feet; relative lengths of Toes I &lt;II &lt;V &lt;III &lt;IV; toes exhibit extensive lateral fringes on both sides; the fringe of the outer margin of Toe V extends to slightly beyond the proximal subarticular tubercle; tarsal flap extensive, distally continuous with fringe on the inner side of Toe I, almost reaching heel; toe tips dilated; toe discs oval from ventral view; disc of Toe I small slightly oval (
<figureCitation id="82DA2A7EFFF9FFF2FD83F8F4E762F8CB" box="[535,622,1858,1882]" captionStart="FIG" captionStartId="5.[142,156,1866,1885]" captionTargetBox="[119,1488,193,1839]" captionTargetId="figure-0@5.[118,1491,191,1840]" captionTargetPageId="5" captionText="FIG. 3.—Hŋlodes caete, CFBH 40524 (holotype), adult male. (A) Dorsal and (B) lateral views of head and ventral views of (C) left hand and (D) left foot. Scale bar ¼ 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7712277" httpUri="https://zenodo.org/record/7712277/files/figure.png" pageId="7" pageNumber="142">Fig. 3D</figureCitation>
); paired scutes on dorsal surfaces of toe discs well developed.
</paragraph>
<paragraph id="1A5E36FBFFF9FFF2FF02F8CBE6CAFBA5" blockId="7.[118,788,873,2001]" lastBlockId="7.[820,1490,873,2001]" pageId="7" pageNumber="142">
Skin almost smooth on dorsum and smooth on flanks and dorsal surfaces of legs; posterior region of body with few small scattered tubercles; ventral surfaces smooth; ventral surfaces of thighs and nearby areas with granular skin. We observed three protuberances at the end of the dorsal posterior region of body, which were formed by colonies of mites (parasites under the skin). Measurements (mm) of 
<typeStatus id="C55A8859FFF9FFF2FCA0FC56E69FFC69" box="[820,915,992,1016]" pageId="7" pageNumber="142" type="holotype">holotype</typeStatus>
are SVL 31.6, HL 12.1, HW 10.4, ED 4.7, TD 2.8, END 2.4, IOD 3.3, IND 4.3, THL 15.8, TBL 17.6, TAL 9.4, and FL 16.4.
</paragraph>
<paragraph id="1A5E36FBFFF9FFF2FCDAFB8CE6E3F9BE" blockId="7.[820,1490,873,2001]" pageId="7" pageNumber="142">
<emphasis id="2895EAE9FFF9FFF2FCDAFB8CE6D1FBC3" bold="true" box="[846,989,1082,1106]" pageId="7" pageNumber="142">Coloration.</emphasis>
—In preservative, the coloration of the 
<typeStatus id="C55A8859FFF9FFF2FCA0FBEEE69FFBE1" box="[820,915,1112,1136]" pageId="7" pageNumber="142" type="holotype">holotype</typeStatus>
is as follows: iris gray; dorsum dark brown with slightly clearer blotches; faint whitish line visible in the groin region; slightly darker-brown lateral stripe extends from tip of snout, through nostril, interrupted by the eye, starting again posterior to the eye, passing over the tympanum, and ending near the insertion of the arm; a slightly clearer brown lateral stripe, below the darker-brown stripe, extends from the tip of the snout to the insertion of the arm; lips gray; dorsal surfaces of arm, hand, thigh, and tibia, and hidden surfaces of thighs dark brown; dorsal surfaces of tarsus and foot whitish with many irregular small brown and whitishbrown blotches; few small dark-brown warts on posterior region of the body and near the coccygeal region; venter whitish, but well covered with many irregular, small, darkbrown blotches, which are more concentrated in the gular region (
<figureCitation id="82DA2A7EFFF9FFF2FC1FF9A1E6EEF9BE" box="[907,994,1559,1583]" captionStart="FIG" captionStartId="4.[166,180,898,917]" captionTargetBox="[142,1515,191,872]" captionTargetId="figure-795@4.[142,1516,191,872]" captionTargetPageId="4" captionText="FIG. 2.—Hŋlodes caete, CFBH 40524 (holotype), adult male in (A) dorsal and (B) ventral views. Scale bar ¼ 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7712275" httpUri="https://zenodo.org/record/7712275/files/figure.png" pageId="7" pageNumber="142">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="1A5E36FBFFF9FFF2FCC0F980E19DF8AD" blockId="7.[820,1490,873,2001]" pageId="7" pageNumber="142">
In life, the colors of the 
<typeStatus id="C55A8859FFF9FFF2FB10F983E1EFF9DC" box="[1156,1251,1589,1613]" pageId="7" pageNumber="142" type="holotype">holotype</typeStatus>
are more vivid and contrasting; however, they are basically the same as described for specimens in preservative, with the following exceptions: iris copper; faint whitish line visible from tip of snout to the anterior corner of the eye; lateral stripe from tip of snout to arm insertion (below the darker brown stripe) whitish-silver; lips pale brown; hidden surfaces of thighs dark brown with black blotches; dark brown transverse bars on thigh, tibia, and tarsus (
<figureCitation id="82DA2A7EFFF9FFF2FBABF892E18EF8AD" box="[1087,1154,1828,1852]" captionStart="FIG" captionStartId="3.[142,156,1909,1928]" captionTargetBox="[118,788,1398,1883]" captionTargetId="figure-805@3.[118,788,1398,1883]" captionTargetPageId="3" captionText="FIG. 1.—Live adult male Hŋlodes caete sp. nov. (holotype CFBH 40524; snout–vent length ¼ 31.6 mm). Photograph taken in Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712273" httpUri="https://zenodo.org/record/7712273/files/figure.png" pageId="7" pageNumber="142">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="1A5E36FBFFF9FFFDFCDAF8F4E7B6FB7B" blockId="7.[820,1490,873,2001]" lastBlockId="8.[142,813,730,2001]" lastPageId="8" lastPageNumber="143" pageId="7" pageNumber="142">
<emphasis id="2895EAE9FFF9FFF2FCDAF8F4E0BBF8CB" bold="true" box="[846,1463,1858,1882]" pageId="7" pageNumber="142">Variation among the type series in preservative.</emphasis>
— Measurements of 
<typeStatus id="C55A8859FFF9FFF2FC6AF8D6E164F8E9" box="[1022,1128,1888,1912]" pageId="7" pageNumber="142" type="paratype">paratypes</typeStatus>
and 
<typeStatus id="C55A8859FFF9FFF2FB36F8D6E00DF8E9" box="[1186,1281,1888,1912]" pageId="7" pageNumber="142" type="holotype">holotype</typeStatus>
(totaling 
<specimenCount id="0CE7FD72FFF9FFF2FAFBF8D6E0DDF8E9" box="[1391,1489,1888,1912]" pageId="7" pageNumber="142" type="male">16 males</specimenCount>
and 
<specimenCount id="0CE7FD72FFF9FFF2FCF1F8CBE6DCF804" box="[869,976,1917,1941]" pageId="7" pageNumber="142" type="female">5 females</specimenCount>
) are presented in 
<tableCitation id="57630340FFF9FFF2FB0AF8CBE1F8F807" box="[1182,1268,1917,1942]" captionStart="TABLE" captionStartId="6.[169,182,188,207]" captionTargetPageId="6" captionText="TABLE 3.—Measurements (mm, reported as ¯X ± 1 SD) of males (n ¼ 16) and females (n ¼ 5) of the type series of Hŋlodes caete sp. nov." httpUri="http://table.plazi.org/id/4E9E6673FFF8FFF3FF3DFF0AE7C7FF76" pageId="7" pageNumber="142" tableUuid="4E9E6673FFF8FFF3FF3DFF0AE7C7FF76">Table 3</tableCitation>
. Females are larger than males in SVL. Females lack nuptial thumb tubercles, vocal sacs, and vocal slits. Fringes are discreet on fingers and toes of females. Dorsum pale brown to grayish-dark brown, but only males possess darker patterns; dorsum ranging from a uniform pattern to patterns with many slightly clearer brown blotches (some blotched individuals may also exhibit medial longitudinal blotches following the vertebral column); thigh and tarsus with 2–5 transverse bars of variable widths; tibia with 3–5 transverse bars of variable widths; transverse bars are visible in females and can be visible, or not, in males; lateral stripe (below the darker-brown stripe) ranges from dark gray to whitish-cream, but only present in males with darker patterns; lips pale brown to gray, but females always with pale brown lips; posterior region of the body and coccygeal region smooth to warty; ventral surfaces vary from cream or white, with few brownish blotches, to being well covered with many dark-grayish or dark-brownish blotches; gular region varies from possessing a creamy-colored pattern, with a medial longitudinal brownish line, to having a blackish pattern which is only present in males.
</paragraph>
</subSubSection>
<caption id="4E9E6673FFF6FFFDFF32FDE5E4FFFD3C" ID-DOI="http://doi.org/10.5281/zenodo.7712287" ID-Zenodo-Dep="7712287" httpUri="https://zenodo.org/record/7712287/files/figure.png" pageId="8" pageNumber="143" startId="8.[166,180,595,614]" targetBox="[142,813,191,568]" targetPageId="8">
<paragraph id="1A5E36FBFFF6FFFDFF32FDE5E4FFFD3C" blockId="8.[142,813,594,685]" pageId="8" pageNumber="143">
FIG. 6.—The holotype locality for 
<taxonomicName id="DDE14D78FFF6FFFDFE43FDE4E75DFDF4" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[471,593,594,613]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFE43FDE4E75DFDF4" box="[471,593,594,613]" italics="true" pageId="8" pageNumber="143">Hŋlodes caete</emphasis>
</taxonomicName>
, a fast-flowing stream of the Rio Camburi at Núcleo Curucutu, Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online.
</paragraph>
</caption>
<subSubSection id="52FB6570FFF6FFFDFF3DFB46E621FA2B" pageId="8" pageNumber="143" type="etymology">
<paragraph id="1A5E36FBFFF6FFFDFF3DFB46E621FA2B" blockId="8.[142,813,730,2001]" pageId="8" pageNumber="143">
<emphasis id="2895EAE9FFF6FFFDFF3DFB46E43FFA99" bold="true" box="[169,307,1264,1288]" pageId="8" pageNumber="143">Etymology.</emphasis>
—The name of the new species, 
<taxonomicName id="DDE14D78FFF6FFFDFD2EFB47E7FDFA99" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[698,753,1265,1288]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFD2EFB47E7FDFA99" box="[698,753,1265,1288]" italics="true" pageId="8" pageNumber="143">caete</emphasis>
</taxonomicName>
, is a noun in apposition that is derived from the combination of two indigenous Tupi words, ‘‘caa’’ and ‘‘ete.’’ The combination expresses ‘‘true forest’’ with, respectively, ‘‘caa’’ meaning forest and ‘‘ete’’ meaning true. Here, 
<taxonomicName id="DDE14D78FFF6FFFDFD3FFADEE7EEFAEE" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[683,738,1384,1407]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFD3FFADEE7EEFAEE" box="[683,738,1384,1407]" italics="true" pageId="8" pageNumber="143">caete</emphasis>
</taxonomicName>
refers to the high preserved forests that harbor the fast streams with clear water in which the new species is known to breed.
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFF6FFFDFF3DFA76E189FD02" pageId="8" pageNumber="143" type="distribution">
<paragraph id="1A5E36FBFFF6FFFDFF3DFA76E46CF91B" blockId="8.[142,813,730,2001]" pageId="8" pageNumber="143">
<emphasis id="2895EAE9FFF6FFFDFF3DFA76E44AFA49" bold="true" box="[169,326,1472,1496]" pageId="8" pageNumber="143">Distribution.</emphasis>
— 
<taxonomicName id="DDE14D78FFF6FFFDFEF5FA76E4F1FA49" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[353,509,1472,1496]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFEF5FA76E4F1FA49" box="[353,509,1472,1496]" italics="true" pageId="8" pageNumber="143">Hŋlodes caete</emphasis>
</taxonomicName>
is known from the crests and slopes in high montane, montane, and submontane dense ombrophilous forests of Serra do Mar in the State of São Paulo (
<figureCitation id="82DA2A7EFFF6FFFDFE9EF9AFE458F9A0" box="[266,340,1561,1585]" captionStart="FIG" captionStartId="8.[166,180,595,614]" captionTargetBox="[142,813,191,568]" captionTargetId="figure-1049@8.[142,814,191,568]" captionTargetPageId="8" captionText="FIG. 6.—The holotype locality for Hŋlodes caete, a fast-flowing stream of the Rio Camburi at Núcleo Curucutu, Parque Estadual da Serra do Mar, municipality of Itanhaém, State of São Paulo, southeastern Brazil. A color version of this figure is available online." figureDoi="http://doi.org/10.5281/zenodo.7712287" httpUri="https://zenodo.org/record/7712287/files/figure.png" pageId="8" pageNumber="143">Figs. 6</figureCitation>
and 
<figureCitation id="82DA2A7EFFF6FFFDFE1FF9AFE496F9A0" box="[395,410,1561,1585]" captionStart="FIG" captionStartId="9.[142,156,1156,1175]" captionTargetBox="[119,1493,194,1130]" captionTargetId="figure-413@9.[118,1493,191,1130]" captionTargetPageId="9" captionText="FIG. 7.—Geographic distribution of Hŋlodes caete and the other three Hŋlodes species having males with nuptial tubercles on the thumb: H. fredi, H. phŋllodes, and H. pipilans. Hŋlodes caete is associated with high montane, montane, and submontane forests in well-preserved areas in the Atlantic Forest of Serra do Mar, southeastern Brazil whereas H. fredi is endemic to an island (Ilha Grande, municipality of Angra dos Reis, State of Rio de Janeiro) and H. pipilans is known only from Serra dos Órgãos (State of Rio de Janeiro). Open symbols indicate type localities. MG ¼ State of Minas Gerais; RJ ¼ State of Rio de Janeiro; and SP ¼ State of São Paulo." figureDoi="http://doi.org/10.5281/zenodo.7712289" httpUri="https://zenodo.org/record/7712289/files/figure.png" pageId="8" pageNumber="143">7</figureCitation>
). The new species is known to occur in the municipalities of Pilar do Sul, Ibiúna, Itanhaém, 
<collectingCountry id="62F6766BFFF6FFFDFC91F981E5E8F9FC" name="Saint Vincent and the Grenadines" pageId="8" pageNumber="143">São Vicente</collectingCountry>
, Santos, Cubatão, 
<collectingRegion id="D825F819FFF6FFFDFE26F9E3E731F9FC" box="[434,573,1621,1645]" country="Saint Vincent and the Grenadines" name="Saint Andrew" pageId="8" pageNumber="143">Santo André</collectingRegion>
(Paranapiacaba), and São Paulo (
<figureCitation id="82DA2A7EFFF6FFFDFE9AF9C4E45EF91B" box="[270,338,1650,1674]" captionStart="FIG" captionStartId="9.[142,156,1156,1175]" captionTargetBox="[119,1493,194,1130]" captionTargetId="figure-413@9.[118,1493,191,1130]" captionTargetPageId="9" captionText="FIG. 7.—Geographic distribution of Hŋlodes caete and the other three Hŋlodes species having males with nuptial tubercles on the thumb: H. fredi, H. phŋllodes, and H. pipilans. Hŋlodes caete is associated with high montane, montane, and submontane forests in well-preserved areas in the Atlantic Forest of Serra do Mar, southeastern Brazil whereas H. fredi is endemic to an island (Ilha Grande, municipality of Angra dos Reis, State of Rio de Janeiro) and H. pipilans is known only from Serra dos Órgãos (State of Rio de Janeiro). Open symbols indicate type localities. MG ¼ State of Minas Gerais; RJ ¼ State of Rio de Janeiro; and SP ¼ State of São Paulo." figureDoi="http://doi.org/10.5281/zenodo.7712289" httpUri="https://zenodo.org/record/7712289/files/figure.png" pageId="8" pageNumber="143">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="1A5E36FBFFF6FFFDFF3AF926E440F8E9" blockId="8.[142,813,730,2001]" pageId="8" pageNumber="143">
The new species is sympatric and syntopic with 
<taxonomicName id="DDE14D78FFF6FFFDFC85F927E5FAF957" baseAuthorityName="Heyer and Cocroft" baseAuthorityYear="1986" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF6FFFDFC85F927E5FAF957" italics="true" pageId="8" pageNumber="143">H. phŋllodes</emphasis>
</taxonomicName>
. We recorded these two species together at the 
<typeStatus id="C55A8859FFF6FFFDFF1AF97AE5B1F975" box="[142,189,1740,1764]" pageId="8" pageNumber="143">type</typeStatus>
locality and in the municipalities of 
<collectingCountry id="62F6766BFFF6FFFDFDD9F97AE7DEF975" box="[589,722,1740,1764]" name="Saint Vincent and the Grenadines" pageId="8" pageNumber="143">São Vicente</collectingCountry>
, Santos, Cubatão, and 
<collectingRegion id="D825F819FFF6FFFDFEA5F95FE4B3F890" box="[305,447,1769,1793]" country="Saint Vincent and the Grenadines" name="Saint Andrew" pageId="8" pageNumber="143">Santo André</collectingRegion>
(Paranapiacaba) of the State of São Paulo. Moreover, we recorded synchronotopic calling activity of the two species in the 
<typeStatus id="C55A8859FFF6FFFDFD92F893E739F8AC" box="[518,565,1829,1853]" pageId="8" pageNumber="143">type</typeStatus>
locality (
<figureCitation id="82DA2A7EFFF6FFFDFD0AF893E7F9F8AC" box="[670,757,1829,1853]" captionStart="FIG" captionStartId="10.[166,180,1296,1315]" captionTargetBox="[143,1516,192,1270]" captionTargetId="figure-364@10.[142,1517,191,1270]" captionTargetPageId="10" captionText="FIG. 8.—(A) Three advertisement calls of an individual of Hŋlodes caete and three advertisement calls of an individual of H. phŋllodes recorded in synchronotopy (upper panel ¼ spectrogram; lower panel ¼ waveform). Calls recorded at the type locality of H. caete on 13 February 2016; air temperature ¼ 21°C and water temperature ¼ 19.3°C. (B) One advertisement call of Hŋlodes caete, CFBH 40524 (holotype), adult male (upper panel ¼ spectrogram; lower panel ¼ waveform). Call recorded on 11 March 2015; air temperature ¼ 22.8°C and water temperature ¼ 21°C." figureDoi="http://doi.org/10.5281/zenodo.7712291" httpUri="https://zenodo.org/record/7712291/files/figure.png" pageId="8" pageNumber="143">Fig. 8A</figureCitation>
; also see audio files associated with the Supplemental Materials available online).
</paragraph>
<paragraph id="1A5E36FBFFF6FFFDFF3DF8C8E189FD02" blockId="8.[142,813,730,2001]" lastBlockId="8.[844,1515,188,2001]" pageId="8" pageNumber="143">
<emphasis id="2895EAE9FFF6FFFDFF3DF8C8E499F807" bold="true" box="[169,405,1918,1942]" pageId="8" pageNumber="143">Advertisement call.</emphasis>
—The advertisement call of 
<taxonomicName id="DDE14D78FFF6FFFDFD46F8C9E620F807" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[722,812,1919,1942]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFD46F8C9E620F807" box="[722,812,1919,1942]" italics="true" pageId="8" pageNumber="143">H. caete</emphasis>
</taxonomicName>
has harmonic structure and the first harmonic is not apparent (
<figureCitation id="82DA2A7EFFF6FFFDFE9CF80FE469F840" box="[264,357,1977,2001]" captionStart="FIG" captionStartId="10.[166,180,1296,1315]" captionTargetBox="[143,1516,192,1270]" captionTargetId="figure-364@10.[142,1517,191,1270]" captionTargetPageId="10" captionText="FIG. 8.—(A) Three advertisement calls of an individual of Hŋlodes caete and three advertisement calls of an individual of H. phŋllodes recorded in synchronotopy (upper panel ¼ spectrogram; lower panel ¼ waveform). Calls recorded at the type locality of H. caete on 13 February 2016; air temperature ¼ 21°C and water temperature ¼ 19.3°C. (B) One advertisement call of Hŋlodes caete, CFBH 40524 (holotype), adult male (upper panel ¼ spectrogram; lower panel ¼ waveform). Call recorded on 11 March 2015; air temperature ¼ 22.8°C and water temperature ¼ 21°C." figureDoi="http://doi.org/10.5281/zenodo.7712291" httpUri="https://zenodo.org/record/7712291/files/figure.png" pageId="8" pageNumber="143">Fig. 8B</figureCitation>
). At 20.8–22.8°C air temperature and 19.2–21°C water temperature, call duration ranges from 1.22– 
<geoCoordinate id="7FD5503CFFF6FFFDFC13FF6CE6C5FF60" box="[903,969,218,242]" degrees="2.72" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="555" value="-2.72">2.72 s</geoCoordinate>
(1.79 ± 
<geoCoordinate id="7FD5503CFFF6FFFDFBA4FF6CE17EFF60" box="[1072,1138,218,242]" degrees="0.38" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="555" value="-0.38">0.38 s</geoCoordinate>
, 
<emphasis id="2895EAE9FFF6FFFDFB14FF6DE182FF63" box="[1152,1166,219,242]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 calls from 
<specimenCount id="0CE7FD72FFF6FFFDFAD9FF6CE0AAFF63" box="[1357,1446,218,242]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); calls occur at intervals ranging from 1.85– 
<geoCoordinate id="7FD5503CFFF6FFFDFB71FF4EE02BFE9E" box="[1253,1319,248,272]" degrees="7.72" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="555" value="-7.72">7.72 s</geoCoordinate>
(3.54 ± 
<geoCoordinate id="7FD5503CFFF6FFFDFA18FF4EE0C2FE9E" box="[1420,1486,248,272]" degrees="1.19" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="555" value="-1.19">1.19 s</geoCoordinate>
, 
<emphasis id="2895EAE9FFF6FFFDFA4FFF4FE0E5FE81" box="[1499,1513,249,272]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 intervals from 
<specimenCount id="0CE7FD72FFF6FFFDFBA8FEA3E19BFEBF" box="[1084,1175,277,302]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); 20–45 notes per call (30 ± 6.67, 
<emphasis id="2895EAE9FFF6FFFDFC1CFE82E69AFEDA" box="[904,918,308,331]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 calls from 
<specimenCount id="0CE7FD72FFF6FFFDFBDAFE85E1AAFEDA" box="[1102,1190,307,331]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
), given at a rate ranging from 14.8–18.3/s (16.6 ± 1.02/s, 
<emphasis id="2895EAE9FFF6FFFDFBEBFEE4E181FEF8" box="[1151,1165,338,361]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 calls from 
<specimenCount id="0CE7FD72FFF6FFFDFAD8FEE7E0A9FEF8" box="[1356,1445,337,361]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); note duration ranges from 
<geoCoordinate id="7FD5503CFFF6FFFDFBD3FED9E18FFE16" box="[1095,1155,367,391]" degrees="0.009" direction="north" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="0.009">0.009</geoCoordinate>
<geoCoordinate id="7FD5503CFFF6FFFDFB17FED9E1C7FE16" box="[1155,1227,367,391]" degrees="0.023" direction="west" orientation="longitude" pageId="8" pageNumber="143" precision="55" value="-0.023">
– 
<geoCoordinate id="7FD5503CFFF6FFFDFB04FED9E1C7FE16" box="[1168,1227,367,391]" degrees="0.023" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="-0.023">0.023</geoCoordinate>
</geoCoordinate>
s (0.016 ± 
<geoCoordinate id="7FD5503CFFF6FFFDFACFFED9E0A0FE16" box="[1371,1452,367,391]" degrees="0.002" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="-0.002">0.002 s</geoCoordinate>
, 
<emphasis id="2895EAE9FFF6FFFDFA28FEC6E0C6FE16" box="[1468,1482,368,391]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 108 notes of 36 calls from 
<specimenCount id="0CE7FD72FFF6FFFDFBEEFE3BE1DFFE34" box="[1146,1235,397,421]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); notes given at intervals ranging from 
<geoCoordinate id="7FD5503CFFF6FFFDFC61FE1CE13DFE53" box="[1013,1073,426,450]" degrees="0.036" direction="north" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="0.036">0.036</geoCoordinate>
<geoCoordinate id="7FD5503CFFF6FFFDFBA5FE1CE175FE53" box="[1073,1145,426,450]" degrees="0.058" direction="west" orientation="longitude" pageId="8" pageNumber="143" precision="55" value="-0.058">
– 
<geoCoordinate id="7FD5503CFFF6FFFDFBAAFE1CE175FE53" box="[1086,1145,426,450]" degrees="0.058" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="-0.058">0.058</geoCoordinate>
</geoCoordinate>
s (0.45 ± 
<geoCoordinate id="7FD5503CFFF6FFFDFA86FE1CE065FE53" box="[1298,1385,426,450]" degrees="0.005" direction="south" orientation="latitude" pageId="8" pageNumber="143" precision="55" value="-0.005">0.005 s</geoCoordinate>
, 
<emphasis id="2895EAE9FFF6FFFDFAEBFE1DE081FE53" box="[1407,1421,427,450]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 108 intervals of 36 calls from 
<specimenCount id="0CE7FD72FFF6FFFDFBE0FE7EE1C3FE71" box="[1140,1231,456,480]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); each note consists of a rising frequency–modulated whistle; the dominant frequency occurs in the third harmonic and ranges from 3937.5– 5062.5 Hz (4640.63 ± 28.47 Hz, 
<emphasis id="2895EAE9FFF6FFFDFB58FD95E1D6FDAB" box="[1228,1242,547,570]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 call notes from 
<specimenCount id="0CE7FD72FFF6FFFDFA48FD94E683FDC9" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
); the first note may be lower, with a dominant frequency range of 3937.5–4687.5 Hz (4283.7 ± 18.01, 
<emphasis id="2895EAE9FFF6FFFDFA2AFDE8E0C0FDE4" box="[1470,1484,606,629]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 36 call notes from 
<specimenCount id="0CE7FD72FFF6FFFDFBB4FDCDE175FD02" box="[1056,1145,635,659]" pageId="8" pageNumber="143" type="male">6 males</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFF6FFFDFCF3FD2FE178FAD0" pageId="8" pageNumber="143" type="biology_ecology">
<paragraph id="1A5E36FBFFF6FFFDFCF3FD2FE178FAD0" blockId="8.[844,1515,188,2001]" pageId="8" pageNumber="143">
<emphasis id="2895EAE9FFF6FFFDFCF3FD2FE1FDFD20" bold="true" box="[871,1265,665,689]" pageId="8" pageNumber="143">Natural history and behavior.</emphasis>
— 
<taxonomicName id="DDE14D78FFF6FFFDFA98FD2FE0BCFD20" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1292,1456,665,689]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFA98FD2FE0BCFD20" box="[1292,1456,665,689]" italics="true" pageId="8" pageNumber="143">Hŋlodes caete</emphasis>
</taxonomicName>
was found along medium and large, fast-torrent streams (width ranging from 
<quantity id="DD199B1EFFF6FFFDFC7AFD63E144FD7C" box="[1006,1096,725,749]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="8" pageNumber="143" unit="m" value="6.0" valueMax="10.0" valueMin="2.0">2–10 m</quantity>
) within a narrow elevational range, from the middle to the top of the mountains of Serra do Mar (about 
<quantity id="DD199B1EFFF6FFFDFC0AFCA6E110FCB9" box="[926,1052,784,808]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="9.0" metricValueMin="4.5" pageId="8" pageNumber="143" unit="m" value="675.0" valueMax="900.0" valueMin="450.0">450–900 m</quantity>
a.s.l.; not recorded at lower elevations). 
<taxonomicName id="DDE14D78FFF6FFFDFCD8FC98E6E1FCD7" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[844,1005,814,838]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFCD8FC98E6E1FCD7" box="[844,1005,814,838]" italics="true" pageId="8" pageNumber="143">Hŋlodes caete</emphasis>
</taxonomicName>
is predominantly diurnal, with its calling activity ending around 1900 h. During the months of breeding and intensive activity, however, we recorded some males calling until 2000 h (
<emphasis id="2895EAE9FFF6FFFDFB19FC3FE197FC31" box="[1165,1179,905,928]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 4). Essentially, males call between September and March, but at lower elevations (~ 
<quantity id="DD199B1EFFF6FFFDFCFEFC75E6BDFC4A" box="[874,945,963,987]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="143" unit="m" value="450.0">450 m</quantity>
a.s.l.) males were recorded calling in July as well. Males called from emergent rocks, from trunks and roots in ravines, from bromeliad leaves in slopes, or from locations concealed among rocks; in all cases, calling males were found in the middle, on the margins, or close to margins of permanent fast streams. 
<taxonomicName id="DDE14D78FFF6FFFDFBEFFBEEE010FBE1" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1147,1308,1112,1136]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFBEFFBEEE010FBE1" box="[1147,1308,1112,1136]" italics="true" pageId="8" pageNumber="143">Hŋlodes caete</emphasis>
</taxonomicName>
called from the ground level up to 
<quantity id="DD199B1EFFF6FFFDFB89FBC0E14BFB1F" box="[1053,1095,1142,1166]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="143" unit="m" value="2.0">2 m</quantity>
when it was perched on vegetation (
<emphasis id="2895EAE9FFF6FFFDFA4FFBC1E0E5FB1F" box="[1499,1513,1143,1166]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 26). When disturbed, calling males dove into the water or hid in crevices among rocks on the stream margins, only to return after a few minutes to the same calling site to begin calling again. At night males and females of 
<taxonomicName id="DDE14D78FFF6FFFDFAD9FB58E0A5FA94" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1357,1449,1262,1285]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFAD9FB58E0A5FA94" box="[1357,1449,1262,1285]" italics="true" pageId="8" pageNumber="143">H. caete</emphasis>
</taxonomicName>
were observed resting on stems and leaves of shrubs (
<quantity id="DD199B1EFFF6FFFDFAEDFABDE0E6FAB2" box="[1401,1514,1291,1315]" metricMagnitude="0" metricUnit="m" metricValue="1.05" metricValueMax="1.6" metricValueMin="0.5" pageId="8" pageNumber="143" unit="m" value="1.05" valueMax="1.6" valueMin="0.5">0.5–1.6 m</quantity>
from the ground; 
<emphasis id="2895EAE9FFF6FFFDFB81FA9CE12FFAD0" box="[1045,1059,1322,1345]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 17).
</paragraph>
</subSubSection>
<subSubSection id="52FB6570FFF6FFFDFCF8FAFEE1AEF840" pageId="8" pageNumber="143" type="discussion">
<paragraph id="1A5E36FBFFF6FFFDFCF8FAFEE1AEF840" blockId="8.[844,1515,188,2001]" pageId="8" pageNumber="143">
We recorded two 
<taxonomicName id="DDE14D78FFF6FFFDFBC0FAFEE1B4FACE" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1108,1208,1352,1375]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFBC0FAFEE1B4FACE" box="[1108,1208,1352,1375]" italics="true" pageId="8" pageNumber="143">H. caete</emphasis>
</taxonomicName>
males performing visual displays. They displayed foot-flagging (
<emphasis id="2895EAE9FFF6FFFDFA91FAD0E01FFAEC" box="[1285,1299,1382,1405]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 2 displays from 
<specimenCount id="0CE7FD72FFF6FFFDFCD8FA35E6A8FA0A" box="[844,932,1411,1435]" pageId="8" pageNumber="143" type="male">2 males</specimenCount>
), leg lifting (
<emphasis id="2895EAE9FFF6FFFDFBA5FA32E133FA0A" box="[1073,1087,1412,1435]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 1 display from 
<specimenCount id="0CE7FD72FFF6FFFDFA9DFA35E05BFA0A" box="[1289,1367,1411,1435]" pageId="8" pageNumber="143" type="male">1 male</specimenCount>
), arm waving (
<emphasis id="2895EAE9FFF6FFFDFCC1FA17E66FFA29" box="[853,867,1441,1464]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 7 displays from 
<specimenCount id="0CE7FD72FFF6FFFDFBDAFA17E1ADFA28" box="[1102,1185,1441,1465]" pageId="8" pageNumber="143" type="male">1 male</specimenCount>
), and mouth gaping (
<emphasis id="2895EAE9FFF6FFFDFA35FA17E0A3FA29" box="[1441,1455,1441,1464]" italics="true" pageId="8" pageNumber="143">n</emphasis>
¼ 1 display from 
<specimenCount id="0CE7FD72FFF6FFFDFC72FA09E134FA47" box="[998,1080,1470,1495]" pageId="8" pageNumber="143" type="male">1 male</specimenCount>
, performed simultaneously with footflagging). See 
<bibRefCitation id="7E704B0AFFF6FFFDFC7DFA6BE1EBFA65" author="Hartmann, M. T. &amp; P. A. Hartmann &amp; C. F. B. Haddad" box="[1001,1255,1500,1524]" pageId="8" pageNumber="143" pagination="203 - 209" refId="ref11018" refString="Hartmann, M. T., P. A. Hartmann, and C. F. B. Haddad. 2006. Repertorio vocal de Hnlodes phnllodes (Amphibia, Anura, Hylodidae). Papeis Avulsos de Zoologia 46: 203 - 209." type="journal article" year="2006">Hartmann et al. (2006)</bibRefCitation>
and 
<bibRefCitation id="7E704B0AFFF6FFFDFAB4FA6AE0E5FA65" author="de Sa, F. P. &amp; J. Zina &amp; C. F. B. Haddad" box="[1312,1513,1500,1524]" pageId="8" pageNumber="143" pagination="0145444" refId="ref10604" refString="de Sa, F. P., J. Zina, and C. F. B. Haddad. 2016. Sophisticated communication in the Brazilian torrent frog Hnlodes japi. PLoS ONE 11: e 0145444." type="journal article" year="2016">de Sá et al. (2016)</bibRefCitation>
for display definitions. At least three of these four displays had already had been recorded for 
<taxonomicName id="DDE14D78FFF6FFFDFB4EF9AFE6C2F9DF" authority="(Hartmann et al. 2006)" baseAuthorityName="Hartmann" baseAuthorityYear="2006" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="phnllodes">
<emphasis id="2895EAE9FFF6FFFDFB4EF9AFE06AF9A1" box="[1242,1382,1560,1584]" italics="true" pageId="8" pageNumber="143">H. phŋllodes</emphasis>
(
<bibRefCitation id="7E704B0AFFF6FFFDFAE2F9AEE6CAF9DF" author="Hartmann, M. T. &amp; P. A. Hartmann &amp; C. F. B. Haddad" pageId="8" pageNumber="143" pagination="203 - 209" refId="ref11018" refString="Hartmann, M. T., P. A. Hartmann, and C. F. B. Haddad. 2006. Repertorio vocal de Hnlodes phnllodes (Amphibia, Anura, Hylodidae). Papeis Avulsos de Zoologia 46: 203 - 209." type="journal article" year="2006">Hartmann et al. 2006</bibRefCitation>
)
</taxonomicName>
. On 
<date id="6E5F103BFFF6FFFDFB90F980E1C2F9DF" box="[1028,1230,1590,1614]" pageId="8" pageNumber="143" value="2015-12-02">2 December 2015</date>
(between 1600 and 1610 h) we also observed a partial courtship of 
<taxonomicName id="DDE14D78FFF6FFFDFAA8F9E2E096F9FA" authorityName="Malagoli &amp; de Sá &amp; Canedo &amp; Haddad" authorityYear="2017" box="[1340,1434,1620,1643]" class="Amphibia" family="Cycloramphidae" genus="Hnlodes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="143" phylum="Chordata" rank="species" species="caete">
<emphasis id="2895EAE9FFF6FFFDFAA8F9E2E096F9FA" box="[1340,1434,1620,1643]" italics="true" pageId="8" pageNumber="143">H. caete</emphasis>
</taxonomicName>
in the municipality of 
<collectingCountry id="62F6766BFFF6FFFDFC6AF9C7E189F918" box="[1022,1157,1649,1673]" name="Saint Vincent and the Grenadines" pageId="8" pageNumber="143">São Vicente</collectingCountry>
, State of São Paulo, before the pair was collected. We noticed the pair and began to observe them when the female (CFBH 40531) jumped on the same rock where the male was calling (CFBH 40533) at the margin of a fast-flowing stream. The male and female stayed side by side with their heads oriented in opposite directions. The male called continuously (inflating vocal sacs on both sides concomitantly), and the female touched the male̕s dorsum with her right arm. The male immediately turned in the direction of the female and emitted a low, short call (this nonrecorded call had fewer notes than the advertisement call) and displayed arm lifting.
</paragraph>
</subSubSection>
</treatment>
</document>