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<document id="AA4F1863A58513C661054ED91EE4B8C0" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed &amp; Barbero, Enrico" docDate="2019" docId="8861532B885B182BFF760C36FC8CC3E8" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus (Eremonthophagus) sticticus Harold 1867" docType="treatment" docVersion="13" lastPageNumber="10" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="9" updateTime="1698751869822" updateUser="ExternalLinkService">
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<mods:title id="193C59D0B6F6CACC37FD07F283B82971">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<mods:namePart id="B45D3C5F945725C89022EFA790952D2F">Ziani, Stefano</mods:namePart>
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<mods:namePart id="94F7D12D7238BAE893E02710CF0D687A">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
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<heading id="5B3F5551885B182AFF760C36FCE8C230" bold="true" box="[151,843,1377,1404]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="C7C899BE885B182AFF760C36FCE8C230" ID-CoL="74MKT" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[151,843,1377,1404]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sticticus" subGenus="Eremonthophagus">
<emphasis id="32BC3E2F885B182AFF760C36FCE8C230" bold="true" box="[151,843,1377,1404]" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFF760C36FE9FC237" bold="true" box="[151,316,1377,1404]" italics="true" pageId="8" pageNumber="9">Onthophagus</emphasis>
(
<emphasis id="32BC3E2F885B182AFEAC0C36FD8DC237" bold="true" box="[333,558,1377,1404]" italics="true" pageId="8" pageNumber="9">Eremonthophagus</emphasis>
)
<emphasis id="32BC3E2F885B182AFDDE0C36FD02C230" bold="true" box="[575,673,1377,1403]" italics="true" pageId="8" pageNumber="9">sticticus</emphasis>
<bibRefCitation id="64599FCC885B182AFD490C36FCE8C230" author="Harold, E." box="[680,843,1377,1403]" pageId="8" pageNumber="9" pagination="23 - 59" refId="ref37904" refString="Harold, E. (1867) Beitrage zur Kenntniss der Gattung Onthophagus vom Herausgeber. I. Theil: Australier und Afrikaner. Coleopterologische Hefte, 2, 23 - 59." type="journal article" year="1867">Harold, 1867</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="48D2B1B6885B182AFF760CD2FF4FC2D4" box="[151,236,1412,1439]" pageId="8" pageNumber="9" type="description">
<paragraph id="0077E23D885B182AFF760CD2FF4FC2D4" blockId="8.[151,843,1377,1439]" box="[151,236,1412,1439]" pageId="8" pageNumber="9">
[
<figureCitation id="98F3FEB8885B182AFF7F0CD2FF47C2D4" box="[158,228,1412,1439]" captionStart="FIGURES 27" captionStartId="10.[151,264,1869,1894]" captionTargetBox="[306,1253,180,1830]" captionTargetId="figure@10.[271,1316,166,1849]" captionTargetPageId="10" captionText="FIGURES 27. Onthophagini of Arabian Peninsula. 2, Cleptocaccobius viridicollis (Fåhraeus in Boheman, 1857): YE (Sanaa) Wadi Dhahr; 3, Digitonthophagus eucatta Génier in Génier &amp; Moretto, 2017: SA (Asir) Wadi Marabah; 4, Euonthophagus carbonarius (Klug, 1855) sensu lato: YE (Sanaa) Wadi Dhahr; 5, Onthophagus (Eremonthophagus) infuscatus Klug, 1845: SA (Asir) 10 km N Wadi Reem; 6, Onthophagus (Eremonthophagus) sticticus Harold, 1867: OM (Dhofar) Ash-Shuwayiyah; 7, Onthophagus (Eremonthophagus) transcaspicus König, 1889: OM (Ad Dakhiliyah) Al Ayn. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373044/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
]
</paragraph>
</subSubSection>
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<paragraph id="0077E23D885B182AFF760C9BFAE0C13C" blockId="8.[151,1437,1484,2015]" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFF760C9BFECCC2AC" bold="true" box="[151,367,1484,1511]" pageId="8" pageNumber="9">Literature quotes</emphasis>
.
<collectingCountry id="78DFA2AD885B182AFE650C9BFD85C2AF" box="[388,550,1484,1511]" name="Saudi Arabia" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFE650C9BFE65C2AF" bold="true" box="[388,454,1484,1511]" pageId="8" pageNumber="9">SAUDI</emphasis>
<taxonomicName id="C7C899BE885B182AFE320C9BFD85C2AF" baseAuthorityName="Zunino" baseAuthorityYear="1979" box="[467,550,1484,1511]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F885B182AFE320C9BFD85C2AF" bold="true" box="[467,550,1484,1511]" pageId="8" pageNumber="9">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
<emphasis id="32BC3E2F885B182AFDC70C9AFD8CC2AC" bold="true" box="[550,559,1485,1511]" pageId="8" pageNumber="9">:</emphasis>
<taxonomicName id="C7C899BE885B182AFDA60C9AFD3AC2AC" authority="Harold 1867" authorityName="Harold" authorityYear="1867" baseAuthorityName="Zunino" baseAuthorityYear="1979" box="[583,665,1485,1511]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
” (
<bibRefCitation id="64599FCC885B182AFD590C9AFCF1C2A3" author="Harold, E." box="[696,850,1485,1512]" pageId="8" pageNumber="9" pagination="23 - 59" refId="ref37904" refString="Harold, E. (1867) Beitrage zur Kenntniss der Gattung Onthophagus vom Herausgeber. I. Theil: Australier und Afrikaner. Coleopterologische Hefte, 2, 23 - 59." type="journal article" year="1867">Harold 1867</bibRefCitation>
).
<collectingRegion id="C20C2CDF885B182AFC8D0C9AFB9FC2AC" box="[876,1084,1485,1511]" country="Saudi Arabia" name="Makkah" pageId="8" pageNumber="9">Makkah Province</collectingRegion>
, “Djedda” (dOrbigny 1900); “Buraiman”; “Taif”; “
<collectingRegion id="C20C2CDF885B182AFE750CA6FE44C140" box="[404,487,1521,1547]" country="Saudi Arabia" name="Makkah" pageId="8" pageNumber="9">Mecca</collectingRegion>
”; “Al Lith”; “Shiaera [Al Shaira], antilocust camp north of Qunfidha” (Scott in
<bibRefCitation id="64599FCC885B182AFA880CA6FEA7C164" author="Paulian, R." pageId="8" pageNumber="9" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
).
<collectingRegion id="C20C2CDF885B182AFEF60F42FE44C164" box="[279,487,1557,1583]" country="Saudi Arabia" name="Al Madinah" pageId="8" pageNumber="9">Madinah Province</collectingRegion>
, “
<quantity id="C7304FD8885B182AFE1C0F42FDE6C164" box="[509,581,1557,1583]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="km" value="16.0">16 km</quantity>
W Badr Hunayn”. Hejaz, “N Hedjaz” (
<bibRefCitation id="64599FCC885B182AFC1D0F42FB31C164" author="Zunino, M." box="[1020,1170,1557,1583]" pageId="8" pageNumber="9" pagination="408 - 416" refId="ref40252" refString="Zunino, M. (1981) Insects of Saudi Arabia Coleoptera: Fam. Scarabaeidae Tribe Onthophagini. Fauna of Saudi Arabia, 3, 408 - 416." type="journal article" year="1981">Zunino 1981</bibRefCitation>
). Yemen: without locality (Scott in
<bibRefCitation id="64599FCC885B182AFEC00F6EFE1BC118" author="Paulian, R." box="[289,440,1593,1619]" pageId="8" pageNumber="9" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
). “Nord Yemen” (
<bibRefCitation id="64599FCC885B182AFD9E0F6EFCBBC118" author="Paulian, R." box="[639,792,1593,1619]" pageId="8" pageNumber="9" pagination="141 - 154" refId="ref39235" refString="Paulian, R. (1980) Insects of Saudi Arabia. Coleoptera: Scarabaeoidea (1 re contribution). Fauna of Saudi Arabia, 2, 141 - 154." type="journal article" year="1980">Paulian 1980</bibRefCitation>
). Oman: Muscat Governorate, “Muscat” (Scott in
<bibRefCitation id="64599FCC885B182AFAA90F6EFF70C13C" author="Paulian, R." pageId="8" pageNumber="9" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
). Without locality (
<bibRefCitation id="64599FCC885B182AFE4B0F0AFDF0C13C" author="Janikova, M." box="[426,595,1629,1655]" pageId="8" pageNumber="9" refId="ref38111" refString="Janikova, M. (1998) Coleoptera. Beetles Checklist. Al Nahda Printing Press, Muscat, 39 pp." type="book" year="1998">Janikova 1998</bibRefCitation>
).
<collectingCountry id="78DFA2AD885B182AFD870F0AFCD3C13F" box="[614,880,1628,1655]" name="United Arab Emirates" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFD870F0AFCD3C13F" bold="true" box="[614,880,1628,1655]" pageId="8" pageNumber="9">
<subScript id="9C4CE078885B182AFD870F0AFD1BC13F" attach="left" box="[614,696,1629,1655]" fontSize="7" pageId="8" pageNumber="9">UNITED</subScript>
ARAB EMIRATES
</emphasis>
</collectingCountry>
: without locality (
<bibRefCitation id="64599FCC885B182AFBDF0F0AFA94C13C" author="Gillett, M. P. T. &amp; Gillett, C. P. D. T." box="[1086,1335,1628,1655]" pageId="8" pageNumber="9" pagination="352 - 356" refId="ref37609" refString="Gillett, M. P. T. &amp; Gillett, C. P. D. T. (2005) Insects &amp; other arthropods, pp. 168 - 194. Species checklist, In: Aspinall, S. &amp; Hellyer, P. (Eds.), The Emirates. A natural history. P. Vine, Trident Press, London, pp. 352 - 356." type="book chapter" year="2005">Gillett &amp; Gillett 2005</bibRefCitation>
).
</paragraph>
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<emphasis id="32BC3E2F885B182AFF260FD7FE0FC1D0" bold="true" box="[199,428,1664,1691]" pageId="8" pageNumber="9">Material examined</emphasis>
.
<materialsCitation id="B0A0E860885B182AFE5C0FD7FC50C1D0" ID-GBIF-Occurrence="2350486142" box="[445,1011,1664,1691]" collectionCode="JSCS" country="Saudi Arabia" location="SAUDI ARABIA" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Jizan Province">
<collectingCountry id="78DFA2AD885B182AFE5C0FD7FDF8C1D3" box="[445,603,1664,1691]" name="Saudi Arabia" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFE5C0FD7FE5CC1D3" bold="true" box="[445,511,1664,1691]" pageId="8" pageNumber="9">SAUDI</emphasis>
<taxonomicName id="C7C899BE885B182AFDE60FD7FDF8C1D3" baseAuthorityName="Zunino" baseAuthorityYear="1979" box="[519,603,1664,1691]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="32BC3E2F885B182AFDE60FD7FDF8C1D3" bold="true" box="[519,603,1664,1691]" pageId="8" pageNumber="9">ARABIA</emphasis>
</taxonomicName>
</collectingCountry>
:
<specimenCount id="16CE29B4885B182AFD8C0FD6FD05C1D0" box="[621,678,1665,1691]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
with no further data (
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)
</materialsCitation>
.
<materialsCitation id="B0A0E860885B182AFBE00FD6FA9FC1F4" ID-GBIF-Occurrence="2350486140" collectingDate="2017-02-17" collectionCode="KSMA" collectorName="U. M. Abu El-Ghiet &amp; T. M. Elsheikh" country="Saudi Arabia" latitude="16.860626" location="Sajid" longLatPrecision="1" longitude="41.932564" municipality="Farasan islands" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Jizan">
<collectingRegion id="C20C2CDF885B182AFBE00FD6FB09C1D0" box="[1025,1194,1665,1691]" country="Saudi Arabia" name="Jizan" pageId="8" pageNumber="9">Jizan Province</collectingRegion>
,
<collectingMunicipality id="E0137847885B182AFB580FD6FAC8C1D0" box="[1209,1387,1665,1691]" pageId="8" pageNumber="9">Farasan islands</collectingMunicipality>
, Al-
<location id="0517B4E6885B182AFF760FF2FF70C1F4" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFF760FF2FF70C1F4" box="[151,211,1701,1727]" country="Saudi Arabia" latitude="16.860626" longLatPrecision="1" longitude="41.932564" municipality="Farasan islands" name="Sajid" pageId="8" pageNumber="9" stateProvince="Jizan">Sajid</location>
, (
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,
<geoCoordinate id="65FC84FA885B182AFE6E0FF3FD84C1F4" box="[399,551,1700,1727]" degrees="41.932564" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="41.932564">41.932564°E</geoCoordinate>
),
<date id="7476C4FD885B182AFDDD0FF2FD13C1F4" box="[572,688,1701,1727]" pageId="8" pageNumber="9" value="2017-02-17">
<collectingDate id="64323D15885B182AFDDD0FF2FD13C1F4" box="[572,688,1701,1727]" pageId="8" pageNumber="9" value="2017-02-17">17.ii.2017</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFD5C0FF2FC34C1F4" box="[701,919,1701,1727]" pageId="8" pageNumber="9">U.M. Abu El-Ghiet</collectorName>
&amp;
<collectorName id="AD3D87EB885B182AFC5F0FF2FBC4C1F4" box="[958,1127,1701,1727]" pageId="8" pageNumber="9">T. M. Elsheikh</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFB400FF2FB6EC1F4" box="[1185,1229,1701,1727]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
(
<collectionCode id="66D97AF8885B182AFB3C0FF2FA97C1F4" box="[1245,1332,1701,1727]" pageId="8" pageNumber="9">KSMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFAA80FF2FCFCC1A8" ID-GBIF-Occurrence="2350486098" collectingDate="2017-10-10" collectionCode="KSMA" collectorName="Abu El-Ghiet &amp; T. M. Elsheikh" country="Saudi Arabia" pageId="8" pageNumber="9" specimenCount="3" specimenCount-male="3" stateProvince="Jizan Province">
<emphasis id="32BC3E2F885B182AFAA80FF2FA36C1F4" box="[1353,1429,1701,1727]" italics="true" pageId="8" pageNumber="9">ibidem</emphasis>
,
<date id="7476C4FD885B182AFF760F9EFEAAC1A8" box="[151,265,1737,1763]" pageId="8" pageNumber="9" value="2017-10-10">
<collectingDate id="64323D15885B182AFF760F9EFEAAC1A8" box="[151,265,1737,1763]" pageId="8" pageNumber="9" value="2017-10-10">10.x.2017</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFEF50F9EFE0FC1A8" box="[276,428,1737,1763]" pageId="8" pageNumber="9">Abu El-Ghiet</collectorName>
&amp;
<collectorName id="AD3D87EB885B182AFE300F9EFDDBC1A8" box="[465,632,1737,1763]" pageId="8" pageNumber="9">T. M. Elsheikh</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFD500F9EFD53C1A8" box="[689,752,1737,1763]" pageId="8" pageNumber="9" type="male">3 ♂♂</specimenCount>
(
<collectionCode id="66D97AF8885B182AFCE10F9EFCF4C1A8" box="[768,855,1737,1763]" pageId="8" pageNumber="9">KSMA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885B182AFC8B0F9EFB1DC04C" ID-GBIF-Occurrence="2350486145" collectingDate="1944-12-04" collectionCode="NHMUK" collectorName="A. R. Waterston" country="Saudi Arabia" county="Hedjas" location="Jidda" municipality="Buraiman" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Makkah" typeStatus="paratype">
<collectingRegion id="C20C2CDF885B182AFC8B0F9EFB90C1A8" box="[874,1075,1737,1763]" country="Saudi Arabia" name="Makkah" pageId="8" pageNumber="9">Makkah Province</collectingRegion>
,
<collectingCounty id="E9169AB1885B182AFBA10F9EFB32C1A8" box="[1088,1169,1737,1763]" pageId="8" pageNumber="9">Hedjas</collectingCounty>
,
<collectingMunicipality id="E0137847885B182AFB7D0F9EFAAEC1A8" box="[1180,1293,1737,1763]" pageId="8" pageNumber="9">Buraiman</collectingMunicipality>
,
<quantity id="C7304FD8885B182AFAF80F9EFAD9C1A8" box="[1305,1402,1737,1763]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="8" pageNumber="9" unit="mi" value="12.0">12 miles</quantity>
N. of
<location id="0517B4E6885B182AFF570FBAFF57C04C" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFF570FBAFF57C04C" box="[182,244,1773,1799]" country="Saudi Arabia" county="Hedjas" municipality="Buraiman" name="Jidda" pageId="8" pageNumber="9" stateProvince="Makkah">Jidda</location>
,
<date id="7476C4FD885B182AFF1E0FBAFED0C04C" box="[255,371,1773,1799]" pageId="8" pageNumber="9" value="1944-12-04">
<collectingDate id="64323D15885B182AFF1E0FBAFED0C04C" box="[255,371,1773,1799]" pageId="8" pageNumber="9" value="1944-12-04">4.xii.1944</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFE9C0FBAFD92C04C" box="[381,561,1773,1799]" pageId="8" pageNumber="9">A. R. Waterston</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFD880FBAFD30C04C" box="[617,659,1773,1799]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
(
<typeStatus id="DF735C9F885B182AFD430FBAFCA7C04C" box="[674,772,1773,1799]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
of
<taxonomicName id="C7C899BE885B182AFCCB0FBAFB9FC04C" authority="Frey, 1962" authorityName="Frey" authorityYear="1962" box="[810,1084,1773,1799]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="saudensis">
<emphasis id="32BC3E2F885B182AFCCB0FBAFC19C04C" box="[810,954,1773,1799]" italics="true" pageId="8" pageNumber="9">O. saudensis</emphasis>
<bibRefCitation id="64599FCC885B182AFC200FBAFB9FC04C" author="Frey, G." box="[961,1084,1773,1799]" pageId="8" pageNumber="9" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey, 1962</bibRefCitation>
</taxonomicName>
,
<collectionCode id="66D97AF8885B182AFBA60FBAFB15C04C" box="[1095,1206,1773,1799]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFB280FBAFC48C060" ID-GBIF-Occurrence="2350486090" collectingDate="1944-12-08" collectionCode="NHMB" collectorName="A. R. Waterston" country="Saudi Arabia" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Makkah Province" typeStatus="holotype">
<emphasis id="32BC3E2F885B182AFB280FBAFAB6C04C" box="[1225,1301,1773,1799]" italics="true" pageId="8" pageNumber="9">ibidem</emphasis>
,
<date id="7476C4FD885B182AFAC50FBAFA3BC04C" box="[1316,1432,1773,1799]" pageId="8" pageNumber="9" value="1944-12-08">
<collectingDate id="64323D15885B182AFAC50FBAFA3BC04C" box="[1316,1432,1773,1799]" pageId="8" pageNumber="9" value="1944-12-08">8.xii.1944</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFF760E46FEF3C060" box="[151,336,1809,1835]" pageId="8" pageNumber="9">A. R. Waterston</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFE6C0E46FE19C060" box="[397,442,1809,1835]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
(
<typeStatus id="DF735C9F885B182AFE2D0E46FD93C060" box="[460,560,1809,1835]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C7C899BE885B182AFDBA0E46FCDFC060" authority="Frey, 1962" authorityName="Frey" authorityYear="1962" box="[603,892,1809,1835]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="saudiensis">
<emphasis id="32BC3E2F885B182AFDBA0E46FD56C060" box="[603,757,1809,1835]" italics="true" pageId="8" pageNumber="9">O. saudiensis</emphasis>
<bibRefCitation id="64599FCC885B182AFD1E0E46FCDFC060" author="Frey, G." box="[767,892,1809,1835]" pageId="8" pageNumber="9" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey, 1962</bibRefCitation>
</taxonomicName>
,
<collectionCode id="66D97AF8885B182AFC6B0E46FC40C060" box="[906,995,1809,1835]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="9">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885B182AFC190E46FDD9C004" ID-GBIF-Occurrence="2350486141" collectionCode="NHMUK" collectorName="C. Rathjens" country="Yemen" elevation="2400" location="Sana'a Governorate" pageId="8" pageNumber="9" specimenCount="1" stateProvince="San'a'" typeStatus="holotype">
<collectingCountry id="78DFA2AD885B182AFC190E46FBE7C063" box="[1016,1092,1809,1835]" name="Yemen" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFC190E46FBE7C063" bold="true" box="[1016,1092,1809,1835]" pageId="8" pageNumber="9">YEMEN</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF885B182AFBB70E46FA98C060" box="[1110,1339,1809,1835]" country="Yemen" name="San'a'" pageId="8" pageNumber="9">Sanaa Governorate</collectingRegion>
,
<collectingRegion id="C20C2CDF885B182AFAA80E46FA3BC060" box="[1353,1432,1809,1835]" country="Yemen" name="San'a'" pageId="8" pageNumber="9">Sanaa</collectingRegion>
,
<quantity id="C7304FD8885B182AFF760E62FF4CC004" box="[151,239,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="9" unit="m" value="2400.0">
<elevation id="8BE5050E885B182AFF760E62FF4CC004" box="[151,239,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="9" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
<collectorName id="AD3D87EB885B182AFF1D0E62FE23C004" box="[252,384,1845,1871]" pageId="8" pageNumber="9">C. Rathjens</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFE5B0E62FE52C004" box="[442,497,1845,1871]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFDE00E62FDD1C004" box="[513,626,1845,1871]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="9" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFD670E62FF5EC038" ID-GBIF-Occurrence="2350486204" collectingDate="2005-10-07" collectionCode="NMPC" collectorName="D. Kral" country="Yemen" elevation="2350" location="Mafhaq" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Sana'a" typeStatus="holotype">
<location id="0517B4E6885B182AFD670E62FD43C004" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFD670E62FD43C004" box="[646,736,1845,1871]" country="Yemen" name="Mafhaq" pageId="8" pageNumber="9" stateProvince="Sana'a">Mafhaq</location>
,
<quantity id="C7304FD8885B182AFD0D0E63FCE1C004" box="[748,834,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="9" unit="m" value="2350.0">
<elevation id="8BE5050E885B182AFD0D0E63FCE1C004" box="[748,834,1844,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="9" unit="m" value="2350.0">2350 m</elevation>
</quantity>
(150903″N 435716″E),
<date id="7476C4FD885B182AFB840E63FB6AC004" box="[1125,1225,1844,1871]" pageId="8" pageNumber="9" value="2005-10-07">
<collectingDate id="64323D15885B182AFB840E63FB6AC004" box="[1125,1225,1844,1871]" pageId="8" pageNumber="9" value="2005-10-07">7.x.2005</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFB370E62FA88C004" box="[1238,1323,1845,1871]" pageId="8" pageNumber="9">D. Král</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFA840E62FA3EC004" box="[1381,1437,1845,1871]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFF7E0E0EFF56C038" box="[159,245,1881,1907]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="9">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885B182AFEE80E0FFE84C0DC" ID-GBIF-Occurrence="2350486205" collectingDate="1982-05-22" collectionCode="JSCS" collectorName="M. D. Gallagher" country="Oman" location="Sohar" municipality="Batinah" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Shamal al Batinah" typeStatus="holotype">
<collectingCountry id="78DFA2AD885B182AFEE80E0FFEEEC03B" box="[265,333,1880,1907]" name="Oman" pageId="8" pageNumber="9">
<emphasis id="32BC3E2F885B182AFEE80E0FFEEEC03B" bold="true" box="[265,333,1880,1907]" pageId="8" pageNumber="9">OMAN</emphasis>
</collectingCountry>
:
<collectingRegion id="C20C2CDF885B182AFEBD0E0EFDCCC038" box="[348,623,1881,1907]" country="Oman" name="Shamal al Batinah" pageId="8" pageNumber="9">Al Batinah Governorate</collectingRegion>
,
<collectingMunicipality id="E0137847885B182AFD9D0E0EFD70C038" box="[636,723,1881,1907]" pageId="8" pageNumber="9">Batinah</collectingMunicipality>
b.
<location id="0517B4E6885B182AFD180E0EFC99C038" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFD180E0EFC99C038" box="[761,826,1881,1907]" country="Oman" municipality="Batinah" name="Sohar" pageId="8" pageNumber="9" stateProvince="Shamal al Batinah">Sohar</location>
(2425N 5642E), 18/
<date id="7476C4FD885B182AFBD20E0EFB00C038" box="[1075,1187,1881,1907]" pageId="8" pageNumber="9" value="1982-05-22">
<collectingDate id="64323D15885B182AFBD20E0EFB00C038" box="[1075,1187,1881,1907]" pageId="8" pageNumber="9" value="1982-05-22">22.v.1982</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFB510E0EFAC9C038" box="[1200,1386,1881,1907]" pageId="8" pageNumber="9">M. D. Gallagher</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFF760E2AFF6DC0DC" box="[151,206,1917,1943]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFF3C0E2AFEBCC0DC" box="[221,287,1917,1943]" pageId="8" pageNumber="9">JSCS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885B182AFED00E2AFB7BC0DC" ID-GBIF-Occurrence="2350486207" box="[305,1240,1916,1943]" collectingDate="1992-02-09" collectionCode="ONHM" country="Oman" location="Harasis" municipality="Jiddat" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Al Wusta" typeStatus="holotype">
<collectingRegion id="C20C2CDF885B182AFED00E2AFD93C0DC" box="[305,560,1917,1943]" country="Oman" name="Al Wusta" pageId="8" pageNumber="9">Al Wusta Governorate</collectingRegion>
,
<collectingMunicipality id="E0137847885B182AFDDA0E2AFDDDC0DC" box="[571,638,1917,1943]" pageId="8" pageNumber="9">Jiddat</collectingMunicipality>
al
<location id="0517B4E6885B182AFD400E2AFD56C0DC" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFD400E2AFD56C0DC" box="[673,757,1917,1943]" country="Oman" municipality="Jiddat" name="Harasis" pageId="8" pageNumber="9" stateProvince="Al Wusta">Harasis</location>
(1918N 5707E,
<date id="7476C4FD885B182AFC5D0E2AFB81C0DC" box="[956,1058,1917,1943]" pageId="8" pageNumber="9" value="1992-02-09">
<collectingDate id="64323D15885B182AFC5D0E2AFB81C0DC" box="[956,1058,1917,1943]" pageId="8" pageNumber="9" value="1992-02-09">9.ii.1992</collectingDate>
</date>
,
<specimenCount id="16CE29B4885B182AFBCF0E2AFBC6C0DC" box="[1070,1125,1917,1943]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFB950E2AFB73C0DC" box="[1140,1232,1917,1943]" pageId="8" pageNumber="9">ONHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFB030E2AFDB2C0F0" ID-GBIF-Occurrence="2350486085" collectingDate="1990-02-25" collectionCode="SOIC" country="Oman" location="Yalooni" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Al Wusta Governorate" typeStatus="holotype">
<location id="0517B4E6885B182AFB030E2AFA9AC0DC" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFB030E2AFA9AC0DC" box="[1250,1337,1917,1943]" country="Oman" name="Yalooni" pageId="8" pageNumber="9" stateProvince="Al Wusta Governorate">Yalooni</location>
(1956N 5706E),
<date id="7476C4FD885B182AFF1F0EF7FED1C0F0" box="[254,370,1952,1979]" pageId="8" pageNumber="9" value="1990-02-25">
<collectingDate id="64323D15885B182AFF1F0EF7FED1C0F0" box="[254,370,1952,1979]" pageId="8" pageNumber="9" value="1990-02-25">25.ii.1990</collectingDate>
</date>
,
<specimenCount id="16CE29B4885B182AFE9F0EF6FE16C0F0" box="[382,437,1953,1979]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFE250EF6FDA9C0F0" box="[452,522,1953,1979]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="8" pageNumber="9">SOIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFDFC0EF6FCCCC0F0" ID-GBIF-Occurrence="2350486086" box="[541,879,1953,1979]" collectingDate="1992-11" collectionCode="SOIC" country="Oman" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Al Wusta Governorate" typeStatus="holotype">
ibidem,
<date id="7476C4FD885B182AFD980EF6FD73C0F0" box="[633,720,1953,1979]" pageId="8" pageNumber="9" value="1992-11">
<collectingDate id="64323D15885B182AFD980EF6FD73C0F0" box="[633,720,1953,1979]" pageId="8" pageNumber="9" value="1992-11">xi.1992</collectingDate>
</date>
,
<specimenCount id="16CE29B4885B182AFD3D0EF6FCB0C0F0" box="[732,787,1953,1979]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFCC30EF6FCCBC0F0" box="[802,872,1953,1979]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="8" pageNumber="9">SOIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182AFC9A0EF6FE51C094" ID-GBIF-Occurrence="2350486210" collectingDate="1998-04-19" collectionCode="SOIC" collectorName="M. D. Gallagher" country="Oman" location="Huqf" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Al Wusta Governorate" typeStatus="holotype">
<location id="0517B4E6885B182AFC9A0EF6FC16C0F0" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFC9A0EF6FC16C0F0" box="[891,949,1953,1979]" country="Oman" name="Huqf" pageId="8" pageNumber="9" stateProvince="Al Wusta Governorate">Huqf</location>
(202542″N 572625″E), 18/
<date id="7476C4FD885B182AFB180EF6FAD3C0F0" box="[1273,1392,1953,1979]" pageId="8" pageNumber="9" value="1998-04-19">
<collectingDate id="64323D15885B182AFB180EF6FAD3C0F0" box="[1273,1392,1953,1979]" pageId="8" pageNumber="9" value="1998-04-19">19.iv.1998</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFA9D0EF6FE84C094" pageId="8" pageNumber="9">M. D. Gallagher</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFEBE0E92FE35C094" box="[351,406,1989,2015]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885B182AFE440E92FE48C094" box="[421,491,1989,2015]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="8" pageNumber="9">SOIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B0A0E860885B182BFE1F0E92FF4EC7F8" ID-GBIF-Occurrence="2350486143" collectingDate="1983-05-13" collectionCode="SOIC" collectorName="M. D. Gallagher" country="Oman" lastPageId="9" lastPageNumber="10" location="Sawairah" municipality="Sahil Al Tazir" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Al Wusta Governorate" typeStatus="holotype">
<collectingMunicipality id="E0137847885B182AFE1F0E92FD39C094" box="[510,666,1989,2015]" pageId="8" pageNumber="9">Sahil Al Tazir</collectingMunicipality>
,
<location id="0517B4E6885B182AFD440E92FCAEC094" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885B182AFD440E92FCAEC094" box="[677,781,1989,2015]" country="Oman" municipality="Sahil Al Tazir" name="Sawairah" pageId="8" pageNumber="9" stateProvince="Al Wusta Governorate">Sawairah</location>
(1808N 5632E) 10/
<date id="7476C4FD885B182AFC1C0E92FBC8C094" box="[1021,1131,1989,2015]" pageId="8" pageNumber="9" value="1983-05-13">
<collectingDate id="64323D15885B182AFC1C0E92FBC8C094" box="[1021,1131,1989,2015]" pageId="8" pageNumber="9" value="1983-05-13">13.v.1983</collectingDate>
</date>
,
<collectorName id="AD3D87EB885B182AFB960E92FA8DC094" box="[1143,1326,1989,2015]" pageId="8" pageNumber="9">M. D. Gallagher</collectorName>
leg.
<specimenCount id="16CE29B4885B182AFA870E92FA3FC094" box="[1382,1436,1989,2015]" pageId="8" pageNumber="9" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885A182BFF7E09CEFF46C7F8" box="[159,229,153,179]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="9" pageNumber="10">SOIC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885A182BFF1B09CEFA9EC7F8" ID-GBIF-Occurrence="2350486311" box="[250,1341,153,179]" collectingDate="2017-11-24" collectionCode="SZCM" collectorName="T. Lackner" country="Oman" location="Ash-Shuwayiyah" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Zufar" typeStatus="holotype">
<collectingRegion id="C20C2CDF885A182BFF1B09CEFE41C7F8" box="[250,482,153,179]" country="Oman" name="Zufar" pageId="9" pageNumber="10">Dhofar Governorate</collectingRegion>
,
<location id="0517B4E6885A182BFE0C09CEFD12C7F8" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885A182BFE0C09CEFD12C7F8" box="[493,689,153,179]" country="Oman" name="Ash-Shuwayiyah" pageId="9" pageNumber="10" stateProvince="Zufar">Ash-Shuwayiyah</location>
, on the beach,
<date id="7476C4FD885A182BFC8009CEFC7FC7F8" box="[865,988,153,179]" pageId="9" pageNumber="10" value="2017-11-24">
<collectingDate id="64323D15885A182BFC8009CEFC7FC7F8" box="[865,988,153,179]" pageId="9" pageNumber="10" value="2017-11-24">24.xi.2017</collectingDate>
</date>
,
<collectorName id="AD3D87EB885A182BFC0809CEFBC7C7F8" box="[1001,1124,153,179]" pageId="9" pageNumber="10">T. Lackner</collectorName>
leg
<specimenCount id="16CE29B4885A182BFB7909CEFB72C7F8" box="[1176,1233,153,179]" pageId="9" pageNumber="10" type="generic">1 ex.</specimenCount>
(
<collectionCode id="66D97AF8885A182BFB0309CEFA97C7F8" box="[1250,1332,153,179]" pageId="9" pageNumber="10">SZCM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B0A0E860885A182BFAAB09CEFA90C79C" ID-GBIF-Occurrence="2350486075" collectingDate="1996-04-05" collectionCode="SOIC" collectorName="M. D. Gallagher" country="Oman" elevation="20" location="Fins" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Masqat" typeStatus="holotype">
<collectingRegion id="C20C2CDF885A182BFAAB09CEFE85C79C" country="Oman" name="Masqat" pageId="9" pageNumber="10">Masqat Governorate</collectingRegion>
, near
<location id="0517B4E6885A182BFE8909EAFE38C79C" LSID="urn:lsid:plazi:treatment:8861532B885B182BFF760C36FC8CC3E8:0517B4E6885A182BFE8909EAFE38C79C" box="[360,411,189,215]" country="Oman" name="Fins" pageId="9" pageNumber="10" stateProvince="Masqat">Fins</location>
,
<quantity id="C7304FD8885A182BFE4609EBFDAFC79C" box="[423,524,188,215]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="9" pageNumber="10" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">
<elevation id="8BE5050E885A182BFE4609EBFDAFC79C" box="[423,524,188,215]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="9" pageNumber="10" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">1525 m</elevation>
</quantity>
, (225653N 591117E), 4/
<date id="7476C4FD885A182BFCD109EBFC3BC79C" box="[816,920,188,215]" pageId="9" pageNumber="10" value="1996-04-05">
<collectingDate id="64323D15885A182BFCD109EBFC3BC79C" box="[816,920,188,215]" pageId="9" pageNumber="10" value="1996-04-05">5.iv.1996</collectingDate>
</date>
,
<collectorName id="AD3D87EB885A182BFC4209EAFBF9C79C" box="[931,1114,189,215]" pageId="9" pageNumber="10">M. D. Gallagher</collectorName>
leg. 8 exx. (
<collectionCode id="66D97AF8885A182BFB0709EAFA8FC79C" box="[1254,1324,189,215]" country="Oman" httpUri="http://grbio.org/cool/txkx-ji9v" name="Natural History Museum, National Insect Collection" pageId="9" pageNumber="10">SOIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D885A182BFF2609B7FAC2C60F" blockId="9.[151,1437,153,1187]" pageId="9" pageNumber="10">
<emphasis id="32BC3E2F885A182BFF2609B7FEFAC7B0" bold="true" box="[199,345,224,251]" pageId="9" pageNumber="10">Distribution</emphasis>
.
<emphasis id="32BC3E2F885A182BFE8A09B7FDEEC7B0" bold="true" box="[363,589,224,251]" pageId="9" pageNumber="10">Arabian Peninsula</emphasis>
:
<collectingCountry id="78DFA2AD885A182BFDBE09B6FD00C7B0" box="[607,675,225,251]" name="Oman" pageId="9" pageNumber="10">Oman</collectingCountry>
and
<collectingCountry id="78DFA2AD885A182BFD3E09B6FC8EC7B0" box="[735,813,225,251]" name="Yemen" pageId="9" pageNumber="10">Yemen</collectingCountry>
(Scott
<emphasis id="32BC3E2F885A182BFC6509B5FC39C7B0" box="[900,922,226,251]" italics="true" pageId="9" pageNumber="10">in</emphasis>
<bibRefCitation id="64599FCC885A182BFC4509B6FB9EC7B0" author="Paulian, R." box="[932,1085,225,251]" pageId="9" pageNumber="10" pagination="141 - 155" refId="ref39189" refString="Paulian, R. (1948) Expedition to South-West Arabia 1937 - 8. 12. Coleoptera, Scarabaeidae: Trogidae, Geotrupinae, Dynamopinae, Hybosorinae, Coprinae, Aphodiinae. British Museum (Natural History), 1, 141 - 155." type="journal article" year="1948">Paulian 1948</bibRefCitation>
),
<collectingCountry id="78DFA2AD885A182BFBB509B6FB4BC7B0" box="[1108,1256,225,251]" name="Saudi Arabia" pageId="9" pageNumber="10">
Saudi
<taxonomicName id="C7C899BE885A182BFB7D09B6FB4BC7B0" authority="(Harold 1867)" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[1180,1256,225,251]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFB1B09B6FA33C7B7" author="Harold, E." box="[1274,1424,225,252]" pageId="9" pageNumber="10" pagination="23 - 59" refId="ref37904" refString="Harold, E. (1867) Beitrage zur Kenntniss der Gattung Onthophagus vom Herausgeber. I. Theil: Australier und Afrikaner. Coleopterologische Hefte, 2, 23 - 59." type="journal article" year="1867">Harold 1867</bibRefCitation>
),
<collectingCountry id="78DFA2AD885A182BFF760852FE31C654" box="[151,402,261,287]" name="United Arab Emirates" pageId="9" pageNumber="10">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFE430852FD07C654" author="Gillett, M. P. T. &amp; Gillett, C. P. D. T." box="[418,676,260,287]" pageId="9" pageNumber="10" pagination="352 - 356" refId="ref37609" refString="Gillett, M. P. T. &amp; Gillett, C. P. D. T. (2005) Insects &amp; other arthropods, pp. 168 - 194. Species checklist, In: Aspinall, S. &amp; Hellyer, P. (Eds.), The Emirates. A natural history. P. Vine, Trident Press, London, pp. 352 - 356." type="book chapter" year="2005">Gillett &amp; Gillett 2005</bibRefCitation>
).
<emphasis id="32BC3E2F885A182BFD580852FCF6C654" bold="true" box="[697,853,260,287]" pageId="9" pageNumber="10">North Africa</emphasis>
:
<collectingCountry id="78DFA2AD885A182BFC840852FC1FC654" box="[869,956,261,287]" name="Algeria" pageId="9" pageNumber="10">Algeria</collectingCountry>
,
<collectingCountry id="78DFA2AD885A182BFC2B0852FBB3C654" box="[970,1040,261,287]" name="Egypt" pageId="9" pageNumber="10">Egypt</collectingCountry>
,
<collectingCountry id="78DFA2AD885A182BFBFF0852FB25C654" box="[1054,1158,261,287]" name="Morocco" pageId="9" pageNumber="10">Morocco</collectingCountry>
,
<collectingCountry id="78DFA2AD885A182BFB720852FB45C654" box="[1171,1254,261,287]" name="Tunisia" pageId="9" pageNumber="10">Tunisia</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFB160852FA33C654" author="Baraud, J." box="[1271,1424,260,287]" pageId="9" pageNumber="10" refId="ref36685" refString="Baraud, J. (1985) Coleopteres Scarabaeoidea, Faune du Nord de l'Afrique, du Maroc au Sinai. Editions Lechevalier, Paris, 650 pp." type="book" year="1985">Baraud 1985</bibRefCitation>
).
<emphasis id="32BC3E2F885A182BFF76087FFF40C608" bold="true" box="[151,227,296,323]" pageId="9" pageNumber="10">Africa</emphasis>
:
<collectingCountry id="78DFA2AD885A182BFF13087EFEE2C608" box="[242,321,297,323]" name="Eritrea" pageId="9" pageNumber="10">Eritrea</collectingCountry>
,
<collectingCountry id="78DFA2AD885A182BFEAD087EFE0BC608" box="[332,424,297,323]" name="Somalia" pageId="9" pageNumber="10">Somalia</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFE56087EFDC7C60F" author="Balthasar, V." box="[439,612,297,324]" pageId="9" pageNumber="10" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar 1963</bibRefCitation>
),
<collectingCountry id="78DFA2AD885A182BFD98087EFD61C608" box="[633,706,297,323]" name="Kenya" pageId="9" pageNumber="10">Kenya</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFD31087EFC68C608" author="Tagliaferri, F. &amp; Moretto, P. &amp; Tarasov, S." box="[720,971,297,323]" pageId="9" pageNumber="10" pagination="1 - 31" refId="ref39627" refString="Tagliaferri, F., Moretto, P. &amp; Tarasov, S. (2012) Essai sur la systematique et la phylogenie des Onthophagus Latreille, 1802, d'Afrique tropicale appartenant au septieme groupe de d'Orbigny. Description d'un sous-genre nouveau et de trois especes nouvelles (Coleoptera, Scarabaeoidea, Onthophagini). Catharsius La Revue, 6, 1 - 31." type="journal article" year="2012">
Tagliaferri
<emphasis id="32BC3E2F885A182BFCAE087DFC23C608" box="[847,896,297,323]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 2012
</bibRefCitation>
).
<emphasis id="32BC3E2F885A182BFC3C087FFBB1C608" bold="true" box="[989,1042,296,323]" pageId="9" pageNumber="10">Asia</emphasis>
:
<collectingCountry id="78DFA2AD885A182BFBC0087EFBEDC608" box="[1057,1102,297,323]" name="Iran" pageId="9" pageNumber="10">Iran</collectingCountry>
(
<bibRefCitation id="64599FCC885A182BFBBD087EFAF6C60F" author="Ziani, S. &amp; Bezdek, A." box="[1116,1365,297,324]" pageId="9" pageNumber="10" pagination="180 - 204" refId="ref39999" refString="Ziani, S. &amp; Bezdek, A. (2016) Scarabaeidae, subfamily Scarabaeinae, tribe Onthophagini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill, Leiden-Boston, pp. 180 - 204." type="book chapter" year="2016">Ziani &amp; Bezděk 2016</bibRefCitation>
).
</paragraph>
<paragraph id="0077E23D885A182BFF26081AFA96C5CC" blockId="9.[151,1437,153,1187]" pageId="9" pageNumber="10">
<emphasis id="32BC3E2F885A182BFF26081AFE97C62C" bold="true" box="[199,308,333,359]" pageId="9" pageNumber="10">Remarks</emphasis>
.
<taxonomicName id="C7C899BE885A182BFEA3081AFDF7C62C" authorityName="Frey" authorityYear="1962" box="[322,596,333,359]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saudiensis">
<emphasis id="32BC3E2F885A182BFEA3081AFDF7C62C" box="[322,596,333,359]" italics="true" pageId="9" pageNumber="10">Onthophagus saudiensis</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="64599FCC885A182BFCC1081AFC06C623" author="Frey, G." box="[800,933,333,360]" pageId="9" pageNumber="10" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey (1962)</bibRefCitation>
from two specimens from
<collectingRegion id="C20C2CDF885A182BFB31081AFA3BC62C" box="[1232,1432,333,359]" country="Saudi Arabia" name="Makkah" pageId="9" pageNumber="10">Makkah Province</collectingRegion>
,
<collectingCountry id="78DFA2AD885A182BFF760826FE84C6C0" box="[151,295,369,395]" name="Saudi Arabia" pageId="9" pageNumber="10">
Saudi
<taxonomicName id="C7C899BE885A182BFF3B0826FE84C6C0" baseAuthorityName="Harold" baseAuthorityYear="1867" box="[218,295,369,395]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="35" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
</collectingCountry>
, and placed in the dOrbigny (1913)s 29
<superScript id="F7BD4F75885A182BFD000827FD4DC635" attach="left" box="[737,750,368,382]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
group, that Frey himself called “29. Gruppe
<emphasis id="32BC3E2F885A182BFB310826FA93C6C0" box="[1232,1328,369,395]" italics="true" pageId="9" pageNumber="10">ochreatu</emphasis>
s dOrb.”. Actually, for
<taxonomicName id="C7C899BE885A182BFEC608C2FDCEC6E4" authority="d'Orbigny, 1898" authorityName="d'Orbigny" authorityYear="1898" box="[295,621,405,431]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ochreatus">
<emphasis id="32BC3E2F885A182BFEC608C2FE11C6E4" box="[295,434,405,431]" italics="true" pageId="9" pageNumber="10">O. ochreatus</emphasis>
dOrbigny, 1898
</taxonomicName>
and others species, almost thirty years before, Balthasar (1935) erected the subgenus
<taxonomicName id="C7C899BE885A182BFEE508EEFE65C698" authorityName="Balthasar" authorityYear="1963" box="[260,454,441,467]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Micronthophagus">
<emphasis id="32BC3E2F885A182BFEE508EEFE65C698" box="[260,454,441,467]" italics="true" pageId="9" pageNumber="10">Micronthophagus</emphasis>
</taxonomicName>
. For these reasons
<bibRefCitation id="64599FCC885A182BFD7808EEFC99C698" author="Zunino, M." box="[665,826,441,467]" pageId="9" pageNumber="10" pagination="408 - 416" refId="ref40252" refString="Zunino, M. (1981) Insects of Saudi Arabia Coleoptera: Fam. Scarabaeidae Tribe Onthophagini. Fauna of Saudi Arabia, 3, 408 - 416." type="journal article" year="1981">Zunino (1981)</bibRefCitation>
, not knowing
<emphasis id="32BC3E2F885A182BFC3608EEFBD2C698" box="[983,1137,441,467]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="C7C899BE885A182BFC3608EEFBCFC698" authorityName="Frey" authorityYear="1962" box="[983,1132,441,467]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saudiensis">O. saudiensis</taxonomicName>
,
</emphasis>
tentatively placed this species in the subgenus
<taxonomicName id="C7C899BE885A182BFE96088AFD4AC6BC" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" box="[375,745,477,503]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Micronthophagus">
<emphasis id="32BC3E2F885A182BFE96088AFD9AC6BC" box="[375,569,477,503]" italics="true" pageId="9" pageNumber="10">Micronthophagus</emphasis>
<bibRefCitation id="64599FCC885A182BFDDE088AFD4AC6BC" author="Balthasar, V." box="[575,745,477,503]" pageId="9" pageNumber="10" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar, 1963</bibRefCitation>
</taxonomicName>
. As said above, the type series is constituted by two specimens, identified as males by
<bibRefCitation id="64599FCC885A182BFE6C0B56FDACC557" author="Frey, G." box="[397,527,513,540]" pageId="9" pageNumber="10" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey (1962)</bibRefCitation>
but actually two females. The author clearly stated (
<bibRefCitation id="64599FCC885A182BFBB20B56FB69C550" author="Frey, G." box="[1107,1226,513,539]" pageId="9" pageNumber="10" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey 1962</bibRefCitation>
, p. 619): “Type im Britischen Museum,
<typeStatus id="DF735C9F885A182BFE600B72FE40C574" box="[385,483,549,575]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
im meinem Museum” [Type in British Museum,
<typeStatus id="DF735C9F885A182BFBEC0B72FBCCC574" box="[1037,1135,549,575]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
in my Museum], discriminating between “Type” [=
<typeStatus id="DF735C9F885A182BFE210B1FFD86C528" box="[448,549,584,611]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
] and “
<typeStatus id="DF735C9F885A182BFD8A0B1EFD75C528" box="[619,726,585,611]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
”. Therefore, article 73.1.1. of the Code (
<bibRefCitation id="64599FCC885A182BFB760B1EFAB9C528" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[1175,1306,585,611]" pageId="9" pageNumber="10" refId="ref38072" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
) is met and the
<typeStatus id="DF735C9F885A182BFF210B3AFE81C5CC" box="[192,290,621,647]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is the specimen housed in BMNH, while specimen deposited in Freys collection is a
<typeStatus id="DF735C9F885A182BFB310B3AFA91C5CC" box="[1232,1330,621,647]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="0077E23D885A182BFF260BC6FDA8C5E0" blockId="9.[151,1437,153,1187]" box="[199,523,657,683]" pageId="9" pageNumber="10">They are labelled as follows:</paragraph>
<paragraph id="0077E23D885A182BFF260BE2FE32C4C8" blockId="9.[151,1437,153,1187]" pageId="9" pageNumber="10">
<materialsCitation id="B0A0E860885A182BFF260BE2FE32C4C8" ID-GBIF-Occurrence="2350486308" collectingDate="1944-12-08" collectionCode="NHMB" collectorName="A. R. Waterston. &amp; P. Typus" determinerName="G. Frey" location="Mus" pageId="9" pageNumber="10" specimenCount="2" specimenCount-female="2" stateProvince="Mus" typeStatus="holotype">
<typeStatus id="DF735C9F885A182BFF260BE2FE91C584" box="[199,306,693,719]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
, fixed by original designation (
<collectionCode id="66D97AF8885A182BFD7A0BE2FD55C584" box="[667,758,693,719]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="9" pageNumber="10">NHMB</collectionCode>
): 1st, white with a horizontal yellow line, printed in black: “HEJAZ / Buraiman. /
<quantity id="C7304FD8885A182BFE780B8EFE5AC5B8" box="[409,505,729,755]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="9" pageNumber="10" unit="mi" value="12.0">12 miles</quantity>
N. of Jidda. /
<date id="7476C4FD885A182BFD760B8EFCAFC5B8" box="[663,780,729,755]" pageId="9" pageNumber="10" value="1944-12-08">
<collectingDate id="64323D15885A182BFD760B8EFCAFC5B8" box="[663,780,729,755]" pageId="9" pageNumber="10" value="1944-12-08">8.xii.1944</collectingDate>
</date>
. /
<collectorName id="AD3D87EB885A182BFCC20B8EFC7DC5B8" box="[803,990,729,755]" pageId="9" pageNumber="10">A. R. Waterston.</collectorName>
/ B. M. 1947-349.”; 2nd, red, printed in black: “
<typeStatus id="DF735C9F885A182BFF0D0BAAFE37C45C" box="[236,404,765,791]" pageId="9" pageNumber="10" type="paratype">PARATYPUS</typeStatus>
”; 3rd, white, handwritten in blue and printed in black: “
<collectorName id="AD3D87EB885A182BFC1D0BAAFBF8C45C" box="[1020,1115,765,791]" pageId="9" pageNumber="10">P. Typus</collectorName>
/
<taxonomicName id="C7C899BE885A182BFB910BAAFAA6C45C" authorityName="Latreille" authorityYear="1802" box="[1136,1285,765,791]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
/ saudiensis / n sp / det.
<determinerName id="F5AB0EF9885A182BFEEA0A76FEC2C470" box="[267,353,801,827]" pageId="9" pageNumber="10">G. Frey</determinerName>
, 1961”; 4th, white, printed and handwritten in black: “
<taxonomicName id="C7C899BE885A182BFC200A76FB14C470" authority="(Erem.)" baseAuthorityName="Erem." box="[961,1207,801,827]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Onthophagus (Erem.)</taxonomicName>
/ sticticus HAR.
<specimenCount id="16CE29B4885A182BFA990A76FA2EC470" box="[1400,1421,801,827]" pageId="9" pageNumber="10" type="female"></specimenCount>
/ J. SCHEUERN det. 98”; 5th, red handwritten in black: “comb. with / TYPE
<specimenCount id="16CE29B4885A182BFC100A12FBA5C414" box="[1009,1030,837,863]" pageId="9" pageNumber="10" type="female"></specimenCount>
/
<taxonomicName id="C7C899BE885A182BFBFA0A12FB7FC414" authority="FREY" authorityName="Frey" authorityYear="1962" box="[1051,1244,837,863]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saudiensis">saudiensis FREY</taxonomicName>
/ Brit.
<collectingRegion id="C20C2CDF885A182BFACA0A12FAC3C414" box="[1323,1376,837,863]" country="Turkey" name="Mus" pageId="9" pageNumber="10">Mus</collectingRegion>
. 2.99 / J. SCHEUERN vid”.
</materialsCitation>
</paragraph>
<paragraph id="0077E23D885A182BFF260ADAFA8AC310" blockId="9.[151,1437,153,1187]" pageId="9" pageNumber="10">
<materialsCitation id="B0A0E860885A182BFF260ADAFA86C310" ID-GBIF-Occurrence="2350486202" collectingDate="1944-12-04" collectionCode="NHMUK" collectorName="A. R. Waterston." determinerName="G. Frey &amp; S. Ziani" pageId="9" pageNumber="10" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="DF735C9F885A182BFF260ADAFE8AC4EC" box="[199,297,909,935]" pageId="9" pageNumber="10" type="paratype">Paratype</typeStatus>
(
<collectionCode id="66D97AF8885A182BFEDD0ADAFE0EC4EC" box="[316,429,909,935]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="10" type="Museum">NHMUK</collectionCode>
): 1st, white with a horizontal yellow line, printed in black: “HEJAZ / Buraiman. /
<quantity id="C7304FD8885A182BFA610ADAFF77C480" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="9" pageNumber="10" unit="mi" value="12.0">12 miles</quantity>
N. of Jidda. /
<date id="7476C4FD885A182BFE980AE6FE4EC480" box="[377,493,945,971]" pageId="9" pageNumber="10" value="1944-12-04">
<collectingDate id="64323D15885A182BFE980AE6FE4EC480" box="[377,493,945,971]" pageId="9" pageNumber="10" value="1944-12-04">4.xii.1944</collectingDate>
</date>
. /
<collectorName id="AD3D87EB885A182BFDE90AE6FD66C480" box="[520,709,945,971]" pageId="9" pageNumber="10">A. R. Waterston.</collectorName>
/ B. M. 1947-349.”; 2nd, round, white, encircled of red, printed in black: “Type”; 3rd, red, printd in black: “TYPUS”; 4th, white, handwritten in blue and printd in black: “Typus
<specimenCount id="16CE29B4885A182BFF760AAEFF0FC358" box="[151,172,1017,1043]" pageId="9" pageNumber="10" type="male"></specimenCount>
/
<taxonomicName id="C7C899BE885A182BFF230AAEFEF4C358" box="[194,343,1017,1043]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
/ saudiensis / n sp / det.
<determinerName id="F5AB0EF9885A182BFD8B0AAEFD63C358" box="[618,704,1017,1043]" pageId="9" pageNumber="10">G. Frey</determinerName>
, 1961”; 5th, white, handwritten in blue: “Synonymous / with /
<taxonomicName id="C7C899BE885A182BFA600AAEFF5AC37C" authorityName="Harold" authorityYear="1867" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sticticus">O. sticticus</taxonomicName>
”; 6th, white, printed and handwritten in black: “
<taxonomicName id="C7C899BE885A182BFCEE0D4AFBA7C37C" authority="(Erem.)" baseAuthorityName="Erem." box="[783,1028,1053,1079]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Onthophagus (Erem.)</taxonomicName>
/ sticticus HAR.
<specimenCount id="16CE29B4885A182BFB240D4AFB79C37C" box="[1221,1242,1053,1079]" pageId="9" pageNumber="10" type="female"></specimenCount>
/ J. SCHEUERN det. 98”; 7th, white, handwritten and printed in black: “
<taxonomicName id="C7C899BE885A182BFCE00D16FC3AC310" box="[769,921,1089,1115]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
/ sticticus HAR. /
<determinerName id="F5AB0EF9885A182BFB8B0D16FB61C310" box="[1130,1218,1089,1115]" pageId="9" pageNumber="10">S. Ziani</determinerName>
det. 19”
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D885A182BFF260D32FC8CC3E8" blockId="9.[151,1437,153,1187]" pageId="9" pageNumber="10">
After the examination of both the specimens, we can propose the new synonymy
<taxonomicName id="C7C899BE885A182BFBA80D32FF6CC3E8" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="32BC3E2F885A182BFBA80D32FAE1C334" box="[1097,1346,1125,1151]" italics="true" pageId="9" pageNumber="10">Onthophagus sticticus</emphasis>
<bibRefCitation id="64599FCC885A182BFAA60D32FF6CC3E8" author="Harold, E." pageId="9" pageNumber="10" pagination="23 - 59" refId="ref37904" refString="Harold, E. (1867) Beitrage zur Kenntniss der Gattung Onthophagus vom Herausgeber. I. Theil: Australier und Afrikaner. Coleopterologische Hefte, 2, 23 - 59." type="journal article" year="1867">Harold, 1867</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="C7C899BE885A182BFF0C0DDEFDDDC3E8" authority="Frey, 1962" authorityName="Frey" authorityYear="1962" box="[237,638,1161,1187]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="saudiensis">
<emphasis id="32BC3E2F885A182BFF0C0DDEFDA3C3E8" box="[237,512,1161,1187]" italics="true" pageId="9" pageNumber="10">Onthophagus saudiensis</emphasis>
<bibRefCitation id="64599FCC885A182BFDE60DDEFDDDC3E8" author="Frey, G." box="[519,638,1161,1187]" pageId="9" pageNumber="10" pagination="616 - 622" refId="ref37396" refString="Frey, G. (1962) Neue Onthophagus - Arten (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 13 (2), 616 - 622." type="journal article" year="1962">Frey, 1962</bibRefCitation>
</taxonomicName>
<emphasis id="32BC3E2F885A182BFD640DDEFC84C3E8" bold="true" box="[645,807,1161,1187]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="298F8354885A182BFD640DDEFC84C3E8" box="[645,807,1161,1187]" pageId="9" pageNumber="10">new synonym</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>