An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae) Author Ziani, Stefano Author Abdel-Dayem, Mahmoud Saleh Author Aldhafer, Hathal Mohammed Author Barbero, Enrico text Zootaxa 2019 2019-08-21 4658 1 1 36 journal article 25961 10.11646/zootaxa.4658.1.1 31d6e081-95c3-4086-a45a-c11555dda3aa 1175-5326 3373040 ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21 Onthophagus ( Eremonthophagus ) sticticus Harold, 1867 [ Fig. 6 ] Literature quotes . SAUDI ARABIA : “ Arabia ” ( Harold 1867 ). Makkah Province , “Djedda” (d’Orbigny 1900); “Buraiman”; “Ta’if”; “ Mecca ”; “Al Lith”; “Shi’aera [Al Shaira], antilocust camp north of Qunfidha” (Scott in Paulian 1948 ). Madinah Province , “ 16 km W Badr Hunayn”. Hejaz, “N Hedjaz” ( Zunino 1981 ). Yemen: without locality (Scott in Paulian 1948 ). “Nord Yemen” ( Paulian 1980 ). Oman: Muscat Governorate, “Muscat” (Scott in Paulian 1948 ). Without locality ( Janikova 1998 ). UNITED ARAB EMIRATES : without locality ( Gillett & Gillett 2005 ). Material examined . SAUDI ARABIA : 1 ex. with no further data ( JSCS ) . Jizan Province , Farasan islands , Al- Sajid , ( 16.860626°N , 41.932564°E ), 17.ii.2017 , U.M. Abu El-Ghiet & T. M. Elsheikh leg. 1 ♀ ( KSMA ) ; ibidem , 10.x.2017 , Abu El-Ghiet & T. M. Elsheikh leg. 3 ♂♂ ( KSMA ) . Makkah Province , Hedjas , Buraiman , 12 miles N. of Jidda , 4.xii.1944 , A. R. Waterston leg. 1 ♀ ( paratype of O. saudensis Frey, 1962 , NHMUK ) ; ibidem , 8.xii.1944 , A. R. Waterston leg. 1 ♀ ( holotype of O. saudiensis Frey, 1962 , NHMB ) . YEMEN : Sana’a Governorate , Sana’a , 2400 m , C. Rathjens leg. 1 ex. ( NHMUK ) ; Mafhaq , 2350 m (1509′03″N 4357′16″E), 7.x.2005 , D. Král leg. 1 ex. ( NMPC ) . OMAN : Al Batinah Governorate , Batinah b. Sohar (2425′N 5642′E), 18/ 22.v.1982 , M. D. Gallagher leg. 1 ex. ( JSCS ) . Al Wusta Governorate , Jiddat al Harasis (1918′N 5707′E, 9.ii.1992 , 1 ex. ( ONHM ) ; Yalooni (1956′N 5706′E), 25.ii.1990 , 1 ex. ( SOIC ) ; ibidem, xi.1992 , 1 ex. ( SOIC ) ; Huqf (2025′42″N 5726′25″E), 18/ 19.iv.1998 , M. D. Gallagher leg. 1 ex. ( SOIC ) ; Sahil Al Tazir , Sawairah (1808′N 5632′E) 10/ 13.v.1983 , M. D. Gallagher leg. 1 ex. ( SOIC ) . Dhofar Governorate , Ash-Shuwayiyah , on the beach, 24.xi.2017 , T. Lackner leg 1 ex. ( SZCM ) . Masqat Governorate , near Fins , 15–25 m , (2256′53N 5911′17E), 4/ 5.iv.1996 , M. D. Gallagher leg. 8 exx. ( SOIC ) . Distribution . Arabian Peninsula : Oman and Yemen (Scott in Paulian 1948 ), Saudi Arabia ( Harold 1867 ), United Arab Emirates ( Gillett & Gillett 2005 ). North Africa : Algeria , Egypt , Morocco , Tunisia ( Baraud 1985 ). Africa : Eritrea , Somalia ( Balthasar 1963 ), Kenya ( Tagliaferri et al . 2012 ). Asia : Iran ( Ziani & Bezděk 2016 ). Remarks . Onthophagus saudiensis was described by Frey (1962) from two specimens from Makkah Province , Saudi Arabia , and placed in the d’Orbigny (1913)’s 29 th group, that Frey himself called “29. Gruppe ochreatu s d’Orb.”. Actually, for O. ochreatus d’Orbigny, 1898 and others species, almost thirty years before, Balthasar (1935) erected the subgenus Micronthophagus . For these reasons Zunino (1981) , not knowing O. saudiensis , tentatively placed this species in the subgenus Micronthophagus Balthasar, 1963 . As said above, the type series is constituted by two specimens, identified as males by Frey (1962) but actually two females. The author clearly stated ( Frey 1962 , p. 619): “Type im Britischen Museum, Paratype im meinem Museum” [Type in British Museum, Paratype in my Museum], discriminating between “Type” [= holotype ] and “ Paratype ”. Therefore, article 73.1.1. of the Code ( ICZN 1999 ) is met and the holotype is the specimen housed in BMNH, while specimen deposited in Frey’s collection is a paratype . They are labelled as follows: Holotype , fixed by original designation ( NHMB ): 1st, white with a horizontal yellow line, printed in black: “HEJAZ / Buraiman. / 12 miles N. of Jidda. / 8.xii.1944 . / A. R. Waterston. / B. M. 1947-349.”; 2nd, red, printed in black: “ PARATYPUS ”; 3rd, white, handwritten in blue and printed in black: “ P. Typus / Onthophagus / saudiensis / n sp / det. G. Frey , 1961”; 4th, white, printed and handwritten in black: “ Onthophagus (Erem.) / sticticus HAR. ♀ / J. SCHEUERN det. 98”; 5th, red handwritten in black: “comb. with / TYPE ♀ / saudiensis FREY / Brit. Mus . 2.99 / J. SCHEUERN vid”. Paratype ( NHMUK ): 1st, white with a horizontal yellow line, printed in black: “HEJAZ / Buraiman. / 12 miles N. of Jidda. / 4.xii.1944 . / A. R. Waterston. / B. M. 1947-349.”; 2nd, round, white, encircled of red, printed in black: “Type”; 3rd, red, printd in black: “TYPUS”; 4th, white, handwritten in blue and printd in black: “Typus ♂ / Onthophagus / saudiensis / n sp / det. G. Frey , 1961”; 5th, white, handwritten in blue: “Synonymous / with / O. sticticus ”; 6th, white, printed and handwritten in black: “ Onthophagus (Erem.) / sticticus HAR. ♀ / J. SCHEUERN det. 98”; 7th, white, handwritten and printed in black: “ Onthophagus / sticticus HAR. / S. Ziani det. 19” . After the examination of both the specimens, we can propose the new synonymy Onthophagus sticticus Harold, 1867 = Onthophagus saudiensis Frey, 1962 new synonym .