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<mods:title id="C699C46F3DAA24D2A14D55B6C62C30D4">Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="0F5CE9E178CBDC6474305C6A9BCF6A02">Fei, Xu-Dong</mods:namePart>
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8.
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<emphasis id="C48FEADFE3572F7AFF0CFDD7FE95035D" bold="true" box="[181,370,520,547]" italics="true" pageId="15" pageNumber="488">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE3572F7AFE3AFDD7FDDF035C" bold="true" box="[387,568,520,546]" italics="true" pageId="15" pageNumber="488">Eumorimotous</emphasis>
)
<emphasis id="C48FEADFE3572F7AFDF0FDD7FCA3035D" bold="true" box="[585,836,520,547]" italics="true" pageId="15" pageNumber="488">laevigatus laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE3572F7AFCEDFDD7FC12035C" author="Fauvel, A." box="[852,1013,520,546]" pageId="15" pageNumber="488" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
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First record from
<collectingCountry id="8EEC765DE3572F7AFEC0FD8FFE210314" box="[377,454,592,618]" name="China" pageId="15" pageNumber="488">China</collectingCountry>
and
<collectingCountry id="8EEC765DE3572F7AFDBAFD8FFD8A0314" box="[515,621,592,618]" name="Vietnam" pageId="15" pageNumber="488">Vietnam</collectingCountry>
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<paragraph id="F64436CDE3572F7AFF2EFDABFE6F03F1" blockId="15.[151,621,592,655]" box="[151,392,628,655]" pageId="15" pageNumber="488">
(
<figureCitation id="6EC02A48E3572F7AFF26FDABFEC103F1" box="[159,294,628,655]" captionStart="FIGURE 9" captionStartId="15.[151,250,1945,1969]" captionTargetBox="[184,1407,998,1920]" captionTargetId="figure-252@15.[173,1415,990,1922]" captionTargetPageId="15" captionText="FIGURE 9. Eccoptolonthus (Eumorimotous) laevigatus laevigatus (Fauvel, 1895) A. Habitus; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. underside of paramere; E. stylus of tergite IX of male; F. sternite VIII of male; G. tergite X of male; H. sternite IX of male. Scale bars, A=2 mm; B, C, D, E, F, G, H= 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4640497" httpUri="https://zenodo.org/record/4640497/files/figure.png" pageId="15" pageNumber="488">Figs. 9AH</figureCitation>
,
<figureCitation id="6EC02A48E3572F7AFE88FDAAFE6703F1" box="[305,384,629,655]" captionStart="FIGURE 10" captionStartId="16.[151,250,1941,1965]" captionTargetBox="[203,1372,181,1912]" captionTargetId="figure-25@16.[202,1385,181,1917]" captionTargetPageId="16" captionText="FIGURE 10. Eccoptolonthus (Eumorimotous) laevigatus laevigatus (Fauvel, 1895) A. median lobe of aedeagus, dorsal (parameral side) view; B. underside of paramere; C. tergite X of male; D. sternite VIII of male; E. sternite IX of male; F. stylus of tergite IX of male. Scale bars, A= 0.14 mm; B, C, E= 0.2 mm; D= 0.34 mm; F= 0.27 mm." figureDoi="http://doi.org/10.5281/zenodo.4640501" httpUri="https://zenodo.org/record/4640501/files/figure.png" pageId="15" pageNumber="488">10AF</figureCitation>
)
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<bibRefCitation id="926A4B3CE3572F7AFF2EFD66FEA803AF" author="Fauvel, A." box="[151,335,697,721]" pageId="15" pageNumber="488" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895: 259</bibRefCitation>
(
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<emphasis id="C48FEADFE3572F7AFEE7FD65FE5903AF" box="[350,446,698,721]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
,
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locality:
<collectingCountry id="8EEC765DE3572F7AFDEFFD66FD5003AF" box="[598,695,697,721]" name="Myanmar" pageId="15" pageNumber="488">Bírmanie</collectingCountry>
, Carin Cheba,
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; Pulo-Penang;
<collectingCountry id="8EEC765DE3572F7AFBD0FD66FB2503AF" box="[1129,1218,697,721]" name="Malaysia" pageId="15" pageNumber="488">Malacca</collectingCountry>
;
<collectingCountry id="8EEC765DE3572F7AFB75FD66FAD103AF" box="[1228,1334,697,721]" name="Singapore" pageId="15" pageNumber="488">Singapore</collectingCountry>
; Sumatra; Borneo; Java);
<bibRefCitation id="926A4B3CE3572F7AFEDCFD07FD250391" author="Bernhauer, M. &amp; Schubert, K." box="[357,706,727,752]" pageId="15" pageNumber="488" pagination="289 - 408" refId="ref14553" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Pars 57. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer &amp; Schubert, 1914: 364</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFD6AFD07FCD40391" authorityName="Fauvel" authorityYear="1874" box="[723,819,728,751]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFD6AFD07FCD40391" box="[723,819,728,751]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, catalog);
<bibRefCitation id="926A4B3CE3572F7AFC18FD07FB81038E" author="Gridelli, E." box="[929,1126,728,752]" pageId="15" pageNumber="488" pagination="170 - 201" refId="ref15186" refString="Gridelli, E. (1924) Sesto contributo allo studio degli Staphylinini appunti di morfologia e sistematica del genere Hesperus Fauv. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 51, 170 - 201." type="journal article" year="1924">Gridelli, 1924: 183</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFBCFFD07FB310391" authorityName="Fauvel" authorityYear="1874" box="[1142,1238,728,751]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFBCFFD07FB310391" box="[1142,1238,728,751]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, characters);
<bibRefCitation id="926A4B3CE3572F7AFADAFD07FE8E0270" author="Bernhauer, M." pageId="15" pageNumber="488" pagination="1 - 28" refId="ref14486" refString="Bernhauer, M. (1928) Dr. E. Mjobergs Zoological Collections from Sumatra. 8. Staphylinidae. Arkiv for Zoologi, (A), 19 (19), 1 - 28. [1927]" type="journal article" year="1928">Bernhauer, 1928: 17</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFEC2FD28FE3C0270" authorityName="Fauvel" authorityYear="1874" box="[379,475,759,782]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFEC2FD28FE3C0270" box="[379,475,759,782]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, Sumatra);
<bibRefCitation id="926A4B3CE3572F7AFDEFFD29FCCC0270" author="Cameron, M." box="[598,811,758,782]" pageId="15" pageNumber="488" pagination="1 - 443" refId="ref14719" refString="Cameron, M. (1932) The Fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. III. Taylor &amp; Francis London, 1 - 443." type="book chapter" year="1932">Cameron, 1932: 157</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFC85FD28FC7B0270" authorityName="Fauvel" authorityYear="1874" box="[828,924,759,782]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFC85FD28FC7B0270" box="[828,924,759,782]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, characters);
<bibRefCitation id="926A4B3CE3572F7AFB93FD29FAC20270" author="Scheerpeltz, O." box="[1066,1317,758,782]" pageId="15" pageNumber="488" pagination="989 - 1500" refId="ref16140" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Eds.), Coleopterorum Catalogus. 6 (129). Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933: 1372</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFA8FFD28FA710270" authorityName="Fauvel" authorityYear="1874" box="[1334,1430,759,782]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFA8FFD28FA710270" box="[1334,1430,759,782]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, catalog);
<bibRefCitation id="926A4B3CE3572F7AFE91FCCAFE1B0253" author="Cameron, M." box="[296,508,789,813]" pageId="15" pageNumber="488" pagination="387" refId="ref14753" refString="Cameron, M. (1933) Fauna Sumatrensis. Staphylinidae. Tijdschrift voor Entomologie, 76, 387" type="journal article" year="1933">Cameron, 1933: 390</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFDB5FCC9FD8B0253" authorityName="Fauvel" authorityYear="1874" box="[524,620,790,813]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFDB5FCC9FD8B0253" box="[524,620,790,813]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, Sumatra);
<bibRefCitation id="926A4B3CE3572F7AFD5EFCCAFC330253" author="Scheerpeltz, O." box="[743,980,789,813]" pageId="15" pageNumber="488" pagination="93 - 371" refId="ref16265" refString="Scheerpeltz, O. (1965) Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except Megalopsidiinae et Steninae). Arkiv for Zoologi, 17, 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965: 108</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFC5DFCC9FBA30253" authorityName="Fauvel" authorityYear="1874" box="[996,1092,790,813]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFC5DFCC9FBA30253" box="[996,1092,790,813]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
,
<collectingCountry id="8EEC765DE3572F7AFBEAFCCAFB7A0253" box="[1107,1181,789,813]" name="Myanmar" pageId="15" pageNumber="488">Burma</collectingCountry>
);
<bibRefCitation id="926A4B3CE3572F7AFB16FCCAFA7B0253" author="Scheerpeltz, O." box="[1199,1436,789,813]" pageId="15" pageNumber="488" pagination="150 - 197" refId="ref16302" refString="Scheerpeltz, O. (1971) Studien an den Arten der Gattung Hesperus Fauvel (Col. Staphylinidae). Entomologische Arbeiten aus dem Museum G. Frey, 22, 150 - 197." type="journal article" year="1971">Scheerpeltz, 1971: 159</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFF69FCEBFED70235" authorityName="Fauvel" authorityYear="1874" box="[208,304,820,843]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFF69FCEBFED70235" box="[208,304,820,843]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, characters);
<bibRefCitation id="926A4B3CE3572F7AFE04FCEBFD850235" author="Frank, J. H." box="[445,610,819,844]" pageId="15" pageNumber="488" pagination="41" refId="ref15147" refString="Frank, J. H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agricultural Experiment Station Bulletin, 824, 41." type="journal article" year="1982">Frank, 1982: 36</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFDCAFCEBFD340235" authorityName="Fauvel" authorityYear="1874" box="[627,723,820,843]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFDCAFCEBFD340235" box="[627,723,820,843]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, list of parasites);
<bibRefCitation id="926A4B3CE3572F7AFC2EFCEBFB900235" author="Herman, L. H." box="[919,1143,819,844]" pageId="15" pageNumber="488" pagination="2819 - 2949" refId="ref15354" refString="Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta: Coleoptera): 1758 to the end of the second millennium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2819 - 2949. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman, 2001a: 2684</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFB31FCEBFB0F0235" authorityName="Fauvel" authorityYear="1874" box="[1160,1256,820,843]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFB31FCEBFB0F0235" box="[1160,1256,820,843]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, catalog); Chani-Posse, 2018: 34 (transferred to
<taxonomicName id="31FB4D4EE3572F7AFDABFC8DFD570214" authorityName="Bernhauer" authorityYear="1912" box="[530,688,850,874]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFDABFC8DFD570214" box="[530,688,850,874]" italics="true" pageId="15" pageNumber="488">Eccoptolonthus</emphasis>
</taxonomicName>
);
<bibRefCitation id="926A4B3CE3572F7AFD71FC8CFC640214" author="Newton, A. F." box="[712,899,850,874]" pageId="15" pageNumber="488" pagination="2000" refId="ref16076" refString="Newton, A. F. (2019) StaphBase: Staphyliniformia World Catalog Database. Version Nov 2018. Online. In: Species 2000 &amp; ITIS Catalogue of Life, 2019 Annual Checklist. Available from: https: // www. catalogueoflife. org / col / details / database / id / 204 (accessed 15 April 2020)." type="journal article" year="2019">Newton, 2019: 20</bibRefCitation>
(catalogued);
<bibRefCitation id="926A4B3CE3572F7AFBA2FC8DFB000214" author="Hayashi, Y." box="[1051,1255,850,874]" pageId="15" pageNumber="488" pagination="135 - 142" refId="ref15304" refString="Hayashi, Y. (2020) A New Subgenus of the Genus Eccoptolonthus Bernhauer, with Descriptions of a New Species and a New Subspecies (Coleoptera, Staphylinidae, Philonthina). Elytra, Tokyo, New Series, 10 (1), 135 - 142." type="journal article" year="2020">Hayashi, 2020: 136</bibRefCitation>
(
<taxonomicName id="31FB4D4EE3572F7AFB41FC8DFA710214" authorityName="Bernhauer" authorityYear="1912" box="[1272,1430,850,874]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFB41FC8DFA710214" box="[1272,1430,850,874]" italics="true" pageId="15" pageNumber="488">Eccoptolonthus</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="31FB4D4EE3572F7AFE95FCADFE2602F7" authorityName="Hayashi" authorityYear="2020" box="[300,449,882,905]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="subGenus" subGenus="Eumorimotous">
<emphasis id="C48FEADFE3572F7AFE95FCADFE2602F7" box="[300,449,882,905]" italics="true" pageId="15" pageNumber="488">Eumorimotous</emphasis>
</taxonomicName>
,
<typeStatus id="2940886FE3572F7AFE77FCAEFE1D02F7" box="[462,506,881,905]" pageId="15" pageNumber="488">type</typeStatus>
species).
</paragraph>
<paragraph id="F64436CDE3572F7AFF2EFC50FDB802B8" blockId="15.[151,1437,697,966]" pageId="15" pageNumber="488">
<emphasis id="C48FEADFE3572F7AFF2EFC50FF2802D9" bold="true" box="[151,207,911,935]" pageId="15" pageNumber="488">Syn.:</emphasis>
<taxonomicName id="31FB4D4EE3572F7AFF6FFC4FFCC902D9" authority="Gridelli, 1924: 185" authorityName="Gridelli" authorityPageNumber="185" authorityYear="1924" box="[214,814,911,936]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="variety" species="laevigatus" variety="sarawakensis">
<emphasis id="C48FEADFE3572F7AFF6FFC4FFE4102D6" box="[214,422,912,936]" italics="true" pageId="15" pageNumber="488">Hesperus laevigatus</emphasis>
var.
<emphasis id="C48FEADFE3572F7AFE63FC4FFD8402D6" box="[474,611,912,936]" italics="true" pageId="15" pageNumber="488">sarawakensis</emphasis>
<bibRefCitation id="926A4B3CE3572F7AFDD3FC4FFCC902D9" author="Gridelli, E." box="[618,814,911,936]" pageId="15" pageNumber="488" pagination="170 - 201" refId="ref15186" refString="Gridelli, E. (1924) Sesto contributo allo studio degli Staphylinini appunti di morfologia e sistematica del genere Hesperus Fauv. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 51, 170 - 201." type="journal article" year="1924">Gridelli, 1924: 185</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="31FB4D4EE3572F7AFC87FC4FFC7902D9" authorityName="Fauvel" authorityYear="1874" box="[830,926,912,935]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="genus">
<emphasis id="C48FEADFE3572F7AFC87FC4FFC7902D9" box="[830,926,912,935]" italics="true" pageId="15" pageNumber="488">Hesperus</emphasis>
</taxonomicName>
, variety of
<taxonomicName id="31FB4D4EE3572F7AFBAFFC4FFB9902D6" authorityName="Fauvel" authorityYear="1895" box="[1046,1150,912,936]" class="Insecta" family="Staphylinidae" genus="Hesperus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="C48FEADFE3572F7AFBAFFC4FFB9902D6" box="[1046,1150,912,936]" italics="true" pageId="15" pageNumber="488">laevigatus</emphasis>
</taxonomicName>
,
<typeStatus id="2940886FE3572F7AFB35FC4FFB5F02D6" box="[1164,1208,912,936]" pageId="15" pageNumber="488">type</typeStatus>
locality: Borneo,
<collectingCountry id="8EEC765DE3572F7AFACAFC4FFEE602B8" name="Malaysia" pageId="15" pageNumber="488">Sarawak</collectingCountry>
); Herman, 2001: 2684 (synonym).
</paragraph>
</subSubSection>
<caption id="A2846645E3572F7AFF2EF846FBD80687" ID-DOI="http://doi.org/10.5281/zenodo.4640497" ID-Zenodo-Dep="4640497" httpUri="https://zenodo.org/record/4640497/files/figure.png" pageId="15" pageNumber="488" startId="15.[151,250,1945,1969]" targetBox="[184,1407,998,1920]" targetPageId="15">
<paragraph id="F64436CDE3572F7AFF2EF846FBD80687" blockId="15.[151,1437,1945,2042]" pageId="15" pageNumber="488">
<emphasis id="C48FEADFE3572F7AFF2EF846FEF306CC" bold="true" box="[151,276,1945,1970]" pageId="15" pageNumber="488">FIGURE 9.</emphasis>
<taxonomicName id="31FB4D4EE3572F7AFEA5F846FB9506CC" authority="laevigatus (Fauvel, 1895) A. Habitus" authorityName="laevigatus (Fauvel" authorityYear="1895" baseAuthorityName="Fauvel" baseAuthorityYear="1895" box="[284,1138,1945,1970]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="488" phylum="Arthropoda" rank="subSpecies" species="laevigatus" subGenus="Eumorimotous" subSpecies="laevigatus">
<emphasis id="C48FEADFE3572F7AFEA5F846FC1F06CF" bold="true" box="[284,1016,1945,1970]" pageId="15" pageNumber="488">
<emphasis id="C48FEADFE3572F7AFEA5F846FE5806CF" bold="true" box="[284,447,1945,1969]" italics="true" pageId="15" pageNumber="488">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE3572F7AFE76F845FD8C06CC" bold="true" box="[463,619,1946,1970]" italics="true" pageId="15" pageNumber="488">Eumorimotous</emphasis>
)
<emphasis id="C48FEADFE3572F7AFDC2F846FCB106CF" bold="true" box="[635,854,1945,1969]" italics="true" pageId="15" pageNumber="488">laevigatus laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE3572F7AFCDDF846FC1606CF" author="Fauvel, A." box="[868,1009,1945,1969]" pageId="15" pageNumber="488" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
</emphasis>
A. Habitus
</taxonomicName>
; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. underside of paramere; E. stylus of tergite IX of male; F. sternite VIII of male; G. tergite X of male; H. sternite IX of male. Scale bars, A=2 mm; B, C, D, E, F, G, H= 0.2 mm.
</paragraph>
</caption>
<caption id="A2846645E3482F65FF2EF84AFBC1068A" ID-DOI="http://doi.org/10.5281/zenodo.4640501" ID-Zenodo-Dep="4640501" httpUri="https://zenodo.org/record/4640501/files/figure.png" pageId="16" pageNumber="489" startId="16.[151,250,1941,1965]" targetBox="[203,1372,181,1912]" targetPageId="16">
<paragraph id="F64436CDE3482F65FF2EF84AFBC1068A" blockId="16.[151,1437,1940,2037]" pageId="16" pageNumber="489">
<emphasis id="C48FEADFE3482F65FF2EF84AFC1106D2" bold="true" box="[151,1014,1940,1965]" pageId="16" pageNumber="489">
FIGURE 10.
<taxonomicName id="31FB4D4EE3482F65FE9DF84BFC1106D2" authority="laevigatus (Fauvel, 1895)" authorityName="laevigatus (Fauvel" authorityYear="1895" baseAuthorityName="Fauvel" baseAuthorityYear="1895" box="[292,1014,1940,1965]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="489" phylum="Arthropoda" rank="subSpecies" species="laevigatus" subGenus="Eumorimotous" subSpecies="laevigatus">
<emphasis id="C48FEADFE3482F65FE9DF84BFE2006D2" bold="true" box="[292,455,1940,1964]" italics="true" pageId="16" pageNumber="489">Eccoptolonthus</emphasis>
(
<emphasis id="C48FEADFE3482F65FE6CF84AFD9606D3" bold="true" box="[469,625,1941,1965]" italics="true" pageId="16" pageNumber="489">Eumorimotous</emphasis>
)
<emphasis id="C48FEADFE3482F65FD39F84BFCBE06D2" bold="true" box="[640,857,1940,1964]" italics="true" pageId="16" pageNumber="489">laevigatus laevigatus</emphasis>
(
<bibRefCitation id="926A4B3CE3482F65FCDCF84BFC0806D2" author="Fauvel, A." box="[869,1007,1940,1964]" pageId="16" pageNumber="489" pagination="180 - 286" refId="ref15110" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286. https: // doi. org / 10.1017 / S 102688120005892 X" type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
)
</taxonomicName>
</emphasis>
A. median lobe of aedeagus, dorsal (parameral side) view; B. underside of paramere; C. tergite X of male; D. sternite VIII of male; E. sternite IX of male; F. stylus of tergite IX of male. Scale bars, A= 0.14 mm; B, C, E= 0.2 mm; D= 0.34 mm; F= 0.27 mm.
</paragraph>
</caption>
<subSubSection id="BEE16546E3492F64FF2EFF49FA710063" pageId="17" pageNumber="490" type="materials_examined">
<paragraph id="F64436CDE3492F64FF2EFF49FA710063" blockId="17.[151,1436,150,357]" pageId="17" pageNumber="490">
<emphasis id="C48FEADFE3492F64FF2EFF49FE6C01CE" bold="true" box="[151,395,150,176]" pageId="17" pageNumber="490">Materials examined.</emphasis>
<materialsCitation id="46933C90E3492F64FE2BFF49FABC01AB" ID-GBIF-Occurrence="3063281352" collectingDate="2010-10-07" collectionCode="IZ-CAS" collectorName="Lv Liang &amp; Zhang Xi" country="China" elevation="982" latitude="21.28795" location="Longmen village" longLatPrecision="8" longitude="101.5315" municipality="Mengla county" pageId="17" pageNumber="490" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Yunnan">
<emphasis id="C48FEADFE3492F64FE2BFF49FDB601CF" bold="true" box="[402,593,150,177]" pageId="17" pageNumber="490">
<collectingCountry id="8EEC765DE3492F64FE2BFF49FE3801CE" box="[402,479,150,176]" name="China" pageId="17" pageNumber="490">China</collectingCountry>
:
<collectingRegion id="343FF82FE3492F64FE53FF48FDAB01CF" box="[490,588,151,177]" country="China" name="Yunnan" pageId="17" pageNumber="490">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="E0FDFD44E3492F64FDE1FF48FD7E01CF" box="[600,665,151,177]" pageId="17" pageNumber="490" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="E0FDFD44E3492F64FD1CFF48FD2801CF" box="[677,719,151,177]" pageId="17" pageNumber="490" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="1620ACB7E3492F64FD65FF48FC6101CF" box="[732,902,151,177]" pageId="17" pageNumber="490">Mengla county</collectingMunicipality>
,
<location id="F3246016E3492F64FC28FF48FBB501CF" LSID="urn:lsid:plazi:treatment:7E5287DBE3572F64FF2EFDD7FEA8001B:F3246016E3492F64FC28FF48FBB501CF" box="[913,1106,151,177]" country="China" latitude="21.28795" longLatPrecision="8" longitude="101.5315" municipality="Mengla county" name="Longmen village" pageId="17" pageNumber="490" stateProvince="Yunnan">Longmen village</location>
, the edge of rice field, rotting banana flower (
<geoCoordinate id="93CF500AE3492F64FEF3FF64FE0701AB" box="[330,480,187,213]" degrees="21" direction="north" minutes="17.277" orientation="latitude" pageId="17" pageNumber="490" precision="1" value="21.28795">21°17.277N</geoCoordinate>
,
<geoCoordinate id="93CF500AE3492F64FE54FF64FD9801AB" box="[493,639,187,213]" degrees="101" direction="east" minutes="31.89" orientation="longitude" pageId="17" pageNumber="490" precision="9" value="101.5315">101°31.89E</geoCoordinate>
),
<quantity id="31039B28E3492F64FD2FFF64FD0601AB" box="[662,737,187,213]" metricMagnitude="2" metricUnit="m" metricValue="9.82" pageId="17" pageNumber="490" unit="m" value="982.0">
<elevation id="7DD6D1FEE3492F64FD2FFF64FD0601AB" box="[662,737,187,213]" metricMagnitude="2" metricUnit="m" metricValue="9.82" pageId="17" pageNumber="490" unit="m" value="982.0">982 m</elevation>
</quantity>
,
<date id="8245100DE3492F64FD56FF64FCBD01AB" box="[751,858,187,213]" pageId="17" pageNumber="490" value="2010-10-07">
<collectingDate id="9201E9E5E3492F64FD56FF64FCBD01AB" box="[751,858,187,213]" pageId="17" pageNumber="490" value="2010-10-07">7.X.2010</collectingDate>
</date>
, collected by
<collectorName id="5B0E531BE3492F64FC42FF64FB8001AB" box="[1019,1127,187,213]" pageId="17" pageNumber="490">Lv Liang</collectorName>
,
<collectorName id="5B0E531BE3492F64FBCCFF64FB0501AB" box="[1141,1250,187,213]" pageId="17" pageNumber="490">Zhang Xi</collectorName>
(
<collectionCode id="90EAAE08E3492F64FB4AFF64FAB301AB" box="[1267,1364,187,213]" pageId="17" pageNumber="490">IZ-CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="46933C90E3492F64FAD0FF64FB0F0187" ID-GBIF-Occurrence="3063281313" collectionCode="VI" collectorName="Meng Lingzeng" country="China" elevation="729" location="Naban river" pageId="17" pageNumber="490" specimenCount="1" specimenCount-male="1">
<specimenCount id="E0FDFD44E3492F64FAD0FF64FA7001AB" box="[1385,1431,187,213]" pageId="17" pageNumber="490" type="male">1 ♂</specimenCount>
,
<collectorName id="5B0E531BE3492F64FF2EFF00FEE60187" box="[151,257,223,249]" pageId="17" pageNumber="490">Jinghong</collectorName>
,
<location id="F3246016E3492F64FEB4FF00FE740187" LSID="urn:lsid:plazi:treatment:7E5287DBE3572F64FF2EFDD7FEA8001B:F3246016E3492F64FEB4FF00FE740187" box="[269,403,223,249]" country="China" name="Naban river" pageId="17" pageNumber="490">Naban river</location>
reserve,
<collectorName id="5B0E531BE3492F64FE43FF00FDA40187" box="[506,579,223,249]" pageId="17" pageNumber="490">Naban</collectorName>
tea factory (
<collectorName id="5B0E531BE3492F64FD6BFF00FCFC0187" box="[722,795,223,249]" pageId="17" pageNumber="490">Forest</collectorName>
) (22°1581N, 10°66543E),
<quantity id="31039B28E3492F64FBD9FF00FB4D0187" box="[1120,1194,223,249]" metricMagnitude="2" metricUnit="m" metricValue="7.29" pageId="17" pageNumber="490" unit="m" value="729.0">
<elevation id="7DD6D1FEE3492F64FBD9FF00FB4D0187" box="[1120,1194,223,249]" metricMagnitude="2" metricUnit="m" metricValue="7.29" pageId="17" pageNumber="490" unit="m" value="729.0">729 m</elevation>
</quantity>
,
<date id="8245100DE3492F64FB0EFF00FB0F0187" box="[1207,1256,223,249]" pageId="17" pageNumber="490" value="2009-06-06">
6.
<collectionCode id="90EAAE08E3492F64FB71FF00FB0F0187" box="[1224,1256,223,249]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="490">VI</collectionCode>
</date>
</materialsCitation>
.2009, collected by
<collectorName id="5B0E531BE3492F64FF03FEDCFE8A0063" box="[186,365,259,285]" pageId="17" pageNumber="490">Meng Lingzeng</collectorName>
(
<collectionCode id="90EAAE08E3492F64FEC2FEDCFE3B0063" box="[379,476,259,285]" pageId="17" pageNumber="490">IZ-CAS</collectionCode>
);
<materialsCitation id="46933C90E3492F64FE56FEDDFA750063" ID-GBIF-Occurrence="3063281344" box="[495,1426,258,285]" collectingDate="1937-07" collectionCode="IZ-CAS" collectorName="A. De. Cooman" country="Vietnam" location="Tonkin Hoa-binh" pageId="17" pageNumber="490" specimenCount="1" specimenCount-male="1">
<emphasis id="C48FEADFE3492F64FE56FEDDFDB90062" bold="true" box="[495,606,258,284]" pageId="17" pageNumber="490">
<collectingCountry id="8EEC765DE3492F64FE56FEDDFDBE0062" box="[495,601,258,284]" name="Vietnam" pageId="17" pageNumber="490">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="E0FDFD44E3492F64FDDCFEDCFD760063" box="[613,657,259,285]" pageId="17" pageNumber="490" type="male">1 ♂</specimenCount>
,
<location id="F3246016E3492F64FD25FEDCFCB80063" LSID="urn:lsid:plazi:treatment:7E5287DBE3572F64FF2EFDD7FEA8001B:F3246016E3492F64FD25FEDCFCB80063" box="[668,863,259,285]" country="Vietnam" name="Tonkin Hoa-binh" pageId="17" pageNumber="490">Tonkin Hoa-binh</location>
,
<date id="8245100DE3492F64FCD3FEDCFC350063" box="[874,978,259,285]" pageId="17" pageNumber="490" value="1937-07">
<collectingDate id="9201E9E5E3492F64FCD3FEDCFC350063" box="[874,978,259,285]" pageId="17" pageNumber="490" value="1937-07">VII.1937</collectingDate>
</date>
, collected by
<collectorName id="5B0E531BE3492F64FBD2FEDCFAFC0063" box="[1131,1307,259,285]" pageId="17" pageNumber="490">A. De. Cooman</collectorName>
(
<collectionCode id="90EAAE08E3492F64FA93FEDCFA6C0063" box="[1322,1419,259,285]" pageId="17" pageNumber="490">IZ-CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEE16546E3492F64FF7EFEF9FEA8001B" pageId="17" pageNumber="490" type="distribution">
<paragraph id="F64436CDE3492F64FF7EFEF9FEA8001B" blockId="17.[151,1436,150,357]" pageId="17" pageNumber="490">
<emphasis id="C48FEADFE3492F64FF7EFEF9FE86003E" bold="true" box="[199,353,294,320]" pageId="17" pageNumber="490">Distribution.</emphasis>
<collectingCountry id="8EEC765DE3492F64FED0FEF8FE4B003F" box="[361,428,295,321]" name="China" pageId="17" pageNumber="490">China</collectingCountry>
(
<collectingRegion id="343FF82FE3492F64FE05FEF8FDFF003F" box="[444,536,295,321]" country="China" name="Yunnan" pageId="17" pageNumber="490">Yunnan</collectingRegion>
),
<collectingCountry id="8EEC765DE3492F64FD92FEF8FD6A003F" box="[555,653,295,321]" name="Vietnam" pageId="17" pageNumber="490">Vietnam</collectingCountry>
,
<collectingCountry id="8EEC765DE3492F64FD23FEF8FD34003F" box="[666,723,295,321]" name="Laos" pageId="17" pageNumber="490">Laos</collectingCountry>
,
<collectingCountry id="8EEC765DE3492F64FD59FEF8FCA8003F" box="[736,847,295,321]" name="Myanmar" pageId="17" pageNumber="490">Myanmar</collectingCountry>
,
<collectingCountry id="8EEC765DE3492F64FCE2FEF8FC25003F" box="[859,962,295,321]" name="Malaysia" pageId="17" pageNumber="490">Malaysia</collectingCountry>
(
<collectingRegion id="343FF82FE3492F64FC6BFEF8FBCE003F" box="[978,1065,295,321]" country="Malaysia" name="Pulau Pinang" pageId="17" pageNumber="490">Penang</collectingRegion>
,
<collectingCountry id="8EEC765DE3492F64FB8CFEF8FB7D003C" box="[1077,1178,295,322]" name="Malaysia" pageId="17" pageNumber="490">Sarawak</collectingCountry>
),
<collectingCountry id="8EEC765DE3492F64FB17FEF8FAC5003F" box="[1198,1314,295,321]" name="Singapore" pageId="17" pageNumber="490">Singapore</collectingCountry>
,
<collectingCountry id="8EEC765DE3492F64FA96FEF8FA7B003F" box="[1327,1436,295,321]" name="Indonesia" pageId="17" pageNumber="490">Indonesia</collectingCountry>
(
<collectingRegion id="343FF82FE3492F64FF26FE94FF32001B" box="[159,213,331,357]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="490">Java</collectingRegion>
,
<collectingRegion id="343FF82FE3492F64FF59FE94FEA4001B" box="[224,323,331,357]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="490">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>