Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China Author Fei, Xu-Dong 0000-0003-0386-5918 Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. & feixudong @ ioz. ac. cn; https: // orcid. org / 0000 - 0003 - 0386 - 5918 feixudong@ioz.ac.cn Author Zhou, Hong-Zhang Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. text Zootaxa 2021 2021-03-26 4949 3 473 498 journal article 7571 10.11646/zootaxa.4949.3.3 39300142-6cea-4253-80bb-00856fdd5f95 1175-5326 4640469 C22ACA1A-4856-4979-BD7A-DFE37949C2D4 8. Eccoptolonthus ( Eumorimotous ) laevigatus laevigatus ( Fauvel, 1895 ) First record from China and Vietnam ( Figs. 9A–H , 10A–F ) Fauvel, 1895: 259 ( Hesperus , type locality: Bírmanie , Carin Cheba, 900–1100 m ; Pulo-Penang; Malacca ; Singapore ; Sumatra; Borneo; Java); Bernhauer & Schubert, 1914: 364 ( Hesperus , catalog); Gridelli, 1924: 183 ( Hesperus , characters); Bernhauer, 1928: 17 ( Hesperus , Sumatra); Cameron, 1932: 157 ( Hesperus , characters); Scheerpeltz, 1933: 1372 ( Hesperus , catalog); Cameron, 1933: 390 ( Hesperus , Sumatra); Scheerpeltz, 1965: 108 ( Hesperus , Burma ); Scheerpeltz, 1971: 159 ( Hesperus , characters); Frank, 1982: 36 ( Hesperus , list of parasites); Herman, 2001a: 2684 ( Hesperus , catalog); Chani-Posse, 2018: 34 (transferred to Eccoptolonthus ); Newton, 2019: 20 (catalogued); Hayashi, 2020: 136 ( Eccoptolonthus , subgenus Eumorimotous , type species). Syn.: Hesperus laevigatus var. sarawakensis Gridelli, 1924: 185 ( Hesperus , variety of laevigatus , type locality: Borneo, Sarawak ); Herman, 2001: 2684 (synonym). FIGURE 9. Eccoptolonthus ( Eumorimotous ) laevigatus laevigatus ( Fauvel, 1895 ) A. Habitus ; B. aedeagus, dorsal (parameral side) view; C. aedeagus, lateral view; D. underside of paramere; E. stylus of tergite IX of male; F. sternite VIII of male; G. tergite X of male; H. sternite IX of male. Scale bars, A=2 mm; B, C, D, E, F, G, H= 0.2 mm. FIGURE 10. Eccoptolonthus ( Eumorimotous ) laevigatus laevigatus ( Fauvel, 1895 ) A. median lobe of aedeagus, dorsal (parameral side) view; B. underside of paramere; C. tergite X of male; D. sternite VIII of male; E. sternite IX of male; F. stylus of tergite IX of male. Scale bars, A= 0.14 mm; B, C, E= 0.2 mm; D= 0.34 mm; F= 0.27 mm. Materials examined. China : Yunnan : 3 ♂♂ , 1 ♀ , Mengla county , Longmen village , the edge of rice field, rotting banana flower ( 21°17.277’N , 101°31.89’E ), 982 m , 7.X.2010 , collected by Lv Liang , Zhang Xi ( IZ-CAS ) ; 1 ♂ , Jinghong , Naban river reserve, Naban tea factory ( Forest ) (22°1581’N, 10°66543’E), 729 m , 6. VI .2009, collected by Meng Lingzeng ( IZ-CAS ); Vietnam : 1 ♂ , Tonkin Hoa-binh , VII.1937 , collected by A. De. Cooman ( IZ-CAS ) . Distribution. China ( Yunnan ), Vietnam , Laos , Myanmar , Malaysia ( Penang , Sarawak ), Singapore , Indonesia ( Java , Sumatra ).