136 lines
15 KiB
XML
136 lines
15 KiB
XML
<document id="BFDFDFC3F18B02FA53D02E74E28A4958" ID-DOI="10.1186/s13358-024-00307-8" ID-ISSN="1664-2384" ID-PMC="PMC10991048" ID-PubMed="38584612" ID-Zenodo-Dep="12005092" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1718654698973" checkinUser="felipe" docAuthor="Pohle, Alexander & Klug, Christian" docDate="2024" docId="761887DBFFA62C2A1301FB3BFD113742" docLanguage="en" docName="SwissJPalaeontol.143.14.pdf" docOrigin="Swiss Journal of Palaeontology (14) 143 (1)" docSource="http://dx.doi.org/10.1186/s13358-024-00307-8" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Trematoceratidae Zakharov 1996" docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="8A21FFA3FFB22C3F102CFFA9FFEF3307" masterDocTitle="Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea" masterLastPageNumber="32" masterPageNumber="1" pageNumber="21" updateTime="1721670897711" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="22CCA04F86EADC9C6DD3AA2F3C436EDF">Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea</mods:title>
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<mods:namePart id="92AF0F2EFFA7AFAF496660E13BACE974">Pohle, Alexander</mods:namePart>
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<mods:affiliation id="F44AAF8FD01889421AE523B75C107C7D">Institut fÜr Geologie, Mineralogie und Geophysik, Ruhr-Universität Bochum, 44801 Bochum, Germany & Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="1E6637B057CC0F5EFA3C733F0639857F">Klug, Christian</mods:namePart>
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<mods:affiliation id="6C238B430301093D51594C3CF789BD22">Paläontologisches Institut und Museum, Universität ZÜrich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:title id="7DB3EC7EBE08A80E3EE59BEF9823561B">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="54FE7AFD123731D1DC71FE3D720C73AA">2024</mods:date>
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<mods:number id="6C30D9FB4480D9AC59BAD23B9C59F5A5">2024-04-03</mods:number>
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<treatment id="761887DBFFA62C2A1301FB3BFD113742" LSID="urn:lsid:plazi:treatment:761887DBFFA62C2A1301FB3BFD113742" httpUri="http://treatment.plazi.org/id/761887DBFFA62C2A1301FB3BFD113742" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="B6AB6546FFA62C2B1301FB3BFB1837AD" box="[813,1271,1170,1194]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="FE0E36CDFFA62C2B1301FB3BFB1837AD" blockId="20.[813,1271,1170,1194]" box="[813,1271,1170,1194]" pageId="20" pageNumber="21">
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<heading id="A54681A1FFA62C2B1301FB3BFB1837AD" box="[813,1271,1170,1194]" fontSize="36" level="2" pageId="20" pageNumber="21" reason="3">
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Family
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<taxonomicName id="39B14D4EFFA62C2B1350FB3BFB1837AD" authority="Zakharov, 1996" authorityName="Zakharov" authorityYear="1996" box="[892,1271,1170,1194]" family="Trematoceratidae" kingdom="Animalia" order="Pseudorthocerida" pageId="20" pageNumber="21" rank="family" subClass="Orthoceratoidea">
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<emphasis id="CCC5EADFFFA62C2B1350FB3BFBA537AD" bold="true" box="[892,1098,1170,1194]" pageId="20" pageNumber="21">Trematoceratidae</emphasis>
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Zakharov, 1996
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6AB6546FFA62C2B1301FB7AFA80356C" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="FE0E36CDFFA62C2B1301FB7AFA80356C" blockId="20.[813,1437,1235,1643]" pageId="20" pageNumber="21">
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<emphasis id="CCC5EADFFFA62C2B1301FB7AFC4F37EC" bold="true" box="[813,928,1235,1259]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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: Relatively small orthocones without any indication of conch curvature with the possible exception of the apicalmost parts. Expansion rate constant, around 5° with relatively little variation. Siphuncle central or very slightly subcentral, with straight or slightly expanded siphuncular segments and short ortho- or suborthochoanitic septal necks that lack the inner prismatic layer and have a reduced outer prismatic layer. Cameral deposits present, petal- or star-shaped. Endosiphuncular deposits parietal, possibly restricted to the septal necks and early ontogenetic stages. Embryonic shell where known comparatively large, up to
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<quantity id="39499B28FFA62C2B140CF99AFB8F354C" box="[1056,1120,1587,1611]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="mm" value="2.0">2 mm</quantity>
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in diameter with blunt conical apex and cicatrix (modified after Zakharov, 1996).
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</paragraph>
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</subSubSection>
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<subSubSection id="B6AB6546FFA62C2B1301F93CFC51342B" pageId="20" pageNumber="21" type="description">
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<paragraph id="FE0E36CDFFA62C2B1301F93CFC51342B" blockId="20.[813,1436,1684,1837]" pageId="20" pageNumber="21">
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<emphasis id="CCC5EADFFFA62C2B1301F93CFC0035AA" bold="true" box="[813,1007,1685,1709]" pageId="20" pageNumber="21">Included genera</emphasis>
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:
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<taxonomicName id="39B14D4EFFA62C2B142DF93CFAA335AB" authority="Eichwald, 1851" authorityName="Eichwald" authorityYear="1851" box="[1025,1356,1684,1709]" class="Cephalopoda" family="Orthoceratidae" genus="Trematoceras" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthocerida" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
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<emphasis id="CCC5EADFFFA62C2B142DF93CFB7A35AB" box="[1025,1173,1685,1708]" italics="true" pageId="20" pageNumber="21">Trematoceras</emphasis>
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Eichwald, 1851
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</taxonomicName>
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;
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<taxonomicName id="39B14D4EFFA62C2B1573F93CFB4735CA" authority="Schastlivceva, 1981" authorityName="Schastlivceva" authorityYear="1981" class="Cephalopoda" family="Orthoceratidae" genus="Paratrematoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
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<emphasis id="CCC5EADFFFA62C2B1573F93CFC5535CB" italics="true" pageId="20" pageNumber="21">Paratrematoceras</emphasis>
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<bibRefCitation id="9A204B3CFFA62C2B13E7F91CFB4735CA" author="Schastlivceva, N. P." box="[971,1192,1717,1741]" pageId="20" pageNumber="21" pagination="76 - 82" refId="ref26976" refString="Schastlivceva, N. P. (1981). O sistematicheskom polozhenii triasovykh ortozeratoidej yuga SSSR [On the systematic position of Triassic orthoceratoids from the southern USSR]. Byulleten' Moskovskogo obshestva ispytatelej prirody. Otdel Geologicheskij, 56, 76 - 82. (in Russian)." type="journal article" year="1981">Schastlivceva, 1981</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="39B14D4EFFA62C2B14ECF91CFBF335EA" authority="Schastlivceva, 1986" authorityName="Schastlivceva" authorityYear="1986" class="Cephalopoda" genus="Pseudotemperoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
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<emphasis id="CCC5EADFFFA62C2B14ECF91CFA7335CA" box="[1216,1436,1717,1741]" italics="true" pageId="20" pageNumber="21">Pseudotemperoceras</emphasis>
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<bibRefCitation id="9A204B3CFFA62C2B1301F97CFBF335EA" author="Schastlivceva, N. P." box="[813,1052,1749,1773]" pageId="20" pageNumber="21" pagination="122 - 129" refId="ref27027" refString="Schastlivceva, N. P. (1986). Nekotorye triasovye ortozeratidy i nautilidy Severo- Vostoka SSSR [Some Triassic orthoceratoids and nautilids from the northeastern USSR]. Byulleten' Moskovskogo obshestva ispytatelej prirody. Otdel Geologicheskij, 61, 122 - 129. (in Russian)." type="journal article" year="1986">Schastlivceva, 1986</bibRefCitation>
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</taxonomicName>
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. Doubtful:
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<taxonomicName id="39B14D4EFFA62C2B14FFF97CFB5E340A" authority="Tongtherm & Nabhitabhata, 2018" authorityName="Tongtherm & Nabhitabhata" authorityYear="2018" genus="Phatthalungoceras" pageId="20" pageNumber="27" rank="genus">
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<emphasis id="CCC5EADFFFA62C2B14FFF97CFA7335EA" box="[1235,1436,1749,1773]" italics="true" pageId="20" pageNumber="21">Phatthalungoceras</emphasis>
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Tongtherm & Nabhitabhata, 2018
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</taxonomicName>
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;
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<taxonomicName id="39B14D4EFFA62C2B14E8F95CFC57342A" authority="Doguzhaeva, 1994" authorityName="Doguzhaeva" authorityYear="1994" class="Cephalopoda" family="Boggyoceratidae" genus="Zhuravlevia" higherTaxonomySource="CoL" kingdom="Animalia" order="Mixosiphonata" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
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<emphasis id="CCC5EADFFFA62C2B14E8F95CFAA6340A" box="[1220,1353,1781,1805]" italics="true" pageId="20" pageNumber="21">Zhuravlevia</emphasis>
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Doguzhaeva, 1994
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6AB6546FFA72C2A10BBFF42FE5030E4" pageId="21" pageNumber="22" type="discussion">
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<paragraph id="FE0E36CDFFA72C2A10BBFF42FD9731A4" blockId="21.[151,776,235,995]" pageId="21" pageNumber="22">
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<emphasis id="CCC5EADFFFA72C2A10BBFF42FF123204" bold="true" box="[151,253,235,259]" pageId="21" pageNumber="22">Remarks</emphasis>
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: Several characters, such as the embryonic shell or the shape of the cameral deposits are known only from a limited number of taxa. However, due to the high consistency in other characters that are usually considered to be more variable between species (e.g., apical angle and shell sculpture), we assume that the variability in these lesser-known characters was likely low as well. Of course, new evidence for distinctly different morphologies may overturn our hypothesis but until then, we propose that the best approach is to classify all Mesozoic orthoceratoids within a single family. This includes the Early Cretaceous
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<taxonomicName id="39B14D4EFFA72C2A10D6FDE2FD8E3164" authority="Doguzhaeva, 1994" authorityName="Doguzhaeva" authorityYear="1994" box="[250,609,587,611]" class="Cephalopoda" family="Boggyoceratidae" genus="Zhuravlevia" higherTaxonomySource="CoL" kingdom="Animalia" order="Mixosiphonata" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
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<emphasis id="CCC5EADFFFA72C2A10D6FDE2FE903164" box="[250,383,587,611]" italics="true" pageId="21" pageNumber="22">Zhuravlevia</emphasis>
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Doguzhaeva, 1994
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</taxonomicName>
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, although we include it here only tentatively due to the 90 million years gap between it and Triassic trematoceratids.
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</paragraph>
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<paragraph id="FE0E36CDFFA72C2A1081FD02FE5030E4" blockId="21.[151,776,235,995]" pageId="21" pageNumber="22">
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Like the Triassic orthoceratoids, there have been very few studies on Permian taxa. It is thus conceivable that the
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<taxonomicName id="39B14D4EFFA72C2A10EEFD42FE6C3004" authorityName="Zakharov" authorityYear="1996" box="[194,387,747,771]" family="Trematoceratidae" pageId="21" pageNumber="21" rank="family">Trematoceratidae</taxonomicName>
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can be extended into the Permian. Nevertheless, the few available studies indicate that orthoceratoid diversity (or at least disparity) was comparatively higher during the Permian (see, e.g., Sweet, 1964;
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<bibRefCitation id="9A204B3CFFA72C2A10F7FCC2FE7C3084" author="Niko, S. & Nishida, T. & Nakazawa, K." box="[219,403,875,899]" pageId="21" pageNumber="22" refId="ref26011" refString="Niko, S., Nishida, T., & Nakazawa, K. (2000). Orthoconic cephalopods from" type="book" year="2000">Niko et al., 2000</bibRefCitation>
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; Yang et al., 2021;
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<bibRefCitation id="9A204B3CFFA72C2A1246FCC2FF2430A4" author="Niko, S. & Ehiro, M." pageId="21" pageNumber="22" pagination="1 - 6" refId="ref25910" refString="Niko, S., & Ehiro, M. (2023). Middle Permian orthoconic nautiloids from the Takakurayama Formation in the Yaguki area, Fukushima Prefecture, Northeast Japan. Bulletin of the Tohoku University Museum, 22, 1 - 6." type="journal article" year="2023">Niko & Ehiro, 2023</bibRefCitation>
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). Thus, a possible extension of the
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<taxonomicName id="39B14D4EFFA72C2A126AFC22FCE830A4" authorityName="Zakharov" authorityYear="1996" box="[582,775,907,931]" family="Trematoceratidae" pageId="21" pageNumber="21" rank="family">Trematoceratidae</taxonomicName>
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to include Permian taxa must await a better phylogenetic understanding of the latter.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6AB6546FFA72C2A10BBFBA5FD113742" pageId="21" pageNumber="22" type="distribution">
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<paragraph id="FE0E36CDFFA72C2A10BBFBA5FD113742" blockId="21.[151,775,1036,1093]" pageId="21" pageNumber="22">
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<emphasis id="CCC5EADFFFA72C2A10BBFBA5FEC93723" bold="true" box="[151,294,1036,1060]" pageId="21" pageNumber="22">Occurrence:</emphasis>
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Global; Early–Late Triassic (Induan? Olenekian–Norian, Rhaetian?), Early Cretaceous? (Aptian?).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |