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<document id="085CF7819D38B9F32EDCC53FE166313B" ID-CLB-Dataset="56018" ID-DOI="10.11646/zootaxa.5125.5.2" ID-GBIF-Dataset="6a677d3b-0f91-4f11-8cf2-64f4ad2a47d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6457259" ID-ZooBank="39EBA798-29B1-4450-9085-A4FB6D1DB331" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649834539484" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. &amp; Palatov, Dmitry M." docDate="2022" docId="721787E39442FFFBF5C962B3FBFD6ABC" docLanguage="en" docName="zootaxa.5125.5.2.pdf" docOrigin="Zootaxa 5125 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa dragani Makarchenko et Semenchenko 2022, sp. nov." docType="treatment" docUuid="B3C5EFC4-7AE3-4419-B2AF-DD40873B467B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="487" masterDocId="8E2EFF9B9440FFFFF55E6618FFBE6F7D" masterDocTitle="Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes" masterLastPageNumber="512" masterPageNumber="483" pageNumber="485" updateTime="1699328882303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FCF84CC6E19B66D69012A7795E95D9A6">Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes</mods:title>
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<mods:namePart id="49559D8A9C415399634E2A8CFDDB67C6">Makarchenko, Eugenyi A.</mods:namePart>
<mods:affiliation id="44FC60F252F578B94EC497EA50AAE5D1">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia</mods:affiliation>
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<mods:namePart id="7EFFE3EAD414F62BF4F7C3B6FF4DCB0C">Semenchenko, Alexander A.</mods:namePart>
<mods:affiliation id="002C1BEE50E89CF2559D2BB252A78965">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia</mods:affiliation>
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<mods:namePart id="12DAA2B461314BCC3B7A9021B71116BA">Palatov, Dmitry M.</mods:namePart>
<mods:nameIdentifier id="1D3100FD411399C19EEA8567484D27FB" type="ORCID">0000-0002-8826-9316</mods:nameIdentifier>
<mods:affiliation id="5FFC3FCB266DF8D5823BD549E36839C1">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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<treatment id="721787E39442FFFBF5C962B3FBFD6ABC" ID-DOI="http://doi.org/10.5281/zenodo.6458129" ID-GBIF-Taxon="194591613" ID-Zenodo-Dep="6458129" LSID="urn:lsid:zoobank.org:act:B3C5EFC4-7AE3-4419-B2AF-DD40873B467B" httpUri="http://treatment.plazi.org/id/721787E39442FFFBF5C962B3FBFD6ABC" lastPageId="4" lastPageNumber="487" pageId="2" pageNumber="485">
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<taxonomicName id="3DBE4D769442FFFDF5C962B3FD4F6BB9" ID-CoL="9J2PP" authority="Makarchenko et Semenchenko" authorityName="Makarchenko et Semenchenko" authorityYear="2022" box="[151,753,1194,1221]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="dragani" status="sp. nov.">
<emphasis id="C8CAEAE79442FFFDF5C962B3FED96BB8" bold="true" box="[151,359,1194,1221]" italics="true" pageId="2" pageNumber="485">Diamesa dragani</emphasis>
Makarchenko
<emphasis id="C8CAEAE79442FFFDF77B62B3FD846BB8" bold="true" box="[549,570,1195,1221]" italics="true" pageId="2" pageNumber="485">et</emphasis>
Semenchenko
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,
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</emphasis>
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http://zoobank.org/NomenclaturalActs/
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<paragraph id="FA0136F59442FFFDF5C962EBFED86A73" blockId="2.[151,1144,1194,1294]" box="[151,358,1267,1294]" pageId="2" pageNumber="485">
(
<figureCitation id="62852A709442FFFDF5C162EBFEB56A70" box="[159,267,1267,1293]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="2" pageNumber="485">Figs. 18</figureCitation>
,
<figureCitation id="62852A709442FFFDF44962EBFE8B6A70" box="[279,309,1267,1293]" captionStart="FIGURES 3036" captionStartId="13.[151,264,1503,1527]" captionTargetBox="[151,1436,758,1472]" captionTargetId="figure-286@13.[151,1436,758,1480]" captionTargetPageId="13" captionText="FIGURES 3036. Wings of Diamesa dragani sp. nov. (30), D. kownackii sp. nov. (31), D. maisaraensis sp. nov. (32), D. marinskiyi sp. nov. (33) and D. steinboecki Goetghebuer from Swiss Alps (34), French Alps (35) and Pamir Mountains (36)." figureDoi="http://doi.org/10.5281/zenodo.6457271" httpUri="https://zenodo.org/record/6457271/files/figure.png" pageId="2" pageNumber="485">30</figureCitation>
,
<figureCitation id="62852A709442FFFDF41F62EBFEE06A73" box="[321,350,1267,1294]" captionStart="FIGURES 6874" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 6874. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (6970), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of AiguillesRouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="2" pageNumber="485">68</figureCitation>
)
</paragraph>
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<subSubSection id="B2A4657E9442FFFDF5C96323FC326AE0" pageId="2" pageNumber="485" type="materials_examined">
<paragraph id="FA0136F59442FFFDF5C96323FC326AE0" blockId="2.[151,1437,1338,2013]" pageId="2" pageNumber="485">
<emphasis id="C8CAEAE79442FFFDF5C96323FEF66A29" bold="true" box="[151,328,1338,1365]" pageId="2" pageNumber="485">Type material.</emphasis>
<materialsCitation id="4AD63CA89442FFFDF40F6323FE816AE0" ID-GBIF-Occurrence="3751071315" collectingDate="2020-10-04" collectorName="S. Dragan." country="Russia" county="Republic" elevation="2063" latitude="51.71467" location="Sayan Mountains" longLatPrecision="1" longitude="89.8845" municipality="Tashtypskyi District" pageId="2" pageNumber="485" specimenCount="1" stateProvince="Khakasiya" typeStatus="holotype">
<typeStatus id="250588579442FFFDF40F6323FE026A28" box="[337,444,1339,1365]" pageId="2" pageNumber="485" type="holotype">Holotype</typeStatus>
, adult male,
<collectingCountry id="82A976659442FFFDF70C6323FD046A28" box="[594,698,1339,1365]" name="Russia" pageId="2" pageNumber="485">RUSSIA</collectingCountry>
:
<collectingCounty id="13604E799442FFFDF7996323FC926A28" box="[711,812,1339,1365]" pageId="2" pageNumber="485">Republic</collectingCounty>
of
<collectingRegion id="387AF8179442FFFDF6096323FC736A28" box="[855,973,1339,1365]" country="Russia" name="Khakasiya" pageId="2" pageNumber="485">Khakassia</collectingRegion>
,
<collectingMunicipality id="1A65AC8F9442FFFDF6846323FB7C6A28" box="[986,1218,1339,1365]" pageId="2" pageNumber="485">Tashtypskyi District</collectingMunicipality>
,
<location id="FF61602E9442FFFDF1916323FA266A28" LSID="urn:lsid:plazi:treatment:721787E39442FFFBF5C962B3FBFD6ABC:FF61602E9442FFFDF1916323FA266A28" box="[1231,1432,1339,1365]" country="Russia" county="Republic" latitude="51.71467" longLatPrecision="1" longitude="89.8845" municipality="Tashtypskyi District" name="Sayan Mountains" pageId="2" pageNumber="485" stateProvince="Khakasiya">Sayan Mountains</location>
, unnamed stream of
<location id="FF61602E9442FFFDF42D6347FD896A04" LSID="urn:lsid:plazi:treatment:721787E39442FFFBF5C962B3FBFD6ABC:FF61602E9442FFFDF42D6347FD896A04" box="[371,567,1375,1401]" country="Russia" county="Republic" latitude="51.71467" longLatPrecision="1" longitude="89.8845" municipality="Tashtypskyi District" name="Bolshoi On River" pageId="2" pageNumber="485" stateProvince="Khakasiya">Bolshoi On River</location>
basin, altitude
<elevation id="7193D1C69442FFFDF7BE6347FC596A04" box="[736,999,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.063" pageId="2" pageNumber="485" unit="m" value="2063.0">
<quantity id="3D469B109442FFFDF7BE6347FC8B6A04" box="[736,821,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.063" pageId="2" pageNumber="485" unit="m" value="2063.0">2063 m</quantity>
above sea level
</elevation>
,
<date id="8E0010359442FFFDF6AC6347FBE26A04" box="[1010,1116,1375,1401]" pageId="2" pageNumber="485" value="2020-10-04">
<collectingDate id="9E44E9DD9442FFFDF6AC6347FBE26A04" box="[1010,1116,1375,1401]" pageId="2" pageNumber="485" value="2020-10-04">4.X.2020</collectingDate>
</date>
,
<geoCoordinate id="9F8A50329442FFFDF1366347FB426A04" box="[1128,1276,1374,1401]" degrees="51.714667" direction="north" orientation="latitude" pageId="2" pageNumber="485" precision="1" value="51.71467">N 51.714667</geoCoordinate>
,
<geoCoordinate id="9F8A50329442FFFDF0596347FA266A04" box="[1287,1432,1374,1401]" degrees="89.884500" direction="east" orientation="longitude" pageId="2" pageNumber="485" precision="1" value="89.8845">E 89.884500</geoCoordinate>
, leg.
<collectorName id="574B53239442FFFDF599639BFE816AE0" box="[199,319,1411,1437]" pageId="2" pageNumber="485">S. Dragan.</collectorName>
</materialsCitation>
<materialsCitation id="4AD63CA89442FFFDF41B639BFC366AE0" ID-GBIF-Occurrence="3751071316" box="[325,904,1411,1437]" collectingDate="2020-10-04" collectorName="S. Dragan." country="Russia" county="Republic" elevation="2063" latitude="51.71467" location="Sayan Mountains" longLatPrecision="1" longitude="89.8845" municipality="Tashtypskyi District" pageId="2" pageNumber="485" specimenCount="6" specimenCount-adult="6" stateProvince="Khakasiya" typeStatus="paratype">
<typeStatus id="250588579442FFFDF41B639BFE086AE0" box="[325,438,1411,1437]" pageId="2" pageNumber="485" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ECB8FD7C9442FFFDF49F639BFDB06AE0" box="[449,526,1411,1437]" pageId="2" pageNumber="485" type="adult">6 adult</specimenCount>
males, the same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9442FFFDF59963BEFAF36A98" pageId="2" pageNumber="485" type="etymology">
<paragraph id="FA0136F59442FFFDF59963BEFAF36A98" blockId="2.[151,1437,1338,2013]" pageId="2" pageNumber="485">
<emphasis id="C8CAEAE79442FFFDF59963BEFE186ABD" bold="true" box="[199,422,1446,1472]" pageId="2" pageNumber="485">Derivatio nominis.</emphasis>
The species is named in honor of Dr. Dragan Sergey Viktorovich from Katanov Khakas State University (Abakan, Republic of
<collectingRegion id="387AF8179442FFFDF71263D3FD7C6A98" box="[588,706,1483,1509]" country="Russia" name="Khakasiya" pageId="2" pageNumber="485">Khakassia</collectingRegion>
,
<collectingCountry id="82A976659442FFFDF79363D3FCA76A9B" box="[717,793,1483,1510]" name="Russia" pageId="2" pageNumber="485">Russia</collectingCountry>
) who collected
<typeStatus id="250588579442FFFDF69263D3FC426A98" box="[972,1020,1483,1509]" pageId="2" pageNumber="485">type</typeStatus>
material in Sayan Mountains.
</paragraph>
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<subSubSection id="B2A4657E9442FFFBF59963F6FE526B6D" lastPageId="4" lastPageNumber="487" pageId="2" pageNumber="485" type="description">
<paragraph id="FA0136F59442FFFDF59963F6FEEF6975" blockId="2.[151,1437,1338,2013]" box="[199,337,1518,1544]" pageId="2" pageNumber="485">
<emphasis id="C8CAEAE79442FFFDF59963F6FEEF6975" bold="true" box="[199,337,1518,1544]" pageId="2" pageNumber="485">Description</emphasis>
</paragraph>
<paragraph id="FA0136F59442FFFDF599600AFA296950" blockId="2.[151,1437,1338,2013]" box="[199,1431,1554,1582]" pageId="2" pageNumber="485">
<emphasis id="C8CAEAE79442FFFDF599600AFEFA6950" bold="true" box="[199,324,1554,1581]" italics="true" pageId="2" pageNumber="485">Adult male</emphasis>
(n = 7, except when otherwise stated). Total length
<quantity id="3D469B109442FFFDF623600AFBBA6950" box="[893,1028,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="2" pageNumber="485" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
. Total length/wing length 0.811.20.
</paragraph>
<paragraph id="FA0136F59442FFFDF599602FFEF76908" blockId="2.[151,1437,1338,2013]" pageId="2" pageNumber="485">Coloration. Dark brown to black. Head, thorax and abdomen dark brown to black. Legs black. Wings grey, with brownish veins.</paragraph>
<paragraph id="FA0136F59442FFFDF5996067FEE268E8" blockId="2.[151,1437,1338,2013]" pageId="2" pageNumber="485">
Head (
<figureCitation id="62852A709442FFFDF44C6067FEC069E4" box="[274,382,1663,1689]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="2" pageNumber="485">Figs. 12</figureCitation>
). Eyes hairy, reniform. Temporal setae including 67 frontals, 36 orbitals, 510 verticals and 34 postorbitals. Clypeus with 46 setae.Antenna with 7 flagellomeres and reduced plume of setae (
<figureCitation id="62852A709442FFFDF1B060BBFA8C69C0" box="[1262,1330,1699,1725]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
); number and length of these setae on 16 flagellomeres respectively: 12 (2628 μm), 12 (1832 μm), 3 (3244 μm), 12 (2432 μm), 2 (4044 μm), 4 (4048 μm); terminal flagellomere with 24 setae, 3248 μm long in basal part and with 2 subapical setae, 2024 μm long. Length of 17 flagellomeres (μm): 96100, 4048, 3236, 2428, 2428, 2024, 120128; AR 0.480.52. Palpomere length (μm): 2840, 6872, 8492, 6468, 88100. Palpomere
<quantity id="3D469B109442FFFDF01E612BFAD56830" box="[1344,1387,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="485" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 1215 μm. Head width/palpal length 1.01.29. Antennal length/palpal length 1.071.09.
</paragraph>
<paragraph id="FA0136F59442FFFDF5996187FBC468A0" blockId="2.[151,1437,1338,2013]" pageId="2" pageNumber="485">Thorax. Antepronotum with 811 ventrolateral setae, 4452 μm long. Dorsocentrals 46, 5672 μm long; prealars 56, 4860 μm long. Scutellum with 1820 setae. AAII with 45 setae, 36 μm long.</paragraph>
<paragraph id="FA0136F59443FFFCF599668FFA266FA8" blockId="3.[151,1437,150,321]" pageId="3" pageNumber="486">
Wing (
<figureCitation id="62852A709443FFFCF44C668FFEDB6FCC" box="[274,357,151,177]" captionStart="FIGURES 3036" captionStartId="13.[151,264,1503,1527]" captionTargetBox="[151,1436,758,1472]" captionTargetId="figure-286@13.[151,1436,758,1480]" captionTargetPageId="13" captionText="FIGURES 3036. Wings of Diamesa dragani sp. nov. (30), D. kownackii sp. nov. (31), D. maisaraensis sp. nov. (32), D. marinskiyi sp. nov. (33) and D. steinboecki Goetghebuer from Swiss Alps (34), French Alps (35) and Pamir Mountains (36)." figureDoi="http://doi.org/10.5281/zenodo.6457271" httpUri="https://zenodo.org/record/6457271/files/figure.png" pageId="3" pageNumber="486">Fig. 30</figureCitation>
). Length
<quantity id="3D469B109443FFFCF492668FFDCE6FCC" box="[460,624,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.44" metricValueMax="2.72" metricValueMin="2.16" pageId="3" pageNumber="486" unit="mm" value="2.44" valueMax="2.72" valueMin="2.16">2.162.72 mm</quantity>
, width
<quantity id="3D469B109443FFFCF79E668EFCDA6FCC" box="[704,868,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.8" metricValueMin="8.5" pageId="3" pageNumber="486" unit="mm" value="0.865" valueMax="0.88" valueMin="0.85">0.850.88 mm</quantity>
. Costal extension 6076 µm long. Anal lobe rounded-angular. Squama with 37 setae, 3244 μm long. R and R
<subScript id="663A34B09443FFFCF66166D5FCF96FA6" attach="left" box="[831,839,205,219]" fontSize="6" pageId="3" pageNumber="486">1</subScript>
with 712 setae, R
<subScript id="663A34B09443FFFCF14066D5FB866FA6" attach="left" box="[1054,1080,205,219]" fontSize="6" pageId="3" pageNumber="486">4+5</subScript>
with 1315. RM/MCu 1.82.0.
</paragraph>
<paragraph id="FA0136F59443FFFCF59966C7FEEB6E3C" blockId="3.[151,1437,150,321]" pageId="3" pageNumber="486">
Legs. Spur of front tibia 3640 µm long. Spurs of mid tibia 3644 µm and 4044 µm long. Spurs of hind tibia 6472 µm and 4044 µm long. Hind tibial comb with 1620 setae. Length (μm) and proportions of leg segments are as in
<tableCitation id="B73C034E9443FFFCF5A5673FFEF16E3C" box="[251,335,295,321]" captionStart="TABLE 1" captionStartId="4.[151,239,151,175]" captionTargetPageId="4" captionText="TABLE 1. Lengths (in µm) and proportions of leg segments of Diamesa dragani sp. nov., male (n=5)" httpUri="http://table.plazi.org/id/AEC1667D9444FFFBF5C9668FFB666FD2" pageId="3" pageNumber="486" tableUuid="AEC1667D9444FFFBF5C9668FFB666FD2">Table 1</tableCitation>
.
</paragraph>
<caption id="AEC1667D9443FFFCF5C9615DFA9A68D8" ID-DOI="http://doi.org/10.5281/zenodo.6457261" ID-Zenodo-Dep="6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="3" pageNumber="486" startId="3.[151,264,1861,1885]" targetBox="[151,1435,345,1834]" targetPageId="3">
<paragraph id="FA0136F59443FFFCF5C9615DFA9A68D8" blockId="3.[151,1437,1861,1957]" pageId="3" pageNumber="486">
<emphasis id="C8CAEAE79443FFFCF5C9615DFE846820" bold="true" box="[151,314,1861,1885]" pageId="3" pageNumber="486">FIGURES 18.</emphasis>
<taxonomicName id="3DBE4D769443FFFCF461615EFE516820" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[319,495,1861,1885]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="dragani" status="sp. nov.">
<emphasis id="C8CAEAE79443FFFCF461615EFE516820" box="[319,495,1861,1885]" italics="true" pageId="3" pageNumber="486">Diamesa dragani</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79443FFFCF4AA615DFDFA6820" bold="true" box="[500,580,1861,1885]" pageId="3" pageNumber="486">
<taxonomicNameLabel id="D3F9579C9443FFFCF4AA615DFDFA6820" box="[500,580,1861,1885]" pageId="3" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="C8CAEAE79443FFFCF7D3615DFD1E6820" bold="true" box="[653,672,1861,1885]" pageId="3" pageNumber="486">1,</emphasis>
head;
<emphasis id="C8CAEAE79443FFFCF7BC615DFD4A6820" bold="true" box="[738,756,1861,1885]" pageId="3" pageNumber="486">2,</emphasis>
antenna;
<emphasis id="C8CAEAE79443FFFCF608615DFCD66820" bold="true" box="[854,872,1861,1885]" pageId="3" pageNumber="486">3,</emphasis>
tergite IX;
<emphasis id="C8CAEAE79443FFFCF685615DFC506820" bold="true" box="[987,1006,1861,1885]" pageId="3" pageNumber="486">4,</emphasis>
hypopygium in dorsal view;
<emphasis id="C8CAEAE79443FFFCF04D615DFA996820" bold="true" box="[1299,1319,1861,1885]" pageId="3" pageNumber="486">5,</emphasis>
gonocoxite and gonostylus;
<emphasis id="C8CAEAE79443FFFCF41F6171FEEA68FC" bold="true" box="[321,340,1897,1921]" pageId="3" pageNumber="486">6,</emphasis>
apex of gonostylus;
<emphasis id="C8CAEAE79443FFFCF7706171FDFF68FC" bold="true" box="[558,577,1897,1921]" pageId="3" pageNumber="486">7,</emphasis>
anal point in lateral view;
<emphasis id="C8CAEAE79443FFFCF6026171FCD168FC" bold="true" box="[860,879,1897,1921]" pageId="3" pageNumber="486">8,</emphasis>
transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band.
</paragraph>
</caption>
<caption id="AEC1667D9444FFFBF5C9668FFB666FD2" ID-Table-UUID="AEC1667D9444FFFBF5C9668FFB666FD2" box="[151,1240,151,176]" httpUri="http://table.plazi.org/id/AEC1667D9444FFFBF5C9668FFB666FD2" pageId="4" pageNumber="487" startId="4.[151,239,151,175]" targetBox="[151,1361,191,576]" targetIsTable="true" targetPageId="4">
<paragraph id="FA0136F59444FFFBF5C9668FFB666FD2" blockId="4.[151,1240,151,176]" box="[151,1240,151,176]" pageId="4" pageNumber="487">
<emphasis id="C8CAEAE79444FFFBF5C9668FFEB46FD2" bold="true" box="[151,266,151,175]" pageId="4" pageNumber="487">TABLE 1.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="3DBE4D769444FFFBF6616680FC476FD2" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[831,1017,151,175]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="dragani" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF6616680FC476FD2" box="[831,1017,151,175]" italics="true" pageId="4" pageNumber="487">Diamesa dragani</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF15E668FFBEB6FD2" bold="true" box="[1024,1109,151,175]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF15E668FFBEB6FD2" box="[1024,1109,151,175]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n=5)
</paragraph>
</caption>
<paragraph id="FA0136F59444FFFBF5AF66A7FB426D47" pageId="4" pageNumber="487">
<table id="88BEC45594440000F5C966A7FAEF6D3D" box="[151,1361,191,576]" colsContinueIn="7.[151,1365,1237,1622]" gridcols="8" gridrows="9" pageId="4" pageNumber="487">
<tr id="448E34B794440000F5C966A7FAEF6FA3" box="[151,1361,191,222]" gridrow="0" pageId="4" pageNumber="487" rowspan-0="1">
<th id="075F5DCB94440000F5AF66A7FE306FA3" box="[241,398,191,222]" gridcol="1" gridrow="0" pageId="4" pageNumber="487">fe</th>
<th id="075F5DCB94440000F4F566A7FDA36FA3" box="[427,541,191,222]" gridcol="2" gridrow="0" pageId="4" pageNumber="487">ti</th>
<th id="075F5DCB94440000F70B66A7FD7F6FA3" box="[597,705,191,222]" gridcol="3" gridrow="0" pageId="4" pageNumber="487">ta1</th>
<th id="075F5DCB94440000F64C66A7FCD56FA3" box="[786,875,191,222]" gridcol="4" gridrow="0" pageId="4" pageNumber="487">ta2</th>
<th id="075F5DCB94440000F6DC66A7FBA66FA3" box="[898,1048,191,222]" gridcol="5" gridrow="0" pageId="4" pageNumber="487">ta3</th>
<th id="075F5DCB94440000F12E66A7FB6F6FA3" box="[1136,1233,191,222]" gridcol="6" gridrow="0" pageId="4" pageNumber="487">ta4</th>
<th id="075F5DCB94440000F1A766A7FAEF6FA3" box="[1273,1361,191,222]" gridcol="7" gridrow="0" pageId="4" pageNumber="487">ta 5</th>
</tr>
<tr id="448E34B794440000F5C966FFFAEF6E78" box="[151,1361,231,261]" gridrow="1" pageId="4" pageNumber="487">
<th id="075F5DCB94440000F5C966FFFF056E78" box="[151,187,231,261]" gridcol="0" gridrow="1" pageId="4" pageNumber="487">P1</th>
<td id="075F5DCB94440000F5AF66FFFE306E78" box="[241,398,231,261]" gridcol="1" gridrow="1" pageId="4" pageNumber="487">13941689</td>
<td id="075F5DCB94440000F4F566FFFDA36E78" box="[427,541,231,261]" gridcol="2" gridrow="1" pageId="4" pageNumber="487">12301394</td>
<td id="075F5DCB94440000F70B66FFFD7F6E78" box="[597,705,231,261]" gridcol="3" gridrow="1" pageId="4" pageNumber="487">738787</td>
<td id="075F5DCB94440000F64C66FFFCD56E78" box="[786,875,231,261]" gridcol="4" gridrow="1" pageId="4" pageNumber="487">312344</td>
<td id="075F5DCB94440000F6DC66FFFBA66E78" box="[898,1048,231,261]" gridcol="5" gridrow="1" pageId="4" pageNumber="487">180197</td>
<td id="075F5DCB94440000F12E66FFFB6F6E78" box="[1136,1233,231,261]" gridcol="6" gridrow="1" pageId="4" pageNumber="487">8298</td>
<td id="075F5DCB94440000F1A766FFFAEF6E78" box="[1273,1361,231,261]" gridcol="7" gridrow="1" pageId="4" pageNumber="487">115131</td>
</tr>
<tr id="448E34B794440000F5C96708FAEF6E53" box="[151,1361,272,302]" gridrow="2" pageId="4" pageNumber="487">
<th id="075F5DCB94440000F5C96708FF056E53" box="[151,187,272,302]" gridcol="0" gridrow="2" pageId="4" pageNumber="487">P 2</th>
<td id="075F5DCB94440000F5AF6708FE306E53" box="[241,398,272,302]" gridcol="1" gridrow="2" pageId="4" pageNumber="487">12791542</td>
<td id="075F5DCB94440000F4F56708FDA36E53" box="[427,541,272,302]" gridcol="2" gridrow="2" pageId="4" pageNumber="487">9841132</td>
<td id="075F5DCB94440000F70B6708FD7F6E53" box="[597,705,272,302]" gridcol="3" gridrow="2" pageId="4" pageNumber="487">410459</td>
<td id="075F5DCB94440000F64C6708FCD56E53" box="[786,875,272,302]" gridcol="4" gridrow="2" pageId="4" pageNumber="487">197213</td>
<td id="075F5DCB94440000F6DC6708FBA66E53" box="[898,1048,272,302]" gridcol="5" gridrow="2" pageId="4" pageNumber="487">131148</td>
<td id="075F5DCB94440000F12E6708FB6F6E53" box="[1136,1233,272,302]" gridcol="6" gridrow="2" pageId="4" pageNumber="487">6698</td>
<td id="075F5DCB94440000F1A76708FAEF6E53" box="[1273,1361,272,302]" gridcol="7" gridrow="2" pageId="4" pageNumber="487">98115</td>
</tr>
<tr id="448E34B794440000F5C96720FAEF6E2B" box="[151,1361,312,342]" gridrow="3" pageId="4" pageNumber="487">
<th id="075F5DCB94440000F5C96720FF056E2B" box="[151,187,312,342]" gridcol="0" gridrow="3" pageId="4" pageNumber="487">P3</th>
<td id="075F5DCB94440000F5AF6720FE306E2B" box="[241,398,312,342]" gridcol="1" gridrow="3" pageId="4" pageNumber="487">13941689</td>
<td id="075F5DCB94440000F4F56720FDA36E2B" box="[427,541,312,342]" gridcol="2" gridrow="3" pageId="4" pageNumber="487">12141410</td>
<td id="075F5DCB94440000F70B6720FD7F6E2B" box="[597,705,312,342]" gridcol="3" gridrow="3" pageId="4" pageNumber="487">754787</td>
<td id="075F5DCB94440000F64C6720FCD56E2B" box="[786,875,312,342]" gridcol="4" gridrow="3" pageId="4" pageNumber="487">402443</td>
<td id="075F5DCB94440000F6DC6720FBA66E2B" box="[898,1048,312,342]" gridcol="5" gridrow="3" pageId="4" pageNumber="487">213230</td>
<td id="075F5DCB94440000F12E6720FB6F6E2B" box="[1136,1233,312,342]" gridcol="6" gridrow="3" pageId="4" pageNumber="487">98</td>
<td id="075F5DCB94440000F1A76720FAEF6E2B" box="[1273,1361,312,342]" gridcol="7" gridrow="3" pageId="4" pageNumber="487">115131</td>
</tr>
<tr id="448E34B794440000F5C96799FAEF6EE4" box="[151,1361,385,409]" gridrow="4" pageId="4" pageNumber="487">
<th id="075F5DCB94440000F5C96799FAEF6EE4" box="[151,1361,385,409]" colspan="8" colspanRight="7" gridcol="0" gridrow="4" pageId="4" pageNumber="487">continued</th>
</tr>
<tr id="448E34B794440000F5C967B2FAEF6EBF" box="[151,1361,426,450]" gridrow="5" pageId="4" pageNumber="487" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-7="1">
<td id="075F5DCB94440000F5AF67B2FE306EBF" box="[241,398,426,450]" gridcol="1" gridrow="5" pageId="4" pageNumber="487">LR</td>
<td id="075F5DCB94440000F70B67B2FD7F6EBF" box="[597,705,426,450]" gridcol="3" gridrow="5" pageId="4" pageNumber="487">BV</td>
<td id="075F5DCB94440000F6DC67B2FBA66EBF" box="[898,1048,426,450]" gridcol="5" gridrow="5" pageId="4" pageNumber="487">SV</td>
<td id="075F5DCB94440000F12E67B2FB6F6EBF" box="[1136,1233,426,450]" gridcol="6" gridrow="5" pageId="4" pageNumber="487">BR</td>
</tr>
<tr id="448E34B794440000F5C967CAFAEF6E8D" box="[151,1361,466,496]" gridrow="6" pageId="4" pageNumber="487" rowspan-2="1" rowspan-4="1">
<th id="075F5DCB94440000F5C967CAFF056E8D" box="[151,187,466,496]" gridcol="0" gridrow="6" pageId="4" pageNumber="487">P 1</th>
<td id="075F5DCB94440000F5AF67CAFE306E8D" box="[241,398,466,496]" gridcol="1" gridrow="6" pageId="4" pageNumber="487">0.560.60</td>
<td id="075F5DCB94440000F70B67CAFD7F6E8D" box="[597,705,466,496]" gridcol="3" gridrow="6" pageId="4" pageNumber="487">4.465.25</td>
<td id="075F5DCB94440000F6DC67CAFBA66E8D" box="[898,1048,466,496]" gridcol="5" gridrow="6" pageId="4" pageNumber="487">3.563.92</td>
<td id="075F5DCB94440000F12E67CAFAEF6E8D" box="[1136,1361,466,496]" colspan="2" colspanRight="1" gridcol="6" gridrow="6" pageId="4" pageNumber="487">1.51.7</td>
</tr>
<tr id="448E34B794440000F5C967E2FAEF6D65" box="[151,1361,506,536]" gridrow="7" pageId="4" pageNumber="487" rowspan-2="1" rowspan-4="1">
<th id="075F5DCB94440000F5C967E2FF056D65" box="[151,187,506,536]" gridcol="0" gridrow="7" pageId="4" pageNumber="487">P2</th>
<td id="075F5DCB94440000F5AF67E2FE306D65" box="[241,398,506,536]" gridcol="1" gridrow="7" pageId="4" pageNumber="487">0.400.43</td>
<td id="075F5DCB94440000F70B67E2FD7F6D65" box="[597,705,506,536]" gridcol="3" gridrow="7" pageId="4" pageNumber="487">4.965.60</td>
<td id="075F5DCB94440000F6DC67E2FBA66D65" box="[898,1048,506,536]" gridcol="5" gridrow="7" pageId="4" pageNumber="487">5.315.83</td>
<td id="075F5DCB94440000F12E67E2FAEF6D65" box="[1136,1361,506,536]" colspan="2" colspanRight="1" gridcol="6" gridrow="7" pageId="4" pageNumber="487">1.51.7</td>
</tr>
<tr id="448E34B794440000F5C9643AFAEF6D3D" box="[151,1361,546,576]" gridrow="8" pageId="4" pageNumber="487" rowspan-2="1" rowspan-4="1">
<th id="075F5DCB94440000F5C9643AFF056D3D" box="[151,187,546,576]" gridcol="0" gridrow="8" pageId="4" pageNumber="487">P3</th>
<td id="075F5DCB94440000F5AF643AFE306D3D" box="[241,398,546,576]" gridcol="1" gridrow="8" pageId="4" pageNumber="487">0.530.62</td>
<td id="075F5DCB94440000F70B643AFD7F6D3D" box="[597,705,546,576]" gridcol="3" gridrow="8" pageId="4" pageNumber="487">3.874.29</td>
<td id="075F5DCB94440000F6DC643AFBA66D3D" box="[898,1048,546,576]" gridcol="5" gridrow="8" pageId="4" pageNumber="487">3.494.11</td>
<td id="075F5DCB94440000F12E643AFAEF6D3D" box="[1136,1361,546,576]" colspan="2" colspanRight="1" gridcol="6" gridrow="8" pageId="4" pageNumber="487">1.72.0</td>
</tr>
</table>
</paragraph>
<paragraph id="FA0136F59444FFFBF5996472FCE26C90" blockId="4.[151,1437,618,1473]" pageId="4" pageNumber="487">
Hypopygium (
<figureCitation id="62852A709444FFFBF4336472FE626DF9" box="[365,476,618,644]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Figs. 38</figureCitation>
). Tergite IX with 814 setae, 1624 µm long and with narrow (46 µm), pale, and naked anal point, 120126 µm long which in lateral view apically curved upwards (
<figureCitation id="62852A709444FFFBF1676496FBC36DD5" box="[1081,1149,654,680]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Fig. 7</figureCitation>
); posterior edge of tergite IX almost straight, anal-lateral angles are rounded and slightly projecting (
<figureCitation id="62852A709444FFFBF68D64AAFBAB6DB1" box="[979,1045,690,716]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Fig. 3</figureCitation>
); anal tergal bands X-type; median tergal band 1620 µm long. Laterosternite IX with 712 setae, 2024 µm long. Transverse sternopodeme (TSA) triangular, peaked (
<figureCitation id="62852A709444FFFBF43164E2FE676C69" box="[367,473,762,788]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Figs. 4, 8</figureCitation>
), 120128 µm high, 156176 µm wide at the base; TSA height/TSA width 0.700.77. Aedeagal lobe as in
<figureCitation id="62852A709444FFFBF4DA6506FE776C45" box="[388,457,798,824]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Fig. 8</figureCitation>
, weakly chitinized, apically with small digitiform process; phallapodeme sclerotized, 210232 µm long. Gonocoxite 400443 µm long, in middle part of dorso-internal position with small flattened transverse lobe, 98100 µm long and 2832 µm wide, directed backwards, and with large longitudinal lobe in middle, anterior outer margin of which is rounded or rounded-angular and slightly projecting (
<figureCitation id="62852A709444FFFBF19F6592FAB86CD8" box="[1217,1286,906,933]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Fig. 5</figureCitation>
); Gonostylus 197236 µm long, slightly curved, megaseta in form of wide terminal spine 812 μm long, next to it there is tooth and two setae of 1216 length µm long (
<figureCitation id="62852A709444FFFBF70365CAFD1F6C90" box="[605,673,978,1005]" captionStart="FIGURES 18" captionStartId="3.[151,264,1861,1885]" captionTargetBox="[151,1435,345,1834]" captionTargetId="figure-103@3.[151,1435,345,1837]" captionTargetPageId="3" captionText="FIGURES 18. Diamesa dragani sp. nov., male. 1, head; 2, antenna; 3, tergite IX; 4, hypopygium in dorsal view; 5, gonocoxite and gonostylus; 6, apex of gonostylus; 7, anal point in lateral view; 8, transverse sternapodeme, phallapodeme and aedeagal lobe. ATB—anterolateral anal tergal bands, MTB—median anal tergal band, PTB—posterolateral anal tergal band." figureDoi="http://doi.org/10.5281/zenodo.6457261" httpUri="https://zenodo.org/record/6457261/files/figure.png" pageId="4" pageNumber="487">Fig. 6</figureCitation>
). HR 2.012.25.
</paragraph>
<paragraph id="FA0136F59444FFFBF59965EEFE526B6D" blockId="4.[151,1437,618,1473]" box="[199,492,1013,1040]" pageId="4" pageNumber="487">
<emphasis id="C8CAEAE79444FFFBF59965EEFEBA6B6D" bold="true" box="[199,260,1014,1040]" italics="true" pageId="4" pageNumber="487">Pupa</emphasis>
and
<emphasis id="C8CAEAE79444FFFBF46465EDFECB6B6D" bold="true" box="[314,373,1013,1040]" italics="true" pageId="4" pageNumber="487">larva</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9444FFFBF5996202FC2C6A4D" pageId="4" pageNumber="487" type="diagnosis">
<paragraph id="FA0136F59444FFFBF5996202FC2C6A4D" blockId="4.[151,1437,618,1473]" pageId="4" pageNumber="487">
<emphasis id="C8CAEAE79444FFFBF5996202FEFC6B49" bold="true" box="[199,322,1050,1076]" pageId="4" pageNumber="487">Diagnosis.</emphasis>
The male of
<taxonomicName id="3DBE4D769444FFFBF4866203FDEA6B49" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[472,596,1050,1076]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="dragani" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF4866203FDEA6B49" box="[472,596,1050,1076]" italics="true" pageId="4" pageNumber="487">D. dragani</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF7056202FD0B6B49" bold="true" box="[603,693,1050,1076]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF7056202FD0B6B49" box="[603,693,1050,1076]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most closely related in the subgroup to the
<taxonomicName id="3DBE4D769444FFFBF1E36203FACD6B49" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[1213,1395,1051,1076]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="maisaraensis" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF1E36203FACD6B49" box="[1213,1395,1051,1076]" italics="true" pageId="4" pageNumber="487">D. maisaraensis</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF0256202FF766B25" bold="true" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF0256202FF766B25" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="3DBE4D769444FFFBF5A36227FE2A6B25" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[253,404,1086,1112]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="marinskiyi" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF5A36227FE2A6B25" box="[253,404,1086,1112]" italics="true" pageId="4" pageNumber="487">D. marinskiyi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF4C56226FE456B25" bold="true" box="[411,507,1086,1112]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF4C56226FE4B6B25" box="[411,501,1086,1112]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
which have anal tergal bands X-type, but the new species is a little shorter than others (
<quantity id="3D469B109444FFFBF599627AFEED6B01" box="[199,339,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="4" pageNumber="487" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.12.5 mm</quantity>
), antenna with 7 flagellomeres, AR 0.480.52, dorsocentrals 46, scutellars 1820, femur of front leg 13941689 µm long, median tergal band 1620 µm long, anal point 230126 µm long. The males of
<taxonomicName id="3DBE4D769444FFFBF065629FFF486BB9" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="maisaraensis" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF065629FFF486BB9" italics="true" pageId="4" pageNumber="487">D. maisaraensis</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF5A262B2FEEA6BB9" bold="true" box="[252,340,1194,1220]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF5A262B2FEEA6BB9" box="[252,340,1194,1220]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="3DBE4D769444FFFBF4D662B3FDA16BB9" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[392,543,1194,1220]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="marinskiyi" status="sp. nov.">
<emphasis id="C8CAEAE79444FFFBF4D662B3FDA16BB9" box="[392,543,1194,1220]" italics="true" pageId="4" pageNumber="487">D. marinskiyi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79444FFFBF77B62B2FDC06BB9" bold="true" box="[549,638,1194,1220]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="D3F9579C9444FFFBF77B62B2FDC06BB9" box="[549,638,1194,1220]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have the following characteristics respectively: total length
<quantity id="3D469B109444FFFBF04962B2FA236BB9" box="[1303,1437,1194,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="4" pageNumber="487" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.73.2 mm</quantity>
and
<quantity id="3D469B109444FFFBF59862D6FEF06B95" box="[198,334,1230,1257]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="4" pageNumber="487" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
, antenna with 8 flagellomeres, AR 0.390.42 and 0.320.39, dorsocentrals 68 and 810, scutellars 3950 and 4446, femur of front leg length 18801960 µm and 16241476 µm, median tergal band length 2832 µm and 2024 µm, anal point length 100120 µm and 104116 µm.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9444FFFBF5996322FD5A6AE1" pageId="4" pageNumber="487" type="biology_ecology">
<paragraph id="FA0136F59444FFFBF5996322FD5A6AE1" blockId="4.[151,1437,618,1473]" pageId="4" pageNumber="487">
<emphasis id="C8CAEAE79444FFFBF5996322FE926A29" bold="true" box="[199,300,1338,1364]" pageId="4" pageNumber="487">Ecology.</emphasis>
Adult males were collected from stones and boulders in mountain stream, located at an altitude of
<quantity id="3D469B109444FFFBF5C96346FF536A05" box="[151,237,1374,1400]" metricMagnitude="3" metricUnit="m" metricValue="2.063" pageId="4" pageNumber="487" unit="m" value="2063.0">2063 m</quantity>
, at a flow rate of
<quantity id="3D469B109444FFFBF4F36346FE506A05" box="[429,494,1374,1400]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="487" unit="m" value="0.6">0.6 m</quantity>
/s, with water temperatures 0.4°C, pH 7.92, general mineralization
<quantity id="3D469B109444FFFBF18A6346FAA76A05" box="[1236,1305,1374,1400]" metricMagnitude="-5" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="487" unit="g" value="0.02">0.02 g</quantity>
/l, electrical conductivity 0.03 mS (pers. comm. by S.V. Dragan).
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9444FFFBF59963BEFBFD6ABC" box="[199,1091,1446,1473]" pageId="4" pageNumber="487" type="distribution">
<paragraph id="FA0136F59444FFFBF59963BEFBFD6ABC" blockId="4.[151,1437,618,1473]" box="[199,1091,1446,1473]" pageId="4" pageNumber="487">
<emphasis id="C8CAEAE79444FFFBF59963BEFEDF6ABD" bold="true" box="[199,353,1446,1472]" pageId="4" pageNumber="487">Distribution.</emphasis>
Known only from the
<typeStatus id="250588579444FFFBF73E63BEFD2E6ABD" box="[608,656,1446,1472]" pageId="4" pageNumber="487">type</typeStatus>
locality in Sayan Mountains (
<figureCitation id="62852A709444FFFBF6BC63BEFB886ABC" box="[994,1078,1446,1473]" captionStart="FIGURES 6874" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 6874. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (6970), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of AiguillesRouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="4" pageNumber="487">Fig. 68</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>