treatments-xml/data/71/03/80/710380D184D06689752525EC98837CC5.xml

264 lines
31 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="764DA3D962EC738417CF9D8802BD443D" ID-CLB-Dataset="35544" ID-DOI="http://dx.doi.org/10.3897/zookeys.162.2370" ID-GBIF-Dataset="f8228b70-5cf8-4846-b2b2-91aec46ab0a5" ID-PMC="PMC3253663" ID-Pensoft-Pub="1313-2970-162-59" ID-PubMed="22303126" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-162-59" ModsDocOrigin="ZooKeys 162" ModsDocTitle="A conspectus on the Canacidae (Diptera) of Brazil" checkinTime="1451249400868" checkinUser="pensoft" docAuthor="Mathis, Wayne N. &amp; Marinoni, Luciane" docDate="2012" docId="710380D184D06689752525EC98837CC5" docLanguage="en" docName="ZooKeys 162: 59-92" docOrigin="ZooKeys 162" docSource="http://dx.doi.org/10.3897/zookeys.162.2370" docTitle="Canacidae Jones" docType="treatment" docVersion="7" lastPageNumber="62" masterDocId="0C201957F3148E5FFFE2FF8834486856" masterDocTitle="A conspectus on the Canacidae (Diptera) of Brazil" masterLastPageNumber="92" masterPageNumber="59" pageNumber="61" updateTime="1732729939142" updateUser="ExternalLinkService">
<mods:mods id="524758740ACFFEA8ECD85A239B3387CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0A19432061FC9B3D0A039CD09DC8E684">
<mods:title id="CE2F816848D0F43EE052111E828C481A">A conspectus on the Canacidae (Diptera) of Brazil</mods:title>
</mods:titleInfo>
<mods:name id="5179B6239B1644B8EC0B4A6D4F78CFFD" type="personal">
<mods:role id="CF953D849701134B93A40921C18767EA">
<mods:roleTerm id="33373825E4064EC3DC4D50629DF9E1DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2BB25AEAF4AA379EBBFC0CC054DCC0D5">Mathis, Wayne N.</mods:namePart>
</mods:name>
<mods:name id="6466A0B51C7E63A838FCEF6F60416D3F" type="personal">
<mods:role id="BD68D247D9D5F1F9B7602D0215B6B6D3">
<mods:roleTerm id="58C157BF80E2B6F6CCE101D5F1F90B46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8C01EEA1AC5AC12EB3FD3981C0EAB63A">Marinoni, Luciane</mods:namePart>
</mods:name>
<mods:typeOfResource id="4A10AE4F95C2192BFF6B7A146641D429">text</mods:typeOfResource>
<mods:relatedItem id="65773C55BE0407907F32BEF6758B5F42" type="host">
<mods:titleInfo id="1F8C31091CF37E2960FA42923A5EE275">
<mods:title id="D53B744864DC79B3756411494BECEC2A">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="3E50C959C54C53872C79A34149444F56">
<mods:date id="A086D84DFE9D03814B922609E5FD9EEC">2012</mods:date>
<mods:detail id="8BC2C24F860A43AE97319233010F14EF" type="volume">
<mods:number id="3128B5C3F04360ACABB544581A164382">162</mods:number>
</mods:detail>
<mods:extent id="2B0C0635D6D342D007EC94221B5483D3" unit="page">
<mods:start id="97DE035EF2C6E566DCDACF53191713C2">59</mods:start>
<mods:end id="BE9129D6356402AF8E4D11EF59613886">92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0F2C4F2D3DF63474F004AFE9B0FE89D0">
<mods:url id="366B1A7FA9DD2A3D33804812055BBE19">http://dx.doi.org/10.3897/zookeys.162.2370</mods:url>
</mods:location>
<mods:classification id="27BA71CE627BDF9A2AD723218CD2EE22">journal article</mods:classification>
<mods:identifier id="74E583A73AB2EE09FD173503393E4AD6" type="DOI">http://dx.doi.org/10.3897/zookeys.162.2370</mods:identifier>
<mods:identifier id="0737BBDBE3F72F53A8FFF1E292FDAE13" type="Pensoft-Pub">1313-2970-162-59</mods:identifier>
</mods:mods>
<treatment id="710380D184D06689752525EC98837CC5" ID-GBIF-Taxon="152032960" LSID="urn:lsid:plazi:treatment:710380D184D06689752525EC98837CC5" httpUri="http://treatment.plazi.org/id/710380D184D06689752525EC98837CC5" lastPageId="4" lastPageNumber="62" pageId="2" pageNumber="61" scope_order="Diptera">
<subSubSection id="383742AB45589DBCC403F16F3715FA47" pageId="2" pageNumber="61" type="nomenclature">
<paragraph id="BF532E1B37847EA7C68F80580915CB37" pageId="2" pageNumber="61">
Family
<taxonomicName id="74C651F58436B53EB5FEE9CDCB3B90A7" ID-CoL="622J4" ID-ENA="1262362" LSID="http://species-id.net/wiki/Canacidae" authority="Jones" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="family">Canacidae Jones</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="49FDEFD90383957256D7F5BE13062BA8" pageId="2" pageNumber="61" type="reference_group">
<paragraph id="FBE16BF49ABE6225241FD2BFEF617EB1" pageId="2" pageNumber="61">
<taxonomicName id="F627AB8E1594CB1CA6F1DFD437945CC4" genus="Canacenae" lsidName="Canacenae" pageId="2" pageNumber="61" rank="genus">Canacenae</taxonomicName>
<bibRefCitation id="236A22D50272BD1AD23F46CE98F9393E" author="Jones, BJ" journalOrPublisher="University of California Publications in Entomology" pageId="24" pageNumber="83" pagination="153 - 198" title="Catalogue of the Ephydridae, with bibliography and description of new species." volume="1" year="1906">Jones 1906</bibRefCitation>
: 170, 198 [as a subfamily of
<taxonomicName id="B2895335C1728807473A671492E84717" class="Insecta" family="Ephydridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="family">Ephydridae</taxonomicName>
, incorrect formation of the family-group name]. Type genus:
<taxonomicName id="127D17AC2DA788D4DF7B08E4FA024E57" class="Insecta" family="Canacidae" genus="Canace" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canace" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Canace</taxonomicName>
Haliday 1837.
</paragraph>
<paragraph id="BC15F2377FAB51B84F9F20252731F64D" pageId="2" pageNumber="61">
<taxonomicName id="2EAAC74F1E558DB0E7552BD0275F61AB" family="Canaceidae" lsidName="" pageId="2" pageNumber="61" rank="family">Canaceidae</taxonomicName>
.
<bibRefCitation id="C4CDB47852A61C68B3E27EEC80DD396F" pageId="2" pageNumber="61">Hendel 1916</bibRefCitation>
: 297 [incorrect formation of the family-group name].
<bibRefCitation id="45E1EEEA13F123678A17001E637E8399" author="Wirth, WW" journalOrPublisher="Bishop Museum" pageId="27" pageNumber="86" pagination="245 - 275" title="A revision of the dipterous family Canaceidae. Occasional Papers of Bernice P." volume="20" year="1951">Wirth 1951</bibRefCitation>
: 245-275 [revision]; 1975: 1-5 [Neotropical catalog]; 1987: 1079-1083 [North American manual].
</paragraph>
<paragraph id="59ADD74C612A119BBCC02CCA5868E2C3" pageId="2" pageNumber="61">
<taxonomicName id="42E3AF7CDC76D22E847D6CCFC6991B82" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
.
<bibRefCitation id="85979F769DE7ABBB5E1C03D1E120FCA2" author="Enderlein, G" journalOrPublisher="Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin" pageId="23" pageNumber="82" pagination="235 - 250" title="Dipterologica, III." volume="1935" year="1935">Enderlein 1935</bibRefCitation>
: 235.
<bibRefCitation id="5EABF64A726E88CCE45D2D17D407072F" author="Mathis, WN" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="25" pageNumber="84" pagination="1 - 29" title="Studies of Canacidae (Diptera), I: Suprageneric revision of the family, with revisions of new Tribe Dynomiellini and new genus Isocanace." volume="347" year="1982">Mathis 1982</bibRefCitation>
: 1-29 [classification].
<bibRefCitation id="3136D420FC04A6B4EA2B2F0144F67357" author="Buck, M" journalOrPublisher="Systematic Entomology" pageId="23" pageNumber="82" pagination="377 - 404" title="A new family and genus of acalypterate flies from the Neotropical Region, with a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora)." url="doi: 10.1111/j.1365-3113.2006.00328.x" volume="31" year="2006">Buck 2006</bibRefCitation>
: 391-392 [familial status].
<bibRefCitation id="4ED4AA5E6800E7B6DA5173107C846F20" author="Munari, L" journalOrPublisher="Zootaxa" pageId="26" pageNumber="85" pagination="1 - 84" title="World catalog of the family Canacidae (including Tethinidae) (Diptera), with keys to the supraspecific taxa." volume="2471" year="2010">Munari and Mathis 2010</bibRefCitation>
: 1-84 [world catalog].
</paragraph>
<paragraph id="BFF23ED3B0A84588A19AEF81CD42B3F0" pageId="2" pageNumber="61">
<taxonomicName id="F1081F019EF32125A8666F7C0D78CFEC" family="Tethinidae" lsidName="" pageId="2" pageNumber="61" rank="family">Tethinidae</taxonomicName>
<bibRefCitation id="69D0B07501806CF2A0BB86D93891C023" pageId="2" pageNumber="61">Hendel 1916</bibRefCitation>
: 297; 1917: 45. Type genus:
<taxonomicName id="F3664B33CDBEEE382DBF3E798D45B89A" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Tethina</taxonomicName>
Haliday.
<bibRefCitation id="D2640B3E204E12F7A9AFC62FBD6C371F" pageId="2" pageNumber="61">Foster 1976b</bibRefCitation>
: 1-4 [Neotropical catalog].
<bibRefCitation id="F9EF4626519119058F43B940048FBDFB" author="Mathis, WN" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="25" pageNumber="84" pagination="1 - 27" title="World catalog of the family Tethinidae (Diptera)." url="doi: 10.5479/si.00810282.584" volume="584" year="1996">Mathis and Munari 1996</bibRefCitation>
: 1-27 [world catalog].
<bibRefCitation id="5CB6D1B8937083874B8DC27B3A25D36C" author="McAlpine, DK" journalOrPublisher="Records of the Australian Museum" pageId="25" pageNumber="84" pagination="27 - 64" title="The surge flies (Diptera, Canacidae, Zaleinae) of Australasia and notes on tethinids-canacid morphology and relationships." url="doi: 10.3853/j.0067-1975.59.2007.1468" volume="57" year="2007">McAlpine 2007</bibRefCitation>
: 42 [synonymy].
</paragraph>
</subSubSection>
<subSubSection id="C0ED88B8CD5A1BB97D46ACDA34872301" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="diagnosis">
<paragraph id="BC2FBAB4AB6DFA11E6922DC5119EDDC0" pageId="2" pageNumber="61">Diagnosis.</paragraph>
<paragraph id="BC6173EB15D854AF9635F5DD3A35B61F" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
The family
<taxonomicName id="4280CA46857910240D43D561351514DD" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
, sensu lato, is distinguished from other families of the
<taxonomicName id="B4699C64EA9D4F8C663B01259EC0639E" genus="Carnoidea" lsidName="Carnoidea" pageId="2" pageNumber="61" rank="genus">Carnoidea</taxonomicName>
by the following combination of characters: Exclusively or tending to occur in saline habitats (secondarily in freshwater habitats). Minute to moderately small flies, length 0.91-5.0 mm. Head: Postocellar setae developed (absent or reduced in some
<taxonomicName id="6A8F6FBF1B078101C29DC522DAEE8A6A" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
); dorsal fronto-orbital seta lateroclinate; oral vibrissae weakly differentiated, except for
<taxonomicName id="DEAC5ADFFC1BA35883F1319054B9FCF8" class="Insecta" family="Canacidae" genus="Dasyrhicnoessa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyrhicnoessa" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Dasyrhicnoessa</taxonomicName>
Hendel species. Arista dorsal. Face sometimes characterized by 2 shiny protuberances laterad to the facial cavity, just above vibrissal pore (
<taxonomicName id="205F28A7DE4CC23C153F8F9C627DF4CD" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Tethina</taxonomicName>
,
<taxonomicName id="F4C421AEBEAFE5984A03AAB32269B533" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudorhicnoessa" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Pseudorhicnoessa</taxonomicName>
) or nearby (
<taxonomicName id="45FB28B663CEF153978F3B59E4CCF602" class="Insecta" family="Canacidae" genus="Afrotethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afrotethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Afrotethina</taxonomicName>
,
<taxonomicName id="C7A8172F8AA93267ADDF009BF912D23B" class="Insecta" family="Canacidae" genus="Horaismoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horaismoptera" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Horaismoptera</taxonomicName>
); face strongly depressed and short (
<taxonomicName id="F56086D198EB964534E95513F26ED77A" class="Insecta" family="Canacidae" genus="Dasyrhicnoessa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dasyrhicnoessa" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Dasyrhicnoessa</taxonomicName>
,
<taxonomicName id="D015754A8958C992B35A4C81B943133B" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Horaismopterinae">Horaismopterinae</taxonomicName>
) or with medial carina (
<taxonomicName id="FF676EFF0765BB33E3C7A6F2FE3E3AF2" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Tethina</taxonomicName>
) or even distinctly convex (
<taxonomicName id="3BF92DE6EF1140B2FD5C2687E41C4BF9" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
). Gena bare, except for ventral or nearly ventral row of setae (peristomal setae), or even with a few anaclinate, strong setae (
<taxonomicName id="27ADD968A5E8D8F4B4EC14C2F531404E" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
). Buccal parts generally strongly sclerotized in
<taxonomicName id="F8DD5468023DCA44FB8B0471B999B52C" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
. Thorax: Precoxal bridge developed. Prescutellar acrostichal setae developed; presutural dorsocentral setae differentiated; anepisternum with 2-3 developed posterior setae, bearing enlarged, dorsally curved seta at posterodorsal corner; usually 1 katepisternal seta present; proepisternal seta developed. Wing generally hyaline, bearing fine, dense microtrichia; subcosta weakened apically, close to vein R1; vein A1 short (except in the sub-Antarctic genus
<taxonomicName id="9D80464CB59A18B2BF8B1D2D831FA071" class="Insecta" family="Canacidae" genus="Apetaenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apetaenus" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Apetaenus</taxonomicName>
); vein A2 long, present as a fold. Abdomen: Pregenital sclerites of male short and fused; male tergite 6 fused with sternite 8, forming a usually symmetrical (except in some species of
<taxonomicName id="3B4ECC296F6111C434F67D6143DD71DB" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Tethina</taxonomicName>
), pregenital sclerite; male sternite 7 lost; postgonites firmly connected laterally to base of phallapodeme, distinctly anterior to basiphallus; hypandrium forming a sheath or phallic mantle around the postgonite and basiphallus; epandrium bearing 1-2 pairs of surstyli ventrally, sometimes anterior surstylus lacking (
<taxonomicName id="31C9FEFD2CA26A806A767FF884893D1A" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
,
<taxonomicName id="0349BBF56A262CC62FB9F28DCF7BC9B3" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina" order="Diptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Tethina</taxonomicName>
); posterior surstylus partially articulated or fused with epandrium; inner basal corner of surstylus connected to broad interparameral sclerite; cercus very short to exceptionally developed (
<taxonomicName id="C9F15BDB2DAFF0FD9242893C2565A5BC" lsidName="" pageId="2" pageNumber="61" rank="subFamily" subFamily="Horaismopterinae">Horaismopterinae</taxonomicName>
); postabdomen of female more or less telescopically retractile; 2 sclerotized spermathecae variable in shape, below with a narrower
<pageBreakToken id="1DA2467F1B67211575AF1CFD50C09864" pageId="3" pageNumber="62" start="start">cylindrical</pageBreakToken>
extension into the spermathecal duct; cercus subcylindrical to compressed, or even tapered distally, sometimes bearing stout to pointed, spinelike setulae.
</paragraph>
</subSubSection>
<subSubSection id="8FD38F4C0C8841E20C77819D2CEB1159" pageId="3" pageNumber="62" type="discussion">
<paragraph id="227007DFDC377FC43616004135FF71F8" pageId="3" pageNumber="62">Discussion.</paragraph>
<paragraph id="5E9E75B95F92FBA607B5C388F65B4D5A" pageId="3" pageNumber="62">
Our concept of
<taxonomicName id="F079D809BDF09551043DC23126D9FC30" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
includes what had been considered as two families, the
<taxonomicName id="ECC6EA829E5A3012C8E3F56A96C7216E" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
and
<taxonomicName id="2597524BDF36AEB8332AE7A13086FCDB" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
. At the familial level,
<bibRefCitation id="32E72200CD2123DF4554E8EEAFE3A85E" pageId="3" pageNumber="62">J. F. McAlpine (1989</bibRefCitation>
: 1472) identified five synapomorphies that link
<taxonomicName id="1425A1C50E07F3944E49761CDF229D33" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
with
<taxonomicName id="8FB78877B565E34C19F39F326FC51DE9" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
and noted that &quot;...these are clear indications of a sister-group relationship between them ... and may even indicate that they are subgroups of a single family.&quot; Other authors (
<bibRefCitation id="B56B12E136C555312F107AA2C3F56236" pageId="3" pageNumber="62">Hennig 1958</bibRefCitation>
;
<bibRefCitation id="28E7A6162B6B2BBC5F6FF2033DB7A1A0" pageId="3" pageNumber="62">Griffiths 1972</bibRefCitation>
;
<bibRefCitation id="1902E9D1203912AD2FAE755FE8BE59FD" author="McAlpine, DK" journalOrPublisher="Memoirs of the Entomological Society of Washington" pageId="25" pageNumber="84" pagination="108 - 117" title="A new genus of Australian littoral flies (Diptera:? Canacidae)." volume="10" year="1982">McAlpine 1982</bibRefCitation>
;
<bibRefCitation id="091687F0E0C72A4CDB61621994006EBF" author="Freidberg, A" journalOrPublisher="Entomologica Scandinavica" pageId="24" pageNumber="83" pagination="447 - 457" title="A study of Zaleinae, a taxon transitional between Canacidae and Tethinidae (Diptera), with the description of a new genus and species." url="doi: 10.1163/187631295X00107" volume="26" year="1995">Freidberg 1995</bibRefCitation>
) have also suggested a relationship with the family
<taxonomicName id="D146FE04F4A7520CFAB42A64AFFAFE1C" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
, and
<bibRefCitation id="7AB52DF143EB8F710E7A8A4F86A90905" pageId="3" pageNumber="62">Griffiths (1972)</bibRefCitation>
further noted some affinities with the
<taxonomicName id="DCB89AA517D04E5977BC5763A2344AF5" class="Insecta" family="Chloropidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
and
<taxonomicName id="1FF17DE0ECAC77909FABB7DB774EDE90" class="Insecta" family="Milichiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Milichiidae</taxonomicName>
. According to J. F.
<normalizedToken id="F6866F891ABDA5DEB81BB1F957D283C8" originalValue="McAlpines">McAlpine's</normalizedToken>
(1989) cladogram, which included an analysis of 25 characters for the families
<taxonomicName id="8C6D2413CA009ABA5517CE68A84A8C19" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
and
<taxonomicName id="47A23B5115CEA895A44E278E75E2DEB8" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
, the superfamily
<taxonomicName id="E77702591816FA8AAE75759432B65D9C" genus="Carnoidea" lsidName="Carnoidea" pageId="3" pageNumber="62" rank="genus">Carnoidea</taxonomicName>
(=
<taxonomicName id="FF2F84A7A38FA49445526808CE7C85E6" genus="Chloropoidea" lsidName="Chloropoidea" pageId="3" pageNumber="62" rank="genus">Chloropoidea</taxonomicName>
) comprises the families with the following relationships in parenthetic notation: ((
<taxonomicName id="0B5050AB98904E7A508EABD6CDDF6273" class="Insecta" family="Australimyzidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Australimyzidae</taxonomicName>
,
<taxonomicName id="617C5506372F115E416560F44420DE70" class="Insecta" family="Braulidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Braulidae</taxonomicName>
)
<taxonomicName id="093FD1E0759316A219CF92A4A35B2FCC" class="Insecta" family="Carnidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Carnidae</taxonomicName>
)((
<taxonomicName id="A2D28B53CF7CF51BCC1B7F45788BF673" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
,
<taxonomicName id="C0CAA9E843F809A0F800BCBA683EED68" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
)((
<taxonomicName id="59CAF62B765947E54A1D866A00FB4AAC" class="Insecta" family="Milichiidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Milichiidae</taxonomicName>
,
<taxonomicName id="37195C4C275140B102F2C1CC38622AF8" family="Risidae" lsidName="" pageId="3" pageNumber="62" rank="family">Risidae</taxonomicName>
) ((
<taxonomicName id="8537E4DE7B4C5A646A4E868DA7F8A652" class="Insecta" family="Cryptochetidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Cryptochetidae</taxonomicName>
,
<taxonomicName id="EB638FA2C2F878CE34273747DEB16A71" class="Insecta" family="Chloropidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
)))).
</paragraph>
<paragraph id="CF10BD69BB0591631BF777CB5C21F7F3" pageId="3" pageNumber="62">
More recently,
<bibRefCitation id="A858160B7817E8BE3B7C8E070A70DF6D" author="Buck, M" journalOrPublisher="Systematic Entomology" pageId="23" pageNumber="82" pagination="377 - 404" title="A new family and genus of acalypterate flies from the Neotropical Region, with a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora)." url="doi: 10.1111/j.1365-3113.2006.00328.x" volume="31" year="2006">Buck (2006)</bibRefCitation>
and D. K.
<bibRefCitation id="8C59177073EAF671E1F5FA704BF49B4A" author="McAlpine, DK" journalOrPublisher="Records of the Australian Museum" pageId="25" pageNumber="84" pagination="27 - 64" title="The surge flies (Diptera, Canacidae, Zaleinae) of Australasia and notes on tethinids-canacid morphology and relationships." url="doi: 10.3853/j.0067-1975.59.2007.1468" volume="57" year="2007">McAlpine (2007)</bibRefCitation>
provided rather compelling character evidence, substantiating that these two families are closely associated, and more specifically that the
<taxonomicName id="528BBAAD70B02C39F0D25513F189E37F" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
sensu stricto are an included lineage within the
<taxonomicName id="FA0DDC37279D1D7402D9BC2DAE979819" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
. Thus, not to include the
<taxonomicName id="33DC5429F7ABC3FCE2BB6157DFD39F91" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
within the
<taxonomicName id="370AEB9E098DDF48161E0254B9A75607" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
would render the
<taxonomicName id="43E19C1C9DDDAAE782215A5D335F6A3D" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
as a paraphyletic family. Buck and D. K. McAlpine cited ten synapomorphies that corroborate the monophyly of the family
<taxonomicName id="7AFBC6197A7969546E4AD91543FB4577" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
sensu lato (the family-group name
<taxonomicName id="ED514B90AC1480136AF22C0545BF5B24" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
is older than
<taxonomicName id="0A3CBBAB10FE222BE01C67B29573FD61" family="Tethinidae" lsidName="" pageId="3" pageNumber="62" rank="family">Tethinidae</taxonomicName>
). These synapomorphies are (only derived state cited): (1) Precoxal bridge present; (2) anepisternum with enlarged, dorsally curved setae at posteroventral corner; (3) vein A2 long, present as a fold; (4) male sternite 6 reduced and divided medially; (5) male tergite 6 fused with sternite 8, forming a symmetrical pregenital sclerite; (6) male sternite 7 lost; (7) postgonites firmly connected laterally to base of phallapodeme, distinctly anterior to basiphallus; (8) hypandrium forming a sheath or phallic mantle around the postgonite and basiphallus; (9) cuticle of larva with covering of fine spicules, and (10) halobiontic in habitat preference, secondarily in freshwater habitats.
<bibRefCitation id="D06CFC7E621FC5EF49C9DD44C254934D" author="Buck, M" journalOrPublisher="Systematic Entomology" pageId="23" pageNumber="82" pagination="377 - 404" title="A new family and genus of acalypterate flies from the Neotropical Region, with a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora)." url="doi: 10.1111/j.1365-3113.2006.00328.x" volume="31" year="2006">Buck (2006)</bibRefCitation>
further suggested that the sister group to
<taxonomicName id="66CE1FDFAF21E32066B5E9C1E3C04E31" lsidName="" pageId="3" pageNumber="62" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
sensu stricto is the subfamily
<taxonomicName id="026B504582D8B65ADE974BDCE991BA7D" lsidName="" pageId="3" pageNumber="62" rank="subFamily" subFamily="Apetaeninae">Apetaeninae</taxonomicName>
and not
<taxonomicName id="3E3902C524D506E224C3CD5CFC2FC99A" lsidName="" pageId="3" pageNumber="62" rank="subFamily" subFamily="Zaleinae">Zaleinae</taxonomicName>
and provided four characters as corroborative evidence for this relationship: (1) antennae broadly separated, inserted more or less on protuberant facial tubercles; (2) clypeus distinctly enlarged and produced anteriorly; (3) prementum distinctly emarginated apically; and (4) tentorial arms of head capsule enormously developed and strongly sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="49E1DB22AD302BBA9C514D12414E318E" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="key to subfamilies of canacidae sensu lato from brazil">
<paragraph id="9478EEE8DF8ACD1642A976639A713F75" pageId="3" pageNumber="62">
Key to Subfamilies of
<taxonomicName id="D82D54AC023A1F1A6593B36D320C79CE" class="Insecta" family="Canacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="family">Canacidae</taxonomicName>
sensu lato from Brazil
</paragraph>
<paragraph id="ABE72C4B26B692F329AB26B5EC69AF9B" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<table id="FFDE8D50982F3B41E653B970113E5155" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<tr id="6822A8C63A7F04CE0A83515118980462" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
<td id="FA06A30600994CE82A706EF8EFA84C36" colspan="1" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" rowspan="1">
<taxonomicName id="3D2D1D90F6D6174F11BFD9B3CA5773B3" class="Insecta" family="Canacidae" genus="Tethina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tethina lusitanica" order="Diptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="lusitanica">Tethina lusitanica</taxonomicName>
<pageBreakToken id="6A7AC07F3B2B32A5859510A04FB94EFF" pageId="4" pageNumber="63" start="start">12</pageBreakToken>
</td>
<td id="DDA4F971F729F760BFF7B76342894FDE" colspan="1" pageId="4" pageNumber="63" rowspan="1">
<taxonomicName id="0A75206DA6013F892DEFA07403A90F6A" lsidName="" pageId="4" pageNumber="63" rank="subFamily" subFamily="Tethininae">Tethininae</taxonomicName>
</td>
</tr>
<tr id="8504EAE728C0F9C12B16540FF55809AF" pageId="4" pageNumber="63">
<td id="C1DBD9CB9DBDD4A0FF9960A9541BA80E" colspan="1" pageId="4" pageNumber="63" rowspan="1">12</td>
<td id="73148FA7303CBF6FA5B002045A57F77E" colspan="1" pageId="4" pageNumber="63" rowspan="1">
<taxonomicName id="A76B3BE555443D231B8AEBCFC3A1A79B" lsidName="" pageId="4" pageNumber="63" rank="subFamily" subFamily="Apetaeninae">Apetaeninae</taxonomicName>
</td>
</tr>
<tr id="104488172AA7C6C246AB44E204C52F8C" pageId="4" pageNumber="63">
<td id="13A5B457ED44BA0EADC02007CD7F35B8" colspan="1" pageId="4" pageNumber="63" rowspan="1">12</td>
<td id="8F21CFC3BB7991868DE6FF5BA8E19759" colspan="1" pageId="4" pageNumber="63" rowspan="1">
<taxonomicName id="70BA3A0118C462C110682F42D17BACD9" lsidName="" pageId="4" pageNumber="63" rank="subFamily" subFamily="Canacinae">Canacinae</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>