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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.509.8897" ID-GBIF-Dataset="ce2f4ab9-1ee1-43ff-9f75-952ff0bee647" ID-PMC="PMC4493342" ID-Pensoft-Pub="1313-2970-509-13" ID-PubMed="26175602" ID-ZBK="6DD8D4EB478844E2B34C995D87F2A0DE" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-509-13" ModsDocOrigin="ZooKeys 509" ModsDocTitle="Systematics of the Madagascar Anelosimus spiders: remarkable local richness and endemism, and dual colonization from the Americas" checkinTime="1451244237153" checkinUser="pensoft" docAuthor="Agnarsson, Ingi, Jencik, Brian B., Veve, Giselle M., Hanitriniaina, Sahondra, Agostini, Diego, Goh, Seok Ping, Pruitt, Jonathan &amp; Kuntner, Matjaz" docDate="2015" docId="70C0731CA03DF52D596D4B49322585B2" docLanguage="en" docName="ZooKeys 509: 13-52" docOrigin="ZooKeys 509" docSource="http://dx.doi.org/10.3897/zookeys.509.8897" docTitle="Anelosimus tita Agnarsson, Kuntner &amp; Jencik, sp. n." docType="treatment" docUuid="85C991FF-0021-4D6E-80A0-65813CCD3F5E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="32" masterDocId="CF43D14EFFC46147FFB7FF82FFC12F62" masterDocTitle="Systematics of the Madagascar Anelosimus spiders: remarkable local richness and endemism, and dual colonization from the Americas" masterLastPageNumber="52" masterPageNumber="13" pageNumber="31" updateTime="1668160423771" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Systematics of the Madagascar Anelosimus spiders: remarkable local richness and endemism, and dual colonization from the Americas</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Agnarsson, Ingi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jencik, Brian B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Veve, Giselle M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hanitriniaina, Sahondra</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Agostini, Diego</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goh, Seok Ping</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pruitt, Jonathan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kuntner, Matjaz</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>509</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.509.8897</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.509.8897</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-509-13</mods:identifier>
<mods:identifier type="ZBK">6DD8D4EB478844E2B34C995D87F2A0DE</mods:identifier>
<mods:identifier type="ZooBank">6DD8D4EB478844E2B34C995D87F2A0DE</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152060444" LSID="urn:lsid:zoobank.org:act:85C991FF-0021-4D6E-80A0-65813CCD3F5E" httpUri="http://treatment.plazi.org/id/70C0731CA03DF52D596D4B49322585B2" lastPageId="19" lastPageNumber="32" pageId="18" pageNumber="31">
<subSubSection pageId="18" pageNumber="31" type="multiple">
<paragraph pageId="18" pageNumber="31">Taxon classification Animalia Araneae Theridiidae</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="31" type="nomenclature">
<paragraph pageId="18" pageNumber="31">
<taxonomicName LSID="http://zoobank.org/85C991FF-0021-4D6E-80A0-65813CCD3F5E" authority="Agnarsson, Kuntner &amp; Jencik" class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus tita" order="Araneae" pageId="18" pageNumber="31" phylum="Arthropoda" rank="species" species="tita">Anelosimus tita Agnarsson, Kuntner &amp; Jencik</taxonomicName>
<taxonomicNameLabel pageId="18" pageNumber="31">sp. n.</taxonomicNameLabel>
Fig. 10
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="type material">
<paragraph pageId="19" pageNumber="32">
<pageBreakToken pageId="19" pageNumber="32" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="19" pageNumber="32">
Holotype female from Ambohitantely Special Reserve (
<geoCoordinate direction="south" orientation="latitude" precision="55" value="-18.161">18.161°S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="47.302">47.302°E</geoCoordinate>
), 1500 m alt, Analamanga region, Ankazobe district, Madagascar, 28.iv.2008, montane forest, col. Agnarsson and Kuntner, in NMNH.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="other material">
<paragraph pageId="19" pageNumber="32">Other material.</paragraph>
<paragraph pageId="19" pageNumber="32">Only known from holotype.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="etymology">
<paragraph pageId="19" pageNumber="32">Etymology.</paragraph>
<paragraph pageId="19" pageNumber="32">
The species is a noun in apposition named in the honor of the first
<normalizedToken originalValue="authors">author's</normalizedToken>
mother-in-law Yadira Collado Ulloa, affectionately known to her grandchildren as
<normalizedToken originalValue="Tita">'Tita'</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="diagnosis">
<paragraph pageId="19" pageNumber="32">Diagnosis.</paragraph>
<paragraph pageId="19" pageNumber="32">
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus tita" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="tita">Anelosimus tita</taxonomicName>
can be diagnosed from other Madagascan
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="genus">Anelosimus</taxonomicName>
by the triangular shape of the septum (Fig. 10E) and on the basis of the following unique mtDNA nucleotide substitutions at the following standard DNA barcode alignment positions: T (30), G (37), T (80), T (81), T(82), A(83), G (109), G (214), T(220), G (319), T (328), T (586), T (625), G (873), G (883), A (903), G (919). It can also be readily diagnosed from most other
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="genus">Anelosimus</taxonomicName>
based the following partially shared nucleotide substitutions, and all other species by their unique combination: T (22, except
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus hookeri" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="hookeri">Anelosimus hookeri</taxonomicName>
and
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus huxleyi" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="huxleyi">Anelosimus huxleyi</taxonomicName>
), T (121, except
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus nazariani" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="nazariani">Anelosimus nazariani</taxonomicName>
), G (190, except some
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus huxleyi" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="huxleyi">Anelosimus huxleyi</taxonomicName>
), G (262, except
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus nazariani" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="nazariani">Anelosimus nazariani</taxonomicName>
), T (532, except some
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus huxleyi" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="huxleyi">Anelosimus huxleyi</taxonomicName>
), T (553, except
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus andasibe" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="andasibe">Anelosimus andasibe</taxonomicName>
).
</paragraph>
<caption pageId="19" pageNumber="32">
<paragraph pageId="19" pageNumber="32">
Figure 10.
<taxonomicName class="Arachnida" family="Theridiidae" genus="Anelosimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anelosimus tita" order="Araneae" pageId="19" pageNumber="32" phylum="Arthropoda" rank="species" species="tita">Anelosimus tita</taxonomicName>
:
<normalizedToken originalValue="AD">A-D</normalizedToken>
female lateral, dorsal, ventral, and ventral abdomen E epigynum ventral F epigynum cleared, dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="description">
<paragraph pageId="19" pageNumber="32">Description.</paragraph>
<paragraph pageId="19" pageNumber="32">Female: Total length 3.87 Cephalothorax 1.9 long, 1.34 wide, 1.09 high, brown. Sternum 1.02 long, 0.87 wide, extending half way between coxae IV, orange. Abdomen 2.44 long, 1.68 wide, 1.33 high. Mixed pattern of white, grey, and black. Eyes subequal in size about 0.11 in diameter. Clypeus height about 2 times one AME diameter. Chelicerae with one large tooth, three denticles prolaterally. Leg I femur 2.01, patella 0.66, tibia 2.02, metatarsus 1.84, tarsus 0.91. Leg formula 3421, with leg 1 significantly longer than leg 2. Legs alternating between light orange and dark brown bands. 3-4 small trichobothria dorsally on tibia, 3 on tibia 1. 3 or 4 dorsal trichobothria on female palpal tibia.</paragraph>
<paragraph pageId="19" pageNumber="32">Variation: only known from holotype.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="distribution">
<paragraph pageId="19" pageNumber="32">Distribution.</paragraph>
<paragraph pageId="19" pageNumber="32">Only known from type locality.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="natural history">
<paragraph pageId="19" pageNumber="32">Natural history.</paragraph>
<paragraph pageId="19" pageNumber="32">Unknown, predicted to be subsocial.</paragraph>
</subSubSection>
</treatment>
</document>