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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1555" ID-GBIF-Dataset="933c9e4c-64f0-4c78-b531-8cdf35cfeed5" ID-PMC="PMC3260768" ID-Pensoft-Pub="1313-2970-130-307" ID-PubMed="22259285" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-307" ModsDocOrigin="ZooKeys 130" ModsDocTitle="New Chironomidae (Diptera) with elongate proboscises from the Late Jurassic of Mongolia" checkinTime="1451249862855" checkinUser="pensoft" docAuthor="Lukashevich, Elena D. &amp; Przhiboro, Andrey A." docDate="2011" docId="7098AD69E04460D58544D5374885AC16" docLanguage="en" docName="ZooKeys 130: 307-322" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1555" docTitle="Podonomius robustus Lukashevich &amp; Przhiboro, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="311" masterDocId="FFF25E04FFF721571631880EFFFE757E" masterDocTitle="New Chironomidae (Diptera) with elongate proboscises from the Late Jurassic of Mongolia" masterLastPageNumber="322" masterPageNumber="307" pageNumber="311" updateTime="1668152268990" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Chironomidae (Diptera) with elongate proboscises from the Late Jurassic of Mongolia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lukashevich, Elena D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Przhiboro, Andrey A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>130</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>307</mods:start>
<mods:end>322</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.130.1555</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1555</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-130-307</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030924" LSID="urn:lsid:zoobank.org:act:189103E1-9FA9-45DE-A4E4-52EF0CE68913" httpUri="http://treatment.plazi.org/id/7098AD69E04460D58544D5374885AC16" lastPageNumber="311" pageId="4" pageNumber="311">
<subSubSection pageId="4" pageNumber="311" type="nomenclature">
<paragraph pageId="4" pageNumber="311">
?
<taxonomicName LSID="urn:lsid:zoobank.org:act:189103E1-9FA9-45DE-A4E4-52EF0CE68913" class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius robustus" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="species" species="robustus">Podonomius robustus</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="311">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="311" type="etymology">
<paragraph pageId="4" pageNumber="311">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="311">
From Latin
<normalizedToken originalValue="“robustus”">&quot;robustus&quot;</normalizedToken>
for stout, after the total habitus.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="311" type="material examined">
<paragraph pageId="4" pageNumber="311">Material examined.</paragraph>
<paragraph pageId="4" pageNumber="311">
Holotype: Part and counterpart of partly preserved female PIN 4270/2254
<normalizedToken originalValue="±">+/-</normalizedToken>
, SW Mongolia, Shar Teg (443/1); Late Jurassic.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="311" type="diagnosis">
<paragraph pageId="4" pageNumber="311">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="311">The new species is distinguished by its medium size (wing length 3.7 mm), well-developed, strongly elongate proboscis, wing with C produced beyond R4+5 and reaching wing tip, strongly sclerotized abdomen and legs, and one spermatheca situated proximally of other two.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="311" type="description">
<paragraph pageId="4" pageNumber="311">Description.</paragraph>
<paragraph pageId="4" pageNumber="311">
Female (Fig. 3). Measurements (mm): Total length 4.1; thorax length 1.5, width ca. 1.0; abdomen length 2.9; wing length 3.7. Total length / wing length 1.1. Coloration. Head, thorax, abdomen, legs uniformly dark. Head 700
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, no less than 1100
<normalizedToken originalValue="μm">μm</normalizedToken>
high with proboscis, 550
<normalizedToken originalValue="μm">μm</normalizedToken>
high to lower eye margin. Eyes large, with wide dorsomedial extension, well-separated by ca. 100
<normalizedToken originalValue="μm">μm</normalizedToken>
. Facets equal. Frontal strip between eye extensions dark-coloured, long, sclerotized. Coronal triangle ca. 100
<normalizedToken originalValue="μm">μm</normalizedToken>
high, coronal suture ca. 70
<normalizedToken originalValue="μm">μm</normalizedToken>
high. Scape ca. 125
<normalizedToken originalValue="μm">μm</normalizedToken>
, pedicel ca. 40
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter. Clypeus ca. 200
<normalizedToken originalValue="μm">μm</normalizedToken>
high, ca. 150
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Proboscis very long, strong (longer than remainder of head), tapering, ca. 550
<normalizedToken originalValue="μm">μm</normalizedToken>
long, ca. 80
<normalizedToken originalValue="μm">μm</normalizedToken>
wide at (visible) apex; apical part with pair of sclerotized blades. Probable palpi no less than 400
<normalizedToken originalValue="μm">μm</normalizedToken>
long, two visible segments elongate, cylindrical, ca. 150
<normalizedToken originalValue="μm">μm</normalizedToken>
long, ca. 30
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Thorax. Scutum 900
<normalizedToken originalValue="μm">μm</normalizedToken>
long; postnotum ca. 250
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Wing clearly longer than abdomen. Vein C produced beyond R4+5 (costal extension ca. 200
<normalizedToken originalValue="μm">μm</normalizedToken>
), reaching wing tip; cell c at r-m level broader than cell r1; cell r5 at R1 tip level almost twice as wide as cell r1; R1 straight, 2/3 as long as R4+5. Vein C, radial veins, M, r-m strong and coloured; other veins thin, pale. Legs. Mid- and hind femora ca. 180
<normalizedToken originalValue="μm">μm</normalizedToken>
wide; tibiae 100-120
<normalizedToken originalValue="μm">μm</normalizedToken>
wide near apex. Apex of mid- or hind tibia with traces of at least 5 spiniform setae. Abdomen. Segments
<normalizedToken originalValue="IIVIII">II-VIII</normalizedToken>
250-300
<normalizedToken originalValue="μm">μm</normalizedToken>
long, 800-900
<normalizedToken originalValue="μm">μm</normalizedToken>
wide. Three rounded sclerotized spermathecae 70-90
<normalizedToken originalValue="μm">μm</normalizedToken>
in diameter, with long necks, largest spermatheca situated proximally of other two. Gonapophysis IX distinctly visible, sclerotized, with notum ca. 200
<normalizedToken originalValue="μm">μm</normalizedToken>
long and rami ca. 50
<normalizedToken originalValue="μm">μm</normalizedToken>
long. Posterior margin of sternite VIII bilobate: probable gonocoxites VIII (gonacoxapodemes?) visible distal to spermathecae. Cerci distinct, elongate-oval, 150
<normalizedToken originalValue="μm">μm</normalizedToken>
long, ca. 70
<normalizedToken originalValue="μm">μm</normalizedToken>
wide.
</paragraph>
<caption pageId="4" pageNumber="311">
<paragraph pageId="4" pageNumber="311">
Figure 3.?
<taxonomicName class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius robustus" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="species" species="robustus">Podonomius robustus</taxonomicName>
sp. n., holotype (a, d head, positive impression b, e abdominal apex, negative impression c, h wing, negative impression f female habitus, negative impression g apex of hind tibia, positive impression; all photos made under alcohol except for f); Shar Teg, J3. Scale bar 0.5 mm
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="311" type="remarks">
<paragraph pageId="4" pageNumber="311">Remarks.</paragraph>
<paragraph pageId="4" pageNumber="311">
Adults of this new species are the largest among
<taxonomicName class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="genus">Podonomius</taxonomicName>
(wing length is similar only in?
<taxonomicName class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius simplex" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="species" species="simplex">Podonomius simplex</taxonomicName>
Kalugina, 1985 (J2, Kubekovo, South Siberia), but in the Siberian species R1 is curved up distally).?
<taxonomicName class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius robustus" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="species" species="robustus">Podonomius robustus</taxonomicName>
sp. n. differs from other species from Shar Teg also in the longer costal extension. Such a long costa extending to the wing tip is an important plesiomorphic character (
<bibRefCitation author="Brundin, L" journalOrPublisher="Zoologica Scripta" pageId="7" pageNumber="314" pagination="139 - 160" title="A Neocomian chironomid and Podonominae-Aphroteniinae (Diptera) in the light of phylogenetics and biogeography." volume="5" year="1976">Brundin 1976</bibRefCitation>
) and may be a character of generic value. Thus, the new species is only tentatively placed in
<taxonomicName class="Insecta" family="Chironomidae" genus="Podonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podonomius" order="Diptera" pageId="4" pageNumber="311" phylum="Arthropoda" rank="genus">Podonomius</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>